705 lines
78 KiB
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705 lines
78 KiB
XML
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<mods:title id="755D62B3B83B34D06F30142224D2401E">Morphological revision of the western Palaearctic species of the genus Copris Geoffroy, 1762 with three foretibial external teeth (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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(
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)
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<emphasis id="B900E1128020FF83FE30FE4A32E4FE40" bold="true" box="[408,478,481,505]" italics="true" pageId="9" pageNumber="8">pueli</emphasis>
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<bibRefCitation id="EFE540F18020FF83FE4EFE4A3070FE40" author="Mollandin de Boissy, R." box="[486,842,481,506]" pageId="9" pageNumber="8" pagination="110 - 113" refId="ref16463" refString="Mollandin de Boissy, R. 1905. Description d'un Copris Geoffroy nouveau d'Algerie (Col.). Bulletin de la Societe entomologique de France: 110 - 113." type="book chapter" year="1905">Mollandin de Boissy, 1905</bibRefCitation>
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(
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<figureCitation id="134F21858020FF83FF61FDA93201FDA3" box="[201,315,514,538]" captionStart-0="Figures 8–11" captionStart-1="Figures 12–13" captionStartId-0="23.[192,269,1549,1571]" captionStartId-1="24.[192,269,1205,1227]" captionTargetBox-0="[257,1363,305,1515]" captionTargetBox-1="[343,1311,336,1175]" captionTargetId-0="figure-71@23.[192,1439,192,1535]" captionTargetId-1="figure-41@24.[192,1439,192,1193]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="Figures 8–11. Copris pueli Molland de Boissy, 1905. 8) Habitus of male (TN-Jendouba prov., Ain Draham). 9) Habitus of female (TN-Jendouba prov., Ain Draham). 10) Head and pronotum of more developed male, posterolateral view (TN- Jendouba prov., Ain Draham). 11) Head and pronotum of less developed male, posterolateral view (syntype, DZ- Kabylie). Photographs by A. Degiovanni." captionText-1="Figures 12–13. Copris pueli Molland de Boissy, 1905. 12) Parameres, dorsal view (DZ-Mt. Edough). 13) Parameres, lateral view (DZ-Mt. Edough). Photographs by A. Degiovanni." figureDoi-0="http://doi.org/10.5281/zenodo.4645825" figureDoi-1="http://doi.org/10.5281/zenodo.4645829" httpUri-0="https://zenodo.org/record/4645825/files/figure.png" httpUri-1="https://zenodo.org/record/4645829/files/figure.png" pageId="9" pageNumber="8">Fig. 8–13</figureCitation>
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)
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<emphasis id="B900E1128020FF83FF68FDE9326FFDE3" box="[192,341,578,602]" italics="true" pageId="9" pageNumber="8">Copris pueli</emphasis>
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<bibRefCitation id="EFE540F18020FF83FEF4FDE931E4FDE3" author="Mollandin de Boissy, R." box="[348,734,577,602]" pageId="9" pageNumber="8" pagination="110 - 113" refId="ref16463" refString="Mollandin de Boissy, R. 1905. Description d'un Copris Geoffroy nouveau d'Algerie (Col.). Bulletin de la Societe entomologique de France: 110 - 113." type="book chapter" year="1905">Mollandin de Boissy, 1905: 110</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FD42FDE930A5FDE3" author="Bedel, L." box="[746,927,578,602]" page="42" pageId="9" pageNumber="8" year="1911">
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<bibRefCitation id="EFE540F18020FF83FD42FDE930A5FDE3" author="Bedel, L." box="[746,927,578,602]" pageId="9" pageNumber="8" pagination="1 - 164" refId="ref14313" refString="Bedel, L. 1911. Faune des Coleopteres du Bassin del la Seine. Scarabeidae. Publications de la Societe Entomologique de France 4 (1): 1 - 164." type="journal article" year="1911">Bedel 1911: 42</bibRefCitation>
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(footnote);
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<treatmentCitation id="0AD51B118020FF83FB82FDE937DBFDE3" author="Gillet, J. J. - E." box="[1066,1249,578,602]" page="28" pageId="9" pageNumber="8" year="1910">
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<bibRefCitation id="EFE540F18020FF83FB82FDE937DBFDE3" author="Gillet, J. J. - E." box="[1066,1249,578,602]" pageId="9" pageNumber="8" pagination="1 - 31" refId="ref15121" refString="Gillet, J. J. - E. 1910. Especes nouvelles du genre Copris et releve synonymique des especes decrites a ce jour. Notes from the Leyden Museum 32 (1): 1 - 31." type="journal article" year="1910">Gillet 1910: 28</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FB46FDE93662FDE0" author="Gillet, J. J. - E." box="[1262,1368,578,602]" page="77" pageId="9" pageNumber="8" year="1911">
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<bibRefCitation id="EFE540F18020FF83FB46FDE93662FDE0" author="Gillet, J. J. - E." box="[1262,1368,578,602]" pageId="9" pageNumber="8" refId="ref15161" refString="Gillet, J. J. - E. 1911. Scarabaeidae: Coprinae I. Pars 38, Vol. 19 (2). In: S. Schenkling (ed.). Coleopterorum Catalogus. W. Junk; Berlin. 100 p." type="book" year="1911">1911: 77</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FACCFDE9328AFDC3" author="Winkler, A." page="1027" pageId="9" pageNumber="8" year="1929">
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<bibRefCitation id="EFE540F18020FF83FACCFDE9328AFDC3" author="Winkler, A." pageId="9" pageNumber="8" refId="ref17599" refString="Winkler, A. 1929. Catalogus Coleopterorum Regionis Palaearcticae (1924 - 1932). Scarabaeidae. p. 1009 - 1136. A. Winkler; Wien 9: 1698 p." type="book" year="1929">Winkler 1929: 1027</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FE13FDC9318CFDC3" author="Balthasar, V." box="[443,694,610,634]" page="107" pageId="9" pageNumber="8" year="1929">
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<bibRefCitation id="EFE540F18020FF83FE13FDC9318CFDC3" author="Balthasar, V." box="[443,694,610,634]" pageId="9" pageNumber="8" pagination="105 - 107" refId="ref14039" refString="Balthasar, V. 1929. Zwei neue Aphodius - Arten (subg. Bodilus) aus Nordafrika nebsteinigen Bemerkungen uber die von mir in Tunesien und Ostalgerien gesammelten Coprophagen. Entomologische Blatter. 25 (2): 105 - 107." type="journal article" year="1929">Balthasar 1929: 107</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FD69FDC930EEFDC3" author="Schatzmayr, A." box="[705,980,610,634]" page="111" pageId="9" pageNumber="8" year="1930">
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<bibRefCitation id="EFE540F18020FF83FD69FDC930EEFDC3" author="Schatzmayr, A." box="[705,980,610,634]" pageId="9" pageNumber="8" pagination="110 - 114" refId="ref17351" refString="Schatzmayr, A. 1930. Risultati scientifici delle spedizioni entomologiche di S. A. S. il Principe Alessandro della Torre e Tasso nell'Africa settentrionale e in Sicilia. Bollettino della Societa Entomologica Italiana 62: 110 - 114." type="journal article" year="1930">Schatzmayr 1930: 111</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FC48FDC9379EFDC3" author="Porta, A." box="[992,1188,610,634]" page="413" pageId="9" pageNumber="8" year="1932">
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<bibRefCitation id="EFE540F18020FF83FC48FDC9379EFDC3" author="Porta, A." box="[992,1188,610,634]" pageId="9" pageNumber="8" refId="ref17145" refString="Porta, A. 1932. Fauna Coleopterorum Italica. V. Rhynchophora - Lamellicornia. Stabilimento Tipografico Piacentino; Piacenza. 476 p." type="book" year="1932">Porta 1932: 413</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FB18FDC936A1FDC3" author="Balthasar, V." box="[1200,1435,609,634]" page="70" pageId="9" pageNumber="8" year="1935">
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<bibRefCitation id="EFE540F18020FF83FB18FDC936A1FDC3" author="Balthasar, V." box="[1200,1435,609,634]" pageId="9" pageNumber="8" refId="ref14082" refString="Balthasar, V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Bestimmungs-Tabellen der europaischen Coleopteren, Troppau 95. 112 p." type="book" year="1935">Balthasar 1935: 70</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FF58FD2932D3FD23" author="Normand, H." box="[240,489,641,666]" page="192" pageId="9" pageNumber="8" year="1936">
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<bibRefCitation id="EFE540F18020FF83FF58FD2932D3FD23" author="Normand, H." box="[240,489,641,666]" pageId="9" pageNumber="8" pagination="355 - 383" refId="ref16690" refString="Normand, H. 1936. Contribution au Catalogue des Coleopteres de la Tunisie. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord 27 (10): 355 - 383." type="journal article" year="1936">Normand 1936: 192</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FE5EFD2931F6FD20" author="Paulian, R." box="[502,716,641,666]" page="59" pageId="9" pageNumber="8" year="1941">
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<bibRefCitation id="EFE540F18020FF83FE5EFD2931F6FD20" author="Paulian, R." box="[502,716,641,666]" pageId="9" pageNumber="8" refId="ref16889" refString="Paulian, R. 1941. Faune de France. 38. Coleopteres Scarabeides. Federation Francaise des Societes de Sciences Naturelles (Ed. Lechevalier; Paris). 240 p." type="book" year="1941">Paulian 1941: 59</bibRefCitation>
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<treatmentCitation id="0AD51B118020FF83FD72FD2930F0FD23" author="Mackauer, M." box="[730,970,641,666]" page="49" pageId="9" pageNumber="8" year="1958">
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<bibRefCitation id="EFE540F18020FF83FD72FD2930F0FD23" author="Mackauer, M." box="[730,970,641,666]" pageId="9" pageNumber="8" pagination="46 - 54" refId="ref16131" refString="Mackauer, M. 1958. Eine Coprophagen-Ausbeute aus dem Noerdlichen Sizilien (Coleopt. Scarabaeidae). Memorie della Societa Entomologica Italiana 37: 46 - 54." type="journal article" year="1958">Mackauer 1958: 49</bibRefCitation>
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<treatmentCitation id="0AD51B118020FF83FC70FD293782FD20" author="Schaefer, L." box="[984,1208,641,666]" page="46" pageId="9" pageNumber="8" year="1958">
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<bibRefCitation id="EFE540F18020FF83FC70FD293782FD20" author="Schaefer, L." box="[984,1208,641,666]" pageId="9" pageNumber="8" pagination="45 - 46" refId="ref17318" refString="Schaefer, L. 1958. Le Copris umbilicatus dans l'Herault (Col. Scarabaeidae). Annales de la Societe d'Horticulture et d'Histoire Naturelle de l'Herault p. 45 - 46." type="book chapter" year="1958">Schaefer 1958: 46</bibRefCitation>
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;
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<treatmentCitation id="0AD51B118020FF83FB6EFD2936A1FD23" author="Paulian, R." box="[1222,1435,641,666]" page="74" pageId="9" pageNumber="8" year="1959">
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<bibRefCitation id="EFE540F18020FF83FB6EFD2936A1FD23" author="Paulian, R." box="[1222,1435,641,666]" pageId="9" pageNumber="8" refId="ref16922" refString="Paulian, R. 1959. Faune de France. 63. Coleopteres Scarabeides. (Deuxieme edition, revue et augmentee). Federation Francaise des Societes de Sciences Naturelles, Libraire de la Faculte des Sciences; Paris. 298 p." type="book" year="1959">Paulian 1959: 74</bibRefCitation>
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<treatmentCitation id="0AD51B118020FF83FF58FD0932D6FD03" author="Balthasar, V." box="[240,492,673,698]" page="334" pageId="9" pageNumber="8" year="1963">
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<bibRefCitation id="EFE540F18020FF83FF58FD0932D6FD03" author="Balthasar, V." box="[240,492,673,698]" pageId="9" pageNumber="8" refId="ref14126" refString="Balthasar, V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 1. 1. Scarabaeinae. 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften; Prag. 391 p." type="book" year="1963">Balthasar 1963: 334</bibRefCitation>
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<treatmentCitation id="0AD51B118020FF83FE51FD0931D5FD03" author="Dellacasa, G." box="[505,751,673,698]" page="139" pageId="9" pageNumber="8" year="1968">
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||
<bibRefCitation id="EFE540F18020FF83FE51FD0931D5FD03" author="Dellacasa, G." box="[505,751,673,698]" pageId="9" pageNumber="8" pagination="135 - 142" refId="ref14916" refString="Dellacasa, G. 1968. Sulle sottospecie del Copris (s. str.) hispanus nell'ambito della fauna italiana (Coleoptera Scarabaeidae). Bollettino della Societa Entomologica Italiana 98 (9 - 10): 135 - 142." type="journal article" year="1968">Dellacasa 1968: 139</bibRefCitation>
|
||
</treatmentCitation>
|
||
(footnote);
|
||
<treatmentCitation id="0AD51B118020FF83FCD4FD0937E2FD03" author="Zaharieva-Stoilova, B." box="[892,1240,674,698]" page="42" pageId="9" pageNumber="8" year="1970">
|
||
<bibRefCitation id="EFE540F18020FF83FCD4FD0937E2FD03" author="Zaharieva-Stoilova, B." box="[892,1240,674,698]" pageId="9" pageNumber="8" pagination="39 - 44" refId="ref17633" refString="Zaharieva-Stoilova, B. 1970. Otnosno Redkiya vid Copris umbilicatus Ab. (Scarabaeidae, Coleoptera). Bulletin de l'Institut de Zoologie et Musee 32: 39 - 44." type="journal article" year="1970">Zaharieva-Stoilova 1970: 42</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FB4DFD093225FD60" author="Petrovitz, R." page="565" pageId="9" pageNumber="8" year="1971">
|
||
<bibRefCitation id="EFE540F18020FF83FB4DFD093225FD60" author="Petrovitz, R." pageId="9" pageNumber="8" pagination="565 - 589" refId="ref17079" refString="Petrovitz, R. 1971. Ergebnisse zoologischer Sammelreisen in der Turkei: Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums 75: 565 - 589." type="journal article" year="1971">Petrovitz 1971: 565</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FE82FD6932CFFD63" author="Baraud, J." box="[298,501,706,730]" page="29" pageId="9" pageNumber="8" year="1977">
|
||
<bibRefCitation id="EFE540F18020FF83FE82FD6932CFFD63" author="Baraud, J." box="[298,501,706,730]" pageId="9" pageNumber="8" pagination="1 - 352" refId="ref14201" refString="Baraud, J. 1977. Coleopteres Scarabaeoidea, Faune de l'Europe occidentale: Belgique, France, Grande- Bretagne, Italie, Peninsule Iberique. Supplement a la Nouvelle Revue d'Entomologie 7 (1): 1 - 352." type="journal article" year="1977">Baraud 1977: 29</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FDA8FD69304FFD63" author="Paulian, R. & J. Baraud" box="[512,885,706,730]" page="241" pageId="9" pageNumber="8" year="1982">
|
||
<bibRefCitation id="EFE540F18020FF83FDA8FD69304FFD63" author="Paulian, R. & J. Baraud" box="[512,885,706,730]" pageId="9" pageNumber="8" refId="ref16966" refString="Paulian, R., and J. Baraud. 1982. Fauna des Coleopteres de France. II Lucanoidea et Scarabaeoidea. Ed. Lechevalier; Paris. 473 p." type="book" year="1982">Paulian and Baraud 1982: 241</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FC28FD693760FD60" author="Baraud, J." box="[896,1114,705,730]" page="256" pageId="9" pageNumber="8" year="1985">
|
||
<bibRefCitation id="EFE540F18020FF83FC28FD693760FD60" author="Baraud, J." box="[896,1114,705,730]" pageId="9" pageNumber="8" refId="ref14242" refString="Baraud, J. 1985. Coleopteres Scarabaeoidea, Faune du Nord de l'Afrique, du Maroc au Sinai. Editions Lechevalier; Paris. 650 p." type="book" year="1985">Baraud 1985: 256</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FBCDFD69367BFD63" author="Kabakov, O. N." box="[1125,1345,705,730]" page="88" pageId="9" pageNumber="8" year="2006">
|
||
<bibRefCitation id="EFE540F18020FF83FBCDFD69367BFD63" author="Kabakov, O. N." box="[1125,1345,705,730]" pageId="9" pageNumber="8" refId="ref15548" refString="Kabakov, O. N. 2006. Plastinchatousye zhuki podsemeystva Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) fauny Rossii i sopredelnykh stran. - Tovarishchestvo Nauchnykh Izdanii KMK; Moskva. 374 p." type="book" year="2006">Kabakov 2006: 88</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FAE4FD6932B5FD40" author="Lobl, I. & F. - T. Krell & D. Kral" page="152" pageId="9" pageNumber="8" year="2006">
|
||
<bibRefCitation id="EFE540F18020FF83FAE4FD6932B5FD40" author="Lobl, I. & F. - T. Krell & D. Kral" pageId="9" pageNumber="8" pagination="151 - 154" refId="ref15923" refString="Lobl, I., F. - T. Krell, and D. Kral. 2006. Scarabaeidae, subfamily Scarabaeinae, tribe Coprini. p. 151 - 154. In: O. Lobl, and A. Smetana (eds.). Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books; Stenstrup. 690 p." type="book chapter" year="2006">Löbl et al. 2006: 152</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FE33FD49300AFD40" author="Maughan, N. & A. Paulian" box="[411,816,737,762]" page="436" pageId="9" pageNumber="8" year="2011">
|
||
<bibRefCitation id="EFE540F18020FF83FE33FD49300AFD40" author="Maughan, N. & A. Paulian" box="[411,816,737,762]" pageId="9" pageNumber="8" pagination="435 - 452" refId="ref16338" refString="Maughan, N., and A. Paulian. 2011. Precisions sur la biogeographie du genre Copris Geoffroy, 1762, dans le sud-est de la France (Coleoptera, Scarabaeidae, Coprini). Bulletin de la Societe Entomologique de France 116 (4): 435 - 452." type="journal article" year="2011">Maughan and Paulian 2011: 436</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FC94FD4937B6FD40" author="Kral, D. & A. Bezdek" box="[828,1164,737,762]" page="169" pageId="9" pageNumber="8" year="2016">
|
||
<bibRefCitation id="EFE540F18020FF83FC94FD4937B6FD40" author="Kral, D. & A. Bezdek" box="[828,1164,737,762]" pageId="9" pageNumber="8" pagination="167 - 170" refId="ref15730" refString="Kral, D., and A. Bezdek. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Coprini. p. 167 - 170. In: I. Lobl, and D. Lobl (eds). Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. Brill; Leiden-Boston. 983 p." type="book chapter" year="2016">Král and Bezděk 2016: 169</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD51B118020FF83FB30FD493640FD40" author="Tonelli & al." box="[1176,1402,737,762]" pageId="9" pageNumber="8" year="2016">
|
||
<bibRefCitation id="EFE540F18020FF83FB30FD493640FD40" author="Tonelli, M. & R. Agoglitta & H. Dawson & M. Zunino" box="[1176,1402,737,762]" pageId="9" pageNumber="8" pagination="161 - 171" refId="ref17546" refString="Tonelli, M., R. Agoglitta, H. Dawson, and M. Zunino. 2016. On the road of dung: hypothetical dispersal routes of dung beetles in the circum-Sicilian volcanic islands. Animal Biodiversity and Conservation 39 (2): 161 - 171 + ix." type="journal article" year="2016">Tonelli et al. 2016</bibRefCitation>
|
||
</treatmentCitation>
|
||
: ii.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6E8B8020FF83FF68FC8931FCFC20" pageId="9" pageNumber="8" type="materials_examined">
|
||
<paragraph id="8BCB3D008020FF83FF68FC89302AFC83" blockId="9.[192,784,801,826]" box="[192,784,801,826]" pageId="9" pageNumber="8">
|
||
<emphasis id="B900E1128020FF83FF68FC89324EFC80" bold="true" box="[192,372,801,826]" pageId="9" pageNumber="8">
|
||
<typeStatus id="54CF83A28020FF83FF68FC89323FFC83" box="[192,261,802,826]" pageId="9" pageNumber="8">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. “Bou-Berak (Kabylie)” [
|
||
<collectingCountry id="F3637D908020FF83FD0BFC89303FFC83" box="[675,773,802,826]" name="Algeria" pageId="9" pageNumber="8">Algeria</collectingCountry>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008020FF83FF68FCC931FCFC20" blockId="9.[192,1440,865,922]" pageId="9" pageNumber="8">
|
||
<emphasis id="B900E1128020FF83FF68FCC932BAFCC0" bold="true" box="[192,384,865,890]" pageId="9" pageNumber="8">Type material</emphasis>
|
||
. Eleven
|
||
<typeStatus id="54CF83A28020FF83FE40FCC9316DFCC3" box="[488,599,866,890]" pageId="9" pageNumber="8" type="syntype">syntypes</typeStatus>
|
||
,
|
||
<specimenCount id="9D72F6898020FF83FDCAFCC93197FCC0" box="[610,685,866,890]" pageId="9" pageNumber="8" type="male">10 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FD4FFCC93035FCC0" box="[743,783,866,890]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
, by original designation. Four probable
|
||
<typeStatus id="54CF83A28020FF83FB5AFCC93665FCC3" box="[1266,1375,866,890]" pageId="9" pageNumber="8" type="syntype">syntypes</typeStatus>
|
||
were examined from the MNHN:
|
||
<specimenCount id="9D72F6898020FF83FDB1FC29316DFC20" box="[537,599,898,922]" pageId="9" pageNumber="8" type="male">3 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FD3DFC293185FC20" box="[661,703,898,922]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6E8B8020FF83FF68FC6A304FFC60" box="[192,885,961,986]" pageId="9" pageNumber="8" type="distribution">
|
||
<paragraph id="8BCB3D008020FF83FF68FC6A304FFC60" blockId="9.[192,885,961,986]" box="[192,885,961,986]" pageId="9" pageNumber="8">
|
||
<emphasis id="B900E1128020FF83FF68FC6A324EFC60" bold="true" box="[192,372,961,985]" pageId="9" pageNumber="8">Distribution.</emphasis>
|
||
<collectingCountry id="F3637D908020FF83FED3FC6932E2FC63" box="[379,472,962,986]" name="Algeria" pageId="9" pageNumber="8">Algeria</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3637D908020FF83FE4CFC693178FC63" box="[484,578,962,986]" name="Tunisia" pageId="9" pageNumber="8">Tunisia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE540F18020FF83FDFAFC693053FC60" author="Kral, D. & A. Bezdek" box="[594,873,961,986]" pageId="9" pageNumber="8" pagination="167 - 170" refId="ref15730" refString="Kral, D., and A. Bezdek. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Coprini. p. 167 - 170. In: I. Lobl, and D. Lobl (eds). Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. Brill; Leiden-Boston. 983 p." type="book chapter" year="2016">Král and Bezděk 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6E8B8020FF83FF68FBAA379FFA40" pageId="9" pageNumber="8" type="materials_examined">
|
||
<paragraph id="8BCB3D008020FF83FF68FBAA319EFAC3" blockId="9.[192,1441,1025,1402]" pageId="9" pageNumber="8">
|
||
<emphasis id="B900E1128020FF83FF68FBAA32F1FBA0" bold="true" box="[192,459,1025,1049]" pageId="9" pageNumber="8">Material examined.</emphasis>
|
||
<materialsCitation id="3B1C375D8020FF83FE78FBAA3661FBA3" ID-GBIF-Occurrence="3776326384" box="[464,1371,1025,1050]" collectionCode="MNHN" collectorName="Kabjlie L. Puel" country="Algeria" location="Bou Berak" pageId="9" pageNumber="8" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="syntype">
|
||
<emphasis id="B900E1128020FF83FE78FBAA3165FBA0" bold="true" box="[464,607,1025,1049]" pageId="9" pageNumber="8">
|
||
<collectingCountry id="F3637D908020FF83FE78FBAA3160FBA0" box="[464,602,1025,1049]" name="Algeria" pageId="9" pageNumber="8">ALGERIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“
|
||
<location id="8EAB6BDB8020FF83FDC7FBA931CAFBA3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FDC7FBA931CAFBA3" box="[623,752,1026,1050]" country="Algeria" name="Bou Berak" pageId="9" pageNumber="8">Bou Berak</location>
|
||
/
|
||
<collectorName id="268158D68020FF83FCABFBA93083FBA3" box="[771,953,1026,1050]" pageId="9" pageNumber="8">Kabjlie L. Puel</collectorName>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FC66FBA93731FBA0" box="[974,1035,1026,1050]" pageId="9" pageNumber="8" type="male">4 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FBECFBA93756FBA0" box="[1092,1132,1026,1050]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<typeStatus id="54CF83A28020FF83FBD3FBA937D0FBA3" box="[1147,1258,1026,1050]" pageId="9" pageNumber="8" type="syntype">syntypes</typeStatus>
|
||
,
|
||
<collectionCode id="ED65A5C58020FF83FB5CFBA93669FBA3" box="[1268,1363,1026,1050]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FACDFBA931C4FB83" ID-GBIF-Occurrence="3776326385" collectionCode="MNHN" collectorName="Kabjlie L. Puel" country="Algeria" location="Bou Berak" pageId="9" pageNumber="8" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="syntype">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FAC7FBA93231FB83" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FAC7FBA93231FB83" country="Algeria" name="Bou Berak" pageId="9" pageNumber="8">Bou Berak</location>
|
||
/
|
||
<collectorName id="268158D68020FF83FEB6FB8932EFFB83" box="[286,469,1058,1082]" pageId="9" pageNumber="8">Kabjlie L. Puel</collectorName>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FE42FB89311DFB80" box="[490,551,1058,1082]" pageId="9" pageNumber="8" type="male">2 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FDC8FB8931B3FB80" box="[608,649,1058,1082]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FD3FFB8931CCFB83" box="[663,758,1058,1082]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FCA0FB8936A1FB83" ID-GBIF-Occurrence="3776326312" box="[776,1435,1057,1082]" collectionCode="MNHN" collectorName="Kabjlie L. Puel" country="Algeria" location="Bou Berak" pageId="9" pageNumber="8" specimenCount="9" specimenCount-female="6" specimenCount-male="3" typeStatus="syntype">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FCBBFB8930AEFB83" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FCBBFB8930AEFB83" box="[787,916,1058,1082]" country="Algeria" name="Bou Berak" pageId="9" pageNumber="8">Bou Berak</location>
|
||
/
|
||
<collectorName id="268158D68020FF83FC0FFB893764FB83" box="[935,1118,1058,1082]" pageId="9" pageNumber="8">Kabjlie L. Puel</collectorName>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FBDCFB89378AFB80" box="[1140,1200,1058,1082]" pageId="9" pageNumber="8" type="male">3 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FB42FB8A361CFB80" box="[1258,1318,1057,1081]" pageId="9" pageNumber="8" type="female">6 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FA9CFB8936A9FB83" box="[1332,1427,1058,1082]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FF68FBE931D9FBE3" ID-GBIF-Occurrence="3776326349" box="[192,739,1090,1114]" collectionCode="MHNG" country="Algeria" location="Bou Berak" pageId="9" pageNumber="8" specimenCount="1" specimenCount-male="1">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FF62FBE93275FBE3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FF62FBE93275FBE3" box="[202,335,1090,1114]" country="Algeria" name="Bou Berak" pageId="9" pageNumber="8">Bou Berak</location>
|
||
/
|
||
<location id="8EAB6BDB8020FF83FEC1FBE9311DFBE3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FEC1FBE9311DFBE3" box="[361,551,1090,1114]" country="Algeria" name="Kabjlie L. Puel" pageId="9" pageNumber="8">Kabjlie L. Puel</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FDE8FBE93156FBE0" box="[576,620,1090,1114]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FDD5FBE931E6FBE3" box="[637,732,1090,1114]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="8" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FD59FBE93791FBE3" ID-GBIF-Occurrence="3776326391" box="[753,1195,1090,1114]" collectionCode="MHNG" country="Algeria" location="Bou Berak" pageId="9" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FD53FBE930B8FBE3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FD53FBE930B8FBE3" box="[763,898,1090,1114]" country="Algeria" name="Bou Berak" pageId="9" pageNumber="8">Bou Berak</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FC33FBE930FDFBE0" box="[923,967,1090,1114]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FBA0FBE9370EFBE0" box="[1032,1076,1090,1114]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FBEDFBE9379EFBE3" box="[1093,1188,1090,1114]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="8" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FB11FBE93085FBC3" ID-GBIF-Occurrence="3776326342" collectionCode="MTD" collectorName="C. Felsche" country="Algeria" location="Algeria" pageId="9" pageNumber="8" specimenCount="1" specimenCount-male="2">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FB6BFBE93627FBE3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FB6BFBE93627FBE3" box="[1219,1309,1090,1114]" country="Algeria" name="Algeria" pageId="9" pageNumber="8">Algeria</location>
|
||
/ Reitter”; “Copris / pueli”; “coll.
|
||
<collectorName id="268158D68020FF83FE65FBC93177FBC3" box="[461,589,1122,1146]" pageId="9" pageNumber="8">C. Felsche</collectorName>
|
||
/ Kauf 20.1918”,
|
||
<specimenCount id="9D72F6898020FF83FC8CFBC93058FBC0" box="[804,866,1122,1146]" pageId="9" pageNumber="8" type="male">2 ♂♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FCDBFBC9308DFBC3" box="[883,951,1122,1146]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="9" pageNumber="8">MTD</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FC63FBC9320AFB23" ID-GBIF-Occurrence="3776326361" collectionCode="MNHN" country="Algeria" location="Massif des Mouzaia" pageId="9" pageNumber="8" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FC7EFBC937F7FBC3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FC7EFBC937F7FBC3" box="[982,1229,1122,1146]" country="Algeria" name="Massif des Mouzaia" pageId="9" pageNumber="8">Massif des Mouzaia</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FB4CFBC93619FBC0" box="[1252,1315,1122,1146]" pageId="9" pageNumber="8" type="male">2 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FAC9FBC9369AFBC0" box="[1377,1440,1122,1146]" pageId="9" pageNumber="8" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FF60FB29321DFB23" box="[200,295,1154,1178]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FE92FB2931C9FB23" ID-GBIF-Occurrence="3776326331" box="[314,755,1154,1178]" collectionCode="MNHN" country="Algeria" location="Mouzaia" pageId="9" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
“les
|
||
<location id="8EAB6BDB8020FF83FEC5FB2932E3FB23" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FEC5FB2932E3FB23" box="[365,473,1154,1178]" country="Algeria" name="Mouzaia" pageId="9" pageNumber="8">Mouzaïa</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FE47FB293123FB20" box="[495,537,1154,1178]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FDFCFB293147FB20" box="[596,637,1154,1178]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FD24FB2931D1FB23" box="[652,747,1154,1178]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FD56FB293713FB23" ID-GBIF-Occurrence="3776326366" box="[766,1065,1154,1178]" collectionCode="MHNG" country="Algeria" location="Mouzaia" pageId="9" pageNumber="8" specimenCount="1" specimenCount-female="1">
|
||
“
|
||
<location id="8EAB6BDB8020FF83FCA1FB29304FFB23" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FCA1FB29304FFB23" box="[777,885,1154,1178]" country="Algeria" name="Mouzaia" pageId="9" pageNumber="8">Mouzaïa</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FC23FB29308EFB20" box="[907,948,1154,1178]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FC6AFB29371BFB23" box="[962,1057,1154,1178]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="8" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FB9CFB2932ADFB03" ID-GBIF-Occurrence="3776326303" collectionCode="MNHN" country="Algeria" location="Mt. Edough" municipality="Algerie" pageId="9" pageNumber="8" specimenCount="51" specimenCount-female="1" specimenCount-male="50">
|
||
“
|
||
<collectingMunicipality id="6BAFA77A8020FF83FB96FB2937ADFB23" box="[1086,1175,1154,1178]" pageId="9" pageNumber="8">Algerie</collectingMunicipality>
|
||
/
|
||
<location id="8EAB6BDB8020FF83FB03FB293604FB23" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FB03FB293604FB23" box="[1195,1342,1154,1178]" country="Algeria" municipality="Algerie" name="Mt. Edough" pageId="9" pageNumber="8">Mt. Edough</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FAFCFB2A369AFB20" box="[1364,1440,1153,1177]" pageId="9" pageNumber="8" type="male">50 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FF5EFB09321AFB00" box="[246,288,1186,1210]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FE98FB0932B5FB03" box="[304,399,1186,1210]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FE0BFB093015FB03" ID-GBIF-Occurrence="3776326343" box="[419,815,1186,1210]" collectionCode="MNHN" country="Algeria" location="Reitter" municipality="Algeria" pageId="9" pageNumber="8" specimenCount="1" specimenCount-male="1">
|
||
“
|
||
<collectingMunicipality id="6BAFA77A8020FF83FE05FB09313DFB03" box="[429,519,1186,1210]" pageId="9" pageNumber="8">Algeria</collectingMunicipality>
|
||
/
|
||
<location id="8EAB6BDB8020FF83FDB6FB09314DFB03" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FDB6FB09314DFB03" box="[542,631,1186,1210]" country="Algeria" municipality="Algeria" name="Reitter" pageId="9" pageNumber="8">Reitter</location>
|
||
”,
|
||
<specimenCount id="9D72F6898020FF83FD26FB093182FB00" box="[654,696,1186,1210]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FD60FB09301DFB03" box="[712,807,1186,1210]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="8">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FC93FB093732FB63" ID-GBIF-Occurrence="3776326308" collectingDate="1988-05-17" collectionCode="MHNG" collectorName="Besuchet & Lobl & Burckh." country="Algeria" county="Gde Kabylie" elevation="1300" location="Adekar" municipality="Foret d'Akfadou" pageId="9" pageNumber="8" specimenCount="2" specimenCount-male="2">
|
||
“
|
||
<collectingCounty id="62AA458C8020FF83FCEDFB0930E7FB03" box="[837,989,1186,1210]" pageId="9" pageNumber="8">Gde Kabylie</collectingCounty>
|
||
/
|
||
<collectingMunicipality id="6BAFA77A8020FF83FC5BFB093787FB03" box="[1011,1213,1186,1210]" pageId="9" pageNumber="8">Forêt d’Akfadou</collectingMunicipality>
|
||
,
|
||
<quantity id="4C8C90E58020FF83FB61FB09363DFB03" box="[1225,1287,1186,1210]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="8" unit="km" value="9.0">9 km</quantity>
|
||
W /
|
||
<location id="8EAB6BDB8020FF83FA96FB0936A1FB03" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FA96FB0936A1FB03" box="[1342,1435,1186,1210]" country="Algeria" county="Gde Kabylie" municipality="Foret d'Akfadou" name="Adekar" pageId="9" pageNumber="8">Adekar</location>
|
||
,
|
||
<quantity id="4C8C90E58020FF83FF68FB693227FB63" box="[192,285,1218,1242]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="8" unit="m" value="1300.0">
|
||
<elevation id="0059DA338020FF83FF68FB693227FB63" box="[192,285,1218,1242]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="8" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
”,
|
||
<date id="FFCA1BC08020FF83FE9EFB693294FB63" box="[310,430,1218,1242]" pageId="9" pageNumber="8" value="1988-05-17">
|
||
<collectingDate id="EF8EE2288020FF83FE9EFB693294FB63" box="[310,430,1218,1242]" pageId="9" pageNumber="8" value="1988-05-17">17.v.1988</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268158D68020FF83FE14FB69310AFB63" box="[444,560,1218,1242]" pageId="9" pageNumber="8">Besuchet</collectorName>
|
||
,
|
||
<collectorName id="268158D68020FF83FD95FB693149FB63" box="[573,627,1218,1242]" pageId="9" pageNumber="8">Löbl</collectorName>
|
||
and
|
||
<collectorName id="268158D68020FF83FD1BFB69302CFB63" box="[691,790,1218,1242]" pageId="9" pageNumber="8">Burckh.</collectorName>
|
||
leg.
|
||
<specimenCount id="9D72F6898020FF83FCFAFB6930ABFB60" box="[850,913,1218,1242]" pageId="9" pageNumber="8" type="male">2 ♂♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FC0AFB69373BFB63" box="[930,1025,1218,1242]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="8" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FBBDFB693637FB43" ID-GBIF-Occurrence="3776326325" collectingDate="1988-05-16" collectionCode="MHNG" collectorName="Besuchet & Lobl & Burckh." country="Algeria" county="Gde Kabylie" elevation="1050" location="Yakouren" municipality="Foret d'Akfadou" pageId="9" pageNumber="8" specimenCount="1" specimenCount-male="1">
|
||
“
|
||
<collectingCounty id="62AA458C8020FF83FB88FB693783FB63" box="[1056,1209,1218,1242]" pageId="9" pageNumber="8">Gde Kabylie</collectingCounty>
|
||
/
|
||
<collectingMunicipality id="6BAFA77A8020FF83FB79FB6936A6FB63" box="[1233,1436,1218,1242]" pageId="9" pageNumber="8">Forêt d’Akfadou</collectingMunicipality>
|
||
,
|
||
<quantity id="4C8C90E58020FF83FF68FB493235FB43" box="[192,271,1250,1274]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="8" unit="km" value="22.0">22 km</quantity>
|
||
E /
|
||
<location id="8EAB6BDB8020FF83FEEEFB4932F8FB43" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FEEEFB4932F8FB43" box="[326,450,1250,1274]" country="Algeria" county="Gde Kabylie" municipality="Foret d'Akfadou" name="Yakouren" pageId="9" pageNumber="8">Yakouren</location>
|
||
,
|
||
<quantity id="4C8C90E58020FF83FE67FB4A3117FB43" box="[463,557,1249,1274]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="8" unit="m" value="1050.0">
|
||
<elevation id="0059DA338020FF83FE67FB4A3117FB43" box="[463,557,1249,1274]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="8" unit="m" value="1050.0">1050 m</elevation>
|
||
</quantity>
|
||
”,
|
||
<date id="FFCA1BC08020FF83FDEFFB4A31FAFB40" box="[583,704,1249,1273]" pageId="9" pageNumber="8" value="1988-05-16">
|
||
<collectingDate id="EF8EE2288020FF83FDEFFB4A31FAFB40" box="[583,704,1249,1273]" pageId="9" pageNumber="8" value="1988-05-16">16.v.1988</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268158D68020FF83FD66FB493078FB43" box="[718,834,1250,1274]" pageId="9" pageNumber="8">Besuchet</collectorName>
|
||
,
|
||
<collectorName id="268158D68020FF83FCF8FB4930BCFB43" box="[848,902,1250,1274]" pageId="9" pageNumber="8">Löbl</collectorName>
|
||
and
|
||
<collectorName id="268158D68020FF83FC6FFB493710FB43" box="[967,1066,1250,1274]" pageId="9" pageNumber="8">Burckh.</collectorName>
|
||
leg.
|
||
<specimenCount id="9D72F6898020FF83FBC0FB4937AFFB40" box="[1128,1173,1250,1274]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FB0EFB49363FFB43" box="[1190,1285,1250,1274]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="8" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FAB3FB493284FAA3" ID-GBIF-Occurrence="3776326335" collectingDate="1985-06" collectionCode="SZCM" country="Algeria" location="Ben Atala" pageId="9" pageNumber="8" specimenCount="1" specimenCount-female="1">
|
||
<location id="8EAB6BDB8020FF83FAB3FB4936A1FB43" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FAB3FB4936A1FB43" box="[1307,1435,1250,1274]" country="Algeria" name="Ben Atala" pageId="9" pageNumber="8">Ben Atala</location>
|
||
,
|
||
<date id="FFCA1BC08020FF83FF68FAAA3226FAA0" box="[192,284,1281,1305]" pageId="9" pageNumber="8" value="1985-06">
|
||
<collectingDate id="EF8EE2288020FF83FF68FAAA3226FAA0" box="[192,284,1281,1305]" pageId="9" pageNumber="8" value="1985-06">vi.1985</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D72F6898020FF83FE81FAA9326EFAA0" box="[297,340,1282,1306]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FECCFAA9328CFAA3" box="[356,438,1282,1306]" pageId="9" pageNumber="8">SZCM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FE63FAA93263FA83" ID-GBIF-Occurrence="3776326347" collectingDate="1986-06-02" collectionCode="SZCM" collectorName="G. Sama" country="Algeria" elevation="1400" location="Afkadou forest" pageId="9" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tizi Ouzou">
|
||
<collectingRegion id="49B0F3E28020FF83FE63FAA931A1FAA0" box="[459,667,1282,1306]" country="Algeria" name="Tizi Ouzou" pageId="9" pageNumber="8">Tizi Ouzou prov.</collectingRegion>
|
||
,
|
||
<location id="8EAB6BDB8020FF83FD0FFAA9305AFAA3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FD0FFAA9305AFAA3" box="[679,864,1282,1306]" country="Algeria" name="Afkadou forest" pageId="9" pageNumber="8" stateProvince="Tizi Ouzou">Afkadou forest</location>
|
||
, 1000/
|
||
<quantity id="4C8C90E58020FF83FC18FAA93737FAA0" box="[944,1037,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="8" unit="m" value="1400.0">
|
||
<elevation id="0059DA338020FF83FC18FAA93737FAA0" box="[944,1037,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="8" unit="m" value="1400.0">1400 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFCA1BC08020FF83FBB2FAAA37B1FAA0" box="[1050,1163,1281,1305]" pageId="9" pageNumber="8" value="1986-06-02">
|
||
<collectingDate id="EF8EE2288020FF83FBB2FAAA37B1FAA0" box="[1050,1163,1281,1305]" pageId="9" pageNumber="8" value="1986-06-02">2.vi.1986</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268158D68020FF83FB31FAA93639FAA3" box="[1177,1283,1282,1306]" pageId="9" pageNumber="8">G. Sama</collectorName>
|
||
leg.
|
||
<specimenCount id="9D72F6898020FF83FA97FAA93650FAA0" box="[1343,1386,1282,1306]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FF68FA8933D7FA80" box="[192,237,1314,1338]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FF56FA89326BFA83" box="[254,337,1314,1338]" pageId="9" pageNumber="8">SZCM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1C375D8020FF83FECFFA8A3623FA83" ID-GBIF-Occurrence="3776326309" box="[359,1305,1313,1338]" collectingDate="1982-06-07" collectionCode="SZCM" collectorName="Slama" country="Tunisia" location="Ain Draham" pageId="9" pageNumber="8" specimenCount="1" specimenCount-female="1" stateProvince="Jendouba">
|
||
<emphasis id="B900E1128020FF83FECFFA8A32DEFA80" bold="true" box="[359,484,1313,1337]" pageId="9" pageNumber="8">
|
||
<collectingCountry id="F3637D908020FF83FECFFA8A32DEFA80" box="[359,484,1313,1337]" name="Tunisia" pageId="9" pageNumber="8">TUNISIA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="49B0F3E28020FF83FE5DFA89318DFA80" box="[501,695,1314,1338]" country="Tunisia" name="Jendouba" pageId="9" pageNumber="8">Jendouba prov.</collectingRegion>
|
||
,
|
||
<location id="8EAB6BDB8020FF83FD6DFA89305EFA83" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FD6DFA89305EFA83" box="[709,868,1314,1338]" country="Tunisia" name="Ain Draham" pageId="9" pageNumber="8" stateProvince="Jendouba">Aïn Draham</location>
|
||
,
|
||
<date id="FFCA1BC08020FF83FCDAFA8930DFFA83" box="[882,997,1314,1338]" pageId="9" pageNumber="8" value="1982-06-07">
|
||
<collectingDate id="EF8EE2288020FF83FCDAFA8930DFFA83" box="[882,997,1314,1338]" pageId="9" pageNumber="8" value="1982-06-07">7.vi.1982</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268158D68020FF83FC5BFA893778FA83" box="[1011,1090,1314,1338]" pageId="9" pageNumber="8">Sláma</collectorName>
|
||
leg.
|
||
<specimenCount id="9D72F6898020FF83FB29FA893797FA80" box="[1153,1197,1314,1338]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FB17FA89362BFA83" box="[1215,1297,1314,1338]" pageId="9" pageNumber="8">SZCM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FA80FA89370CFAE3" ID-GBIF-Occurrence="3776326363" collectingDate="1995-04-09" collectionCode="SZCM" collectorName="S. and R. Ziani" country="Tunisia" location="Ain Draham" municipality="Jendouna prov." pageId="9" pageNumber="8" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
|
||
<collectingMunicipality id="6BAFA77A8020FF83FA80FA89323BFAE0" pageId="9" pageNumber="8">Jendouna prov.</collectingMunicipality>
|
||
,
|
||
<location id="8EAB6BDB8020FF83FEA5FAE93293FAE3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FEA5FAE93293FAE3" box="[269,425,1346,1370]" country="Tunisia" municipality="Jendouna prov." name="Ain Draham" pageId="9" pageNumber="8">Aïn Draham</location>
|
||
,
|
||
<date id="FFCA1BC08020FF83FE1DFAEA3112FAE0" box="[437,552,1345,1369]" pageId="9" pageNumber="8" value="1995-04-09">
|
||
<collectingDate id="EF8EE2288020FF83FE1DFAEA3112FAE0" box="[437,552,1345,1369]" pageId="9" pageNumber="8" value="1995-04-09">9.iv.1995</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268158D68020FF83FD9CFAE931D6FAE3" box="[564,748,1346,1370]" pageId="9" pageNumber="8">S. and R. Ziani</collectorName>
|
||
leg.
|
||
<specimenCount id="9D72F6898020FF83FC8EFAE9305EFAE0" box="[806,868,1346,1370]" pageId="9" pageNumber="8" type="male">3 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FC0AFAE930F6FAE0" box="[930,972,1346,1370]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FC73FAE93717FAE3" box="[987,1069,1346,1370]" pageId="9" pageNumber="8">SZCM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1C375D8020FF83FBE9FAE9319AFAC3" ID-GBIF-Occurrence="3776326377" collectingDate="2013-11-05" collectionCode="SZCM" collectorName="W. Ben Aba" country="Tunisia" location="Ain Essobh" pageId="9" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Jendouba">
|
||
<collectingRegion id="49B0F3E28020FF83FBE9FAE9363BFAE0" box="[1089,1281,1346,1370]" country="Tunisia" name="Jendouba" pageId="9" pageNumber="8">Jendouba prov.</collectingRegion>
|
||
,
|
||
<location id="8EAB6BDB8020FF83FAA5FAE936A1FAE3" LSID="urn:lsid:plazi:treatment:03DD8C168020FF81FF68FE4A3100FE43:8EAB6BDB8020FF83FAA5FAE936A1FAE3" box="[1293,1435,1346,1370]" country="Tunisia" name="Ain Essobh" pageId="9" pageNumber="8" stateProvince="Jendouba">Ain Essobh</location>
|
||
,
|
||
<date id="FFCA1BC08020FF83FF68FACA3209FAC0" box="[192,307,1377,1401]" pageId="9" pageNumber="8" value="2013-11-05">
|
||
<collectingDate id="EF8EE2288020FF83FF68FACA3209FAC0" box="[192,307,1377,1401]" pageId="9" pageNumber="8" value="2013-11-05">5.xi.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268158D68020FF83FE97FAC932EBFAC3" box="[319,465,1378,1402]" pageId="9" pageNumber="8">W. Ben Aba</collectorName>
|
||
leg.
|
||
<specimenCount id="9D72F6898020FF83FDA3FAC9310FFAC0" box="[523,565,1378,1402]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED65A5C58020FF83FDEDFAC931A2FAC3" box="[581,664,1378,1402]" pageId="9" pageNumber="8">SZCM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008020FF83FF58FA093622FA60" blockId="9.[192,1440,1442,1530]" pageId="9" pageNumber="8">
|
||
Furthermore, thanks to the Internet discussion site FEI (Italian Entomologists Forum), I learned of the following unpublished Algerian record:
|
||
<collectingRegion id="49B0F3E28020FF83FD5BFA69309AFA63" box="[755,928,1474,1498]" country="Algeria" name="Mila" pageId="9" pageNumber="8">Mila province</collectingRegion>
|
||
, Hamala,
|
||
<date id="FFCA1BC08020FF83FBB5FA6A3749FA60" box="[1053,1139,1473,1497]" pageId="9" pageNumber="8" value="1985-02">ii.1985</date>
|
||
,
|
||
<specimenCount id="9D72F6898020FF83FBD7FA693793FA60" box="[1151,1193,1474,1498]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FB4FFA69362BFA60" box="[1255,1297,1474,1498]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008020FF83FF58FA49379FFA40" blockId="9.[192,1440,1442,1530]" box="[240,1189,1506,1530]" pageId="9" pageNumber="8">
|
||
I have also examined
|
||
<specimenCount id="9D72F6898020FF83FE54FA49311CFA40" box="[508,550,1506,1530]" pageId="9" pageNumber="8" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898020FF83FDCBFA4931B7FA40" box="[611,653,1506,1530]" pageId="9" pageNumber="8" type="female">1 ♀</specimenCount>
|
||
without locality labels, both in the MNHN.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6E8B8020FF80FF68F98A31B4FE23" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="description">
|
||
<paragraph id="8BCB3D008020FF83FF68F98A30B2F943" blockId="9.[192,1440,1569,1882]" pageId="9" pageNumber="8">
|
||
<emphasis id="B900E1128020FF83FF68F98A3073F983" bold="true" box="[192,841,1569,1594]" pageId="9" pageNumber="8">Variability of cephalic and pronotal armament</emphasis>
|
||
. More developed males (
|
||
<figureCitation id="134F21858020FF83FBD2F98937FAF983" box="[1146,1216,1570,1594]" captionStart="Figures 8–11" captionStartId="23.[192,269,1549,1571]" captionTargetBox="[257,1363,305,1515]" captionTargetId="figure-71@23.[192,1439,192,1535]" captionTargetPageId="23" captionText="Figures 8–11. Copris pueli Molland de Boissy, 1905. 8) Habitus of male (TN-Jendouba prov., Ain Draham). 9) Habitus of female (TN-Jendouba prov., Ain Draham). 10) Head and pronotum of more developed male, posterolateral view (TN- Jendouba prov., Ain Draham). 11) Head and pronotum of less developed male, posterolateral view (syntype, DZ- Kabylie). Photographs by A. Degiovanni." figureDoi="http://doi.org/10.5281/zenodo.4645825" httpUri="https://zenodo.org/record/4645825/files/figure.png" pageId="9" pageNumber="8">Fig. 8</figureCitation>
|
||
) bear a very long, tapering, slightly backward-curved clypeal horn situated forward of the middle of the head and as long as the maximum head width. Anteromedian pronotal prominence is transversely truncated, with upper edge of the declivity rather narrow but a little dilated at its front margin which forms a straight carina interrupted in the middle by a longitudinal groove, 1/5 the width of the whole carina in dorsal view. On each side of the dorsal elevation there is a deep groove, the outer margin produced obliquely upward and forward as a pyramidal prominence (
|
||
<figureCitation id="134F21858020FF83FC8BF9493040F943" box="[803,890,1762,1786]" captionStart="Figures 8–11" captionStartId="23.[192,269,1549,1571]" captionTargetBox="[257,1363,305,1515]" captionTargetId="figure-71@23.[192,1439,192,1535]" captionTargetPageId="23" captionText="Figures 8–11. Copris pueli Molland de Boissy, 1905. 8) Habitus of male (TN-Jendouba prov., Ain Draham). 9) Habitus of female (TN-Jendouba prov., Ain Draham). 10) Head and pronotum of more developed male, posterolateral view (TN- Jendouba prov., Ain Draham). 11) Head and pronotum of less developed male, posterolateral view (syntype, DZ- Kabylie). Photographs by A. Degiovanni." figureDoi="http://doi.org/10.5281/zenodo.4645825" httpUri="https://zenodo.org/record/4645825/files/figure.png" pageId="9" pageNumber="8">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008020FF83FF58F8A932F5F8E3" blockId="9.[192,1440,1569,1882]" pageId="9" pageNumber="8">
|
||
The few less-developed males examined (
|
||
<figureCitation id="134F21858020FF83FD59F8A9307DF8A3" box="[753,839,1794,1818]" captionStart="Figures 8–11" captionStartId="23.[192,269,1549,1571]" captionTargetBox="[257,1363,305,1515]" captionTargetId="figure-71@23.[192,1439,192,1535]" captionTargetPageId="23" captionText="Figures 8–11. Copris pueli Molland de Boissy, 1905. 8) Habitus of male (TN-Jendouba prov., Ain Draham). 9) Habitus of female (TN-Jendouba prov., Ain Draham). 10) Head and pronotum of more developed male, posterolateral view (TN- Jendouba prov., Ain Draham). 11) Head and pronotum of less developed male, posterolateral view (syntype, DZ- Kabylie). Photographs by A. Degiovanni." figureDoi="http://doi.org/10.5281/zenodo.4645825" httpUri="https://zenodo.org/record/4645825/files/figure.png" pageId="9" pageNumber="8">Fig. 11</figureCitation>
|
||
) have a shorter horn, as long as the interocular distance but never female-like, and pronotum with the same structures but less pronounced, although always clearly visible.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FF693274FE83" blockId="10.[192,1440,194,314]" pageId="10" pageNumber="9">
|
||
Females (
|
||
<figureCitation id="134F21858023FF80FECFFF693296FF63" box="[359,428,194,218]" captionStart="Figures 8–11" captionStartId="23.[192,269,1549,1571]" captionTargetBox="[257,1363,305,1515]" captionTargetId="figure-71@23.[192,1439,192,1535]" captionTargetPageId="23" captionText="Figures 8–11. Copris pueli Molland de Boissy, 1905. 8) Habitus of male (TN-Jendouba prov., Ain Draham). 9) Habitus of female (TN-Jendouba prov., Ain Draham). 10) Head and pronotum of more developed male, posterolateral view (TN- Jendouba prov., Ain Draham). 11) Head and pronotum of less developed male, posterolateral view (syntype, DZ- Kabylie). Photographs by A. Degiovanni." figureDoi="http://doi.org/10.5281/zenodo.4645825" httpUri="https://zenodo.org/record/4645825/files/figure.png" pageId="10" pageNumber="9">Fig. 9</figureCitation>
|
||
) have a more or less elevated, transverse clypeal carina truncate at its summit and feebly bicuspid, and a slight, straight, transverse pronotal carina just behind the front margin, with two very low and vague tumescences on either side. In less developed females these tumescences are obliterated.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF68FECA31B4FE23" blockId="10.[192,1440,353,410]" pageId="10" pageNumber="9">
|
||
<emphasis id="B900E1128023FF80FF68FECA327FFEC0" bold="true" box="[192,325,353,377]" pageId="10" pageNumber="9">Aedeagus</emphasis>
|
||
. Parameres in lateral view slender (
|
||
<figureCitation id="134F21858023FF80FCAEFEC93067FEC3" box="[774,861,354,378]" captionStart="Figures 12–13" captionStartId="24.[192,269,1205,1227]" captionTargetBox="[343,1311,336,1175]" captionTargetId="figure-41@24.[192,1439,192,1193]" captionTargetPageId="24" captionText="Figures 12–13. Copris pueli Molland de Boissy, 1905. 12) Parameres, dorsal view (DZ-Mt. Edough). 13) Parameres, lateral view (DZ-Mt. Edough). Photographs by A. Degiovanni." figureDoi="http://doi.org/10.5281/zenodo.4645829" httpUri="https://zenodo.org/record/4645829/files/figure.png" pageId="10" pageNumber="9">Fig. 13</figureCitation>
|
||
). Apex blunt, truncate in dorsal view, and ventrally with some short hairs (
|
||
<figureCitation id="134F21858023FF80FD80FE2931BAFE23" box="[552,640,386,410]" captionStart="Figures 12–13" captionStartId="24.[192,269,1205,1227]" captionTargetBox="[343,1311,336,1175]" captionTargetId="figure-41@24.[192,1439,192,1193]" captionTargetPageId="24" captionText="Figures 12–13. Copris pueli Molland de Boissy, 1905. 12) Parameres, dorsal view (DZ-Mt. Edough). 13) Parameres, lateral view (DZ-Mt. Edough). Photographs by A. Degiovanni." figureDoi="http://doi.org/10.5281/zenodo.4645829" httpUri="https://zenodo.org/record/4645829/files/figure.png" pageId="10" pageNumber="9">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6E8B8023FF81FF68FE693100FE43" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="discussion">
|
||
<paragraph id="8BCB3D008023FF80FF68FE6937EDFDE3" blockId="10.[192,1440,449,1626]" pageId="10" pageNumber="9">
|
||
<emphasis id="B900E1128023FF80FF68FE693206FE63" bold="true" box="[192,316,450,474]" pageId="10" pageNumber="9">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFE540F18023FF80FEE1FE6931A9FE60" author="Mollandin de Boissy, R." box="[329,659,449,474]" pageId="10" pageNumber="9" pagination="110 - 113" refId="ref16463" refString="Mollandin de Boissy, R. 1905. Description d'un Copris Geoffroy nouveau d'Algerie (Col.). Bulletin de la Societe entomologique de France: 110 - 113." type="book chapter" year="1905">Mollandin de Boissy (1905)</bibRefCitation>
|
||
in the original description of
|
||
<taxonomicName id="4C7446838023FF80FC5EFE69376EFE63" authorityName="Mollandin de Boissy" authorityYear="1905" box="[1014,1108,450,474]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="pueli">
|
||
<emphasis id="B900E1128023FF80FC5EFE69376EFE63" box="[1014,1108,450,474]" italics="true" pageId="10" pageNumber="9">C. pueli</emphasis>
|
||
</taxonomicName>
|
||
stated: “Les onze individus (
|
||
<specimenCount id="9D72F6898023FF80FF61FE493239FE40" box="[201,259,482,506]" pageId="10" pageNumber="9" type="male">10 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D72F6898023FF80FEBAFE493207FE40" box="[274,317,482,506]" pageId="10" pageNumber="9" type="female">1 ♀</specimenCount>
|
||
, Collections Puel, J. Clermont, Delfieu et de Boissy) que j’ai eus sous les yeux proviennent tous de Bou-Berak (Kabylie), où ils ont été pris par M. Louis Puel (de Béziers)” [The eleven specimens (
|
||
<specimenCount id="9D72F6898023FF80FF61FD893239FD80" box="[201,259,546,570]" pageId="10" pageNumber="9" type="male">10 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D72F6898023FF80FEBBFD893204FD80" box="[275,318,546,570]" pageId="10" pageNumber="9" type="female">1 ♀</specimenCount>
|
||
, Puel, J. Clermont, Delfieu et de Boissy collections) that I have had under my eyes are all from Bou-Berak (Kabylie), where they were collected by Mr. Louis Puel (de Béziers)].
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FDC9370BFCA3" blockId="10.[192,1440,449,1626]" pageId="10" pageNumber="9">
|
||
I have examined
|
||
<specimenCount id="9D72F6898023FF80FE6FFDC93156FDC3" box="[455,620,610,634]" pageId="10" pageNumber="9" type="generic">18 specimens</specimenCount>
|
||
with a label “Bou Berak / Kabjlie L. Puel”,
|
||
<quantity id="4C8C90E58023FF80FB26FDC937F7FDC3" box="[1166,1229,610,634]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="10" pageNumber="9" unit="in" value="17.0">17 in</quantity>
|
||
MNHN and
|
||
<quantity id="4C8C90E58023FF80FAD8FDC9369AFDC3" box="[1392,1440,610,634]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="9" unit="in" value="1.0">1 in</quantity>
|
||
MHNG. It is not possible, however, to deem all these specimens as included in the
|
||
<typeStatus id="54CF83A28023FF80FB06FD2937D9FD23" box="[1198,1251,642,666]" pageId="10" pageNumber="9">type</typeStatus>
|
||
series, because unfortunately their number and sex (
|
||
<specimenCount id="9D72F6898023FF80FD2DFD0931EBFD00" box="[645,721,674,698]" pageId="10" pageNumber="9" type="male">13 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898023FF80FCA3FD0A3072FD00" box="[779,840,673,697]" pageId="10" pageNumber="9" type="female">5 ♀♀</specimenCount>
|
||
) do not coincide with number and sex (
|
||
<specimenCount id="9D72F6898023FF80FA88FD093656FD00" box="[1312,1388,674,698]" pageId="10" pageNumber="9" type="male">10 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898023FF80FF68FD6933D2FD60" box="[192,232,706,730]" pageId="10" pageNumber="9" type="female">1 ♀</specimenCount>
|
||
) of the specimens used by Mollandin de Boissy to describe the species. This notwithstanding, among these
|
||
<specimenCount id="9D72F6898023FF80FEA1FD493294FD43" box="[265,430,738,762]" pageId="10" pageNumber="9" type="generic">18 specimens</specimenCount>
|
||
I noticed four critical specimens belonging to the Puel collection and preserved in the MNHN, that according to the labels could be part of the
|
||
<typeStatus id="54CF83A28023FF80FC0DFCA930E0FCA3" box="[933,986,770,794]" pageId="10" pageNumber="9">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FC8930BFFC83" blockId="10.[192,1440,449,1626]" box="[240,901,802,826]" pageId="10" pageNumber="9">The first specimen, a male, bears the following labels:</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FCE937E4FCE3" blockId="10.[192,1440,449,1626]" box="[240,1246,834,858]" pageId="10" pageNumber="9">1) White, with black square frame, printed in black “Bou Berak / Kabilie L. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FCC931F3FCC3" blockId="10.[192,1440,449,1626]" box="[240,713,866,890]" pageId="10" pageNumber="9">2) White, printed in black “Coll. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FC29369AFC23" blockId="10.[192,1440,449,1626]" box="[240,1440,898,922]" pageId="10" pageNumber="9">3) Red, with black double square frame, printed in black “Co-Typus” [letters “Co” are handwritten];</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FC0937F7FC03" blockId="10.[192,1440,449,1626]" box="[240,1229,929,954]" pageId="10" pageNumber="9">4) White, printed in black “MUSÉUM PARIS / 1936 / COLL. A. BOUCOMONT”.</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FC49309FFC43" blockId="10.[192,1440,449,1626]" box="[240,933,994,1018]" pageId="10" pageNumber="9">The second specimen, a male, bears the following labels:</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FBA937E4FBA3" blockId="10.[192,1440,449,1626]" box="[240,1246,1026,1050]" pageId="10" pageNumber="9">1) White, with black square frame, printed in black “Bou Berak / Kabilie L. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FB8931EAFB83" blockId="10.[192,1440,449,1626]" box="[240,720,1058,1082]" pageId="10" pageNumber="9">2) White, handwritten in black “Pueli”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FBE931F3FBE3" blockId="10.[192,1440,449,1626]" box="[240,713,1090,1114]" pageId="10" pageNumber="9">3) White, printed in black “Coll. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FBC9372CFBC3" blockId="10.[192,1440,449,1626]" box="[240,1046,1122,1146]" pageId="10" pageNumber="9">
|
||
4) Red, with black double square frame, printed in black “
|
||
<typeStatus id="54CF83A28023FF80FC10FBC9373CFBC3" box="[952,1030,1122,1146]" pageId="10" pageNumber="9">Typus</typeStatus>
|
||
”;
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FB2A37F7FB23" blockId="10.[192,1440,449,1626]" box="[240,1229,1153,1178]" pageId="10" pageNumber="9">5) White, printed in black “MUSÉUM PARIS / 1936 / COLL. A. BOUCOMONT”.</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FB6930AAFB63" blockId="10.[192,1440,449,1626]" box="[240,912,1218,1242]" pageId="10" pageNumber="9">The third specimen, a male, bears the following labels:</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FB4937E4FB43" blockId="10.[192,1440,449,1626]" box="[240,1246,1250,1274]" pageId="10" pageNumber="9">1) White, with black square frame, printed in black “Bou Berak / Kabilie L. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FAA9372CFAA3" blockId="10.[192,1440,449,1626]" box="[240,1046,1282,1306]" pageId="10" pageNumber="9">
|
||
2) Red, with black double square frame, printed in black “
|
||
<typeStatus id="54CF83A28023FF80FC10FAA9373CFAA3" box="[952,1030,1282,1306]" pageId="10" pageNumber="9">Typus</typeStatus>
|
||
”;
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FA8930FAFA83" blockId="10.[192,1440,449,1626]" box="[240,960,1314,1338]" pageId="10" pageNumber="9">
|
||
3) White, handwritten in black “
|
||
<taxonomicName id="4C7446838023FF80FDD6FA8931F4FA83" box="[638,718,1314,1338]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">Copris</taxonomicName>
|
||
/ Pueli / de Boissy”;
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FAE931F3FAE3" blockId="10.[192,1440,449,1626]" box="[240,713,1346,1370]" pageId="10" pageNumber="9">4) White, printed in black “Coll. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FACA37F7FAC3" blockId="10.[192,1440,449,1626]" box="[240,1229,1377,1402]" pageId="10" pageNumber="9">5) White, printed in black “MUSÉUM PARIS / 1936 / COLL. A. BOUCOMONT”.</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FA09308FFA03" blockId="10.[192,1440,449,1626]" box="[240,949,1442,1466]" pageId="10" pageNumber="9">The fourth specimen, a female, bears the following labels:</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FA6937E4FA63" blockId="10.[192,1440,449,1626]" box="[240,1246,1474,1498]" pageId="10" pageNumber="9">1) White, with black square frame, printed in black “Bou Berak / Kabilie L. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58FA49305AFA43" blockId="10.[192,1440,449,1626]" box="[240,864,1506,1530]" pageId="10" pageNumber="9">
|
||
2) White, handwritten in black “
|
||
<taxonomicName id="4C7446838023FF80FDD6FA4931F4FA43" box="[638,718,1506,1530]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">Copris</taxonomicName>
|
||
/ Pueli /
|
||
<specimenCount id="9D72F6898023FF80FC93FA493075FA40" box="[827,847,1506,1529]" pageId="10" pageNumber="9" type="female">♀</specimenCount>
|
||
”;
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58F9A931F3F9A3" blockId="10.[192,1440,449,1626]" box="[240,713,1538,1562]" pageId="10" pageNumber="9">3) White, printed in black “Coll. Puel”;</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58F989372CF983" blockId="10.[192,1440,449,1626]" box="[240,1046,1570,1594]" pageId="10" pageNumber="9">
|
||
4) Red, with black double square frame, printed in black “
|
||
<typeStatus id="54CF83A28023FF80FC10F989373CF983" box="[952,1030,1570,1594]" pageId="10" pageNumber="9">Typus</typeStatus>
|
||
”;
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58F9EA37F7F9E3" blockId="10.[192,1440,449,1626]" box="[240,1229,1601,1626]" pageId="10" pageNumber="9">5) White, printed in black “MUSÉUM PARIS / 1936 / COLL. A. BOUCOMONT”.</paragraph>
|
||
<paragraph id="8BCB3D008023FF80FF58F929327DF883" blockId="10.[192,1440,1666,1914]" pageId="10" pageNumber="9">
|
||
On the basis of these labels, I believe that these four specimens can collectively constitute the name-bearing type of the taxon, therefore they can be considered
|
||
<typeStatus id="54CF83A28023FF80FC47F9093764F903" box="[1007,1118,1698,1722]" pageId="10" pageNumber="9" type="syntype">syntypes</typeStatus>
|
||
, part of the material seen by Mollandin de Boissy. I added to every specimen a red label printed in black: “
|
||
<typeStatus id="54CF83A28023FF80FB30F96937C6F963" box="[1176,1276,1730,1754]" pageId="10" pageNumber="9" type="syntype">Syntype</typeStatus>
|
||
/
|
||
<taxonomicName id="4C7446838023FF80FAA3F969369AF963" authorityName="Mollandin de Boissy" authorityYear="1905" box="[1291,1440,1730,1754]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="pueli">
|
||
<emphasis id="B900E1128023FF80FAA3F969369AF963" box="[1291,1440,1730,1754]" italics="true" pageId="10" pageNumber="9">Copris pueli</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EFE540F18023FF80FF67F9493129F940" author="Mollandin de Boissy, R." box="[207,531,1761,1786]" pageId="10" pageNumber="9" pagination="110 - 113" refId="ref16463" refString="Mollandin de Boissy, R. 1905. Description d'un Copris Geoffroy nouveau d'Algerie (Col.). Bulletin de la Societe entomologique de France: 110 - 113." type="book chapter" year="1905">Mollandin de Boissy, 1905</bibRefCitation>
|
||
/ S. Ziani vidit, 2017”. In my opinion there is no need to designate a
|
||
<typeStatus id="54CF83A28023FF80FAF6F94933C2F8A3" pageId="10" pageNumber="9" type="lectotype">lectotype</typeStatus>
|
||
, the taxonomic status of the species is well known and the nomenclature does not run the risk of instability.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008023FF81FF58F8E930F2FE03" blockId="10.[192,1440,1666,1914]" lastBlockId="11.[192,1440,194,506]" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
|
||
In the collections of the MNHN there are also
|
||
<specimenCount id="9D72F6898023FF80FC9AF8EA30E0F8E3" box="[818,986,1857,1882]" pageId="10" pageNumber="9" type="generic">52 specimens</specimenCount>
|
||
,
|
||
<specimenCount id="9D72F6898023FF80FC4EF8EA370EF8E0" box="[998,1076,1857,1881]" pageId="10" pageNumber="9" type="male">51 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D72F6898023FF80FBDCF8E937A5F8E0" box="[1140,1183,1858,1882]" pageId="10" pageNumber="9" type="female">1 ♀</specimenCount>
|
||
, of
|
||
<taxonomicName id="4C7446838023FF80FB65F8E93617F8E3" authorityName="Mollandin de Boissy" authorityYear="1905" box="[1229,1325,1858,1882]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="pueli">
|
||
<emphasis id="B900E1128023FF80FB65F8E93617F8E3" box="[1229,1325,1858,1882]" italics="true" pageId="10" pageNumber="9">C. pueli</emphasis>
|
||
</taxonomicName>
|
||
collected in
|
||
<collectingCountry id="F3637D908023FF80FF4AF8C93205F8C3" box="[226,319,1890,1914]" name="Algeria" pageId="10" pageNumber="9">Algeria</collectingCountry>
|
||
, at Edough Mountain (Djebel Edough, Province of
|
||
<collectingRegion id="49B0F3E28023FF80FC13F8C93721F8C3" box="[955,1051,1890,1914]" country="Algeria" name="Annaba" pageId="10" pageNumber="9">Annaba</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B0F3E28023FF80FBF4F8C9378DF8C3" box="[1116,1207,1890,1914]" country="Algeria" name="Skikda" pageId="10" pageNumber="9">Skikda</collectingRegion>
|
||
). One male is very different from all the others, with the anterolateral pronotal teeth very weak, the upper ridge of the anteromedian declivity that bears two small and blunt prominences separated at middle by a hollow equal to their diameter, and the cephalic horn slightly dilated and feebly but clearly bifurcate at the extremity, all characters shared by males of
|
||
<emphasis id="B900E1128022FF81FD44FE89304BFE83" box="[748,881,290,314]" italics="true" pageId="11" pageNumber="10">
|
||
<taxonomicName id="4C7446838022FF81FD44FE893057FE83" authorityName="Reitter" authorityYear="1892" box="[748,877,290,314]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="felschei">C. felschei</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
On the other hand, the specimen has elytral striae very crenulated and parameres subquadrate and ciliate at the top, as in
|
||
<taxonomicName id="4C7446838022FF81FB26FEE937D7FEE3" authorityName="Mollandin de Boissy" authorityYear="1905" box="[1166,1261,322,346]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="pueli">
|
||
<emphasis id="B900E1128022FF81FB26FEE937D7FEE3" box="[1166,1261,322,346]" italics="true" pageId="11" pageNumber="10">C. pueli</emphasis>
|
||
</taxonomicName>
|
||
. However, the abdomen was deliberately glued to the prothorax and the pubescence on the inner base of the pronotum and elytra is clearly of two different shades of dark yellow, which is why it is reasonable to suspect that the prothorax and the abdomen belong to two different species.
|
||
</paragraph>
|
||
<paragraph id="8BCB3D008022FF81FF68FE693100FE43" blockId="11.[192,1440,194,506]" pageId="11" pageNumber="10">
|
||
<bibRefCitation id="EFE540F18022FF81FF68FE69329DFE63" author="Schatzmayr, A." box="[192,423,450,474]" pageId="11" pageNumber="10" pagination="110 - 114" refId="ref17351" refString="Schatzmayr, A. 1930. Risultati scientifici delle spedizioni entomologiche di S. A. S. il Principe Alessandro della Torre e Tasso nell'Africa settentrionale e in Sicilia. Bollettino della Societa Entomologica Italiana 62: 110 - 114." type="journal article" year="1930">Schatzmayr (1930)</bibRefCitation>
|
||
assumed that this species was present in the mountains of
|
||
<collectingRegion id="49B0F3E28022FF81FB28FE6937FDFE63" box="[1152,1223,450,474]" country="Italy" name="Sicilia" pageId="11" pageNumber="10">Sicily</collectingRegion>
|
||
, but this assumption has never been confirmed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |