treatments-xml/data/03/DA/D3/03DAD35F6928EF06FE08AB67FEBC2258.xml

<document id="050DEC9D3D4C8CCB7CD368C01C416C05" ID-CLB-Dataset="22089" ID-DOI="10.5852/ejt.2017.330" ID-GBIF-Dataset="14d5ad89-9e69-45f8-bc61-e14ca9deafbd" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589572621568" checkinUser="carolina" docAuthor="Inkhavilay, Khamla, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2017" docId="03DAD35F6928EF06FE08AB67FEBC2258" docLanguage="en" docName="ejt-2017-330.pdf.imf" docOrigin="European Journal of Taxonomy 330" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Amphidromus (Amphidromus) protania Lehmann &amp; Maassen 2004" docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FFE3AB276926EF08FF8BAF3EFFC82563" masterDocTitle="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species" masterLastPageNumber="40" masterPageNumber="1" pageNumber="15" updateTime="1698839926330" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A6246C2CE7D193826F312D90EFC51A31" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="00D38C38F10F697968085637D26FBEB0">
<mods:title id="8FB75F587D233CF394F0B1DB50DC5F83">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="80C846F5792A110B782C9298274943B1" type="personal">
<mods:role id="93A42A8E6F2710C3197D76779E9159A5">
<mods:roleTerm id="83DA93474B47D6700509FB7DCD4DEDAF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86198BD73F5F9C3E01A9BCE79D99E0B2">Inkhavilay, Khamla</mods:namePart>
<mods:nameIdentifier id="AE38630CF2C052EA6B6099F34D2FDB24" type="LSID">2E620B9C-48BA-48E6-A017-FE8ED2AB0E59</mods:nameIdentifier>
<mods:affiliation id="0B6A356515BB6C0249D3D0DF5AD68CCF">Email: kinkhavilay @ gmail. com &amp; urn: lsid: zoobank. org: author: 2 E 620 B 9 C- 48 BA- 48 E 6 - A 017 - FE 8 ED 2 AB 0 E 59</mods:affiliation>
<mods:nameIdentifier id="E72B708D416FBD3D8DF64B037969E60E" type="email">kinkhavilay@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="DAA659B8C5B842B45D5071182DD55B76" type="personal">
<mods:role id="5DA6FE18A85FAE25F490E12532EAB116">
<mods:roleTerm id="095CDAF8A1DBA172764469999ADF7EC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="087FF942554C2B0FEDF1BF92EBCF9B46">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="15962E605A799B12EAF2805621A7DFE3" type="LSID">C2E2FA6B-A3F9-4F33-B447-B59B1BD322D4</mods:nameIdentifier>
<mods:affiliation id="F76BB2D1F1F9101559BB307A953A433B">urn: lsid: zoobank. org: author: C 2 E 2 FA 6 B-A 3 F 9 - 4 F 33 - B 447 - B 59 B 1 BD 322 D 4 &amp; Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="A30644DD64D72B1EAD80153E45811D2E" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="09A09C422255323EFF52F71FE53382E2" type="personal">
<mods:role id="206B09C5A79C8999833016A35C350DC3">
<mods:roleTerm id="DF35BBFA1E7ED30496AADED621A40CB2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA2548BA699A3480B8072B10F05F12C7">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="1A32C914058B3E1CE80E47E9DD661371" type="LSID">AC935098-D901-4F35-A414-4B0D4FE44E79</mods:nameIdentifier>
<mods:affiliation id="BF000BADBFE6A81D53EF317CC46BBFCE">urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="44521FDDC636AA7FA7D9F35C5466A579" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1C33C206F847FF0F164B78E0C31CA342">text</mods:typeOfResource>
<mods:relatedItem id="EE80FFE8BA53B7C3F3248E9B53D5A47D" type="host">
<mods:titleInfo id="558FC1F1BCFCF31DBCFD50396AB337D3">
<mods:title id="DEFCC4D1521B20A69263C009DB7782DD">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3EEBF4CDEECFF7D25BE1CFE97F9B47E1">
<mods:date id="63440A9B6C517BEC6703B570E5002F75">2017</mods:date>
<mods:detail id="7BBABFC0728267CFC9BFFD7613B54BF3" type="pubDate">
<mods:number id="1AA0EEB93833247DA8EA755BFE81D8F0">2017-06-13</mods:number>
</mods:detail>
<mods:detail id="49DA796F6E8FFDF367A9AAB92F338D2C" type="issue">
<mods:number id="CE71DEB7BE43A1C1D8FA7452A4F471A6">330</mods:number>
</mods:detail>
<mods:extent id="DC99FDDDB6351336C7D7B16BA80914CB" unit="page">
<mods:start id="2D2AA5886CE729458D812EF38BD31CF5">1</mods:start>
<mods:end id="BCE631665EE65A53F8F36C10F4F0B1A5">40</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7D00A87745D712A87D9EC9FD3EC7EF29">journal article</mods:classification>
<mods:identifier id="E3C183E7BCFF12317AD48D5608FA9CAC" type="CLB-Dataset">22089</mods:identifier>
<mods:identifier id="9B4F84C2896256C37098F933CA67A8D8" type="DOI">10.5852/ejt.2017.330</mods:identifier>
<mods:identifier id="83D6C50544D4E81521B2B6C284C68577" type="GBIF-Dataset">14d5ad89-9e69-45f8-bc61-e14ca9deafbd</mods:identifier>
<mods:identifier id="DDB37ECBFC89731EF632AE26591256D6" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="B750DA970D1702906CD36FD9CA380FCB" type="Zenodo-Dep">3829744</mods:identifier>
<mods:identifier id="A9D473FF26E84A9D0CE44FE5F0D3FF9C" type="ZooBank">FDFA7FAC-FC92-404B-9058-A426DAB6C693</mods:identifier>
</mods:mods>
<treatment id="03DAD35F6928EF06FE08AB67FEBC2258" ID-DOI="http://doi.org/10.5281/zenodo.3848207" ID-GBIF-Taxon="164161066" ID-Zenodo-Dep="3848207" LSID="urn:lsid:plazi:treatment:03DAD35F6928EF06FE08AB67FEBC2258" httpUri="http://treatment.plazi.org/id/03DAD35F6928EF06FE08AB67FEBC2258" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="C36931C26928EF06FE08AB67FB792117" box="[387,1201,1113,1140]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BCC62496928EF06FE08AB67FB792117" blockId="14.[387,1201,1113,1175]" box="[387,1201,1113,1140]" pageId="14" pageNumber="15">
<heading id="D084D5256928EF06FE08AB67FB792117" box="[387,1201,1113,1140]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C7319CA6928EF06FE08AB67FB792117" ID-CoL="D2NC" authority="Lehmann &amp; Maassen, 2004" authorityName="Lehmann &amp; Maassen" authorityYear="2004" box="[387,1201,1113,1140]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="protania" subGenus="Amphidromus">
<emphasis id="B907BE5B6928EF06FE08AB67FDF82117" bold="true" box="[387,560,1113,1140]" italics="true" pageId="14" pageNumber="15">Amphidromus</emphasis>
<emphasis id="B907BE5B6928EF06FDBCAB67FD3F2110" bold="true" box="[567,759,1113,1140]" pageId="14" pageNumber="15">
(
<emphasis id="B907BE5B6928EF06FDCAAB67FD262117" bold="true" box="[577,750,1113,1140]" italics="true" pageId="14" pageNumber="15">Amphidromus</emphasis>
)
</emphasis>
<emphasis id="B907BE5B6928EF06FD75AB67FCAD2110" bold="true" box="[766,869,1113,1139]" italics="true" pageId="14" pageNumber="15">protania</emphasis>
<bibRefCitation id="EFE21FB86928EF06FCE6AB67FB792117" author="Lehmann H. &amp; Maassen W. J. M." box="[877,1201,1113,1140]" pageId="14" pageNumber="15" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">Lehmann &amp; Maassen, 2004</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36931C26928EF06FD57AB43FC9F21F4" box="[732,855,1149,1175]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BCC62496928EF06FD57AB43FC9F21F4" blockId="14.[387,1201,1113,1175]" box="[732,855,1149,1175]" pageId="14" pageNumber="15">
<figureCitation id="13487ECC6928EF06FD57AB43FCED21F4" box="[732,805,1149,1175]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="14" pageNumber="15">Figs 1</figureCitation>
, 
<figureCitation id="13487ECC6928EF06FCB8AB43FC9F21F4" box="[819,855,1149,1175]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="14" pageNumber="15">4H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36931C26928EF06FF36AB80FB512198" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BCC62496928EF06FF36AB80FB512198" blockId="14.[189,1399,1214,1276]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB6345676928EF06FF36AB80FB512198" pageId="14" pageNumber="15">
<treatmentCitation id="0AD244586928EF06FF36AB80FB8521BB" author="Lehmann H. &amp; Maassen W. J. M." box="[189,1101,1214,1241]" page="20" pageId="14" pageNumber="15" year="2004">
<taxonomicName id="4C7319CA6928EF06FF36AB80FB8521BB" ID-CoL="D2NC" authority="Lehmann &amp; Maassen, 2004: 17 - 20" authorityName="Lehmann &amp; Maassen" authorityPageNumber="17 - 20" authorityYear="2004" box="[189,1101,1214,1241]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="protania" subGenus="Amphidromus">
<emphasis id="B907BE5B6928EF06FF36AB80FEA921BB" box="[189,353,1214,1240]" italics="true" pageId="14" pageNumber="15">Amphidromus</emphasis>
(
<emphasis id="B907BE5B6928EF06FEFDAB80FDD221BB" box="[374,538,1214,1240]" italics="true" pageId="14" pageNumber="15">Amphidromus</emphasis>
) 
<emphasis id="B907BE5B6928EF06FDA5AB81FD5A21BB" box="[558,658,1215,1240]" italics="true" pageId="14" pageNumber="15">protania</emphasis>
<bibRefCitation id="EFE21FB86928EF06FD16AB80FB8521BB" author="Lehmann H. &amp; Maassen W. J. M." box="[669,1101,1214,1241]" pageId="14" pageNumber="15" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">Lehmann &amp; Maassen, 2004: 17–20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1–4. 
<typeStatus id="54C8DCEB6928EF06FB5AAB81FAC321BA" box="[1233,1291,1215,1241]" pageId="14" pageNumber="15">Type</typeStatus>
locality: South 
<collectingCountry id="F36422D96928EF06FEA4ABDCFEA3219F" box="[303,363,1250,1276]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
, 
<collectingRegion id="49B7ACAB6928EF06FEFDABDFFD8D2198" box="[374,581,1249,1275]" country="Laos" name="Salavan" pageId="14" pageNumber="15">Salavan Province</collectingRegion>
, near Ban Donxé Village, E bank of Se Don River.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36931C26928EF06FF36AA1DFB122008" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BCC62496928EF06FF36AA1DFE62205E" blockId="14.[189,426,1315,1341]" box="[189,426,1315,1341]" pageId="14" pageNumber="15">
<heading id="D084D5256928EF06FF36AA1DFE62205E" bold="true" box="[189,426,1315,1341]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B907BE5B6928EF06FF36AA1DFE62205E" bold="true" box="[189,426,1315,1341]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496928EF06FF36AA6FFB122008" blockId="14.[189,1242,1361,1388]" box="[189,1242,1361,1388]" pageId="14" pageNumber="15">
<collectingCountry id="F36422D96928EF06FF36AA6FFEC62008" box="[189,270,1361,1387]" name="Laos" pageId="14" pageNumber="15">
<collectionCode id="ED62FA8C6928EF06FF36AA6FFEC62008" box="[189,270,1361,1387]" pageId="14" pageNumber="15">LAOS</collectionCode>
</collectingCountry>
: Dextral, 
<typeStatus id="54C8DCEB6928EF06FE0BAA6FFE2F2008" box="[384,487,1361,1387]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED62FA8C6928EF06FE7EAA6FFD872008" box="[501,591,1361,1387]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="15" type="Museum">RMNH</collectionCode>
98143, 
<figureCitation id="13487ECC6928EF06FD24AA6FFCC52008" box="[687,781,1361,1387]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="14" pageNumber="15">
Fig. 4 
<collectionCode id="ED62FA8C6928EF06FD7EAA6FFCC52008" box="[757,781,1361,1387]" country="Finland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="14" pageNumber="15" type="Herbarium">H</collectionCode>
</figureCitation>
); 4D shells, 
<typeStatus id="54C8DCEB6928EF06FC15AA6CFBC6200F" box="[926,1038,1362,1388]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED62FA8C6928EF06FB95AA6FFBB02008" box="[1054,1144,1361,1387]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="15" type="Museum">RMNH</collectionCode>
98144).
</paragraph>
</subSubSection>
<subSubSection id="C36931C26928EF06FF36AAADFEBC2258" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BCC62496928EF06FF36AAADFEE720CE" blockId="14.[189,303,1427,1453]" box="[189,303,1427,1453]" pageId="14" pageNumber="15">
<heading id="D084D5256928EF06FF36AAADFEE720CE" bold="true" box="[189,303,1427,1453]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B907BE5B6928EF06FF36AAADFEE720CE" bold="true" box="[189,303,1427,1453]" pageId="14" pageNumber="15">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496928EF06FF36AAFCFEBC2258" blockId="14.[189,1399,1473,1851]" pageId="14" pageNumber="15">
No specimen was found in this survey. 
<taxonomicName id="4C7319CA6928EF06FD16AAFFFC6720B8" authorityName="Lehmann &amp; Maassen" authorityYear="2004" box="[669,943,1473,1499]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="protania">
<emphasis id="B907BE5B6928EF06FD16AAFFFC6720B8" box="[669,943,1473,1499]" italics="true" pageId="14" pageNumber="15">Amphidromus protania</emphasis>
</taxonomicName>
has a very similar shell shape to that the dipterocarp forest species 
<taxonomicName id="4C7319CA6928EF06FD97AADBFD65209D" authorityName="Geret" authorityYear="1912" box="[540,685,1508,1535]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6928EF06FD97AADBFD65209D" box="[540,685,1508,1535]" italics="true" pageId="14" pageNumber="15">A. givenchyi</emphasis>
</taxonomicName>
. The genitalia contain a long and coiled proximal part of the epiphallus, with a relatively short appendix, as in 
<taxonomicName id="4C7319CA6928EF06FC89A936FC5A2341" authorityName="Geret" authorityYear="1912" box="[770,914,1544,1570]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6928EF06FC89A936FC5A2341" box="[770,914,1544,1570]" italics="true" pageId="14" pageNumber="15">A. givenchyi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE21FB86928EF06FC2AA936FB2B2341" author="Lehmann H. &amp; Maassen W. J. M." box="[929,1251,1544,1570]" pageId="14" pageNumber="15" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">
Lehmann &amp; Maassen 
<date id="FFCD44896928EF06FB2EA936FB2B2341" box="[1189,1251,1544,1570]" pageId="14" pageNumber="15" value="2004">2004</date>
</bibRefCitation>
; 
<bibRefCitation id="EFE21FB86928EF06FB64A936FE982325" author="Sutcharit C. &amp; Panha S." pageId="14" pageNumber="15" pagination="1 - 30" refId="ref19880" refString="Sutcharit C. &amp; Panha S. 2006. Taxonomic review of the tree snail Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Thailand and adjacent areas: Subgenus Amphidromus. Journal of Molluscan Studies 72: 1 - 30. https: // doi. org / 10.1093 / mollus / eyi 044" type="journal article" year="2006">
Sutcharit &amp; Panha 
<date id="FFCD44896928EF06FE9BA915FE982325" box="[272,336,1579,1606]" pageId="14" pageNumber="15" value="2006">2006</date>
</bibRefCitation>
). However, the characters distinguishing it from 
<taxonomicName id="4C7319CA6928EF06FC2CA915FBF42326" authorityName="Geret" authorityYear="1912" box="[935,1084,1579,1605]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6928EF06FC2CA915FBF42326" box="[935,1084,1579,1605]" italics="true" pageId="14" pageNumber="15">A. givenchyi</emphasis>
</taxonomicName>
are the relatively smaller shell (height 
<quantity id="4C8BCFAC6928EF06FED5A970FE28230A" box="[350,480,1614,1641]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="14" pageNumber="15" unit="mm" value="28.5" valueMax="31.0" valueMin="26.0">26–31 mm</quantity>
in 
<taxonomicName id="4C7319CA6928EF06FD86A970FD5E230B" authorityName="Lehmann &amp; Maassen" authorityYear="2004" box="[525,662,1614,1640]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="protania">
<emphasis id="B907BE5B6928EF06FD86A970FD5E230B" box="[525,662,1614,1640]" italics="true" pageId="14" pageNumber="15">A. protania</emphasis>
</taxonomicName>
; 
<tableCitation id="C6F157F26928EF06FD23A970FCC9230B" box="[680,769,1614,1640]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="14" pageNumber="15" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
), the yellowish shell color as well as the brownish subsutural and spiral bands on the periphery. Moreover, 
<bibRefCitation id="EFE21FB86928EF06FCEFA94FFB7323E8" author="Lehmann H. &amp; Maassen W. J. M." box="[868,1211,1649,1676]" pageId="14" pageNumber="15" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">
Lehmann &amp; Maassen (
<date id="FFCD44896928EF06FBFEA94FFB7B23E8" box="[1141,1203,1649,1675]" pageId="14" pageNumber="15" value="2004">2004</date>
)
</bibRefCitation>
mentioned that the brownish banding pattern had less variation. However, a brownish spiral band on the periphery can also be observed in juvenile 
<taxonomicName id="4C7319CA6928EF06FD85A986FD5723B1" authorityName="Geret" authorityYear="1912" box="[526,671,1720,1746]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6928EF06FD85A986FD5723B1" box="[526,671,1720,1746]" italics="true" pageId="14" pageNumber="15">A. givenchyi</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFE21FB86928EF06FD54A986FC3623B1" author="Sutcharit C. &amp; Panha S." box="[735,1022,1720,1746]" pageId="14" pageNumber="15" pagination="1 - 30" refId="ref19880" refString="Sutcharit C. &amp; Panha S. 2006. Taxonomic review of the tree snail Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Thailand and adjacent areas: Subgenus Amphidromus. Journal of Molluscan Studies 72: 1 - 30. https: // doi. org / 10.1093 / mollus / eyi 044" type="journal article" year="2006">
Sutcharit &amp; Panha 
<date id="FFCD44896928EF06FC34A986FC3623B1" box="[959,1022,1720,1746]" pageId="14" pageNumber="15" value="2006">2006</date>
</bibRefCitation>
: fig. 4o). Moreover, this species occurs in dipterocarp forest within the range of 
<taxonomicName id="4C7319CA6928EF06FD68A9E5FCBB2396" authorityName="Geret" authorityYear="1912" box="[739,883,1755,1781]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6928EF06FD68A9E5FCBB2396" box="[739,883,1755,1781]" italics="true" pageId="14" pageNumber="15">A. givenchyi</emphasis>
</taxonomicName>
; therefore, it is possibly a shell color variant of 
<taxonomicName id="4C7319CA6928EF06FF55A9C0FEB8227B" authorityName="Geret" authorityYear="1912" box="[222,368,1790,1816]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6928EF06FF55A9C0FEB8227B" box="[222,368,1790,1816]" italics="true" pageId="14" pageNumber="15">A. givenchyi</emphasis>
</taxonomicName>
. Additional specimens for further anatomical and molecular study are desirable to test this hypothesis.
</paragraph>
</subSubSection>
</treatment>
</document>