treatments-xml/data/03/DD/80/03DD801AE541FFB9FF43D234FCB3FD75.xml
2024-06-21 12:22:17 +02:00

350 lines
45 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="500DEE35DFC8F66CBACA78EE2E695FF5" ID-DOI="10.5281/zenodo.190536" ID-GBIF-Dataset="d7ff8abd-bb96-4ea4-b61a-66aa70982a9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="190536" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461114760079" checkinUser="plazi" docAuthor="Pesic, Vladimir &amp; Smit, Harry" docDate="2009" docId="03DD801AE541FFB9FF43D234FCB3FD75" docLanguage="en" docName="zt02240p030.pdf" docOrigin="Zootaxa 2240" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atractides (Atractides) cognatus K. Viets 1935" docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="FFE4F862E555FFA1FFD4D17BFF87FFBA" masterDocTitle="New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from Thailand, Malaysia and Sulawesi (Indonesia), with the description of four new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="21" updateTime="1698594892522" updateUser="plazi">
<mods:mods id="22060783F658C72E9F9989B1CFA5A35B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="37A6FBD7E5EE061F13391C1F1962C2C0">
<mods:title id="30AEE96B904A9E49964B233288E87FB2">New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from Thailand, Malaysia and Sulawesi (Indonesia), with the description of four new species</mods:title>
</mods:titleInfo>
<mods:name id="D9CF0AF780E7E760E75D9821FD5BAC94" type="personal">
<mods:role id="7EBCA7465A5A09D2F631D5C213CACF48">
<mods:roleTerm id="C1E43ABD505C5813C4CEF7070C62D4EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFEC36A8D9B620DF3527308C9D9EA36D">Pesic, Vladimir</mods:namePart>
</mods:name>
<mods:name id="20638C7E3297825411605B7F1B815E2E" type="personal">
<mods:role id="B8541D711CFB122F9407AE6A425302C5">
<mods:roleTerm id="8E9C10F68A2CC37D54B6EE89C5539BB4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E535071E7D8E4E95C4FE3AD47B59A6BC">Smit, Harry</mods:namePart>
</mods:name>
<mods:typeOfResource id="65DD15E9863884305892EFD6A47274EB">text</mods:typeOfResource>
<mods:relatedItem id="33BD82F3A20760E30B0F09F24CD23ABD" type="host">
<mods:titleInfo id="6F0A351D3A376A6834D4D9CA42B93ED5">
<mods:title id="6AB7EC102EABDC5F87CFD4CB6E21162D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EFAE65C3926B6E8D9C2FCA249A5724C4">
<mods:date id="2C39BC65D1030AB5AC55CAA3EAC8AA27">2009</mods:date>
<mods:detail id="597E714271D69C8BFE8FA04545B9795A" type="volume">
<mods:number id="B441CB94937D42730257C871F89B1C01">2240</mods:number>
</mods:detail>
<mods:extent id="214F6C5E86CD89EF56388221BB4688CF" unit="page">
<mods:start id="B9433AA8A6280F06C287A7840046C593">1</mods:start>
<mods:end id="828415D0A31ACA02B496B8CF31F3C578">30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D3B30DBE0982933DED17B44615269440">journal article</mods:classification>
<mods:identifier id="33EA3B36A4B4E8F3E3C015BF0BA16796" type="DOI">10.5281/zenodo.190536</mods:identifier>
<mods:identifier id="117AB48AC85ACF49D97D9E20597F7FC4" type="GBIF-Dataset">d7ff8abd-bb96-4ea4-b61a-66aa70982a9b</mods:identifier>
<mods:identifier id="530E856F3C09B6F8B1B15196E25CD961" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9A6502BFADC4B34F7260718E0BF491C8" type="Zenodo-Dep">190536</mods:identifier>
</mods:mods>
<treatment id="03DD801AE541FFB9FF43D234FCB3FD75" ID-DOI="http://doi.org/10.5281/zenodo.6216160" ID-GBIF-Taxon="119624646" ID-Zenodo-Dep="6216160" LSID="urn:lsid:plazi:treatment:03DD801AE541FFB9FF43D234FCB3FD75" httpUri="http://treatment.plazi.org/id/03DD801AE541FFB9FF43D234FCB3FD75" lastPageId="24" lastPageNumber="25" pageId="20" pageNumber="21">
<subSubSection id="C36E6287E541FFB5FF43D234FD8EFC36" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BCB310CE541FFB5FF43D234FCDEFCD3" blockId="20.[151,857,847,908]" box="[151,857,847,873]" pageId="20" pageNumber="21">
<heading id="D0838660E541FFB5FF43D234FCDEFCD3" bold="true" box="[151,857,847,873]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B900ED1EE541FFB5FF43D234FCDEFCD3" bold="true" box="[151,857,847,873]" pageId="20" pageNumber="21">
<taxonomicName id="4C744A8FE541FFB5FF43D234FD65FCD3" authority="K. Viets, 1935" authorityName="K. Viets" authorityYear="1935" box="[151,738,847,873]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cognatus" subGenus="Atractides">
<emphasis id="B900ED1EE541FFB5FF43D234FE96FCD3" bold="true" box="[151,273,847,873]" italics="true" pageId="20" pageNumber="21">Atractides</emphasis>
(
<emphasis id="B900ED1EE541FFB5FEF6D234FE1BFCD3" bold="true" box="[290,412,847,873]" italics="true" pageId="20" pageNumber="21">Atractides</emphasis>
)
<emphasis id="B900ED1EE541FFB5FE79D234FD9DFCD3" bold="true" box="[429,538,847,873]" italics="true" pageId="20" pageNumber="21">cognatus</emphasis>
(
<bibRefCitation id="EFE54CFDE541FFB5FDFED234FD5DFCD3" author="Viets" box="[554,730,847,873]" pageId="20" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K. Viets, 1935</bibRefCitation>
)
</taxonomicName>
nov. stat.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB310CE541FFB5FF43D209FD8EFC36" blockId="20.[151,857,847,908]" box="[151,521,882,908]" pageId="20" pageNumber="21">
(
<figureCitation id="134F2D89E541FFB5FF4BD209FEBDFC36" box="[159,314,882,908]" captionStart="FIGURE 15 A D" captionStartId="21.[151,255,1586,1610]" captionTargetBox="[167,1417,224,1540]" captionTargetId="figure@21.[151,1436,194,1562]" captionTargetPageId="21" captionText="FIGURE 15 A D. Atractides cognatus K. Viets, 1935, male, stream near Ban Khoom, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190551/files/figure.png" pageId="20" pageNumber="21">Figs. 15AD</figureCitation>
,
<figureCitation id="134F2D89E541FFB5FE93D209FE19FC36" box="[327,414,882,908]" captionStart="FIGURE 16 A C" captionStartId="22.[151,255,1784,1808]" captionTargetBox="[161,1412,629,1746]" captionTargetId="figure@22.[151,1436,619,1760]" captionTargetPageId="22" captionText="FIGURE 16 A C. Atractides cognatus K. Viets, 1935, female (A, C - specimen from stream near Ban Khoom, Thailand; B- specimen from tributary of Tahan River near Lata Berkoh, Malaysia): A = I-Leg- 5 and - 6; B = genital field; C = palp, lateral view. Scale bar = 100 μm." httpUri="https://zenodo.org/record/190552/files/figure.png" pageId="20" pageNumber="21">16AC</figureCitation>
,
<figureCitation id="134F2D89E541FFB5FE7ED209FD85FC36" box="[426,514,882,908]" captionStart="FIGURE 17 A D" captionStartId="23.[151,257,1895,1919]" captionTargetBox="[151,1436,204,1860]" captionTargetId="figure@23.[151,1436,194,1871]" captionTargetPageId="23" captionText="FIGURE 17 A D. Atractides cognatus K. Viets, 1935, female, stream 18 km west of Molibagu, Sulawesi: A = coxal field; B = palp, medial view; C = genital field; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190553/files/figure.png" pageId="20" pageNumber="21">17AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E6287E541FFB5FF43D2BBFC39FC4D" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8BCB310CE541FFB5FF43D2BBFD0FFC6D" blockId="20.[151,958,960,1015]" box="[151,648,960,983]" pageId="20" pageNumber="21">
<taxonomicName id="4C744A8FE541FFB5FF43D2BBFD0FFC6D" authority="K. Viets, 1935" authorityName="K. Viets" authorityYear="1935" box="[151,648,960,983]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="spatiosus" subSpecies="cognatus">
<emphasis id="B900ED1EE541FFB5FF43D2BBFE51FC6D" box="[151,470,960,983]" italics="true" pageId="20" pageNumber="21">Atractides spatiosus cognatus</emphasis>
(
<bibRefCitation id="EFE54CFDE541FFB5FE30D2BBFD07FC6D" author="Viets" box="[484,640,960,983]" pageId="20" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K. Viets, 1935</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BCB310CE541FFB5FF43D29BFC39FC4D" blockId="20.[151,958,960,1015]" box="[151,958,992,1015]" pageId="20" pageNumber="21">
[syn. to
<taxonomicName id="4C744A8FE541FFB5FF39D29BFDA4FC4D" authority="K. Viets, 1935" authorityName="K. Viets" authorityYear="1935" box="[237,547,992,1015]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE541FFB5FF39D29BFEE9FC4D" box="[237,366,992,1015]" italics="true" pageId="20" pageNumber="21">A. spatiosus</emphasis>
(
<bibRefCitation id="EFE54CFDE541FFB5FEA9D29BFD9CFC4D" author="Viets" box="[381,539,992,1015]" pageId="20" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K. Viets, 1935</bibRefCitation>
)
</taxonomicName>
;
<bibRefCitation id="EFE54CFDE541FFB5FDFAD29BFD57FC4D" author="Lundblad" box="[558,720,992,1015]" pageId="20" pageNumber="30" refString="Lundblad, O. (1971) Weitere Beitrage zur Kenntnis der Fliesswassermilben Javas. Arkiv for Zoologi, 23, 293 - 359." type="journal article" year="1971">Lundblad 1971</bibRefCitation>
rejected synonymy]
</paragraph>
</subSubSection>
<subSubSection id="C36E6287E541FFB7FF43D55DFA17FD8E" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BCB310CE541FFB5FF43D55DFCC5FAF5" blockId="20.[151,1437,1062,1939]" pageId="20" pageNumber="21">
<emphasis id="B900ED1EE541FFB5FF43D55DFE8CFBFA" bold="true" box="[151,267,1062,1088]" pageId="20" pageNumber="21">Material.</emphasis>
<collectingCountry id="F363719CE541FFB5FEC6D55DFE29FBFA" box="[274,430,1062,1088]" name="Thailand" pageId="20" pageNumber="21">THAILAND</collectingCountry>
: unnamed creek crossing track from watchtower, Khao Yai NP
<date id="FFCA17CCE541FFB5FB45D55DFA97FBFA" box="[1169,1296,1062,1088]" pageId="20" pageNumber="21" value="2007-11-11">11.xi.2007</date>
, 14°
<geoCoordinate id="EE4057CBE541FFB5FA9FD55DFF2BFBDD" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="55" value="27.533">27.533 N</geoCoordinate>
101°
<geoCoordinate id="EE4057CBE541FFB5FF20D536FEE6FBDD" box="[244,353,1101,1127]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="55" value="20.386">20.386 E</geoCoordinate>
, leg. Smit, 2/6/0 (1/1/0 mounted); fast flowing stream crossing road to Den Ya Kat Station, alt.
<quantity id="4C8C9CE9E541FFB5FF16D50FFE89FB34" box="[194,270,1140,1166]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="20" pageNumber="21" unit="m" value="410.0">410 m</quantity>
, Chiang Dao NP,
<date id="FFCA17CCE541FFB5FE08D50FFDDBFB34" box="[476,604,1140,1166]" pageId="20" pageNumber="21" value="2007-11-23">23.xi.2007</date>
, 19 °
<geoCoordinate id="EE4057CBE541FFB5FD4AD50FFC8DFB34" box="[670,778,1140,1166]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="55" value="19.735">19.735 N</geoCoordinate>
98°
<geoCoordinate id="EE4057CBE541FFB5FC94D50FFC2CFB34" box="[832,939,1140,1166]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="55" value="56.201">56.201 E</geoCoordinate>
, leg. Smit, 0/1/0; stream near Ban Khoom, alt.
<quantity id="4C8C9CE9E541FFB5FF1CD5E1FE9CFB0E" box="[200,283,1178,1204]" metricMagnitude="2" metricUnit="m" metricValue="7.56" pageId="20" pageNumber="21" unit="m" value="756.0">756 m</quantity>
, Doi Ang Kang,
<date id="FFCA17CCE541FFB5FE20D5E1FDF8FB0E" box="[500,639,1178,1204]" pageId="20" pageNumber="21" value="2007-11-20">20.xi.2007</date>
, 19°
<geoCoordinate id="EE4057CBE541FFB5FD16D5E1FCB0FB0E" box="[706,823,1178,1204]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="55" value="56.905">56.905 N</geoCoordinate>
99°
<geoCoordinate id="EE4057CBE541FFB5FCA1D5E1FC6CFB0E" box="[885,1003,1178,1204]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="0.065633334">0 3.938 E</geoCoordinate>
, leg. Smit, 3/8/0 (1/1/0 mounted).
<collectingCountry id="F363719CE541FFB5FF43D5BAFEB3FB61" box="[151,308,1217,1243]" name="Malaysia" pageId="20" pageNumber="21">MALAYSIA</collectingCountry>
: small tributary of Tahan River, Taman Negara NP,
<date id="FFCA17CCE541FFB5FC48D5BAFB90FB61" box="[924,1047,1217,1243]" pageId="20" pageNumber="21" value="2008-02-07">07.ii.2008</date>
, 4°
<geoCoordinate id="EE4057CBE541FFB5FB91D5BAFB33FB61" box="[1093,1204,1217,1243]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="55" value="23.977">23.977 N</geoCoordinate>
102°
<geoCoordinate id="EE4057CBE541FFB5FB28D5BAFAEDFB61" box="[1276,1386,1217,1243]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="55" value="24.162">24.162 E</geoCoordinate>
, alt.
<quantity id="4C8C9CE9E541FFB5FF43D593FF51FAB8" box="[151,214,1256,1282]" metricMagnitude="1" metricUnit="m" metricValue="9.4" pageId="20" pageNumber="21" unit="m" value="94.0">94 m</quantity>
, leg. Smit, 0/1/0; tributary of Tahan River near Lata Berkoh, Taman Negara NP,
<date id="FFCA17CCE541FFB5FB47D593FB79FAB8" box="[1171,1278,1256,1282]" pageId="20" pageNumber="21" value="2009-02-09">9.ii.2009</date>
, 4°
<geoCoordinate id="EE4057CBE541FFB5FAFAD593FA1BFAB8" box="[1326,1436,1256,1282]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="55" value="26.635">26.635 N</geoCoordinate>
102°
<geoCoordinate id="EE4057CBE541FFB5FF02D475FEC5FA92" box="[214,322,1294,1320]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="55" value="23.057">23.057 E</geoCoordinate>
, alt.
<quantity id="4C8C9CE9E541FFB5FEAFD475FE3DFA92" box="[379,442,1294,1320]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="20" pageNumber="21" unit="m" value="82.0">82 m</quantity>
, leg. Smit, 0/1/0 (0/1/0 mounted).
<collectingCountry id="F363719CE541FFB5FC9AD475FC76FA92" box="[846,1009,1294,1320]" name="Indonesia" pageId="20" pageNumber="21">INDONESIA</collectingCountry>
, Sulawesi Utara: stream
<quantity id="4C8C9CE9E541FFB5FAC0D475FAD8FA92" box="[1300,1375,1294,1320]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="21" unit="km" value="18.0">18 km</quantity>
west of Molibagu,
<date id="FFCA17CCE541FFB5FEE3D44EFE32FAF5" box="[311,437,1333,1359]" pageId="20" pageNumber="21" value="1991-04-16">16.iv.1991</date>
, leg. Smit, 0/2/0 (0/1/0 mounted).
</paragraph>
<paragraph id="8BCB310CE541FFB5FF12D420FE13F98D" blockId="20.[151,1437,1062,1939]" pageId="20" pageNumber="21">
<emphasis id="B900ED1EE541FFB5FF12D420FEE0FACF" bold="true" box="[198,359,1371,1397]" pageId="20" pageNumber="21">Morphology.</emphasis>
General features. Integument dorsally striated. Muscle attachment plates unsclerotized. Genital field: Ac in obtuse triangle; excretory pore smooth; Vgl-1 not fused (but occasionally close or fused in males with Vgl-2). Palp with a weak sexual dimorphism; ventral margin of P-2 straight, sword seta of P-4 near distal hair. I-Leg: setae S-1 and -2 distanced, S-1 near the ventral seta, long, pointed, S-2 pointed, strongly enlarged; I-L-6 very long and slender, thickened basally, with parallel dorsal and ventral margins from the centre to the claw pit.
</paragraph>
<paragraph id="8BCB310CE541FFB5FF11D73FFF64F8A5" blockId="20.[151,1437,1062,1939]" pageId="20" pageNumber="21">
Male (from stream near Ban Khoom,
<collectingCountry id="F363719CE541FFB5FD5AD73FFD7BF9E4" box="[654,764,1604,1630]" name="Thailand" pageId="20" pageNumber="21">Thailand</collectingCountry>
; in parentheses specimen from unnamed creek crossing track from watchtower,
<collectingCountry id="F363719CE541FFB5FE65D711FD98F93E" box="[433,543,1642,1668]" name="Thailand" pageId="20" pageNumber="21">Thailand</collectingCountry>
): Idiosoma L/W 403/284; Coxal field (
<figureCitation id="134F2D89E541FFB5FC3DD711FBDFF93E" box="[1001,1112,1642,1668]" captionStart="FIGURE 15 A D" captionStartId="21.[151,255,1586,1610]" captionTargetBox="[167,1417,224,1540]" captionTargetId="figure@21.[151,1436,194,1562]" captionTargetPageId="21" captionText="FIGURE 15 A D. Atractides cognatus K. Viets, 1935, male, stream near Ban Khoom, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190551/files/figure.png" pageId="20" pageNumber="21">Fig. 15A</figureCitation>
): L 252 (222), Cx-3 W 269 (240), Cx-1+2 mL 97 (86), Cx-1+2 lL 163 (144). Palp (
<figureCitation id="134F2D89E541FFB5FCFFD7EAFC1FF911" box="[811,920,1681,1707]" captionStart="FIGURE 15 A D" captionStartId="21.[151,255,1586,1610]" captionTargetBox="[167,1417,224,1540]" captionTargetId="figure@21.[151,1436,194,1562]" captionTargetPageId="21" captionText="FIGURE 15 A D. Atractides cognatus K. Viets, 1935, male, stream near Ban Khoom, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190551/files/figure.png" pageId="20" pageNumber="21">Fig. 15C</figureCitation>
): total L 211 (199), dL: P-1 24 (24), P-2 46 (42), P-3 50 (48), P-4 67 (61), P-5 24 (24); %L: P-1 11.4 (12.1), P-2 21.8 (21.1), P-3 23.7 (24.1), P-4 31.8 (30.7), P-5 11.4 (12.1); L P-2/P-4 0.69 (0.69). Genital field (
<figureCitation id="134F2D89E541FFB5FCB3D7A5FC53F942" box="[871,980,1758,1784]" captionStart="FIGURE 15 A D" captionStartId="21.[151,255,1586,1610]" captionTargetBox="[167,1417,224,1540]" captionTargetId="figure@21.[151,1436,194,1562]" captionTargetPageId="21" captionText="FIGURE 15 A D. Atractides cognatus K. Viets, 1935, male, stream near Ban Khoom, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190551/files/figure.png" pageId="20" pageNumber="21">Fig. 15B</figureCitation>
): L/W 76/92, L Ac-1-3: 27-29: 32-34: 29-30.
</paragraph>
<paragraph id="8BCB310CE541FFB5FF11D657FB07F829" blockId="20.[151,1437,1062,1939]" pageId="20" pageNumber="21">
I-Leg (
<figureCitation id="134F2D89E541FFB5FEC2D657FE03F8FC" box="[278,388,1836,1862]" captionStart="FIGURE 15 A D" captionStartId="21.[151,255,1586,1610]" captionTargetBox="[167,1417,224,1540]" captionTargetId="figure@21.[151,1436,194,1562]" captionTargetPageId="21" captionText="FIGURE 15 A D. Atractides cognatus K. Viets, 1935, male, stream near Ban Khoom, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190551/files/figure.png" pageId="20" pageNumber="21">Fig. 15D</figureCitation>
): I-Leg-5 dL 129 (121), vL 74 (74), dL/vL 1.74 (1.64), HB 32 (31), dL/HB 4.0 (4.0), S-1 L 73 (75), L/
<geoCoordinate id="EE4057CBE541FFB5FEE6D629FE0EF8D6" box="[306,393,1874,1900]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="5555" value="-10.7">W 10.7</geoCoordinate>
(9.9), S-2 L 56 (57), L/
<geoCoordinate id="EE4057CBE541FFB5FD71D629FD69F8D6" box="[677,750,1874,1900]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="5555" value="-5.7">W 5.7</geoCoordinate>
(5.4), distance of sword setae at I-Leg-5 20 (19), L S-1/2 1.3 (1.32); I-Leg-6 L 106 (104), HB 12 (12), L/HB 8.8 (8.7); L I-Leg-5/6 1.22 (1.16).
</paragraph>
<caption id="DF0B6184E540FFB4FF43D749FBC9F9D1" httpUri="https://zenodo.org/record/190551/files/figure.png" pageId="21" pageNumber="22" targetBox="[167,1417,224,1540]" targetPageId="21">
<paragraph id="8BCB310CE540FFB4FF43D749FBC9F9D1" blockId="21.[151,1436,1586,1643]" pageId="21" pageNumber="22">
<emphasis id="B900ED1EE540FFB4FF43D749FED2F9F0" bold="true" box="[151,341,1586,1610]" pageId="21" pageNumber="22">FIGURE 15AD</emphasis>
.
<taxonomicName id="4C744A8FE540FFB4FEB7D749FD67F9F3" authority="K. Viets, 1935" authorityName="K. Viets" authorityYear="1935" box="[355,736,1586,1609]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE540FFB4FEB7D749FDBDF9F3" box="[355,570,1586,1609]" italics="true" pageId="21" pageNumber="22">Atractides cognatus</emphasis>
K. Viets, 1935
</taxonomicName>
, male, stream near Ban Khoom, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg-5 and - 6. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8BCB310CE540FFB4FF11D7E1FD2CF879" blockId="21.[151,1436,1690,2025]" pageId="21" pageNumber="22">
Female (from stream near Ban Khoom,
<collectingCountry id="F363719CE540FFB4FD4AD7E1FC8DF90E" box="[670,778,1690,1716]" name="Thailand" pageId="21" pageNumber="22">Thailand</collectingCountry>
; in parentheses specimen from unnamed creek crossing track from watchtower,
<collectingCountry id="F363719CE540FFB4FE16D7BAFDB3F961" box="[450,564,1729,1755]" name="Thailand" pageId="21" pageNumber="22">Thailand</collectingCountry>
; in square brackets specimen from tributary of Tahan River near Lata Berkoh,
<collectingCountry id="F363719CE540FFB4FF29D79CFEE8F8BB" box="[253,367,1767,1793]" name="Malaysia" pageId="21" pageNumber="22">Malaysia</collectingCountry>
): Idiosoma L/W 694 (894) [509]/519 (709) [372]; Coxal field: L 331 (370) [284], Cx-3 W 400 (441) [356], Cx-1+2 mL 111 (131) [89], Cx-1+2 lL 206 (234) [186]. Palp (
<figureCitation id="134F2D89E540FFB4FB98D675FB3DF892" box="[1100,1210,1806,1832]" captionStart="FIGURE 16 A C" captionStartId="22.[151,255,1784,1808]" captionTargetBox="[161,1412,629,1746]" captionTargetId="figure@22.[151,1436,619,1760]" captionTargetPageId="22" captionText="FIGURE 16 A C. Atractides cognatus K. Viets, 1935, female (A, C - specimen from stream near Ban Khoom, Thailand; B- specimen from tributary of Tahan River near Lata Berkoh, Malaysia): A = I-Leg- 5 and - 6; B = genital field; C = palp, lateral view. Scale bar = 100 μm." httpUri="https://zenodo.org/record/190552/files/figure.png" pageId="21" pageNumber="22">Fig. 16C</figureCitation>
): total L 290 (318) [267], dL: P-1 27 (33) [29], P-2 61 (64) [55], P-3 79 (88) [71], P-4 91 (99) [79], P-5 32 (38) [33]; %L: P-1 9.3 (10.4) [10.9], P-2 21.0 (20.1) [20.6], P-3 27.2 (27.7) [26.6], P-4 31.4 (31.1) [30.0], P-5 11.0 (12.0) [12.4]; L P- 2/P-4 0.67 (0.65) [0.7]. Genital field (
<figureCitation id="134F2D89E540FFB4FD9AD6F9FD3EF826" box="[590,697,1922,1948]" captionStart="FIGURE 16 A C" captionStartId="22.[151,255,1784,1808]" captionTargetBox="[161,1412,629,1746]" captionTargetId="figure@22.[151,1436,619,1760]" captionTargetPageId="22" captionText="FIGURE 16 A C. Atractides cognatus K. Viets, 1935, female (A, C - specimen from stream near Ban Khoom, Thailand; B- specimen from tributary of Tahan River near Lata Berkoh, Malaysia): A = I-Leg- 5 and - 6; B = genital field; C = palp, lateral view. Scale bar = 100 μm." httpUri="https://zenodo.org/record/190552/files/figure.png" pageId="21" pageNumber="22">Fig. 16B</figureCitation>
): L (150)/W (154), genital plate L 114 (108) [85], L Ac-1-3: 37 (32-35) [30]-42 (30-35) [33]-42 (35-36) [34].
</paragraph>
<paragraph id="8BCB310CE540FFB7FF11D6B4FC5FFF45" blockId="21.[151,1436,1690,2025]" lastBlockId="22.[151,1436,152,564]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
I-Leg (
<figureCitation id="134F2D89E540FFB4FECDD6B4FE0DF853" box="[281,394,1999,2025]" captionStart="FIGURE 16 A C" captionStartId="22.[151,255,1784,1808]" captionTargetBox="[161,1412,629,1746]" captionTargetId="figure@22.[151,1436,619,1760]" captionTargetPageId="22" captionText="FIGURE 16 A C. Atractides cognatus K. Viets, 1935, female (A, C - specimen from stream near Ban Khoom, Thailand; B- specimen from tributary of Tahan River near Lata Berkoh, Malaysia): A = I-Leg- 5 and - 6; B = genital field; C = palp, lateral view. Scale bar = 100 μm." httpUri="https://zenodo.org/record/190552/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
): I-Leg-5 dL 194 (210) [171], vL 98 (121) [100], dL/vL 1.98 (1.74) [1.7], HB 44 (48) [43], dL/HB 4.4 (4.4) [4.0], S-1 L 103 (125) [109], L/
<geoCoordinate id="EE4057CBE543FFB7FCC3D1E3FCE7FF08" box="[791,864,152,178]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="-8.5">W 8.5</geoCoordinate>
(10.5) [10.3], S-2 L 69 (83) [65], L/
<geoCoordinate id="EE4057CBE543FFB7FAC7D1E3FADCFF08" box="[1299,1371,152,178]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="-4.1">W 4.1</geoCoordinate>
(4.5) [4.3], distance of sword setae at I-Leg-5 49 (39) [33], L S-1/2 1.5 (1.5) [1.7]; I-Leg-6 L 146 (179) [138], HB 13.6 (14) [12], L/HB 10.7 (13.0) [11.2]; L I-Leg-5/6 1.33 (1.17) [1.24].
</paragraph>
<paragraph id="8BCB310CE543FFB7FF11D077FC09FEF6" blockId="22.[151,1436,152,564]" pageId="22" pageNumber="23">In addition we give measurements of the female from Sulawesi which are suspected to belong to the same species (general differences see below in the “remarks”-section).</paragraph>
<paragraph id="8BCB310CE543FFB7FF11D022FCA9FE5D" blockId="22.[151,1436,152,564]" pageId="22" pageNumber="23">
Female: Idiosoma L/W 806/619. Coxal field (
<figureCitation id="134F2D89E543FFB7FD08D022FCCEFEC9" box="[732,841,345,371]" captionStart="FIGURE 17 A D" captionStartId="23.[151,257,1895,1919]" captionTargetBox="[151,1436,204,1860]" captionTargetId="figure@23.[151,1436,194,1871]" captionTargetPageId="23" captionText="FIGURE 17 A D. Atractides cognatus K. Viets, 1935, female, stream 18 km west of Molibagu, Sulawesi: A = coxal field; B = palp, medial view; C = genital field; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190553/files/figure.png" pageId="22" pageNumber="23">Fig. 17A</figureCitation>
): posterior margin Cx-1 strongly indented; Cx-1+2 mL 97, Cx-1+2 lL. Palp (
<figureCitation id="134F2D89E543FFB7FE6BD0FBFDACFE20" box="[447,555,384,410]" captionStart="FIGURE 17 A D" captionStartId="23.[151,257,1895,1919]" captionTargetBox="[151,1436,204,1860]" captionTargetId="figure@23.[151,1436,194,1871]" captionTargetPageId="23" captionText="FIGURE 17 A D. Atractides cognatus K. Viets, 1935, female, stream 18 km west of Molibagu, Sulawesi: A = coxal field; B = palp, medial view; C = genital field; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190553/files/figure.png" pageId="22" pageNumber="23">Fig. 17B</figureCitation>
): total L 296, dL and %L (in parentheses): P-1 31 (10.5), P-2 64 (21.6), P-3 78 (26.4), P-4 86 (29.1), P-5 37 (12.5); L P-2/P-4 0.74. Genital field (
<figureCitation id="134F2D89E543FFB7FC04D0DDFBBAFE7A" box="[976,1085,422,448]" captionStart="FIGURE 17 A D" captionStartId="23.[151,257,1895,1919]" captionTargetBox="[151,1436,204,1860]" captionTargetId="figure@23.[151,1436,194,1871]" captionTargetPageId="23" captionText="FIGURE 17 A D. Atractides cognatus K. Viets, 1935, female, stream 18 km west of Molibagu, Sulawesi: A = coxal field; B = palp, medial view; C = genital field; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190553/files/figure.png" pageId="22" pageNumber="23">Fig. 17C</figureCitation>
): L/W 136/144, genital plates bean-shaped, L genital plate 100-102, Ac-1-3: 32-33-33.
</paragraph>
<paragraph id="8BCB310CE543FFB7FF11D08FFA17FD8E" blockId="22.[151,1436,152,564]" pageId="22" pageNumber="23">
I-Leg (
<figureCitation id="134F2D89E543FFB7FEC2D08FFE04FDB4" box="[278,387,500,526]" captionStart="FIGURE 17 A D" captionStartId="23.[151,257,1895,1919]" captionTargetBox="[151,1436,204,1860]" captionTargetId="figure@23.[151,1436,194,1871]" captionTargetPageId="23" captionText="FIGURE 17 A D. Atractides cognatus K. Viets, 1935, female, stream 18 km west of Molibagu, Sulawesi: A = coxal field; B = palp, medial view; C = genital field; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190553/files/figure.png" pageId="22" pageNumber="23">Fig. 17D</figureCitation>
): I-Leg-5 dL 188, vL 110, dL/vL 1.7, HB 53, dL/HB 3.6, S-1 L 112, L/
<geoCoordinate id="EE4057CBE543FFB7FB1CD08FFAA6FDB4" box="[1224,1313,500,526]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="-10.6">W 10.6</geoCoordinate>
, S-2 L 74, L/
<geoCoordinate id="EE4057CBE543FFB7FF65D361FF7CFD8E" box="[177,251,538,564]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="-4.4">W 4.4</geoCoordinate>
, distance of sword setae at I-Leg-5 42, L S-1/2 1.5; I-Leg-6 L 155, HB 11, L/HB 14.1; L I-Leg-5/6.
</paragraph>
</subSubSection>
<caption id="DF0B6184E543FFB7FF43D783FE70F8EE" httpUri="https://zenodo.org/record/190552/files/figure.png" pageId="22" pageNumber="23" targetBox="[161,1412,629,1746]" targetPageId="22">
<paragraph id="8BCB310CE543FFB7FF43D783FE70F8EE" blockId="22.[151,1436,1784,1877]" pageId="22" pageNumber="23">
<emphasis id="B900ED1EE543FFB7FF43D783FED4F8AA" bold="true" box="[151,339,1784,1808]" pageId="22" pageNumber="23">FIGURE 16AC</emphasis>
.
<taxonomicName id="4C744A8FE543FFB7FEB4D782FD50F8AA" authority="K. Viets, 1935" authorityName="K. Viets" authorityYear="1935" box="[352,727,1785,1808]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE543FFB7FEB4D782FDB3F8AA" box="[352,564,1785,1808]" italics="true" pageId="22" pageNumber="23">Atractides cognatus</emphasis>
K. Viets, 1935
</taxonomicName>
, female (A, C - specimen from stream near Ban Khoom, Thailand; B- specimen from tributary of Tahan River near Lata Berkoh, Malaysia): A = I-Leg-5 and - 6; B = genital field; C = palp, lateral view. Scale bar = 100 μm.
</paragraph>
</caption>
<subSubSection id="C36E6287E543FFB9FF11D6F8FE58FDE1" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BCB310CE543FFB9FF11D6F8FDAFFE5D" blockId="22.[151,1436,1923,2027]" lastBlockId="24.[151,1437,152,719]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<emphasis id="B900ED1EE543FFB7FF11D6F8FEB9F827" bold="true" box="[197,318,1923,1949]" pageId="22" pageNumber="23">Remarks.</emphasis>
In the original description
<bibRefCitation id="EFE54CFDE543FFB7FDA9D6F8FCACF827" author="Viets" box="[637,811,1923,1949]" pageId="22" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K.Viets (1935)</bibRefCitation>
compared
<taxonomicName id="4C744A8FE543FFB7FC79D6FFFBBFF827" box="[941,1080,1924,1949]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE543FFB7FC79D6FFFBBFF827" box="[941,1080,1924,1949]" italics="true" pageId="22" pageNumber="23">A. cognatus</emphasis>
</taxonomicName>
with
<taxonomicName id="4C744A8FE543FFB7FBAFD6FFFA8FF827" box="[1147,1288,1924,1949]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE543FFB7FBAFD6FFFA8FF827" box="[1147,1288,1924,1949]" italics="true" pageId="22" pageNumber="23">A. spatiosus</emphasis>
</taxonomicName>
, believing it to be a subspecies of the latter. He noticed the following differences: in
<taxonomicName id="4C744A8FE543FFB7FC08D6D1FBEDF879" box="[988,1130,1962,1987]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE543FFB7FC08D6D1FBEDF879" box="[988,1130,1962,1987]" italics="true" pageId="22" pageNumber="23">A. spatiosus</emphasis>
</taxonomicName>
the sword seta of P-4 was inserted halfway between ventral hairs, Cx-3+4 medially indented; in
<taxonomicName id="4C744A8FE543FFB7FC1FD6AAFBD1F850" box="[971,1110,2001,2026]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE543FFB7FC1FD6AAFBD1F850" box="[971,1110,2001,2026]" italics="true" pageId="22" pageNumber="23">A. cognatus</emphasis>
</taxonomicName>
the sword seta of P-4 was close to the distoventral hair, Cx-3+4 medially straight. Later on,
<bibRefCitation id="EFE54CFDE54DFFB9FC4ED1E3FBE7FF08" author="Lundblad" box="[922,1120,152,178]" pageId="24" pageNumber="30" refString="Lundblad, O. (1971) Weitere Beitrage zur Kenntnis der Fliesswassermilben Javas. Arkiv for Zoologi, 23, 293 - 359." type="journal article" year="1971">Lundblad (1971)</bibRefCitation>
showed that differences in shape of posterior coxae and length and direction of medial apodemes of Cx-1, were insignificant and that
<taxonomicName id="4C744A8FE54DFFB9FA57D1C4FE85FF44" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE54DFFB9FA57D1C4FE85FF44" italics="true" pageId="24" pageNumber="25">A. cognatus</emphasis>
</taxonomicName>
should be considered a junior synonym of
<taxonomicName id="4C744A8FE54DFFB9FCD1D19EFC12FF44" box="[773,917,229,254]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE54DFFB9FCD1D19EFC12FF44" box="[773,917,229,254]" italics="true" pageId="24" pageNumber="25">A. spatiosus</emphasis>
</taxonomicName>
. This conclusion, however, contradicted by other morphological evidence.
<taxonomicName id="4C744A8FE54DFFB9FDC4D077FC83FE9F" box="[528,772,268,293]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE54DFFB9FDC4D077FC83FE9F" box="[528,772,268,293]" italics="true" pageId="24" pageNumber="25">Atractides spatiosus</emphasis>
</taxonomicName>
(in parentheses data taken from
<bibRefCitation id="EFE54CFDE54DFFB9FB47D077FABBFE9C" author="Viets" box="[1171,1340,268,294]" pageId="24" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K. Viets 1935</bibRefCitation>
) differs from
<taxonomicName id="4C744A8FE54DFFB9FF0CD049FEE2FEF1" box="[216,357,306,331]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE54DFFB9FF0CD049FEE2FEF1" box="[216,357,306,331]" italics="true" pageId="24" pageNumber="25">A. cognatus</emphasis>
</taxonomicName>
in longer I-Leg-5-6, a more distant S-1 and - 2 on I-Leg-5 (&gt;
<quantity id="4C8C9CE9E54DFFB9FB9AD049FB0DFEF6" box="[1102,1162,306,332]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="24" pageNumber="25" unit="in" value="40.0">40 in</quantity>
both sexes). However, because some important characters such as the S-1 and -2 interspace in the male were lacking in the original description (K
<bibRefCitation id="EFE54CFDE54DFFB9FE9CD0FBFE4CFE20" author="Viets" box="[328,459,384,410]" pageId="24" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">Viets 1935</bibRefCitation>
), our assignment of
<taxonomicName id="4C744A8FE54DFFB9FD69D0FBFCCFFE23" box="[701,840,384,409]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE54DFFB9FD69D0FBFCCFFE23" box="[701,840,384,409]" italics="true" pageId="24" pageNumber="25">A. cognatus</emphasis>
</taxonomicName>
is still provisional. The variability of further male and female specimens of
<taxonomicName id="4C744A8FE54DFFB9FE04D0DCFDE7FE7A" box="[464,608,423,448]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE54DFFB9FE04D0DCFDE7FE7A" box="[464,608,423,448]" italics="true" pageId="24" pageNumber="25">A. cognatus</emphasis>
</taxonomicName>
from the area of
<typeStatus id="54CF8FAEE54DFFB9FCECD0DDFCEAFE7A" box="[824,877,422,448]" pageId="24" pageNumber="25">type</typeStatus>
locality (South Sumatra, Ranau), needs to be examined to clarify the taxonomy.
</paragraph>
<caption id="DF0B6184E542FFB6FF43D61CFBC9F81A" httpUri="https://zenodo.org/record/190553/files/figure.png" pageId="23" pageNumber="24" targetBox="[151,1436,204,1860]" targetPageId="23">
<paragraph id="8BCB310CE542FFB6FF43D61CFBC9F81A" blockId="23.[151,1436,1895,1953]" pageId="23" pageNumber="24">
<emphasis id="B900ED1EE542FFB6FF43D61CFEDFF8C5" bold="true" box="[151,344,1895,1919]" pageId="23" pageNumber="24">FIGURE 17AD</emphasis>
.
<taxonomicName id="4C744A8FE542FFB6FEB2D613FD6DF8C4" authority="K. Viets, 1935" authorityName="K. Viets" authorityYear="1935" box="[358,746,1895,1919]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B900ED1EE542FFB6FEB2D613FDC7F8C5" box="[358,576,1896,1919]" italics="true" pageId="23" pageNumber="24">Atractides cognatus</emphasis>
K. Viets, 1935
</taxonomicName>
, female, stream 18 km west of Molibagu, Sulawesi: A = coxal field; B = palp, medial view; C = genital field; D = I-Leg-5 and - 6. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8BCB310CE54DFFB9FF12D08FFE58FDE1" blockId="24.[151,1437,152,719]" pageId="24" pageNumber="25">
The specimens from Sulawesi agree well with the original description (
<bibRefCitation id="EFE54CFDE54DFFB9FBE8D08FFB6EFDB4" author="Viets" box="[1084,1257,500,526]" pageId="24" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K. Viets 1935</bibRefCitation>
) and with our material from
<collectingCountry id="F363719CE54DFFB9FE96D361FE2BFD8E" box="[322,428,538,564]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
and
<collectingCountry id="F363719CE54DFFB9FE33D361FDDFFD8E" box="[487,600,538,564]" name="Malaysia" pageId="24" pageNumber="25">Malaysia</collectingCountry>
. However, due to the fact that we do not have male specimens, this is only a tentative assignment.
</paragraph>
</subSubSection>
<subSubSection id="C36E6287E54DFFB9FF11D31CFCB3FD75" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BCB310CE54DFFB9FF11D31CFCB3FD75" blockId="24.[151,1437,152,719]" pageId="24" pageNumber="25">
<emphasis id="B900ED1EE54DFFB9FF11D31CFEEAFD3B" bold="true" box="[197,365,615,641]" pageId="24" pageNumber="25">Distribution.</emphasis>
<collectingCountry id="F363719CE54DFFB9FEA2D313FE6AFD38" box="[374,493,616,642]" name="Indonesia" pageId="24" pageNumber="25">Indonesia</collectingCountry>
(Sumatra, E Java:
<taxonomicName id="4C744A8FE54DFFB9FD30D313FA9FFD38" authority="K. Viets 1935" authorityName="K. Viets" authorityYear="1935" box="[740,1304,616,642]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="spatiosus" subSpecies="cognatus">
<emphasis id="B900ED1EE54DFFB9FD30D313FBD1FD3B" box="[740,1110,616,641]" italics="true" pageId="24" pageNumber="25">Atractides spatiosus cognatus</emphasis>
<bibRefCitation id="EFE54CFDE54DFFB9FBBCD313FA9FFD38" author="Viets" box="[1128,1304,616,642]" pageId="24" pageNumber="30" refString="Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement. 13, Tropische Binnengewasser, 5, 484 - 594." type="journal article" year="1935">K. Viets 1935</bibRefCitation>
</taxonomicName>
; Sulawesi:
<taxonomicName id="4C744A8FE54DFFB9FF76D3F4FDB9FD12" authority="Smit 1996" authorityName="Smit" authorityYear="1996" box="[162,574,654,680]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE54DFFB9FF76D3F4FE24FD12" box="[162,419,655,680]" italics="true" pageId="24" pageNumber="25">Atractides spatiosus</emphasis>
<bibRefCitation id="EFE54CFDE54DFFB9FE62D3F5FDB9FD12" author="Smit" box="[438,574,654,680]" pageId="24" pageNumber="30" refString="Smit, H. (1996) Ten new species of water mites from Sulawesi and Waigeo, Indonesia (Acari: Hydrachnellae). Bulletin Zoologisch Museum, Universiteit van Amsterdam, 15 (2), 5 - 19." type="journal article" year="1996">Smit 1996</bibRefCitation>
</taxonomicName>
),
<collectingCountry id="F363719CE54DFFB9FD81D3F5FD4EFD12" box="[597,713,654,680]" name="Malaysia" pageId="24" pageNumber="25">Malaysia</collectingCountry>
(
<taxonomicName id="4C744A8FE54DFFB9FD33D3F4FB17FD12" authority="Wiles 1991" authorityName="Wiles" authorityYear="1991" box="[743,1168,654,680]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B900ED1EE54DFFB9FD33D3F4FC6FFD12" box="[743,1000,655,680]" italics="true" pageId="24" pageNumber="25">Atractides spatiosus</emphasis>
<bibRefCitation id="EFE54CFDE54DFFB9FC28D3F5FB17FD12" author="Wiles" box="[1020,1168,654,680]" pageId="24" pageNumber="30" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32, 41 - 56." type="journal article" year="1991">Wiles 1991</bibRefCitation>
</taxonomicName>
),
<collectingCountry id="F363719CE54DFFB9FB73D3F5FB7BFD12" box="[1191,1276,654,680]" name="Myanmar" pageId="24" pageNumber="25">Burma</collectingCountry>
(
<emphasis id="B900ED1EE54DFFB9FACFD3F4FE07FD74" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C744A8FE54DFFB9FACFD3F4FE83FD74" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spatiosus">Atractides spatiosus</taxonomicName>
f. cognata
</emphasis>
<bibRefCitation id="EFE54CFDE54DFFB9FE46D3CEFDCFFD75" author="Lundblad" box="[402,584,693,719]" pageId="24" pageNumber="30" refString="Lundblad, O. (1969) Indische Wassermilben, hauptsachlich von Hinterindien. Arkiv for Zoologi, 22, 1 - 126." type="journal article" year="1969">Lundblad 1969</bibRefCitation>
). New for
<collectingCountry id="F363719CE54DFFB9FD10D3CEFCB7FD75" box="[708,816,693,719]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>