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<mods:title id="6E2F93A1C6BBB1C6B06D17B2E244F561">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<mods:namePart id="9C9DEFC3C9B52CDBB62B41073962553E">Inkhavilay, Khamla</mods:namePart>
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<mods:namePart id="71FE21B4C7B65B9E290C8624EA36A40B">Sutcharit, Chirasak</mods:namePart>
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<treatment id="03DAD35F692FEF05FEEEA9E5FE292223" ID-DOI="http://doi.org/10.5281/zenodo.3848191" ID-GBIF-Taxon="164161062" ID-Zenodo-Dep="3848191" LSID="urn:lsid:plazi:treatment:03DAD35F692FEF05FEEEA9E5FE292223" httpUri="http://treatment.plazi.org/id/03DAD35F692FEF05FEEEA9E5FE292223" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10">
<subSubSection id="C36931C2692FEF01FEEEA9E5FC542278" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BCC6249692FEF01FEEEA9E5FC542278" blockId="9.[357,1231,1755,1819]" pageId="9" pageNumber="10">
<taxonomicName id="4C7319CA692FEF01FEEEA9E5FB072395" ID-CoL="D2N4" authority="Bavay &amp; Dautzenberg, 1909" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[357,1231,1755,1782]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="pervariabilis" subGenus="Amphidromus">
<emphasis id="B907BE5B692FEF01FEEEA9E5FCB12395" bold="true" box="[357,889,1755,1782]" pageId="9" pageNumber="10">
<emphasis id="B907BE5B692FEF01FEEEA9E5FDDA2395" bold="true" box="[357,530,1755,1782]" italics="true" pageId="9" pageNumber="10">Amphidromus</emphasis>
(
<emphasis id="B907BE5B692FEF01FDA8A9E5FD182395" bold="true" box="[547,720,1755,1782]" italics="true" pageId="9" pageNumber="10">Amphidromus</emphasis>
)
<emphasis id="B907BE5B692FEF01FD6AA9E5FCB12395" bold="true" box="[737,889,1755,1782]" italics="true" pageId="9" pageNumber="10">pervariabilis</emphasis>
</emphasis>
Bavay &amp; Dautzenberg, 1909
</taxonomicName>
<figureCitation id="13487ECC692FEF01FD13A83FFD292278" box="[664,737,1793,1819]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
,
<figureCitation id="13487ECC692FEF01FD64A83EFD342278" box="[751,764,1792,1819]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="9" pageNumber="10">5</figureCitation>
AL;
<tableCitation id="C6F157F2692FEF01FCCEA83FFC542278" box="[837,924,1793,1819]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="9" pageNumber="10" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36931C2692FEF01FF36A87AFC522288" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8BCC6249692FEF01FF36A87AFDD722E0" blockId="9.[189,1399,1860,1923]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB634567692FEF01FF36A87AFDD722E0" pageId="9" pageNumber="10">
<treatmentCitation id="0AD24458692FEF01FF36A87AFC2E223D" author="Bavay A. &amp; Dautzenberg P." box="[189,998,1860,1886]" page="247" pageId="9" pageNumber="10" year="1909">
<taxonomicName id="4C7319CA692FEF01FF36A87AFC2E223D" ID-CoL="D2N4" authority="Bavay &amp; Dautzenberg, 1909 b: 246 - 247" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="246 - 247" authorityYear="1909" box="[189,998,1860,1886]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692FEF01FF36A87AFDCA223D" box="[189,514,1860,1886]" italics="true" pageId="9" pageNumber="10">Amphidromus pervariabilis</emphasis>
<bibRefCitation id="EFE21FB8692FEF01FD80A87AFC2E223D" author="Bavay A. &amp; Dautzenberg P." box="[523,998,1860,1886]" pageId="9" pageNumber="10" pagination="229 - 251" refId="ref18119" refString="Bavay A. &amp; Dautzenberg P. 1909 b [1908]. Molluscorum terrestrium Tonkinorum diagnoses. Journal de Conchyliologie 56: 229 - 251. Available from http: // www. biodiversitylibrary. org / page / 16298225 [accessed 18 May 2017]." type="journal article" year="1909">Bavay &amp; Dautzenberg, 1909b: 246-247</bibRefCitation>
</taxonomicName>
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.
<typeStatus id="54C8DCEB692FEF01FC78A87AFBE5223D" box="[1011,1069,1860,1886]" pageId="9" pageNumber="10">Type</typeStatus>
locality: Ban-Lao, Muong- Kong, Pha-Long, Pac Kha.
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</paragraph>
<paragraph id="8BCC6249692FEF01FF36A892FC522288" blockId="9.[189,1398,1964,2027]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB634567692FEF01FF36A892FC522288" pageId="9" pageNumber="10">
<taxonomicName id="4C7319CA692FEF01FF36A892FE3022A5" ID-CoL="D2N4" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[189,504,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692FEF01FF36A892FE3022A5" box="[189,504,1964,1990]" italics="true" pageId="9" pageNumber="10">Amphidronus pervariabilis</emphasis>
</taxonomicName>
<treatmentCitation id="0AD24458692FEF01FD99A892FC1E22A5" author="Bavay A. &amp; Dautzenberg P." box="[530,982,1964,1990]" page="279" pageId="9" pageNumber="10" year="1909">
<bibRefCitation id="EFE21FB8692FEF01FD99A892FC1E22A5" author="Bavay A. &amp; Dautzenberg P." box="[530,982,1964,1990]" pageId="9" pageNumber="10" pagination="279 - 281" refId="ref18073" refString="Bavay A. &amp; Dautzenberg P. 1909 a. Description de coquilles nouvelles de lIndo-Chine. Journal de Conchyliologie 57: 279 - 281. Available from http: // biodiversitylibrary. org / page / 27393458 [accessed 9 Oct. 2013]." type="journal article" year="1909">Bavay &amp; Dautzenberg 1909a: 279-281</bibRefCitation>
</treatmentCitation>
, pl. 9, figs 110, pl. 10, figs 18. —
<treatmentCitation id="0AD24458692FEF01FF68A8EFFDFC2288" author="Laidlaw F. F. &amp; Solem A." box="[227,564,2000,2027]" page="527" pageId="9" pageNumber="10" year="1961">
<bibRefCitation id="EFE21FB8692FEF01FF68A8EFFDFC2288" author="Laidlaw F. F. &amp; Solem A." box="[227,564,2000,2027]" pageId="9" pageNumber="10" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem 1961: 527</bibRefCitation>
</treatmentCitation>
, 528. —
<treatmentCitation id="0AD24458692FEF01FD14A8EFFC5E2288" author="Richardson L." box="[671,918,2000,2027]" page="15" pageId="9" pageNumber="10" year="1985">
<bibRefCitation id="EFE21FB8692FEF01FD14A8EFFC5E2288" author="Richardson L." box="[671,918,2000,2027]" pageId="9" pageNumber="10" pagination="1 - 479" refId="ref19668" refString="Richardson L. 1985. Camaenidae: catalog of species. Tryonia 12: 1 - 479." type="journal article" year="1985">Richardson 1985: 15</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF0C32C1692CEF02FF36A988FB7F2289" ID-DOI="http://doi.org/10.5281/zenodo.3829754" ID-Zenodo-Dep="3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1718,1744]" targetBox="[218,1375,265,1702]" targetPageId="10">
<paragraph id="8BCC6249692CEF02FF36A988FB7F2289" blockId="10.[189,1399,1718,2027]" pageId="10" pageNumber="11">
<emphasis id="B907BE5B692CEF02FF36A988FEDA23B2" bold="true" box="[189,274,1718,1745]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
Shells of
<taxonomicName id="4C7319CA692CEF02FE16A989FB9523B2" authority="Bavay &amp; Dautzenberg, 1909" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[413,1117,1719,1745]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692CEF02FE16A989FD2F23B2" box="[413,743,1719,1745]" italics="true" pageId="10" pageNumber="11">Amphidromus pervariabilis</emphasis>
Bavay &amp; Dautzenberg, 1909
</taxonomicName>
.
<emphasis id="B907BE5B692CEF02FBFBA988FB6F23B3" bold="true" box="[1136,1191,1718,1744]" pageId="10" pageNumber="11">AB</emphasis>
. Syntype of the nominotypical form (MNHM-IM-2000-2049).
<emphasis id="B907BE5B692CEF02FD74A9E4FCDC2397" bold="true" box="[767,788,1754,1780]" pageId="10" pageNumber="11">C</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FCFDA9E4FC222397" box="[886,1002,1754,1780]" italics="true" pageId="10" pageNumber="11">bifasciata</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059).
<emphasis id="B907BE5B692CEF02FDD9A9C3FDAF2274" bold="true" box="[594,615,1789,1815]" pageId="10" pageNumber="11">D</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FD3FA9C0FCF52274" box="[692,829,1790,1815]" italics="true" pageId="10" pageNumber="11">goniostoma</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058).
<emphasis id="B907BE5B692CEF02FE03A81EFE542259" bold="true" box="[392,412,1824,1850]" pageId="10" pageNumber="11">E</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FE7CA81EFD992259" box="[503,593,1824,1850]" italics="true" pageId="10" pageNumber="11">lilacina</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052).
<emphasis id="B907BE5B692CEF02FF36A87AFF07223D" bold="true" box="[189,207,1860,1886]" pageId="10" pageNumber="11">F</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FE94A87AFEAD223E" box="[287,357,1860,1885]" italics="true" pageId="10" pageNumber="11">minor</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050).
<emphasis id="B907BE5B692CEF02FBE5A87DFB4D223E" bold="true" box="[1134,1157,1859,1885]" pageId="10" pageNumber="11">G</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FB5EA87DFAA1223E" box="[1237,1385,1859,1885]" italics="true" pageId="10" pageNumber="11">monozonalis</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057).
<emphasis id="B907BE5B692CEF02FC25A859FC0D22E2" bold="true" box="[942,965,1895,1921]" pageId="10" pageNumber="11">H</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FB9FA859FB9022E2" box="[1044,1112,1895,1921]" italics="true" pageId="10" pageNumber="11">obesa</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053).
<emphasis id="B907BE5B692CEF02FD38A8B4FD7622C7" bold="true" box="[691,702,1930,1956]" pageId="10" pageNumber="11">I</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FC97A8B5FC4222C7" box="[796,906,1931,1956]" italics="true" pageId="10" pageNumber="11">protracta</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051).
<emphasis id="B907BE5B692CEF02FD88A893FDDA22A4" bold="true" box="[515,530,1965,1991]" pageId="10" pageNumber="11">J</emphasis>
. Var. “
<emphasis id="B907BE5B692CEF02FDF4A893FD1122A4" box="[639,729,1965,1991]" italics="true" pageId="10" pageNumber="11">tricolor</emphasis>
” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054).
<emphasis id="B907BE5B692CEF02FE09A8EEFE732289" bold="true" box="[386,443,2000,2026]" pageId="10" pageNumber="11">KL</emphasis>
. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014).
</paragraph>
</caption>
<caption id="DF0C32C1692DEF03FF36A858FB032289" ID-DOI="http://doi.org/10.5281/zenodo.3829756" ID-Zenodo-Dep="3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1894,1920]" targetBox="[189,1399,348,1849]" targetPageId="11">
<paragraph id="8BCC6249692DEF03FF36A858FB032289" blockId="11.[189,1399,1894,2026]" pageId="11" pageNumber="12">
<emphasis id="B907BE5B692DEF03FF36A858FECF22E3" bold="true" box="[189,263,1894,1920]" pageId="11" pageNumber="12">Fig. 6.</emphasis>
Genitalia of
<taxonomicName id="4C7319CA692DEF03FE13A858FDF422E3" authorityName="Albers" authorityYear="1850" box="[408,572,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="B907BE5B692DEF03FE13A858FDF422E3" box="[408,572,1894,1920]" italics="true" pageId="11" pageNumber="12">Amphidromus</emphasis>
</taxonomicName>
spp.
<emphasis id="B907BE5B692DEF03FDFFA858FD6322E3" bold="true" box="[628,683,1894,1920]" pageId="11" pageNumber="12">AB</emphasis>
.
<taxonomicName id="4C7319CA692DEF03FD3DA858FB5C22E3" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[694,1172,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B692DEF03FD3DA858FC3522E3" box="[694,1021,1894,1920]" italics="true" pageId="11" pageNumber="12">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB8692DEF03FB8AA858FB5C22E3" author="Fulton H. C." box="[1025,1172,1894,1920]" pageId="11" pageNumber="12" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012).
<emphasis id="B907BE5B692DEF03FEFCA892FE7822A5" bold="true" box="[375,432,1964,1990]" pageId="11" pageNumber="12">CD</emphasis>
.
<taxonomicName id="4C7319CA692DEF03FE4BA893FCD522A4" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[448,797,1965,1991]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="syndromoideus" status="sp. nov.">
<emphasis id="B907BE5B692DEF03FE4BA893FCD522A4" box="[448,797,1965,1991]" italics="true" pageId="11" pageNumber="12">Amphidromus syndromoideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340320692DEF03FCAFA893FC4A22A4" box="[804,898,1965,1991]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019).
</paragraph>
</caption>
<subSubSection id="C36931C2692AEF05FF36AE35FD7C2680" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BCC6249692AEF04FF36AE35FE622446" blockId="12.[189,426,267,293]" box="[189,426,267,293]" pageId="12" pageNumber="13">
<heading id="D084D525692AEF04FF36AE35FE622446" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B907BE5B692AEF04FF36AE35FE622446" bold="true" box="[189,426,267,293]" pageId="12" pageNumber="13">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249692AEF04FF61AE04FE512430" blockId="12.[189,1398,313,446]" box="[234,409,313,340]" pageId="12" pageNumber="13">
<emphasis id="B907BE5B692AEF04FF61AE04FE512430" bold="true" box="[234,409,313,340]" pageId="12" pageNumber="13">
<typeStatus id="54C8DCEB692AEF04FF61AE04FEEF2437" box="[234,295,314,340]" pageId="12" pageNumber="13">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BCC6249692AEF04FF36AE63FDF524DE" blockId="12.[189,1398,313,446]" pageId="12" pageNumber="13">
<collectingCountry id="F36422D9692AEF04FF36AE63FE812414" box="[189,329,349,375]" name="Vietnam" pageId="12" pageNumber="13">VIETNAM</collectingCountry>
: 2 shells (
<figureCitation id="13487ECC692AEF04FE37AE62FDC82414" box="[444,512,348,375]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
AB),
<typeStatus id="54C8DCEB692AEF04FDDAAE63FD702414" box="[593,696,349,375]" pageId="12" pageNumber="13" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7319CA692AEF04FD55AE63FC5E2414" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[734,918,349,375]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" sensu="stricto" species="pervariabilis">
<emphasis id="B907BE5B692AEF04FD55AE63FC5E2414" box="[734,918,349,375]" italics="true" pageId="12" pageNumber="13">A. pervariabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340320692AEF04FC16AE63FC1B2414" box="[925,979,349,375]" pageId="12" pageNumber="13" sensu="stricto">s.str.</taxonomicNameLabel>
MNHM-IM-
<date id="FFCD4489692AEF04FBFFAE63FB3A2414" box="[1140,1266,349,375]" pageId="12" pageNumber="13" value="2000" valueMax="2049">2000-2049</date>
, from Ban- Lao [
<geoCoordinate id="EE47048E692AEF04FF70AEBEFEBA24F9" box="[251,370,382,410]" degrees="19" direction="north" minutes="39" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="19.65">19° 39 N</geoCoordinate>
,
<geoCoordinate id="EE47048E692AEF04FEF5AEBEFDC824F9" box="[382,512,382,410]" degrees="103" direction="east" minutes="19" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="103.316666">103° 19 E</geoCoordinate>
], Muong-Kong [
<geoCoordinate id="EE47048E692AEF04FD4EAEBEFCF424F9" box="[709,828,382,410]" degrees="19" direction="north" minutes="33" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="19.55">19° 33 N</geoCoordinate>
,
<geoCoordinate id="EE47048E692AEF04FCC2AEBEFC0324F9" box="[841,971,382,410]" degrees="104" direction="east" minutes="44" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="104.73333">104° 44 E</geoCoordinate>
], Pha-Long [
<geoCoordinate id="EE47048E692AEF04FBE8AEBEFB1324F9" box="[1123,1243,382,410]" degrees="22" direction="north" minutes="48" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="22.8">22° 48 N</geoCoordinate>
,
<geoCoordinate id="EE47048E692AEF04FB63AEBEFAA124F9" box="[1256,1385,382,410]" degrees="104" direction="east" minutes="14" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="104.23333">104° 14 E</geoCoordinate>
], Pac Kha [
<geoCoordinate id="EE47048E692AEF04FEA4AE9DFE6C24DD" box="[303,420,417,446]" degrees="21" direction="north" minutes="00" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="21.0">21° 00 N</geoCoordinate>
,
<geoCoordinate id="EE47048E692AEF04FE3BAE9DFDF924DE" box="[432,561,417,445]" degrees="104" direction="east" minutes="35" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="104.583336">104° 35 E</geoCoordinate>
].
</paragraph>
<paragraph id="8BCC6249692AEF04FF36AEDBFCE12741" blockId="12.[189,1399,484,546]" pageId="12" pageNumber="13">
<collectingCountry id="F36422D9692AEF04FF36AEDBFE81249C" box="[189,329,485,511]" name="Vietnam" pageId="12" pageNumber="13">VIETNAM</collectingCountry>
: 2 shells (
<figureCitation id="13487ECC692AEF04FE49AEDAFDE8249C" box="[450,544,484,511]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="12" pageNumber="13">Fig. 5C</figureCitation>
),
<typeStatus id="54C8DCEB692AEF04FDBCAEDBFD56249C" box="[567,670,485,511]" pageId="12" pageNumber="13" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7319CA692AEF04FD42AEDBFC4B249D" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[713,899,484,511]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692AEF04FD42AEDBFC4B249D" box="[713,899,484,511]" italics="true" pageId="12" pageNumber="13">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692AEF04FC46AEDAFB89249D" box="[973,1089,484,510]" italics="true" pageId="12" pageNumber="13">bifasciata</emphasis>
”, from Ban-Lao, Muong- Kong, Pha-Long, Pac Kha (MNHM-IM-
<date id="FFCD4489692AEF04FD16AD39FCD52741" box="[669,797,519,546]" pageId="12" pageNumber="13" value="2000" valueMax="2059">2000-2059</date>
).
</paragraph>
<paragraph id="8BCC6249692AEF04FF36AD77FDA127E4" blockId="12.[189,1399,583,647]" pageId="12" pageNumber="13">
<collectingCountry id="F36422D9692AEF04FF36AD77FE812700" box="[189,329,585,611]" name="Vietnam" pageId="12" pageNumber="13">VIETNAM</collectingCountry>
: 1 shell (
<figureCitation id="13487ECC692AEF04FE27AD77FDC02707" box="[428,520,585,612]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="12" pageNumber="13">Fig. 5D</figureCitation>
),
<typeStatus id="54C8DCEB692AEF04FD97AD74FDBF2707" box="[540,631,586,612]" pageId="12" pageNumber="13" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C7319CA692AEF04FD10AD74FC9A2700" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[667,850,585,612]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692AEF04FD10AD74FC9A2700" box="[667,850,585,612]" italics="true" pageId="12" pageNumber="13">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692AEF04FC1EAD74FBD62700" box="[917,1054,586,611]" italics="true" pageId="12" pageNumber="13">goniostoma</emphasis>
”, from Phong-Tho,
<geoCoordinate id="EE47048E692AEF04FA8CAD79FABA2707" box="[1287,1394,583,612]" degrees="22" direction="north" minutes="34" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="22.566668">22°34 N</geoCoordinate>
,
<geoCoordinate id="EE47048E692AEF04FF36AD54FEFA27E4" box="[189,306,618,647]" degrees="103" direction="east" minutes="22" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="103.36667">103°22 E</geoCoordinate>
(MNHM-IM-
<date id="FFCD4489692AEF04FE56AD52FD9627E4" box="[477,606,620,647]" pageId="12" pageNumber="13" value="2000" valueMax="2058">2000-2058</date>
).
</paragraph>
<caption id="DF0C32C1692AEF04FF36A821FCC5228A" ID-DOI="http://doi.org/10.5281/zenodo.3829758" ID-Zenodo-Dep="3829758" httpUri="https://zenodo.org/record/3829758/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1823,1849]" targetBox="[196,1393,710,1775]" targetPageId="12">
<paragraph id="8BCC6249692AEF04FF36A821FCC5228A" blockId="12.[189,1399,1823,2025]" pageId="12" pageNumber="13">
<emphasis id="B907BE5B692AEF04FF36A821FEC3225A" bold="true" box="[189,267,1823,1849]" pageId="12" pageNumber="13">Fig. 7.</emphasis>
SEM images of the radula.
<emphasis id="B907BE5B692AEF04FDDCA821FD58225A" bold="true" box="[599,656,1823,1849]" pageId="12" pageNumber="13">AC</emphasis>
.
<taxonomicName id="4C7319CA692AEF04FD2BA821FB41225A" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[672,1161,1823,1849]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B692AEF04FD2BA821FC22225A" box="[672,1002,1823,1849]" italics="true" pageId="12" pageNumber="13">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB8692AEF04FC79A821FB41225A" author="Fulton H. C." box="[1010,1161,1823,1849]" pageId="12" pageNumber="13" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
from Ban Phavong, Khammouan, Laos (CUMZ 7012).
<emphasis id="B907BE5B692AEF04FDFEA87CFD63223F" bold="true" box="[629,683,1858,1884]" pageId="12" pageNumber="13">DF</emphasis>
.
<taxonomicName id="4C7319CA692AEF04FD4BA87CFBEA223F" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[704,1058,1858,1884]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="syndromoideus" status="sp. nov.">
<emphasis id="B907BE5B692AEF04FD4BA87CFBEA223F" box="[704,1058,1858,1884]" italics="true" pageId="12" pageNumber="13">Amphidromus syndromoideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340320692AEF04FBA4A87DFB5A223E" box="[1071,1170,1859,1885]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (CUMZ 7019).
<emphasis id="B907BE5B692AEF04FE9AA85BFE8A221C" bold="true" box="[273,322,1893,1919]" pageId="12" pageNumber="13">GI</emphasis>
.
<taxonomicName id="4C7319CA692AEF04FED9A85BFCE222E3" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[338,810,1893,1920]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="areolatus">
<emphasis id="B907BE5B692AEF04FED9A85BFDA4221C" box="[338,620,1893,1919]" italics="true" pageId="12" pageNumber="13">Amphidromus areolatus</emphasis>
(Pfeiffer, 1861)
</taxonomicName>
from Thad Fek, Attapue, Laos (CUMZ 7023). A, D, G = central tooth with the first to fifth to eighth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by C.
</paragraph>
</caption>
<paragraph id="8BCC6249692BEF05FF36AE32FEB3242F" blockId="13.[189,1398,267,332]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AE32FE812445" box="[189,329,268,294]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
: 2 shells (
<figureCitation id="13487ECC692BEF05FE4DAE35FDEB2445" box="[454,547,267,294]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
),
<typeStatus id="54C8DCEB692BEF05FDB7AE32FD6B2445" box="[572,675,268,294]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7319CA692BEF05FD44AE32FC432446" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[719,907,267,294]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FD44AE32FC432446" box="[719,907,267,294]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692BEF05FC53AE35FBFA2446" box="[984,1074,267,293]" italics="true" pageId="13" pageNumber="14">lilacina</emphasis>
”, from Ban-Lao (MNHM- IM-
<date id="FFCD4489692BEF05FF67AE0FFEA7242F" box="[236,367,305,332]" pageId="13" pageNumber="14" value="2000" valueMax="2052">2000-2052</date>
).
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AE4BFEB324D6" blockId="13.[189,1398,373,437]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AE4BFE8124EC" box="[189,329,373,399]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
: 1 shell (
<figureCitation id="13487ECC692BEF05FE26AE4BFDCC24F3" box="[429,516,373,400]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5F</figureCitation>
),
<typeStatus id="54C8DCEB692BEF05FD92AE4BFDBC24EC" box="[537,628,373,399]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C7319CA692BEF05FD12AE4BFC9824EC" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[665,848,373,399]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FD12AE4BFC9824EC" box="[665,848,373,399]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692BEF05FC1FAE48FC1224EC" box="[916,986,374,399]" italics="true" pageId="13" pageNumber="14">minor</emphasis>
”, from unknown locality (MNHM- IM-2000-2050).
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AEE0FEB3277C" blockId="13.[189,1398,478,543]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AEE0FE81249B" box="[189,329,478,504]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
: 2 shells (
<figureCitation id="13487ECC692BEF05FE3CAEE0FDDB249A" box="[439,531,478,505]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5G</figureCitation>
),
<typeStatus id="54C8DCEB692BEF05FDACAEE1FD46249A" box="[551,654,479,505]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7319CA692BEF05FD3AAEE1FCA0249B" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[689,872,478,505]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FD3AAEE1FCA0249B" box="[689,872,478,505]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692BEF05FC20AEE0FBF7249B" box="[939,1087,478,504]" italics="true" pageId="13" pageNumber="14">monozonalis</emphasis>
”, from Ban-Lao (MNHM- IM-
<date id="FFCD4489692BEF05FF67AD3AFEA7277C" box="[236,367,516,543]" pageId="13" pageNumber="14" value="2000" valueMax="2057">2000-2057</date>
).
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AD78FDA127E4" blockId="13.[189,1399,580,647]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AD78FE812703" box="[189,329,582,608]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
: 1 shell (
<figureCitation id="13487ECC692BEF05FE30AD78FDD42702" box="[443,540,582,609]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5H</figureCitation>
),
<typeStatus id="54C8DCEB692BEF05FDBEAD79FD582702" box="[565,656,583,609]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C7319CA692BEF05FD34AD79FCB32703" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[703,891,582,609]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FD34AD79FCB32703" box="[703,891,582,609]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692BEF05FC43AD78FBC42703" box="[968,1036,582,608]" italics="true" pageId="13" pageNumber="14">obesa</emphasis>
”, from Muong-Bo,
<geoCoordinate id="EE47048E692BEF05FA89AD7AFABA2702" box="[1282,1394,580,609]" degrees="22" direction="north" minutes="24" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="22.4">22°24 N</geoCoordinate>
,
<geoCoordinate id="EE47048E692BEF05FF36AD54FEFA27E5" box="[189,306,618,646]" degrees="102" direction="east" minutes="49" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="102.816666">102°49 E</geoCoordinate>
(MNHM-IM-
<date id="FFCD4489692BEF05FE55AD52FD9527E4" box="[478,605,620,647]" pageId="13" pageNumber="14" value="2000" valueMax="2053">2000-2053</date>
).
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AD90FD12278C" blockId="13.[189,1398,686,751]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AD90FE8127AB" box="[189,329,686,712]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
: 1 shell (
<figureCitation id="13487ECC692BEF05FE24AD90FE3727AA" box="[431,511,686,713]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5I</figureCitation>
),
<typeStatus id="54C8DCEB692BEF05FD9EAD91FDB827AA" box="[533,624,687,713]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C7319CA692BEF05FD1DAD91FC8627AB" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[662,846,686,713]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FD1DAD91FC8627AB" box="[662,846,686,713]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692BEF05FC18AD91FBC927AB" box="[915,1025,687,712]" italics="true" pageId="13" pageNumber="14">protracta</emphasis>
”, from Ban-Lao, Muong-Kong, Pha-Long, Pac Kha (MNHM-IM-
<date id="FFCD4489692BEF05FDC5ADEAFD05278C" box="[590,717,724,751]" pageId="13" pageNumber="14" value="2000" valueMax="2051">2000-2051</date>
).
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AC28FEB32634" blockId="13.[189,1398,790,855]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AC28FE812653" box="[189,329,790,816]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
: 2 shells (
<figureCitation id="13487ECC692BEF05FE4CAC28FDD62652" box="[455,542,790,817]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5J</figureCitation>
),
<typeStatus id="54C8DCEB692BEF05FDB3AC29FD572652" box="[568,671,791,817]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7319CA692BEF05FD47AC29FC412653" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[716,905,790,817]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FD47AC29FC412653" box="[716,905,790,817]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
var. “
<emphasis id="B907BE5B692BEF05FC5DAC28FBF82653" box="[982,1072,790,816]" italics="true" pageId="13" pageNumber="14">tricolor</emphasis>
”, from Ban-Lao (MNHM- IM-
<date id="FFCD4489692BEF05FF67AC02FEB82634" box="[236,368,828,855]" pageId="13" pageNumber="14" value="2000" valueMax="2054">2000-2054</date>
).
</paragraph>
<paragraph id="8BCC6249692BEF05FF61AC40FE6E26FB" blockId="13.[189,1399,894,996]" box="[234,422,894,920]" pageId="13" pageNumber="14">
<emphasis id="B907BE5B692BEF05FF61AC40FE6E26FB" bold="true" box="[234,422,894,920]" pageId="13" pageNumber="14">Other material</emphasis>
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AC9AFD7C2680" blockId="13.[189,1399,894,996]" pageId="13" pageNumber="14">
<collectingCountry id="F36422D9692BEF05FF36AC9AFEC626DD" box="[189,270,932,958]" name="Laos" pageId="13" pageNumber="14">LAOS</collectingCountry>
: 3 shells (
<figureCitation id="13487ECC692BEF05FE07AC9AFE1B26DD" box="[396,467,931,958]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[218,1375,265,1702]" captionTargetId="figure@10.[218,1375,265,1702]" captionTargetPageId="10" captionText="Fig. 5. Shells of Amphidromus pervariabilis Bavay &amp; Dautzenberg, 1909. AB. Syntype of the nominotypical form (MNHM-IM-2000-2049). C. Var. “bifasciata” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2059). D. Var. “goniostoma” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2058). E. Var. “lilacina” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2052). F. Var. “minor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2050). G. Var. “monozonalis” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2057). H. Var. “obesa” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2053). I. Var. “protracta” Bavay &amp; Dautzenberg, 1909, syntype (MNHM-IM-2000-2051). J. Var. “tricolor” Bavay &amp; Dautzenberg, 1909, syntype (MNHM- IM-2000-2054). KL. Specimens from Khua District, Phongsaly, Laos (CUMZ 7014)." figureDoi="http://doi.org/10.5281/zenodo.3829754" httpUri="https://zenodo.org/record/3829754/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
KL), Km 34, road to Ban Namly, Khua District,
<collectingRegion id="49B7ACAB692BEF05FBB1AC9AFB7D26DD" box="[1082,1205,932,958]" country="Laos" name="Phongsali" pageId="13" pageNumber="14">Phongsaly</collectingRegion>
,
<geoCoordinate id="EE47048E692BEF05FB4FAC9AFABA26DD" box="[1220,1394,930,958]" degrees="21" direction="north" minutes="11" orientation="latitude" pageId="13" pageNumber="14" precision="1" seconds="55.5" value="21.19875">21°1155.5″ N</geoCoordinate>
,
<geoCoordinate id="EE47048E692BEF05FF36ACF7FEAD2687" box="[189,357,967,996]" degrees="102" direction="east" minutes="6" orientation="longitude" pageId="13" pageNumber="14" precision="1" seconds="40.2" value="102.11117">102°640.2″ E</geoCoordinate>
,
<quantity id="4C8BCFAC692BEF05FEFAACF7FE742687" box="[369,444,969,996]" metricMagnitude="2" metricUnit="m" metricValue="8.34" pageId="13" pageNumber="14" unit="m" value="834.0">834 m</quantity>
amsl (
<collectionCode id="ED62FA8C692BEF05FD82ACF7FDA82680" box="[521,608,969,995]" country="0" name="Cameroon University, Museum of Zoology" pageId="13" pageNumber="14">CUMZ</collectionCode>
7014).
</paragraph>
</subSubSection>
<subSubSection id="C36931C2692BEF05FF36AB35FE16205B" pageId="13" pageNumber="14" type="description">
<paragraph id="8BCC6249692BEF05FF36AB35FE862146" blockId="13.[189,334,1035,1061]" box="[189,334,1035,1061]" pageId="13" pageNumber="14">
<heading id="D084D525692BEF05FF36AB35FE862146" bold="true" box="[189,334,1035,1061]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B907BE5B692BEF05FF36AB35FE862146" bold="true" box="[189,334,1035,1061]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AB02FE16205B" blockId="13.[189,1399,1084,1336]" pageId="13" pageNumber="14">Shell chirally dimorphic, elongate conical, rather thick and glossy. Spire elongate conical; apex acute, with or without black spot on tip. Whorls 6 to 7 convex to smooth; suture wide and shallow; last whorl rounded to well rounded. Periostracum thin corneous; varix usually absent. Shell color varies from uniform white, yellowish to reddish brown or with narrow to wide brownish peripheral and umbilical bands. Parietal callus thin and transparent. Aperture ovate to oblique elongated; peristome expanded and not reflected; lip white, brownish or purplish. Columella straight, thick or thin and white or brownish. Umbilicus imperforated.</paragraph>
</subSubSection>
<subSubSection id="C36931C2692BEF05FF36AA5FFE292223" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BCC6249692BEF05FF36AA5FFEE72018" blockId="13.[189,303,1377,1403]" box="[189,303,1377,1403]" pageId="13" pageNumber="14">
<heading id="D084D525692BEF05FF36AA5FFEE72018" bold="true" box="[189,303,1377,1403]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B907BE5B692BEF05FF36AA5FFEE72018" bold="true" box="[189,303,1377,1403]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249692BEF05FF36AAAFFBE02094" blockId="13.[189,1399,1425,1527]" pageId="13" pageNumber="14">
This species tend to have a high diversity of shell colors, with eight infra-specific names being provisionally proposed to distinguish the distinct shell shape and color patterns. We examined all the
<typeStatus id="54C8DCEB692BEF05FF36AAE3FF382094" box="[189,240,1501,1527]" pageId="13" pageNumber="14">type</typeStatus>
specimens and recognized them as subspecific entities of one species.
</paragraph>
<paragraph id="8BCC6249692BEF05FF36A921FE292223" blockId="13.[189,1399,1567,1857]" pageId="13" pageNumber="14">
<bibRefCitation id="EFE21FB8692BEF05FF36A921FE212359" author="Laidlaw F. F. &amp; Solem A." box="[189,489,1567,1594]" pageId="13" pageNumber="14" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem (1961)</bibRefCitation>
recognized this species as a synonym of
<taxonomicName id="4C7319CA692BEF05FC56A91EFAF4235A" authority="Fulton, 1899" authorityName="Fulton" authorityYear="1899" box="[989,1340,1567,1594]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="dautzenbergi">
<emphasis id="B907BE5B692BEF05FC56A91EFB5F235A" box="[989,1175,1567,1594]" italics="true" pageId="13" pageNumber="14">A. dautzenbergi</emphasis>
Fulton, 1899
</taxonomicName>
. The latter species exhibits a thin shell, yellowish with a thin faded greenish streak near the umbilicus, a white subsutural band and an elongated ovate aperture, while
<taxonomicName id="4C7319CA692BEF05FCC5A955FBCF23E7" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[846,1031,1642,1669]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FCC5A955FBCF23E7" box="[846,1031,1642,1669]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
has a thick shell, monochrome or with dark brown spiral band and ovate aperture. However, the type locality of
<taxonomicName id="4C7319CA692BEF05FB0DA9AEFA8823C9" authorityName="Fulton" authorityYear="1899" box="[1158,1344,1680,1706]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="dautzenbergi">
<emphasis id="B907BE5B692BEF05FB0DA9AEFA8823C9" box="[1158,1344,1680,1706]" italics="true" pageId="13" pageNumber="14">A. dautzenbergi</emphasis>
</taxonomicName>
was “
<collectingCountry id="F36422D9692BEF05FF4CA98BFEEA23AC" box="[199,290,1717,1743]" name="Vietnam" pageId="13" pageNumber="14">Tonkin</collectingCountry>
”, the historical political division during the French colonial period, and it still is an uncertain boundary. The species is known from the
<typeStatus id="54C8DCEB692BEF05FD2AA9E5FCC02396" box="[673,776,1755,1781]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
only (
<bibRefCitation id="EFE21FB8692BEF05FCDAA9E5FB8B2395" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[849,1091,1755,1782]" pageId="13" pageNumber="14" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit
<emphasis id="B907BE5B692BEF05FC4FA9E2FC3E2396" box="[964,1014,1755,1781]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. 2015
</bibRefCitation>
: fig.
<quantity id="4C8BCFAC692BEF05FBF0A9E5FB522395" box="[1147,1178,1755,1782]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="13" pageNumber="14" unit="g" value="6.0">6g</quantity>
), no specimen was found in the present study, and, therefore, the taxonomic status of
<taxonomicName id="4C7319CA692BEF05FC41A83FFB4C2278" authorityName="Fulton" authorityYear="1899" box="[970,1156,1793,1819]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="dautzenbergi">
<emphasis id="B907BE5B692BEF05FC41A83FFB4C2278" box="[970,1156,1793,1819]" italics="true" pageId="13" pageNumber="14">A. dautzenbergi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7319CA692BEF05FB35A83FFABF2278" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[1214,1399,1793,1819]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B692BEF05FB35A83FFABF2278" box="[1214,1399,1793,1819]" italics="true" pageId="13" pageNumber="14">A. pervariabilis</emphasis>
</taxonomicName>
remains to be confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>