160 lines
18 KiB
XML
160 lines
18 KiB
XML
<document id="2F333B34C4C511E4CD3DE3054676FCC2" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFB7FFF9FF0DBF855B20A00F" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus franziskaeweissbachae Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="350" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="349" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="0159FE43C688DEC4191DBFB5AD909700" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2310039005479C7F4C0E5E78648DC413">
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<mods:title id="C0D6F735BD5F119D8BDECA7FC15ABA0C">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:name id="1A3A4DA6E4C40E2EC8F9ECB70ECBD303" type="personal">
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<mods:namePart id="89253A2FB712B66E1ACDFCD2650BBFF1">Fanti, Fabrizio</mods:namePart>
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<mods:name id="C3910B2362AE568CE7C2E84F2BAABF73" type="personal">
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<mods:roleTerm id="9480144E5A9768085D1C559AE39822CD">Author</mods:roleTerm>
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<mods:namePart id="7AA35F7780BE6E64C2142A1DF3D475B5">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="E0453360CD6F1B253507A1C2390A5609">text</mods:typeOfResource>
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<mods:title id="0DB0E1DC3B3DBDF1E7BA2ADF26D2B8B7">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="093BF303081607DC2876762002A65656">
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<mods:date id="02CC24C3194DCCFD1C83ED00C7368A81">2022</mods:date>
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<mods:detail id="4FF350B362E3E00687BA536219F94CDF" type="volume">
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<mods:number id="281C8EC2A0E061F2FB6FD15ABFCDB660">22</mods:number>
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<mods:number id="63E2F4F3D64C8A42158C67422F30FE75">2</mods:number>
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<mods:start id="F772270EC50E73AF63FD6F1160FA15BB">331</mods:start>
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<mods:classification id="7DEE28C3826B98AB68360A0745031AFC">journal article</mods:classification>
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<mods:identifier id="D780F322AC7756E345C585FA8DAFCB1E" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="5C4FBD0E4CEA11D30DB8E75D6DC0F611" type="ISSN">1407-8619</mods:identifier>
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<mods:identifier id="780EF0233DA8514089832EC532C03154" type="Zenodo-Dep">13203160</mods:identifier>
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<treatment id="03DAD52BFFB7FFF9FF0DBF855B20A00F" ID-DOI="http://doi.org/10.5281/zenodo.13199805" ID-Zenodo-Dep="13199805" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F" httpUri="http://treatment.plazi.org/id/03DAD52BFFB7FFF9FF0DBF855B20A00F" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349">
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<subSubSection id="C36937B6FFB7FFF8FF0DBF855DC5A3BF" box="[151,643,695,719]" pageId="18" pageNumber="349" type="nomenclature">
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<paragraph id="8BCC643DFFB7FFF8FF0DBF855DC5A3BF" blockId="18.[151,643,695,751]" box="[151,643,695,719]" pageId="18" pageNumber="349">
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<heading id="D084D351FFB7FFF8FF0DBF855DC5A3BF" bold="true" box="[151,643,695,719]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="349" reason="4">
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<emphasis id="B907B82FFFB7FFF8FF0DBF855DC5A3BF" bold="true" box="[151,643,695,719]" pageId="18" pageNumber="349">
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<taxonomicName id="4C731FBEFFB7FFF8FF0DBF855D6FA3BF" authorityName="Fanti & Müller" authorityYear="2022" box="[151,553,695,719]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="franziskaeweissbachae" status="sp. nov.">
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<emphasis id="B907B82FFFB7FFF8FF0DBF855D6FA3BF" bold="true" box="[151,553,695,719]" italics="true" pageId="18" pageNumber="349">Sanaungulus franziskaeweissbachae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFB7FFF8FDB5BF855DC5A3BF" box="[559,643,695,719]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFB7FFF8FF0DBFEA5FB2A39F" box="[151,244,728,751]" pageId="18" pageNumber="349" type="description">
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<paragraph id="8BCC643DFFB7FFF8FF0DBFEA5FB2A39F" blockId="18.[151,643,695,751]" box="[151,244,728,751]" pageId="18" pageNumber="349">
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(
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<figureCitation id="134878B8FFB7FFF8FF05BFEA5FAAA39F" box="[159,236,728,751]" captionStart="Fig" captionStartId="18.[151,190,1665,1688]" captionTargetBox="[151,1172,1023,1635]" captionTargetId="figure-344@18.[151,1172,1023,1635]" captionTargetPageId="18" captionText="Fig. 10. Sanaungulus franziskaeweissbachae sp. nov. in Burmese amber. Holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13203195" httpUri="https://zenodo.org/record/13203195/files/figure.png" pageId="18" pageNumber="349">Fig. 10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFB7FFF8FF0DBE3D5C81A058" pageId="18" pageNumber="349" type="materials_examined">
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<paragraph id="8BCC643DFFB7FFF8FF0DBE3D5F98A2BA" blockId="18.[151,643,783,970]" pageId="18" pageNumber="349">
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<materialsCitation id="3B1B6E60FFB7FFF8FF0DBE3D5F98A2BA" collectingDate="2021-12" collectionCode="BUB" county="Probably" location="Catalog" municipality="Burmese" pageId="18" pageNumber="349" specimenCode="BUB 3899" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFB7FFF8FF0DBE3D5E40A257" bold="true" box="[151,262,783,807]" pageId="18" pageNumber="349">
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<typeStatus id="54C8DA9FFFB7FFF8FF0DBE3D5E47A257" box="[151,257,783,807]" pageId="18" pageNumber="349" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFB7FFF8FE8ABE3D5E37A256" box="[272,369,783,806]" pageId="18" pageNumber="349">Probably</collectingCounty>
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male, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFB7FFF8FF35BE025E54A237" box="[175,274,816,839]" pageId="18" pageNumber="349">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFFB7FFF8FEBDBE025E3EA237" box="[295,376,816,839]" country="Myanmar" name="Kachin" pageId="18" pageNumber="349">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFB7FFF8FDB6BE025DC5A237" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FDB6BE025DC5A237" box="[556,643,816,839]" county="Probably" municipality="Burmese" name="Catalog" pageId="18" pageNumber="349" stateProvince="Kachin">Catalog</location>
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number SNSBBSPG
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<date id="FFCD42FDFFB7FFF8FE1EBE635EA3A218" box="[388,485,849,872]" pageId="18" pageNumber="349" value="2021-12">
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<collectingDate id="EF89BB15FFB7FFF8FE1EBE635EA3A218" box="[388,485,849,872]" pageId="18" pageNumber="349" value="2021-12">2021 XII</collectingDate>
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</date>
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<quantity id="4C8BC9D8FFB7FFF8FE71BE635D64A218" box="[491,546,849,872]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" pageId="18" pageNumber="349" unit="in" value="72.0">72 in</quantity>
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the
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<location id="8EAC32E6FFB7FFF8FDD5BE635ED1A2F9" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FDD5BE635ED1A2F9" county="Probably" municipality="Burmese" name="Bayerische Staatssammlung" pageId="18" pageNumber="349" stateProvince="Kachin">Bayerische Staatssammlung</location>
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für
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<location id="8EAC32E6FFB7FFF8FE58BE405D13A2F9" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FE58BE405D13A2F9" box="[450,597,882,905]" county="Probably" municipality="Burmese" name="Palaontologie" pageId="18" pageNumber="349" stateProvince="Kachin">Paläontologie</location>
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und
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<location id="8EAC32E6FFB7FFF8FF0DBEA15FBEA2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FF0DBEA15FBEA2DA" box="[151,248,915,938]" county="Probably" municipality="Burmese" name="Geologie" pageId="18" pageNumber="349" stateProvince="Kachin">Geologie</location>
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(ex
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<location id="8EAC32E6FFB7FFF8FEBEBEA15E38A2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FEBEBEA15E38A2DA" box="[292,382,915,938]" county="Probably" municipality="Burmese" name="Muller's" pageId="18" pageNumber="349" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFB7FFF8FDD6BEA15F97A2BA" collectionCode="BUB" pageId="18" pageNumber="349">BUB 3899</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCC643DFFB7FFF8FD32BDA85CBDA1A2" blockId="18.[680,1172,154,210]" pageId="18" pageNumber="349">
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<emphasis id="B907B82FFFB7FFF8FD32BDA85C07A1C2" bold="true" box="[680,833,154,178]" pageId="18" pageNumber="349">Type locality.</emphasis>
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<collectingCountry id="F36424ADFFB7FFF8FCD0BDA95CF0A1C2" box="[842,950,155,178]" name="Myanmar" pageId="18" pageNumber="349">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFB7FFF8FC5BBDA95B0DA1C2" box="[961,1099,155,178]" country="Myanmar" name="Kachin" pageId="18" pageNumber="349">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFFB7FFF8FD32BDC35C81A058" blockId="18.[680,1172,241,296]" pageId="18" pageNumber="349">
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<emphasis id="B907B82FFFB7FFF8FD32BDC35C7AA079" bold="true" box="[680,828,241,265]" pageId="18" pageNumber="349">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFB7FFF8FD32BC7A5C3BA32F" pageId="18" pageNumber="349" type="diagnosis">
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<paragraph id="8BCC643DFFB7FFF8FD32BC7A5C3BA32F" blockId="18.[680,1172,328,607]" pageId="18" pageNumber="349">
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<emphasis id="B907B82FFFB7FFF8FD32BC7A5CE0A010" bold="true" box="[680,934,328,352]" pageId="18" pageNumber="349">Differential diagnosis.</emphasis>
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<taxonomicName id="4C731FBEFFB7FFF8FC35BC7A5C12A00F" authorityName="Fanti & Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="franziskaeweissbachae" status="sp. nov.">
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<emphasis id="B907B82FFFB7FFF8FC35BC7A5C12A00F" italics="true" pageId="18" pageNumber="349">Sanaungulus franziskaeweissbachae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFB7FFF8FCC0BC5A5CECA00F" box="[858,938,360,383]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
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is very similar to
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<taxonomicName id="4C731FBEFFB7FFF8FBF9BC5A5C78A0EF" authorityName="Fanti & Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="nilsi">
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<emphasis id="B907B82FFFB7FFF8FBF9BC5A5C78A0EF" italics="true" pageId="18" pageNumber="349">Sanaungulus nilsi</emphasis>
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</taxonomicName>
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(described in the present work), but the new species differs based on its more robost second antennomere and much longer metathoracic wings. Based on these characters and others of lesser importance, the
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<specimenCount id="9D75AFB4FFB7FFF8FBBCBF3A5D99A34F" count="2" pageId="18" pageNumber="349" type="generic">two specimens</specimenCount>
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are not considered here as male and female of the same species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFB7FFF9FD32BF4C5EA8A7E2" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349" type="description">
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<paragraph id="8BCC643DFFB7FFF8FD32BF4C5C09A386" blockId="18.[680,1172,638,758]" pageId="18" pageNumber="349">
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<emphasis id="B907B82FFFB7FFF8FD32BF4C5C72A3E6" bold="true" box="[680,820,638,662]" pageId="18" pageNumber="349">Description.</emphasis>
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Adult, winged, slender. Probably male, defined on the basis of the last abdominal segment triangular shaped. Body length
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<quantity id="4C8BC9D8FFB7FFF8FBDABF8D5BD6A3A6" box="[1088,1168,703,726]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="349" unit="mm" value="2.8">2.8 mm</quantity>
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. Entirely brown.
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</paragraph>
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<paragraph id="8BCC643DFFB7FFF9FD32BE275EA8A7E2" blockId="18.[680,1172,789,972]" lastBlockId="19.[113,605,155,1682]" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349">Head elongated, slightly constricted (triangularshaped) behind the eyes, equipped with pubescence and shallow punctation. Eyes prominent, extremely large, elongated, inserted in the lateral part of the head.Mandibles elongated, very robust, strongly curved, falciform, pointed apicallly, without teeth. Maxillary palps 4 segmented with the last palpomere very elongated and securiform. Labial palps 3 segmented, with the last palpomere elongated and securiform.Antennae 11 segmented, filiform, rather robust, short, approximately reaching the half of elytra, antennal insertion far from the eyes; scape robust, very elongated, slightly clubshaped; antennomeres II–III short, thinner than first, antennomere II suboval and more robust than antennomere III; antennomeres IV–IX subequal, filiform, very elongated; antennomere X slightly shorter than previous ones; antennomere XI filiform, rounded at apex; all the antennomeres with short pubescence. Pronotum very elongated, slightly narrower than head, anterior margin slightly roundish, posterior margin slightly undulated and slightly bordered, sides straight and bordered, corners obtuse, pronotum disc pubescent and slighlty convex near the posterior margin. Scutellum triangularshaped. Elytra short which reveals various abdominal segments, at humeri slightly wider than pronotum, posteriorly parallelsided and rather dehiscent apically, rounded apexes, surface pubescent and with a very small rugosity. Metathoracic wings transparent, very long, longer than elytra and surpassing the last abdominal segments. Sternum elongated with posterior margin rounded, sternites narrow and transverse, last tergite elongated with pointed apex which has the margin rounded, last sternite shorter and narrower than last tergite with pointed apex.Legs thin, extremely long, pubescent; coxae extraordinary massive, elongated, very enlarged; trochanters elongated, with rounded apex; femora slightly curved, more robust than tibiae, subcylindrical; tibiae flattened, thin, with a small and obtuse spur near the apex, protibiae very shorter than profemora, meso and metatibiae very longer than meso and metafemora. Tarsal formula 5 5 5; first protarsomere long, about 3.0 times longer than second; second protarsomere robust, enlarged apically; third protarsomere short, bilobed with pointed apex; fourth protarsomere strongly bilobed with a pointed and short lobe at each corners; fifth protarsomere elongated, thin, flat and curved; claws simple, without denticle at the base.</paragraph>
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</subSubSection>
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<caption id="DF0C34B5FFB7FFF8FF0DBBB35BD2A7E8" ID-DOI="http://doi.org/10.5281/zenodo.13203195" ID-Zenodo-Dep="13203195" box="[151,1172,1665,1688]" httpUri="https://zenodo.org/record/13203195/files/figure.png" pageId="18" pageNumber="349" startId="18.[151,190,1665,1688]" targetBox="[151,1172,1023,1635]" targetPageId="18" targetType="figure">
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<paragraph id="8BCC643DFFB7FFF8FF0DBBB35BD2A7E8" blockId="18.[151,1172,1665,1688]" box="[151,1172,1665,1688]" pageId="18" pageNumber="349">
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Fig. 10.
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<taxonomicName id="4C731FBEFFB7FFF8FF6ABBB35D33A7E8" authorityName="Fanti & Müller" authorityYear="2022" box="[240,629,1665,1688]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="franziskaeweissbachae" status="sp. nov.">
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||
<emphasis id="B907B82FFFB7FFF8FF6ABBB35D33A7E8" box="[240,629,1665,1688]" italics="true" pageId="18" pageNumber="349">Sanaungulus franziskaeweissbachae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFB7FFF8FDE1BBB35D8BA7E8" box="[635,717,1665,1688]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber. Holotype, lateral view.
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</paragraph>
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</caption>
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<subSubSection id="C36937B6FFB6FFF9FD19BDA85B4DA078" pageId="19" pageNumber="350" type="etymology">
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<paragraph id="8BCC643DFFB6FFF9FD19BDA85B29A1A2" blockId="19.[643,1135,154,210]" pageId="19" pageNumber="350">
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<emphasis id="B907B82FFFB6FFF9FD19BDA85C46A1C2" bold="true" box="[643,768,154,178]" pageId="19" pageNumber="350">Etymology.</emphasis>
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Named in honor of Franziska Weissbach, dear friend of the second author (Patrick).
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</paragraph>
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<paragraph id="8BCC643DFFB6FFF9FD19BDC35B4DA078" blockId="19.[643,1035,241,265]" box="[643,1035,241,265]" pageId="19" pageNumber="350">
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<emphasis id="B907B82FFFB6FFF9FD19BDC35C60A079" bold="true" box="[643,806,241,265]" pageId="19" pageNumber="350">Syninclusions.</emphasis>
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Air bubbles, detritus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFB6FFF9FD19BC1A5B20A00F" pageId="19" pageNumber="350" type="discussion">
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<paragraph id="8BCC643DFFB6FFF9FD19BC1A5B20A00F" blockId="19.[643,1134,296,383]" pageId="19" pageNumber="350">
|
||
<emphasis id="B907B82FFFB6FFF9FD19BC1A5DB4A030" bold="true" box="[643,754,296,320]" pageId="19" pageNumber="350">Remarks.</emphasis>
|
||
Piece of clear, golden amber that measures 11x10x
|
||
<quantity id="4C8BC9D8FFB6FFF9FCA7BC7A5C38A02F" box="[829,894,328,351]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="350" unit="mm" value="2.0">2 mm</quantity>
|
||
.The inclusion is complete, but well visible only in the lateral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |