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<emphasis id="B907BE5B6907EF29FE2DAC71FD9B2609" bold="true" box="[422,595,847,874]" italics="true" pageId="33" pageNumber="34">Amphidromus</emphasis>
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<bibRefCitation id="EFE21FB86907EF29FC3EAC6EFB462609" author="Mollendorff O. F. von" box="[949,1166,848,874]" pageId="33" pageNumber="34" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
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<figureCitation id="13487ECC6907EF29FD48AC4AFCC326ED" box="[707,779,884,910]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="33" pageNumber="34">Figs 1</figureCitation>
,
<figureCitation id="13487ECC6907EF29FC91AC4AFCFC26ED" box="[794,820,884,910]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. AC. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). DF. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). GH. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). IM. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). JM. Specimens from Ban Phon, Sekong, Laos (CUMZ 70417042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). OP. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). QR. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). ST. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">13</figureCitation>
OR
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locality: Boloven (Boloven Plateau,
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var.
<emphasis id="B907BE5B6907EF29FDD6AB74FD7D2100" box="[605,693,1098,1123]" italics="true" pageId="33" pageNumber="34">varians</emphasis>
<bibRefCitation id="EFE21FB86907EF29FD49AB77FC1D2107" author="Mollendorff O. F. von" box="[706,981,1097,1124]" pageId="33" pageNumber="34" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898: 75</bibRefCitation>
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locality: Boloven (Boloven Plateau,
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<emphasis id="B907BE5B6907EF29FF36AB8EFDDA21A9" box="[189,530,1200,1226]" italics="true" pageId="33" pageNumber="34">Amphidromus haematostoma</emphasis>
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<treatmentCitation id="0AD244586907EF29FDBBAB8EFCF621A9" author="Pilsbry H. A." box="[560,830,1200,1226]" page="182" pageId="33" pageNumber="34" year="1900">
<bibRefCitation id="EFE21FB86907EF29FDBBAB8EFCF621A9" author="Pilsbry H. A." box="[560,830,1200,1226]" pageId="33" pageNumber="34" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 182183</bibRefCitation>
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. —
<treatmentCitation id="0AD244586907EF29FBD7AB8EFECA218C" author="Laidlaw F. F. &amp; Solem A." page="527" pageId="33" pageNumber="34" year="1961">
<bibRefCitation id="EFE21FB86907EF29FBD7AB8EFECA218C" author="Laidlaw F. F. &amp; Solem A." pageId="33" pageNumber="34" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem 1961: 527</bibRefCitation>
</treatmentCitation>
, 625. —
<treatmentCitation id="0AD244586907EF29FEE6ABEAFDAC218D" author="Richardson L." box="[365,612,1236,1263]" page="19" pageId="33" pageNumber="34" year="1985">
<bibRefCitation id="EFE21FB86907EF29FEE6ABEAFDAC218D" author="Richardson L." box="[365,612,1236,1263]" pageId="33" pageNumber="34" pagination="1 - 479" refId="ref19668" refString="Richardson L. 1985. Camaenidae: catalog of species. Tryonia 12: 1 - 479." type="journal article" year="1985">Richardson 1985: 19</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCC62496907EF29FF36ABC7FC432054" blockId="33.[189,1398,1200,1445]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB6345676907EF29FF36ABC7FC432054" pageId="33" pageNumber="34">
<taxonomicName id="4C7319CA6907EF29FF36ABC7FD6A2070" ID-CoL="5M72R" authorityName="Mollendorff" authorityYear="1898" box="[189,674,1273,1299]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="variety" species="haematostoma" variety="viridis">
<emphasis id="B907BE5B6907EF29FF36ABC7FDDE2070" box="[189,534,1273,1299]" italics="true" pageId="33" pageNumber="34">Amphidromus haematostoma</emphasis>
var.
<emphasis id="B907BE5B6907EF29FDDCABC7FD6A2070" box="[599,674,1273,1299]" italics="true" pageId="33" pageNumber="34">viridis</emphasis>
</taxonomicName>
<treatmentCitation id="0AD244586907EF29FD4CABC7FC6A2070" author="Pilsbry H. A." box="[711,930,1273,1299]" page="183" pageId="33" pageNumber="34" year="1900">
<bibRefCitation id="EFE21FB86907EF29FD4CABC7FC6A2070" author="Pilsbry H. A." box="[711,930,1273,1299]" pageId="33" pageNumber="34" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 183</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AD244586907EF29FC52ABC7FB572070" author="Zilch A." box="[985,1183,1273,1300]" page="132" pageId="33" pageNumber="34" year="1953">
<bibRefCitation id="EFE21FB86907EF29FC52ABC7FB572070" author="Zilch A." box="[985,1183,1273,1300]" pageId="33" pageNumber="34" pagination="131 - 140" refId="ref20321" refString="Zilch A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg 10: Mollusca, Pleurodontidae (l). Archiv fur Molluskenkunde 82: 131 - 140." type="journal article" year="1953">Zilch 1953: 132</bibRefCitation>
</treatmentCitation>
, pl. 22, fig. 4. —
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<bibRefCitation id="EFE21FB86907EF29FF58AA23FDEC205B" author="Laidlaw F. F. &amp; Solem A." box="[211,548,1309,1336]" pageId="33" pageNumber="34" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem 1961: 527</bibRefCitation>
</treatmentCitation>
, 670. —
<treatmentCitation id="0AD244586907EF29FD04AA23FC4F2054" author="Richardson L." box="[655,903,1309,1336]" page="19" pageId="33" pageNumber="34" year="1985">
<bibRefCitation id="EFE21FB86907EF29FD04AA23FC4F2054" author="Richardson L." box="[655,903,1309,1336]" pageId="33" pageNumber="34" pagination="1 - 479" refId="ref19668" refString="Richardson L. 1985. Camaenidae: catalog of species. Tryonia 12: 1 - 479." type="journal article" year="1985">Richardson 1985: 19</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BCC62496907EF29FF36AA7CFC4320E3" blockId="33.[189,1398,1200,1445]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB6345676907EF29FF36AA7CFC4320E3" pageId="33" pageNumber="34">
<taxonomicName id="4C7319CA6907EF29FF36AA7CFD65203F" ID-CoL="5M72Q" authorityName="Mollendorff" authorityYear="1898" box="[189,685,1346,1372]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="variety" species="haematostoma" variety="varians">
<emphasis id="B907BE5B6907EF29FF36AA7CFDDD203F" box="[189,533,1346,1372]" italics="true" pageId="33" pageNumber="34">Amphidromus haematostoma</emphasis>
var.
<emphasis id="B907BE5B6907EF29FDDEAA7DFD65203F" box="[597,685,1347,1372]" italics="true" pageId="33" pageNumber="34">varians</emphasis>
</taxonomicName>
<treatmentCitation id="0AD244586907EF29FD5BAA7CFC61203F" author="Pilsbry H. A." box="[720,937,1346,1372]" page="183" pageId="33" pageNumber="34" year="1900">
<bibRefCitation id="EFE21FB86907EF29FD5BAA7CFC61203F" author="Pilsbry H. A." box="[720,937,1346,1372]" pageId="33" pageNumber="34" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 183</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AD244586907EF29FC54AA7CFB6B203F" author="Zilch A." box="[991,1187,1346,1373]" page="132" pageId="33" pageNumber="34" year="1953">
<bibRefCitation id="EFE21FB86907EF29FC54AA7CFB6B203F" author="Zilch A." box="[991,1187,1346,1373]" pageId="33" pageNumber="34" pagination="131 - 140" refId="ref20321" refString="Zilch A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg 10: Mollusca, Pleurodontidae (l). Archiv fur Molluskenkunde 82: 131 - 140." type="journal article" year="1953">Zilch 1953: 132</bibRefCitation>
</treatmentCitation>
, pl. 22, fig. 5. —
<treatmentCitation id="0AD244586907EF29FF58AA58FDEC20E2" author="Laidlaw F. F. &amp; Solem A." box="[211,548,1382,1409]" page="527" pageId="33" pageNumber="34" year="1961">
<bibRefCitation id="EFE21FB86907EF29FF58AA58FDEC20E2" author="Laidlaw F. F. &amp; Solem A." box="[211,548,1382,1409]" pageId="33" pageNumber="34" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem 1961: 527</bibRefCitation>
</treatmentCitation>
, 668. —
<treatmentCitation id="0AD244586907EF29FD04AA58FC4F20E3" author="Richardson L." box="[655,903,1382,1409]" page="19" pageId="33" pageNumber="34" year="1985">
<bibRefCitation id="EFE21FB86907EF29FD04AA58FC4F20E3" author="Richardson L." box="[655,903,1382,1409]" pageId="33" pageNumber="34" pagination="1 - 479" refId="ref19668" refString="Richardson L. 1985. Camaenidae: catalog of species. Tryonia 12: 1 - 479." type="journal article" year="1985">Richardson 1985: 19</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCC62496907EF29FF36AAB5FB8120C6" blockId="33.[189,1398,1200,1445]" box="[189,1097,1419,1445]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB6345676907EF29FF36AAB5FB8120C6" box="[189,1097,1419,1445]" pageId="33" pageNumber="34">
<taxonomicName id="4C7319CA6907EF29FF36AAB5FD7320C6" box="[189,699,1419,1445]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="haematostomus" subGenus="Syndromus">
<emphasis id="B907BE5B6907EF29FF36AAB5FEA920C6" box="[189,353,1419,1445]" italics="true" pageId="33" pageNumber="34">Amphidromus</emphasis>
(
<emphasis id="B907BE5B6907EF29FEFAAAB5FE3A20C6" box="[369,498,1419,1445]" italics="true" pageId="33" pageNumber="34">Syndromus</emphasis>
)
<emphasis id="B907BE5B6907EF29FD8FAAB5FD7320C6" box="[516,699,1419,1445]" italics="true" pageId="33" pageNumber="34">haematostomus</emphasis>
</taxonomicName>
<treatmentCitation id="0AD244586907EF29FD53AAB5FB8D20C6" author="Lehmann H. &amp; Maassen W. J. M." box="[728,1093,1419,1445]" page="20" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="EFE21FB86907EF29FD53AAB5FB8D20C6" author="Lehmann H. &amp; Maassen W. J. M." box="[728,1093,1419,1445]" pageId="33" pageNumber="34" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">Lehmann &amp; Maassen 2004: 20</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36931C26907EF2AFF36AAF3FC9924D1" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BCC62496907EF29FF36AAF3FE622084" blockId="33.[189,426,1485,1511]" box="[189,426,1485,1511]" pageId="33" pageNumber="34">
<heading id="D084D5256907EF29FF36AAF3FE622084" bold="true" box="[189,426,1485,1511]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B907BE5B6907EF29FF36AAF3FE622084" bold="true" box="[189,426,1485,1511]" pageId="33" pageNumber="34">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496907EF29FF61AAC3FE512374" blockId="33.[189,1399,1533,1815]" box="[234,409,1533,1559]" pageId="33" pageNumber="34">
<emphasis id="B907BE5B6907EF29FF61AAC3FE512374" bold="true" box="[234,409,1533,1559]" pageId="33" pageNumber="34">
<typeStatus id="54C8DCEB6907EF29FF61AAC3FEEF2374" box="[234,295,1533,1559]" pageId="33" pageNumber="34">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BCC62496907EF29FF36A91FFB222275" blockId="33.[189,1399,1533,1815]" pageId="33" pageNumber="34">
The species description included two varieties, “var. A.
<emphasis id="B907BE5B6907EF29FCDBA91CFC53235F" box="[848,923,1570,1596]" italics="true" pageId="33" pageNumber="34">viridis</emphasis>
” and “var. B.
<emphasis id="B907BE5B6907EF29FBCEA91CFB552358" box="[1093,1181,1570,1595]" italics="true" pageId="33" pageNumber="34">varians</emphasis>
”, but only one set of shell measurements was given in the original description, of the nominotypical variety. The specimen SMF 7559, labelled as “var. A.
<emphasis id="B907BE5B6907EF29FDAFA954FDA723E7" box="[548,623,1642,1668]" italics="true" pageId="33" pageNumber="34">viridis</emphasis>
”, exactly matches the dimensions given in the original description. Therefore, we believe this implied that the specimens of “var. A.
<emphasis id="B907BE5B6907EF29FC46A9B1FBD023CA" box="[973,1048,1679,1705]" italics="true" pageId="33" pageNumber="34">viridis</emphasis>
” pertain to the type series of
<taxonomicName id="4C7319CA6907EF29FF36A98AFE5F23AE" box="[189,407,1715,1742]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" sensu="stricto" species="haematostomus">
<emphasis id="B907BE5B6907EF29FF36A98AFE5F23AE" box="[189,407,1715,1742]" italics="true" pageId="33" pageNumber="34">A. haematostomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206907EF29FE2AA98AFE0E23AD" box="[417,454,1716,1742]" pageId="33" pageNumber="34" sensu="stricto">s.s.</taxonomicNameLabel>
The specimen SMF 7559 (
<figureCitation id="13487ECC6907EF29FC99A98DFCB723AE" box="[786,895,1715,1741]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. AC. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). DF. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). GH. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). IM. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). JM. Specimens from Ban Phon, Sekong, Laos (CUMZ 70417042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). OP. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). QR. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). ST. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13O</figureCitation>
) was designated as the
<typeStatus id="54C8DCEB6907EF29FB29A98DFAC623AE" box="[1186,1294,1715,1741]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
in
<bibRefCitation id="EFE21FB86907EF29FAB3A98DFEF72391" author="Zilch A." pageId="33" pageNumber="34" pagination="131 - 140" refId="ref20321" refString="Zilch A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg 10: Mollusca, Pleurodontidae (l). Archiv fur Molluskenkunde 82: 131 - 140." type="journal article" year="1953">Zilch (1953: 132</bibRefCitation>
, pl. 22, fig. 4), which is considered to be a valid
<typeStatus id="54C8DCEB6907EF29FCFFA9E6FC282391" box="[884,992,1752,1778]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
designation of the species, and the other specimen from the same lot becomes the
<typeStatus id="54C8DCEB6907EF29FD6EA9C2FC4B2275" box="[741,899,1788,1814]" pageId="33" pageNumber="34" type="paralectotype">paralectotype</typeStatus>
SMF 5760 (1 shell;
<figureCitation id="13487ECC6907EF29FBFEA9C2FB142275" box="[1141,1244,1788,1814]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. AC. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). DF. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). GH. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). IM. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). JM. Specimens from Ban Phon, Sekong, Laos (CUMZ 70417042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). OP. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). QR. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). ST. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13P</figureCitation>
).
</paragraph>
<paragraph id="8BCC62496907EF29FF36A801FE152288" blockId="33.[189,1399,1855,2027]" pageId="33" pageNumber="34">
The other specimens labeled as “var. B.
<emphasis id="B907BE5B6907EF29FD1EA87EFD25223A" box="[661,749,1856,1881]" italics="true" pageId="33" pageNumber="34">varians</emphasis>
” are distinct variants and are, therefore, excluded from the type series of this nominal species (
<collectionCode id="ED62FA8C6907EF29FDF7A85DFD0A221E" box="[636,706,1891,1917]" pageId="33" pageNumber="34">ICZN</collectionCode>
1999: Art. 72.4.1). The specimen SMF 7561 (
<figureCitation id="13487ECC6907EF29FB59A85DFAF2221E" box="[1234,1338,1891,1917]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. AC. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). DF. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). GH. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). IM. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). JM. Specimens from Ban Phon, Sekong, Laos (CUMZ 70417042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). OP. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). QR. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). ST. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13Q</figureCitation>
) was designated as the
<typeStatus id="54C8DCEB6907EF29FE02A8B6FE3D22C1" box="[393,501,1928,1954]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
in
<bibRefCitation id="EFE21FB86907EF29FD93A8B6FD1622C1" author="Zilch A." box="[536,734,1928,1955]" pageId="33" pageNumber="34" pagination="131 - 140" refId="ref20321" refString="Zilch A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg 10: Mollusca, Pleurodontidae (l). Archiv fur Molluskenkunde 82: 131 - 140." type="journal article" year="1953">Zilch (1953: 132</bibRefCitation>
, pl. 22, fig. 5), which is considered to be a valid
<typeStatus id="54C8DCEB6907EF29FA80A8B6FABF22C1" box="[1291,1399,1928,1954]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
designation of “var. B.
<emphasis id="B907BE5B6907EF29FE59A893FDE222A5" box="[466,554,1965,1990]" italics="true" pageId="33" pageNumber="34">varians</emphasis>
”. Another specimen from the same lot became the
<typeStatus id="54C8DCEB6907EF29FB05A892FAE722A5" box="[1166,1327,1964,1990]" pageId="33" pageNumber="34" type="paralectotype">paralectotype</typeStatus>
, SMF 5762 (1 shell;
<figureCitation id="13487ECC6907EF29FEEEA8EFFE182288" box="[357,464,2001,2027]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. AC. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). DF. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). GH. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). IM. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). JM. Specimens from Ban Phon, Sekong, Laos (CUMZ 70417042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). OP. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). QR. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). ST. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13R</figureCitation>
).
</paragraph>
<paragraph id="8BCC62496904EF2AFF61AE35FE6E2446" blockId="34.[189,1399,267,435]" box="[234,422,267,293]" pageId="34" pageNumber="35">
<emphasis id="B907BE5B6904EF2AFF61AE35FE6E2446" bold="true" box="[234,422,267,293]" pageId="34" pageNumber="35">Other material</emphasis>
</paragraph>
<paragraph id="8BCC62496904EF2AFF36AE10FC9924D1" blockId="34.[189,1399,267,435]" pageId="34" pageNumber="35">
<collectingCountry id="F36422D96904EF2AFF36AE10FEC6242B" box="[189,270,302,328]" name="Laos" pageId="34" pageNumber="35">LAOS</collectingCountry>
: 6 +
<specimenCount id="9D75A9C06904EF2AFEC7AE10FE18242B" box="[332,464,302,328]" pageId="34" pageNumber="35" type="juvenile">2 juveniles</specimenCount>
, Boloven Plateau, Pakxong District,
<collectingRegion id="49B7ACAB6904EF2AFC03AE10FBD9242B" box="[904,1041,302,328]" country="Laos" name="Champasak" pageId="34" pageNumber="35">Champasak</collectingRegion>
(
<collectionCode id="ED62FA8C6904EF2AFBA9AE10FBB4242B" box="[1058,1148,302,328]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="34" pageNumber="35" type="Museum">RMNH</collectionCode>
<date id="FFCD44896904EF2AFB0EAE10FB2A242A" box="[1157,1250,302,329]" pageId="34" pageNumber="35" value="50-10-10">101050</date>
); two lots in W.J.M.Maassen Collection (8S)and (12 +
<specimenCount id="9D75A9C06904EF2AFD2BAE6CFCE9240E" box="[672,801,338,365]" pageId="34" pageNumber="35" type="juvenile">2juveniles</specimenCount>
); 1shell,Boloven,Annam (
<collectionCode id="ED62FA8C6904EF2AFBDDAE6CFB0F240F" box="[1110,1223,338,364]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="35" type="Museum">NHMUK</collectionCode>
1899.4.22.74); 15 shells (
<collectionCode id="ED62FA8C6904EF2AFEB2AE4BFEB524EC" box="[313,381,373,399]" country="Germany" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="34" pageNumber="35" type="Museum">ZMB</collectionCode>
, ex. Lehmann Collection), 5 +
<specimenCount id="9D75A9C06904EF2AFD73AE4BFCA624EC" box="[760,878,373,399]" pageId="34" pageNumber="35" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED62FA8C6904EF2AFC0AAE4BFC0D24EC" box="[897,965,373,399]" country="Germany" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="34" pageNumber="35" type="Museum">ZMB</collectionCode>
, ex. Lehmann Collection), in coffee plantation, near Pakxong, Boloven Plateau,
<collectingRegion id="49B7ACAB6904EF2AFD4BAEA6FC8524D1" box="[704,845,408,434]" country="Laos" name="Champasak" pageId="34" pageNumber="35">Champasak</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C36931C26904EF2AFF36AEEBFDF221EB" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BCC62496904EF2AFF36AEEBFEE7248C" blockId="34.[189,303,469,495]" box="[189,303,469,495]" pageId="34" pageNumber="35">
<heading id="D084D5256904EF2AFF36AEEBFEE7248C" bold="true" box="[189,303,469,495]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B907BE5B6904EF2AFF36AEEBFEE7248C" bold="true" box="[189,303,469,495]" pageId="34" pageNumber="35">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496904EF2AFF36AD3AFB3927AC" blockId="34.[189,1399,516,719]" pageId="34" pageNumber="35">
This species is currently known from the
<typeStatus id="54C8DCEB6904EF2AFD17AD3AFD07277D" box="[668,719,516,542]" pageId="34" pageNumber="35">type</typeStatus>
locality only. The shell characters are clearly distinct from all other recognized species. The shell is sinistral and ovate conical. Then apex is acute, with a black spot on the tip. The last whorl is rounded; the color of the shell is a monochrome greenish (faded in old specimen), and there are a yellowish subsutural band and an umbilical area. Then aperture is ovate; the peristome expanded and weakly reflected; the lip is reddish-purple. The parietal callus is reddish-purple and thickened; the columella is also reddish-purple and dilated; the umbilicus imperforated.
</paragraph>
<paragraph id="8BCC62496904EF2AFF36ADCCFE072683" blockId="34.[189,1399,754,993]" pageId="34" pageNumber="35">
<bibRefCitation id="EFE21FB86904EF2AFF36ADCCFE28266E" author="Laidlaw F. F. &amp; Solem A." box="[189,480,754,781]" pageId="34" pageNumber="35" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem (1961)</bibRefCitation>
suggested that this species was probably a junior synonym of
<taxonomicName id="4C7319CA6904EF2AFB3AADCCFAA7266F" authorityName="Fulton" authorityYear="1896" box="[1201,1391,754,780]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6904EF2AFB3AADCCFAA7266F" box="[1201,1391,754,780]" italics="true" pageId="34" pageNumber="35">A. roseolabiatus</emphasis>
</taxonomicName>
. However, after comparing the
<typeStatus id="54C8DCEB6904EF2AFDBDAC28FDA12653" box="[566,617,790,816]" pageId="34" pageNumber="35">type</typeStatus>
specimens of both species and recently collected specimens of
<taxonomicName id="4C7319CA6904EF2AFF36AC07FE482631" authorityName="Fulton" authorityYear="1896" box="[189,384,824,851]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6904EF2AFF36AC07FE482631" box="[189,384,824,851]" italics="true" pageId="34" pageNumber="35">A. roseolabiatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7319CA6904EF2AFE1AAC07FDA82631" authorityName="Mollendorff" authorityYear="1898" box="[401,608,824,851]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="haematostoma">
<emphasis id="B907BE5B6904EF2AFE1AAC07FDA82631" box="[401,608,824,851]" italics="true" pageId="34" pageNumber="35">A. haematostoma</emphasis>
</taxonomicName>
clearly differs from it in having an ovate conical shape, uniform greenish to yellowish shell, yellowish subsutural and umbilical area, thickened parietal callus, lip widely expanded, and reddish-purple lip and parietal callus. Moreover,
<taxonomicName id="4C7319CA6904EF2AFC36ACBEFBB626FA" authorityName="Fulton" authorityYear="1896" box="[957,1150,895,922]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6904EF2AFC36ACBEFBB626FA" box="[957,1150,895,922]" italics="true" pageId="34" pageNumber="35">A. roseolabiatus</emphasis>
</taxonomicName>
exhibits an elongate conical shell, greenish radial streaks, white subsutural band, transparent parietal callus, and lip pinkish or white and expanded.
</paragraph>
<paragraph id="8BCC62496904EF2AFF36AB3DFDF221EB" blockId="34.[189,1399,1027,1160]" pageId="34" pageNumber="35">
<bibRefCitation id="EFE21FB86904EF2AFF36AB3DFDF7217E" author="Lehmann H. &amp; Maassen W. J. M." box="[189,575,1027,1053]" pageId="34" pageNumber="35" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">Lehmann &amp; Maassen (2004: 20)</bibRefCitation>
recorded the genital characters (without illustration) of the topotypic population and mentioned that it had a “very short flagellum and a short conical verge”. These are the distinguishing characters of this subgenus. However, further investigation of the genitalia is necessary to confirm the subgeneric status.
</paragraph>
</subSubSection>
</treatment>
</document>