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<document id="AFAA635568D3711FFB3AA73541881F33" ID-DOI="10.5852/ejt.2024.921.2429" ID-ISSN="2118-9773" ID-Zenodo-Dep="10663874" ID-ZooBank="02633F29-4CDF-4027-BEBF-07AD2F925B42" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707990651991" checkinUser="plazi" docAuthor="Bellvert, Adrià, Dimitrov, Dragomir, Zamani, Alireza &amp; Arnedo, Miquel A." docDate="2024" docId="03DA7943B736D841718B8F7B40BAFE01" docLanguage="en" docName="EJT.2024.921.210-235.pdf" docOrigin="European Journal of Taxonomy 921 (210)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2429/10759" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dysdera kourosh Bellvert, Zamani &amp; Dimitrov 2024, sp. nov." docType="treatment" docUuid="2242E86D-6C29-47C0-883C-A7A549D8039E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="226" masterDocId="FFE3013BB738D85170598D5B445AFFC5" masterDocTitle="Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia" masterLastPageNumber="235" masterPageNumber="210" pageNumber="224" updateTime="1707991084405" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="7C71EAA29AB45D05356859B72C200819" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4791E8C5EF6E6A7DBBFC3BA6E5E1B4DC">
<mods:title id="EBF2F5BF3165A5711D6737B724B4B1D0">Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia</mods:title>
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<mods:namePart id="140CF8F03DED9F4D3BA923EE00259F3D">Bellvert, Adrià</mods:namePart>
<mods:nameIdentifier id="90E34F484E2EA327EB9F7334E18217B7" type="LSID">0ABC3DFE-E982-47F5-A5DA-732688BCDEA9</mods:nameIdentifier>
<mods:affiliation id="56F010D1C619ABB0D2F1B978F69B2727">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
<mods:nameIdentifier id="566AF74F492FC705DD39726E23E1190D" type="email">abellvertba@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="BB9FB72F7F497F88FFA307853E1185E2" type="personal">
<mods:role id="F33ABFA061A08CE559F6C5C88C032328">
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</mods:role>
<mods:namePart id="9D64F4121D034E63A7C3AC9D2923D3BF">Dimitrov, Dragomir</mods:namePart>
<mods:nameIdentifier id="EF3FB602703C2B70BAC2BA35046FD96F" type="LSID">37EC22F1-6348-4A61-B55F-3E74498AE5E9</mods:nameIdentifier>
<mods:affiliation id="AF8082D105F73F11241B2DD60C9BC6DD">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain. &amp; National Museum of Natural Sciences, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria.</mods:affiliation>
<mods:nameIdentifier id="188B34C13721FAE63262CF0B21E3E1AD" type="email">dimitrov.drk@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2971EF7D283A8ABD5671DCC24F560B30" type="personal">
<mods:role id="3D012D4A77689EF77F550BC3B45EBE80">
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</mods:role>
<mods:namePart id="106AE3DE109BFC150F13156D940D24A7">Zamani, Alireza</mods:namePart>
<mods:nameIdentifier id="C62AC7879BD83222ADFC43EA6D52C65E" type="LSID">A21C0B82-E2D9-4402-9408-D6160843DAF4</mods:nameIdentifier>
<mods:affiliation id="379BFCF99753DBACD2EF55B4F2584614">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland.</mods:affiliation>
<mods:nameIdentifier id="37150C383475006A437714C10CE5B1DC" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="2387AE73E2BA9C5A217734625F070670">Arnedo, Miquel A.</mods:namePart>
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<mods:affiliation id="B81489CCF3234A5E06C3BCE23C47612C">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
<mods:nameIdentifier id="2F42500C459A1409449EDD97A464EC39" type="email">marnedo@gmail.com</mods:nameIdentifier>
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<mods:title id="03C5DA50F9FE899CC8BB25DD93728EFE">European Journal of Taxonomy</mods:title>
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<mods:part id="8ED0E10A63A2C8ACA047DB89242BB419">
<mods:date id="F6BF4CC463EA4E3F420015F0A7F8C73F">2024</mods:date>
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<mods:number id="A13758FBD2BF716BEB2B84EA5D80A8AE">2024-02-14</mods:number>
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<mods:number id="DA72E088761216E97CC30FBB1D484FF8">921</mods:number>
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<mods:classification id="48B40684E9E07F7FA7A0A2A8E9A991EA">journal article</mods:classification>
<mods:identifier id="BF1F7FFA677A9210732FAA31CEE42007" type="DOI">10.5852/ejt.2024.921.2429</mods:identifier>
<mods:identifier id="C084D081546591DD9C85AB4F95693C02" type="ISSN">2118-9773</mods:identifier>
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<treatment id="03DA7943B736D841718B8F7B40BAFE01" LSID="urn:lsid:zoobank.org:act:2242E86D-6C29-47C0-883C-A7A549D8039E" httpUri="http://treatment.plazi.org/id/03DA7943B736D841718B8F7B40BAFE01" lastPageId="16" lastPageNumber="226" pageId="14" pageNumber="224">
<subSubSection id="C3699BDEB736D85F718B8F7B4099FD9A" pageId="14" pageNumber="224" type="nomenclature">
<paragraph id="8BCCC855B736D85F718B8F7B4038FDFE" blockId="14.[369,1219,544,642]" box="[466,1122,544,571]" pageId="14" pageNumber="224">
<heading id="D0847F39B736D85F718B8F7B4038FDFE" box="[466,1122,544,571]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="224" reason="2">
<taxonomicName id="4C73B3D6B736D85F718B8F7B405AFDFF" authority="Bellvert, Zamani &amp; Dimitrov" authorityName="Bellvert, Zamani &amp; Dimitrov" authorityYear="2024" box="[466,1024,544,571]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="kourosh" status="sp. nov.">
<emphasis id="B9071447B736D85F718B8F7B46FAFDFE" bold="true" box="[466,672,544,571]" italics="true" pageId="14" pageNumber="224">Dysdera kourosh</emphasis>
Bellvert, Zamani &amp; Dimitrov
</taxonomicName>
<taxonomicNameLabel id="A234A93CB736D85F745E8F7A4038FDFE" box="[1031,1122,545,571]" pageId="14" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BCCC855B736D85F71288F1F4099FD9A" blockId="14.[369,1219,544,642]" box="[369,1219,580,607]" pageId="14" pageNumber="224">
<uri id="FFE2C457B736D85F71288F1F4099FD9A" box="[369,1219,580,607]" pageId="14" pageNumber="224">
urn:lsid:zoobank.org:act:
<uuid id="FFD5F280B736D85F72D68F1F4099FD9A" box="[655,1219,580,607]" pageId="14" pageNumber="224">2242E86D-6C29-47C0-883C-A7A549D8039E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB736D85F72818F334701FD47" box="[728,859,616,642]" pageId="14" pageNumber="224" type="description">
<paragraph id="8BCCC855B736D85F72818F334701FD47" blockId="14.[369,1219,544,642]" box="[728,859,616,642]" pageId="14" pageNumber="224">
<figureCitation id="1348D4D0B736D85F72818F334701FD47" box="[728,859,616,642]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="14" pageNumber="224">Figs 18–20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB736D85F70E48FF14130FC62" pageId="14" pageNumber="224" type="diagnosis">
<paragraph id="8BCCC855B736D85F70E48FF1456CFD01" blockId="14.[189,310,682,708]" box="[189,310,682,708]" pageId="14" pageNumber="224">
<heading id="D0847F39B736D85F70E48FF1456CFD01" bold="true" box="[189,310,682,708]" fontSize="11" level="3" pageId="14" pageNumber="224" reason="3">
<emphasis id="B9071447B736D85F70E48FF1456CFD01" bold="true" box="[189,310,682,708]" pageId="14" pageNumber="224">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B736D85F70E48F824130FC62" blockId="14.[189,1399,729,936]" pageId="14" pageNumber="224">
<taxonomicName id="4C73B3D6B736D85F70E48F8245DCFD36" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[189,390,729,755]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="kourosh" status="sp. nov.">
<emphasis id="B9071447B736D85F70E48F8245DCFD36" box="[189,390,729,755]" italics="true" pageId="14" pageNumber="224">Dysdera kourosh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB736D85F71D68F8145B1FD31" box="[399,491,730,756]" pageId="14" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C73B3D6B736D85F722B8F814758FD36" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[626,770,729,756]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="naouelae" status="sp. nov.">
<emphasis id="B9071447B736D85F722B8F8146DCFD36" box="[626,646,730,755]" italics="true" pageId="14" pageNumber="224">D</emphasis>
.
<emphasis id="B9071447B736D85F72CF8F824758FD36" box="[662,770,729,755]" italics="true" pageId="14" pageNumber="224">naouelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB736D85F73538F814732FD31" box="[778,872,730,756]" pageId="14" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C73B3D6B736D85F732F8F81404CFD36" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[886,1046,729,756]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="mikhailovi">
<emphasis id="B9071447B736D85F732F8F8147D0FD36" box="[886,906,730,755]" italics="true" pageId="14" pageNumber="224">D</emphasis>
.
<emphasis id="B9071447B736D85F73C38F82404CFD36" box="[922,1046,729,755]" italics="true" pageId="14" pageNumber="224">mikhailovi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C73B3D6B736D85F74038F8140BAFD36" authorityName="Zamani, Marusik &amp; Szuts" authorityYear="2023" box="[1114,1248,730,756]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="sagartia">
<emphasis id="B9071447B736D85F74038F814034FD36" box="[1114,1134,730,755]" italics="true" pageId="14" pageNumber="224">D</emphasis>
.
<emphasis id="B9071447B736D85F74278F8140BAFD36" box="[1150,1248,730,755]" italics="true" pageId="14" pageNumber="224">sagartia</emphasis>
</taxonomicName>
by the well-developed, wide crest, interior sclerite significantly wider than the exterior one, and the characteristic lateral margin of the lateral sheet. The male differs by (1) the lateral margin of the lateral sheet pointed dorsally (
<figureCitation id="1348D4D0B736D85F71758E1E4525FC9A" box="[300,383,837,863]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="14" pageNumber="224">Fig 20</figureCitation>
), vs thin, sharp, more sclerotized and almost perpendicular to the embolic division in
<taxonomicName id="4C73B3D6B736D85F70E78E314516FC46" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[190,332,873,900]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="naouelae">
<emphasis id="B9071447B736D85F70E78E314488FC46" box="[190,210,874,899]" italics="true" pageId="14" pageNumber="224">D</emphasis>
.
<emphasis id="B9071447B736D85F70B98E324516FC46" box="[224,332,873,899]" italics="true" pageId="14" pageNumber="224">naouelae</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B736D85F71038E3245EAFC46" box="[346,432,873,899]" captionStart="Figs 9–14. 9–11" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 9–14. 9–11. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 12–14. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="14" pageNumber="224">Fig. 14</figureCitation>
), and significantly wider in
<taxonomicName id="4C73B3D6B736D85F72AA8E3147CBFC46" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[755,913,873,899]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="mikhailovi">
<emphasis id="B9071447B736D85F72AA8E3147CBFC46" box="[755,913,873,899]" italics="true" pageId="14" pageNumber="224">D. mikhailovi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B736D85F73C68E3240BBFC46" author="Fomichev A. A. &amp; Marusik Y. M." box="[927,1249,873,899]" pageId="14" pageNumber="224" pagination="73 - 89" refId="ref11966" refString="Fomichev A. A. &amp; Marusik Y. M. 2021. Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia. Zootaxa 5006 (1): 73 - 89. https: // doi. org / 10.11646 / zootaxa. 5006.1.10" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 46), and
<taxonomicName id="4C73B3D6B736D85F70E48ED54519FC62" authorityName="Zamani, Marusik &amp; Szuts" authorityYear="2023" box="[189,323,910,936]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="sagartia">
<emphasis id="B9071447B736D85F70E48ED54488FC62" box="[189,210,910,935]" italics="true" pageId="14" pageNumber="224">D</emphasis>
.
<emphasis id="B9071447B736D85F70B88ED54519FC62" box="[225,323,910,935]" italics="true" pageId="14" pageNumber="224">sagartia</emphasis>
</taxonomicName>
, and (2) the concave crest (
<figureCitation id="1348D4D0B736D85F72DE8ED64687FC62" box="[647,733,909,935]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="14" pageNumber="224">Fig. 18</figureCitation>
), vs rounded and not notched in the other three species.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB736D85F70E48E8B4722FBF9" pageId="14" pageNumber="224" type="etymology">
<paragraph id="8BCCC855B736D85F70E48E8B451EFC2F" blockId="14.[189,324,976,1002]" box="[189,324,976,1002]" pageId="14" pageNumber="224">
<heading id="D0847F39B736D85F70E48E8B451EFC2F" bold="true" box="[189,324,976,1002]" fontSize="11" level="3" pageId="14" pageNumber="224" reason="3">
<emphasis id="B9071447B736D85F70E48E8B451EFC2F" bold="true" box="[189,324,976,1002]" pageId="14" pageNumber="224">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B736D85F70E48EA44722FBF9" blockId="14.[189,1399,1023,1085]" pageId="14" pageNumber="224">The specific epithet is a noun in apposition after Cyrus the Great – Kourosh in Persian, which translates as Lord of the Sun – the founder of the first Persian empire.</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB736D85F70E4893E4016FB17" pageId="14" pageNumber="224" type="materials_examined">
<paragraph id="8BCCC855B736D85F70E4893E4536FBBB" blockId="14.[189,364,1124,1151]" box="[189,364,1124,1151]" pageId="14" pageNumber="224">
<heading id="D0847F39B736D85F70E4893E4536FBBB" bold="true" box="[189,364,1124,1151]" fontSize="11" level="3" pageId="14" pageNumber="224" reason="3">
<emphasis id="B9071447B736D85F70E4893E4536FBBB" bold="true" box="[189,364,1124,1151]" pageId="14" pageNumber="224">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B736D85F70B389CF4506FB6B" blockId="14.[189,1100,1172,1235]" box="[234,348,1172,1198]" pageId="14" pageNumber="224">
<emphasis id="B9071447B736D85F70B389CF4506FB6B" bold="true" box="[234,348,1172,1198]" pageId="14" pageNumber="224">
<typeStatus id="54C876F7B736D85F70B389CF4506FB6B" box="[234,348,1172,1198]" pageId="14" pageNumber="224" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCCC855B736D85F70E489E34016FB17" blockId="14.[189,1100,1172,1235]" box="[189,1100,1208,1235]" pageId="14" pageNumber="224">
<materialsCitation id="3B1BC208B736D85F70E489E34016FB17" box="[189,1100,1208,1235]" collectingDate="2013-03-25" collectionCode="SMF" country="Iran" latitude="29.75" location="Jahrom" longLatPrecision="735" longitude="52.68" pageId="14" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Fars" typeStatus="holotype">
<collectingCountry id="F36488C5B736D85F70E489E3455FFB17" box="[189,261,1208,1234]" name="Iran" pageId="14" pageNumber="224">IRAN</collectingCountry>
•
<specimenCount id="9D7503DCB736D85F714789E34517FB17" box="[286,333,1208,1234]" count="1" pageId="14" pageNumber="224" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49B706B7B736D85F710089E345A4FB17" box="[345,510,1208,1234]" country="Iran" name="Fars" pageId="14" pageNumber="224">Fars Province</collectingRegion>
,
<location id="8EAC9E8EB736D85F725389E3463EFB17" LSID="urn:lsid:plazi:treatment:03DA7943B736D841718B8F7B40BAFE01:8EAC9E8EB736D85F725389E3463EFB17" box="[522,612,1208,1234]" country="Iran" latitude="29.75" longLatPrecision="735" longitude="52.68" name="Jahrom" pageId="14" pageNumber="224" stateProvince="Fars">Jahrom</location>
;
<geoCoordinate id="EE47AE92B736D85F722989E34687FB17" box="[624,733,1208,1235]" degrees="29.75" direction="north" orientation="latitude" pageId="14" pageNumber="224" precision="555" value="29.75">29.75° N</geoCoordinate>
,
<geoCoordinate id="EE47AE92B736D85F72B089E34709FB17" box="[745,851,1208,1235]" degrees="52.68" direction="east" orientation="longitude" pageId="14" pageNumber="224" precision="555" value="52.68">52.68° E</geoCoordinate>
;
<date id="FFCDEE95B736D85F730689E3405AFB17" box="[863,1024,1208,1235]" pageId="14" pageNumber="224" value="2013-03-25">
<collectingDate id="EF89177DB736D85F730689E3405AFB17" box="[863,1024,1208,1235]" pageId="14" pageNumber="224" value="2013-03-25">25 Mar. 2013</collectingDate>
</date>
;
<collectionCode id="ED625090B736D85F745589E34012FB17" box="[1036,1096,1208,1234]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="224">SMF</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB736D85E70E489A14563FBFB" lastPageId="15" lastPageNumber="225" pageId="14" pageNumber="224" type="description">
<paragraph id="8BCCC855B736D85F70E489A14514FAD1" blockId="14.[189,334,1274,1300]" box="[189,334,1274,1300]" pageId="14" pageNumber="224">
<heading id="D0847F39B736D85F70E489A14514FAD1" bold="true" box="[189,334,1274,1300]" fontSize="11" level="3" pageId="14" pageNumber="224" reason="3">
<emphasis id="B9071447B736D85F70E489A14514FAD1" bold="true" box="[189,334,1274,1300]" pageId="14" pageNumber="224">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B736D85F70B388724592FA86" blockId="14.[187,1400,1321,1671]" box="[234,456,1321,1347]" pageId="14" pageNumber="224">
<emphasis id="B9071447B736D85F70B388724570FA86" bold="true" box="[234,298,1321,1347]" pageId="14" pageNumber="224">Male</emphasis>
(
<figureCitation id="1348D4D0B736D85F71608872459AFA86" box="[313,448,1321,1347]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="14" pageNumber="224">Figs 18–20</figureCitation>
)
</paragraph>
<paragraph id="8BCCC855B736D85F70E488164705F942" blockId="14.[187,1400,1321,1671]" pageId="14" pageNumber="224">PROSOMA. 2.77 long; maximum width 2.11; minimum width 1.37. Red, frontally darker, becoming lighter towards the back; smooth with some small black grains mainly at the front. Anterior border roughly round, about ½ of carapace length; anterior lateral borders convergent; rounded at maximum dorsal width, posterior lateral borders rounded; posterior margin narrow, straight. Eye diameters:AME 0.13; PLE 0.10; PME 0.10; AMEs on edge of the anterior border, separated from one another by about 1 diameter, close to PLEs; PMEs very close to one another, less than ¼ of PME diameter from PLEs. Labium trapezoid-shaped, its base wider than its distal part, borders slightly curved; longer than wide at base; semicircular groove at the tip. Sternum orange, darkened on borders; very slightly wrinkled, mainly between legs and anterior border; uniformly covered in slender black setae.</paragraph>
<paragraph id="8BCCC855B736D85F70E48BF44122F82D" blockId="14.[187,1400,1709,2025]" pageId="14" pageNumber="224">CHELICERAE. 1.26 long, about ½ of carapace length in dorsal view; fang 1.12 long; paturon dorsal and ventral side completely covered with piligerous granulations. Cheliceral inner groove medium-size, about ⅖ of cheliceral length; armed with three teeth and lamina at base; D =B&gt;M; D triangular, located near segment tip; B close to basal lamina; M close to B. Legs orange. Lengths of leg segments: fe1 2.36; pa1 1.45; ti1 2.00; me1 2.06; ta1 0.56; total 8.43; fe2 2.04; pa2 1.29; ti2 1.73; me2 1.92; ta2 0.49; total 7.48; fe3 1.59; pa3 0.89; ti3 1.08; me3 1.57; ta3 0.51; total 5.64; fe4 2.04; pa4 1.07; ti4 1.59; me4 1.98; total 6.69; fe Pdp 1.38; pa Pdp 0.73; ti Pdp 0.63; ta Pdp 0.70; total 3.45; leg formula: 1&gt;2&gt;4&gt;3. Spination: palp: one spine on pa, three spines on fe distal internal; leg1 two terminal spines on forward margin; leg2 one terminal spine on the forward margin. Fe3d spines in two rows; forward 2; backward 1; pa3</paragraph>
<paragraph id="8BCCC855B737D85E70E48C52412FFE33" blockId="15.[187,1400,265,503]" pageId="15" pageNumber="225">1 ventral; tb3d spines arranged in two bands; proximal 1.0.1; medial-proximal 0; medial-distal 0; distal 1.0.1.; tb3v spines arranged in three bands; proximal 0.1.0; medial-proximal 1.1.0; medial-distal 0; distal 1.0.0; with two terminal spines. Fe4d spines in two rows; forward 1; backward 5; pa4 1 ventral medial; tb4d spines arranged in two bands; proximal 1.0.1; medial-proximal 0; medial-distal 0; distal 1.0.1; tb4v spines arranged in three bands; proximal 0.1.1; medial-proximal 1.1.0; medial-distal 0; distal 1.0.0; with two terminal spines. Dorsal side of the frontal legs covered with small piligerous grains; ventral side of the pedipalp covered with small piligerous grains. Claws with 8 teeth or less; hardly larger than claw width.</paragraph>
<paragraph id="8BCCC855B737D85E70E48F454792FD9E" blockId="15.[189,1398,541,603]" pageId="15" pageNumber="225">OPISTHOSOMA. 2.54 long; cream-colored. Abdominal dorsal setae 0.02 long; thick, slightly curved, compressed, blunt, tip not enlarged; uniformly, thickly distributed.</paragraph>
<paragraph id="8BCCC855B737D85E70E48FD9450CFC1C" blockId="15.[188,1401,642,985]" pageId="15" pageNumber="225">
PALP (
<figureCitation id="1348D4D0B737D85E715F8FD945D7FD59" box="[262,397,642,668]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="15" pageNumber="225">Figs 18–20</figureCitation>
). T slightly shorter than ED; external distal border straight; internal sloped backward. ED the same T axis in lateral view, internal distal border not expanded. IS wider than ES, continuous to tip (
<figureCitation id="1348D4D0B737D85E70B28F92451BFD26" box="[235,321,713,739]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="15" pageNumber="225">Fig. 19</figureCitation>
). ED tip sloped towards back in lateral view. C present, long, slightly concave at the middle, seen from lateral view (
<figureCitation id="1348D4D0B737D85E718F8FB7463BFCC3" box="[470,609,748,774]" captionStart="Figs 18–20" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 18–20. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="15" pageNumber="225">Figs 18, 20</figureCitation>
); distal border round, the posterior border with a small sclerotized apophysis, markedly expanded, perpendicular to ED. AC absent. LF absent. L well developed; external border sclerotized, not folded, with processus-like basal lateral apophysis. LA absent. F absent. AL present, very poorly developed; proximal border in posterior view fused with DH. P not fused to T; perpendicular to T in lateral view; lateral length from ⅖ to ½ of T width; ridge present, parallel to T; not expanded, upper margin smooth, distally ridge-like expanded, back margin slightly folded towards the internal side.
</paragraph>
<paragraph id="8BCCC855B737D85E70B3895A451FFBDE" blockId="15.[189,325,1025,1086]" box="[234,325,1025,1051]" pageId="15" pageNumber="225">
<emphasis id="B9071447B737D85E70B3895A451FFBDE" bold="true" box="[234,325,1025,1051]" pageId="15" pageNumber="225">Female</emphasis>
</paragraph>
<paragraph id="8BCCC855B737D85E70E4897F4563FBFB" blockId="15.[189,325,1025,1086]" box="[189,313,1060,1086]" pageId="15" pageNumber="225">Unknown.</paragraph>
</subSubSection>
<caption id="DF0C98DDB737D85E70E48AF04794F82C" ID-DOI="http://doi.org/10.5281/zenodo.10663892" ID-Zenodo-Dep="10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="15" pageNumber="225" startId="15.[189,241,1963,1989]" targetBox="[194,1398,1197,1925]" targetPageId="15">
<paragraph id="8BCCC855B737D85E70E48AF04794F82C" blockId="15.[188,1399,1963,2025]" pageId="15" pageNumber="225">
<emphasis id="B9071447B737D85E70E48AF0451DF800" bold="true" box="[189,327,1963,1989]" pageId="15" pageNumber="225">Figs 18–20.</emphasis>
Male
<taxonomicName id="4C73B3D6B737D85E71D68AF047F4F800" authority="Bellvert, Zamani &amp; Dimitrov" authorityName="Bellvert, Zamani &amp; Dimitrov" authorityYear="2024" box="[399,942,1963,1989]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="225" phylum="Arthropoda" rank="species" species="kourosh" status="sp. nov.">
<emphasis id="B9071447B737D85E71D68AF0460FF800" box="[399,597,1963,1989]" italics="true" pageId="15" pageNumber="225">Dysdera kourosh</emphasis>
Bellvert, Zamani &amp; Dimitrov
</taxonomicName>
<taxonomicNameLabel id="A234A93CB737D85E73ED8AF74057F803" box="[948,1037,1964,1990]" pageId="15" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
(SMF), left palp.
<emphasis id="B9071447B737D85E74808AF040ACF800" bold="true" box="[1241,1270,1963,1989]" pageId="15" pageNumber="225">18</emphasis>
. Prolateral view.
<emphasis id="B9071447B737D85E715A8A95457AF82D" bold="true" box="[259,288,1998,2024]" pageId="15" pageNumber="225">19</emphasis>
. Anterior view.
<emphasis id="B9071447B737D85E71858A9545A3F82D" bold="true" box="[476,505,1998,2024]" pageId="15" pageNumber="225">20</emphasis>
. Retrolateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3699BDEB728D84170E48C534797FE96" box="[189,973,264,339]" pageId="16" pageNumber="226" type="distribution">
<paragraph id="8BCCC855B728D84170E48C53450CFEE7" blockId="16.[189,342,264,290]" box="[189,342,264,290]" pageId="16" pageNumber="226">
<heading id="D0847F39B728D84170E48C53450CFEE7" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="16" pageNumber="226" reason="3">
<emphasis id="B9071447B728D84170E48C53450CFEE7" bold="true" box="[189,342,264,290]" pageId="16" pageNumber="226">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B728D84170E48C624797FE96" blockId="16.[189,973,312,339]" box="[189,973,312,339]" pageId="16" pageNumber="226">
Known only from the
<typeStatus id="54C876F7B728D841719B8C6245AFFE96" box="[450,501,313,339]" pageId="16" pageNumber="226">type</typeStatus>
locality in
<collectingRegion id="49B706B7B728D84172208C624744FE97" box="[633,798,312,339]" country="Iran" name="Fars" pageId="16" pageNumber="226">Fars Province</collectingRegion>
, southern
<collectingCountry id="F36488C5B728D84173CF8C624793FE96" box="[918,969,313,339]" name="Iran" pageId="16" pageNumber="226">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84170E48C2040BAFE01" box="[189,1248,379,453]" pageId="16" pageNumber="226" type="discussion">
<paragraph id="8BCCC855B728D84170E48C204575FE50" blockId="16.[189,303,379,405]" box="[189,303,379,405]" pageId="16" pageNumber="226">
<heading id="D0847F39B728D84170E48C204575FE50" bold="true" box="[189,303,379,405]" fontSize="11" level="3" pageId="16" pageNumber="226" reason="3">
<emphasis id="B9071447B728D84170E48C204575FE50" bold="true" box="[189,303,379,405]" pageId="16" pageNumber="226">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B728D84170E48CF140BAFE01" blockId="16.[189,1248,426,453]" box="[189,1248,426,453]" pageId="16" pageNumber="226">
The species belongs to the
<emphasis id="B9071447B728D84171A38CF14605FE01" box="[506,607,426,452]" italics="true" pageId="16" pageNumber="226">aculeata</emphasis>
group (sensu
<bibRefCitation id="EFE2B5A4B728D841735D8CF14089FE01" author="Deeleman-Reinhold C. L. &amp; Deeleman P. R." box="[772,1235,426,452]" pageId="16" pageNumber="226" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman 1988</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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