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<document id="C2E0D90E8900777F77FB9FA3CDAF331B" ID-CLB-Dataset="55466" ID-DOI="10.5281/zenodo.5169525" ID-GBIF-Dataset="4910761a-706e-4f53-a5a8-6c72379bdbc6" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169525" ID-ZooBank="DA6CBFE5-927E-45B6-9D05-69AC97AF7B76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628277966057" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2017" docId="03DA133F751EB13A4BC2FA0288EE6892" docLanguage="en" docName="InsectaMundi.0592.1-16.pdf" docOrigin="Insecta Mundi 2017 (592)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ganelius oberndorferi" docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FFE36B47751BB13D4B02FFA38A0B6E57" masterDocTitle="Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="4" updateTime="1699056540890" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="35CDD8D1CACDD27D39723D69451D1156" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D688CCC9909E12FD8FB3C16267834C82">
<mods:title id="0F9C9D61ACDAAF4E46633B338C6849AA">Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
</mods:titleInfo>
<mods:name id="78BAAB6C2668FA298D761345B6A91D79" type="personal">
<mods:role id="0B62D3174B83339966D3BD15E0488606">
<mods:roleTerm id="CC1CB84550E48CF50CE99C6CBEB7A171">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9CE9012D2E1EE6E725B8B80D4DC4709">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="BBFF9170E753CD1C16328A0E55442CD3">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
<mods:nameIdentifier id="D7FDB539B061A87DAB3B3FDF6C3F8FC6" type="email">mjpaulsen@unl.edu</mods:nameIdentifier>
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<mods:title id="2967BFE29F54064D76E40D2FC6C48B60">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="370CFD22DD794380F42E4087E995281D">
<mods:date id="AC346ADB088616CF2CB5C01CFA480E80">2017</mods:date>
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<mods:number id="6A3B70B4F44E62612EEC351EA40A586A">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="DF1D8516FF5410A28BDAF3DDF00C300B" type="volume">
<mods:number id="64EE8F58BDE02613734D2741A45EA3E4">2017</mods:number>
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<mods:number id="F72B0ECD60A2C7930D3E230F89A6D1BB">592</mods:number>
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<mods:end id="98C432698E84BF7079E8AA2F49B9AF58">16</mods:end>
</mods:extent>
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<mods:classification id="A152A048BF5FFC943D7E2957CE9166DC">journal article</mods:classification>
<mods:identifier id="99510D78BFDD5C6A95686E79522C1DE8" type="CLB-Dataset">55466</mods:identifier>
<mods:identifier id="703DBE57392BC32A1A839394AAA2AAA3" type="DOI">10.5281/zenodo.5169525</mods:identifier>
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<mods:identifier id="1D77C8C09EACDB3DB6F185DC201E567B" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="298459D931E51C57DF46D6E079E0663D" type="Zenodo-Dep">5169525</mods:identifier>
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<treatment id="03DA133F751EB13A4BC2FA0288EE6892" ID-DOI="http://doi.org/10.5281/zenodo.6567174" ID-GBIF-Taxon="195537733" ID-Zenodo-Dep="6567174" LSID="urn:lsid:plazi:treatment:03DA133F751EB13A4BC2FA0288EE6892" httpUri="http://treatment.plazi.org/id/03DA133F751EB13A4BC2FA0288EE6892" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
<subSubSection id="C369F1A2751EB1384BC2FA0288826BED" box="[192,649,1441,1466]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BCCA229751EB1384BC2FA0288826BED" blockId="5.[192,649,1441,1466]" box="[192,649,1441,1466]" pageId="5" pageNumber="4">
<heading id="D0841545751EB1384BC2FA0288826BED" bold="true" box="[192,649,1441,1466]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<taxonomicName id="4C73D9AA751EB1384BC2FA0288826BED" ID-CoL="3F8ZT" authority="(Nonfried)" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[192,649,1441,1466]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384BC2FA0288826BED" bold="true" box="[192,649,1441,1466]" pageId="5" pageNumber="4">
<emphasis id="B9077E3B751EB1384BC2FA028BFB6BEE" bold="true" box="[192,496,1441,1465]" italics="true" pageId="5" pageNumber="4">Ganelius oberndorferi</emphasis>
(Nonfried)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369F1A2751EB1384BC2FA478EAD684A" pageId="5" pageNumber="4" type="reference_group">
<paragraph id="8BCCA229751EB1384BC2FA4789DF6BAB" blockId="5.[192,1190,1508,1565]" box="[192,980,1508,1532]" pageId="5" pageNumber="4">
<treatmentCitationGroup id="AB638507751EB1384BC2FA4789DF6BAB" box="[192,980,1508,1532]" pageId="5" pageNumber="4">
<treatmentCitation id="0AD28438751EB1384BC2FA4788B76BAB" author="Nonfried, A. F." box="[192,700,1508,1532]" page="118" pageId="5" pageNumber="4" year="1892">
<taxonomicName id="4C73D9AA751EB1384BC2FA4788B76BAB" ID-CoL="6RXVP" authority="Nonfried 1892: 118" authorityName="Nonfried" authorityPageNumber="118" authorityYear="1892" box="[192,700,1508,1532]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384BC2FA478BC16BAB" box="[192,458,1508,1532]" italics="true" pageId="5" pageNumber="4">Nigidius oberndorferi</emphasis>
<bibRefCitation id="EFE2DFD8751EB1384AD3FA4788B76BAB" author="Nonfried, A. F." box="[465,700,1508,1532]" pageId="5" pageNumber="4" pagination="105 - 111" refId="ref7558" refString="Nonfried, A. F. 1892. Beitrage zur Coleopterenfauna von Africa und Madagascar. Entomologische Nachrichten 18: 105 - 111." type="journal article" year="1892">Nonfried 1892: 118</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original combination)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCA229751EB1384BC2F9A78EAD684A" blockId="5.[192,1190,1508,1565]" box="[192,1190,1540,1565]" pageId="5" pageNumber="4">
<treatmentCitationGroup id="AB638507751EB1384BC2F9A78EAD684A" box="[192,1190,1540,1565]" pageId="5" pageNumber="4">
<treatmentCitation id="0AD28438751EB1384BC2F9A788B9684A" author="Benesh, B." box="[192,690,1540,1565]" page="44" pageId="5" pageNumber="4" year="1943">
<taxonomicName id="4C73D9AA751EB1384BC2F9A788B9684A" ID-CoL="6RXVN" authority="Benesh, 1943: 44" authorityName="Benesh" authorityPageNumber="44" authorityYear="1943" box="[192,690,1540,1565]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="passaliformis">
<emphasis id="B9077E3B751EB1384BC2F9A78BD2684B" box="[192,473,1540,1564]" italics="true" pageId="5" pageNumber="4">Nigidius passaliformis</emphasis>
<bibRefCitation id="EFE2DFD8751EB1384AE2F9A788B9684A" author="Benesh, B." box="[480,690,1540,1565]" pageId="5" pageNumber="4" pagination="37 - 49" refId="ref7324" refString="Benesh, B. 1943. Descriptions of some new exotic species of stag beetles. Psyche 50: 37 - 49." type="journal article" year="1943">Benesh, 1943: 44</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original combination),
<emphasis id="B9077E3B751EB13848DAF9A68EAD684A" bold="true" box="[984,1190,1541,1565]" pageId="5" pageNumber="4">new synonymy</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C369F1A2751EB13A4BC2F9E5884D6F6D" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BCCA229751EB1384BC2F9E5884C6896" blockId="5.[192,1442,1606,1926]" pageId="5" pageNumber="4">
<materialsCitation id="3B1BA874751EB1384BC2F9E588486896" ID-GBIF-Occurrence="3776282320" collectionCode="MNHN" country="Madagascar" determinerName="M. J. Paulsen" location="Madagascar" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<emphasis id="B9077E3B751EB1384BC2F9E58B806809" bold="true" box="[192,395,1606,1630]" pageId="5" pageNumber="4">Type material.</emphasis>
<typeStatus id="54C81C8B751EB1384A90F9E48BF36808" box="[402,504,1607,1631]" pageId="5" pageNumber="4" type="neotype">Neotype</typeStatus>
male of
<taxonomicName id="4C73D9AA751EB1384960F9E589EA6809" authority="Nonfried" authorityName="Nonfried" authorityYear="1892" box="[610,993,1606,1630]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384960F9E589676809" box="[610,876,1606,1630]" italics="true" pageId="5" pageNumber="4">Nigidius oberndorferi</emphasis>
Nonfried
</taxonomicName>
(
<figureCitation id="1348BEAC751EB13848F0F9E58E316809" box="[1010,1082,1606,1630]" captionStart="Figures 5–8" captionStartId="6.[192,288,1875,1897]" captionTargetBox="[208,1419,355,1726]" captionTargetId="figure-48@6.[191,1439,311,1741]" captionTargetPageId="6" captionText="Figures 5–8. Ganelius oberndorferi (Nonfried, 1892). 5) Male neotype, dorsal habitus. 6) Female, dorsal habitus. 7) Neotype labels. 8) Male genitalia, flagellum, 41 mm." figureDoi="http://doi.org/10.5281/zenodo.5169529" httpUri="https://zenodo.org/record/5169529/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
),
<collectionCode id="ED623AEC751EB1384F53F9E48EA46809" box="[1105,1199,1607,1630]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="4">MNHN</collectionCode>
,
<emphasis id="B9077E3B751EB1384FB9F9E58FAB6809" bold="true" box="[1211,1440,1606,1630]" pageId="5" pageNumber="4">here designated,</emphasis>
labeled (
<figureCitation id="1348BEAC751EB1384A28F9C48B786828" box="[298,371,1639,1663]" captionStart="Figures 5–8" captionStartId="6.[192,288,1875,1897]" captionTargetBox="[208,1419,355,1726]" captionTargetId="figure-48@6.[191,1439,311,1741]" captionTargetPageId="6" captionText="Figures 5–8. Ganelius oberndorferi (Nonfried, 1892). 5) Male neotype, dorsal habitus. 6) Female, dorsal habitus. 7) Neotype labels. 8) Male genitalia, flagellum, 41 mm." figureDoi="http://doi.org/10.5281/zenodo.5169529" httpUri="https://zenodo.org/record/5169529/files/figure.png" pageId="5" pageNumber="4">Fig. 7</figureCitation>
): a) “
<collectingCountry id="F364E2B9751EB1384AB7F9C488406828" box="[437,587,1639,1663]" pageId="5" pageNumber="4">Madagascar</collectingCountry>
/ en|au Nossi Bé / Dr. Sicard”; b) on red paper “
<taxonomicName id="4C73D9AA751EB1384FA4F9C48F1A6828" box="[1190,1297,1639,1663]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384FA4F9C48F1A6828" box="[1190,1297,1639,1663]" italics="true" pageId="5" pageNumber="4">Nigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384E28F9C48AE268F7" authorityName="Nonfried" authorityYear="1892" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384E28F9C48AE268F7" italics="true" pageId="5" pageNumber="4">oberndorferi</emphasis>
</taxonomicName>
<specimenCount id="9D7569A0751EB1384BF0F92B8B0D68C8" box="[242,262,1672,1695]" pageId="5" pageNumber="4" type="male">♂</specimenCount>
/
<bibRefCitation id="EFE2DFD8751EB1384A1FF92B8BDE68F7" author="Nonfried, A. F." box="[285,469,1672,1696]" pageId="5" pageNumber="4" pagination="105 - 111" refId="ref7558" refString="Nonfried, A. F. 1892. Beitrage zur Coleopterenfauna von Africa und Madagascar. Entomologische Nachrichten 18: 105 - 111." type="journal article" year="1892">Nonfried, 1892</bibRefCitation>
/
<typeStatus id="54C81C8B751EB1384AEFF92B887F68F7" box="[493,628,1672,1696]" pageId="5" pageNumber="4" type="neotype">NEOTYPE</typeStatus>
/ des. M.J. Paulsen 2017”; b) “
<taxonomicName id="4C73D9AA751EB13848ECF92B8E5768F7" authorityName="Benesh" authorityYear="1955" box="[1006,1116,1672,1696]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB13848ECF92B8E5768F7" box="[1006,1116,1672,1696]" italics="true" pageId="5" pageNumber="4">Ganelius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384F76F92B8F0068F7" authorityName="Nonfried" authorityYear="1892" box="[1140,1291,1672,1696]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384F76F92B8F0068F7" box="[1140,1291,1672,1696]" italics="true" pageId="5" pageNumber="4">oberndorferi</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFE2DFD8751EB1384E29F92B8AF66896" author="Nonfried, A. F." pageId="5" pageNumber="4" pagination="105 - 111" refId="ref7558" refString="Nonfried, A. F. 1892. Beitrage zur Coleopterenfauna von Africa und Madagascar. Entomologische Nachrichten 18: 105 - 111." type="journal article" year="1892">Nonfried, 1892</bibRefCitation>
) / det.
<determinerName id="7E104EED751EB1384A53F90A8BF96896" box="[337,498,1705,1729]" pageId="5" pageNumber="4">M.J. Paulsen</determinerName>
2017”
</materialsCitation>
.
</paragraph>
<paragraph id="8BCCA229751EB1384BF2F969897969D1" blockId="5.[192,1442,1606,1926]" pageId="5" pageNumber="4">
<materialsCitation id="3B1BA874751EB1384BF2F969896569D1" ID-GBIF-Occurrence="3776282312" collectorName="B. Benesh" country="Madagascar" determinerName="H. Bomans" location="Madagascar" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54C81C8B751EB1384BF2F9698B5468B5" box="[240,351,1738,1762]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
female of
<taxonomicName id="4C73D9AA751EB1384AE4F969896368B5" authority="Benesh" authorityName="Benesh" authorityYear="1943" box="[486,872,1738,1762]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="passaliformis">
<emphasis id="B9077E3B751EB1384AE4F969890B68B5" box="[486,768,1738,1762]" italics="true" pageId="5" pageNumber="4">Nigidius passaliformis</emphasis>
Benesh
</taxonomicName>
, Museum of Comparative Zoology, labeled: a) handwritten, “West Africa / coll. Harvard”; b) handwritten, “
<originalDetermination id="A86D1AD5751EB13848AFF9498F466955" box="[941,1357,1770,1795]" pageId="5" pageNumber="4">
<taxonomicName id="4C73D9AA751EB13848AFF9498E026955" authorityName="De Haan" authorityYear="1833" box="[941,1033,1770,1794]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB13848AFF9498E026955" box="[941,1033,1770,1794]" italics="true" pageId="5" pageNumber="4">Eudora</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384F23F9498ECC6955" authorityName="Benesh" authorityYear="1943" box="[1057,1223,1770,1794]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="passaliformis">
<emphasis id="B9077E3B751EB1384F23F9498ECC6955" box="[1057,1223,1770,1794]" italics="true" pageId="5" pageNumber="4">passaliformis</emphasis>
</taxonomicName>
/ (Benesh)
</originalDetermination>
”; c) on red paper, “HoloTYPE /
<collectorName id="2686C7FF751EB1384AE4F8AF886F6974" box="[486,612,1803,1827]" pageId="5" pageNumber="4">B. Benesh</collectorName>
”; d) on red paper, “M.C.Z. / Type / 25913”; e) handwritten, “
<taxonomicName id="4C73D9AA751EB1384E37F8A88FAB6974" box="[1333,1440,1803,1827]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384E37F8A88FAB6974" box="[1333,1440,1803,1827]" italics="true" pageId="5" pageNumber="4">Nigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384BCDF88F8B7E6913" authorityName="Benesh" authorityYear="1943" box="[207,373,1836,1860]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="passaliformis">
<emphasis id="B9077E3B751EB1384BCDF88F8B7E6913" box="[207,373,1836,1860]" italics="true" pageId="5" pageNumber="4">passaliformis</emphasis>
</taxonomicName>
/ Benesh”; f) on orange paper, “Allotype” (sic); g) “
<determinerName id="7E104EED751EB13848E6F88F8E676913" box="[996,1132,1836,1860]" pageId="5" pageNumber="4">H. Bomans</determinerName>
det. 196 [72] / [=
<specimenCount id="9D7569A0751EB1384E45F88E8F506913" box="[1351,1371,1837,1860]" pageId="5" pageNumber="4" type="female">♀</specimenCount>
<taxonomicName id="4C73D9AA751EB1384E61F88E8AF16932" authorityName="Benesh" authorityYear="1955" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384E61F88E8AF16932" italics="true" pageId="5" pageNumber="4">Ganelius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384A11F8EE8BFB6932" box="[275,496,1869,1893]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB1384A11F8EE8BFB6932" box="[275,496,1869,1893]" italics="true" pageId="5" pageNumber="4">madagascariensis</emphasis>
</taxonomicName>
/ Cast.]”; h) handwritten, “Origin certainly / inaccurate – / This insect seen only in
<collectingCountry id="F364E2B9751EB1384A1FF8CD8BB869D1" box="[285,435,1902,1926]" pageId="5" pageNumber="4">Madagascar</collectingCountry>
/ !!”. Images studied (
<bibRefCitation id="EFE2DFD8751EB13849BDF8CD896C69D1" author="Perkins, P." box="[703,871,1902,1926]" pageId="5" pageNumber="4" refId="ref7582" refString="Perkins, P. 2010. MCZ Type Database, Harvard University. (Available at ~ http: // http: // 140.247.96.247 / mcz /. Last accessed November 2017.)" type="url" year="2010">Perkins 2010</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="DF0CF2A1751DB13B4BC2F8F0892869D2" ID-DOI="http://doi.org/10.5281/zenodo.5169529" ID-Zenodo-Dep="5169529" httpUri="https://zenodo.org/record/5169529/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,288,1875,1897]" targetBox="[208,1419,355,1726]" targetPageId="6">
<paragraph id="8BCCA229751DB13B4BC2F8F0892869D2" blockId="6.[192,1440,1875,1926]" pageId="6" pageNumber="5">
<emphasis id="B9077E3B751DB13B4BC2F8F08B5F693F" bold="true" box="[192,340,1875,1897]" pageId="6" pageNumber="5">Figures 5–8.</emphasis>
<taxonomicName id="4C73D9AA751DB13B4A58F8F08902693E" authority="(Nonfried, 1892)" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[346,777,1875,1897]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751DB13B4A58F8F08847693F" box="[346,588,1875,1896]" italics="true" pageId="6" pageNumber="5">Ganelius oberndorferi</emphasis>
(
<bibRefCitation id="EFE2DFD8751DB13B495BF8F08909693E" author="Nonfried, A. F." box="[601,770,1875,1897]" pageId="6" pageNumber="5" pagination="105 - 111" refId="ref7558" refString="Nonfried, A. F. 1892. Beitrage zur Coleopterenfauna von Africa und Madagascar. Entomologische Nachrichten 18: 105 - 111." type="journal article" year="1892">Nonfried, 1892</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B9077E3B751DB13B4811F8F08921693E" bold="true" box="[787,810,1875,1897]" pageId="6" pageNumber="5">5)</emphasis>
Male neotype, dorsal habitus.
<emphasis id="B9077E3B751DB13B4F7CF8F08E9E693E" bold="true" box="[1150,1173,1875,1897]" pageId="6" pageNumber="5">6)</emphasis>
Female, dorsal habitus.
<emphasis id="B9077E3B751DB13B4BC2F8CC8ADC69D2" bold="true" box="[192,215,1903,1925]" pageId="6" pageNumber="5">7)</emphasis>
Neotype labels.
<emphasis id="B9077E3B751DB13B4A8CF8CC8BAD69D2" bold="true" box="[398,422,1903,1925]" pageId="6" pageNumber="5">8)</emphasis>
Male genitalia, flagellum, 41 mm.
</paragraph>
</caption>
<paragraph id="8BCCA229751CB13A4BC2FF61884D6F6D" blockId="7.[192,1441,193,314]" pageId="7" pageNumber="6">
<emphasis id="B9077E3B751CB13A4BC2FF618BB66E8E" bold="true" box="[192,445,193,218]" pageId="7" pageNumber="6">Non-type material</emphasis>
(
<figureCitation id="1348BEAC751CB13A4ACFFF6188286E8D" box="[461,547,194,218]" captionStart="Figures 33–37" captionStartId="17.[192,298,1697,1721]" captionTargetBox="[233,1386,399,1638]" captionTargetId="figure-36@17.[192,1440,364,1666]" captionTargetPageId="17" captionText="Figures 33–37. Species distributions. 33) Ganelius madagascariensis. 34) G. oberndorferi. 35) G. gnamptus. 36) G. zombi. 37) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169541" httpUri="https://zenodo.org/record/5169541/files/figure.png" pageId="7" pageNumber="6">Fig. 34</figureCitation>
).
<materialsCitation id="3B1BA874751CB13A4939FF628F436E8D" ID-GBIF-Occurrence="3776282346" box="[571,1352,193,218]" collectionCode="MNHN" location="Ambanja" pageId="7" pageNumber="6" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Antsiranana">
<emphasis id="B9077E3B751CB13A4939FF62891C6E8E" bold="true" box="[571,791,193,217]" pageId="7" pageNumber="6">MADAGASCAR:</emphasis>
<collectingRegion id="49B76CCB751CB13A481DFF6189F96E8D" box="[799,1010,194,218]" country="Madagascar" name="Antsiranana" pageId="7" pageNumber="6">ANTSIRANANA</collectingRegion>
:
<location id="8EACF4F2751CB13A48FFFF618E666E8D" LSID="urn:lsid:plazi:treatment:03DA133F751EB13A4BC2FA0288EE6892:8EACF4F2751CB13A48FFFF618E666E8D" box="[1021,1133,194,218]" name="Ambanja" pageId="7" pageNumber="6" stateProvince="Antsiranana">Ambanja</location>
(
<specimenCount id="9D7569A0751CB13A4F7FFF618EA86E8D" box="[1149,1187,194,218]" pageId="7" pageNumber="6" type="male">2♂</specimenCount>
,
<specimenCount id="9D7569A0751CB13A4FB2FF618ED86E8D" box="[1200,1235,194,218]" pageId="7" pageNumber="6" type="female">1♀</specimenCount>
;
<collectionCode id="ED623AEC751CB13A4FE0FF618F4A6E8D" box="[1250,1345,194,218]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="6">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA874751CB13A4E57FF618B9C6EAD" ID-GBIF-Occurrence="3776282332" collectionCode="MNHN" pageId="7" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
Ambilobe (
<specimenCount id="9D7569A0751CB13A4A00FF418B2E6EAD" box="[258,293,226,250]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
,
<collectionCode id="ED623AEC751CB13A4A33FF418B9B6EAD" box="[305,400,226,250]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="6">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA874751CB13A4AA0FF418F116EAD" ID-GBIF-Occurrence="3776282302" box="[418,1306,226,250]" collectionCode="MHNG" isTailEnumeration="true" location="Passandava Bay" municipality="Fort Ambodimadiro" pageId="7" pageNumber="6" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Antsiranana">
“Ambodinadiro” (
<collectingMunicipality id="6BA83853751CB13A4971FF4189636EAD" box="[627,872,226,250]" pageId="7" pageNumber="6">Fort Ambodimadiro</collectingMunicipality>
,
<location id="8EACF4F2751CB13A4871FF418E366EAD" LSID="urn:lsid:plazi:treatment:03DA133F751EB13A4BC2FA0288EE6892:8EACF4F2751CB13A4871FF418E366EAD" box="[883,1085,226,250]" municipality="Fort Ambodimadiro" name="Passandava Bay" pageId="7" pageNumber="6">Passandava Bay</location>
) (
<specimenCount id="9D7569A0751CB13A4F57FF418E736EAD" box="[1109,1144,226,250]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
,
<specimenCount id="9D7569A0751CB13A4F86FF418EAC6EAD" box="[1156,1191,226,250]" pageId="7" pageNumber="6" type="female">1♀</specimenCount>
;
<collectionCode id="ED623AEC751CB13A4FB6FF418F196EAD" box="[1204,1298,226,250]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="6" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA874751CB13A4E26FF418B6B6F4D" ID-GBIF-Occurrence="3776282319" collectionCode="MNHN" location="Ankarana" pageId="7" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
<location id="8EACF4F2751CB13A4E26FF418FAB6EAD" LSID="urn:lsid:plazi:treatment:03DA133F751EB13A4BC2FA0288EE6892:8EACF4F2751CB13A4E26FF418FAB6EAD" box="[1316,1440,226,250]" name="Ankarana" pageId="7" pageNumber="6" stateProvince="Antsiranana">Ankarana</location>
(
<specimenCount id="9D7569A0751CB13A4BC8FEA18AE66F4D" box="[202,237,258,282]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
;
<collectionCode id="ED623AEC751CB13A4BFBFEA18B536F4D" box="[249,344,258,282]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="6">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA874751CB13A4A68FEA0884A6F6D" ID-GBIF-Occurrence="3776282305" collectionCode="MNHN, MJPC, IRSNB" county="Nosy Be" location="Maroantsetra" municipality="Foret d'Anobohazo" pageId="7" pageNumber="6" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="Mahajanga">
<collectingCounty id="62ADDAA5751CB13A4A68FEA08BC56F4D" box="[362,462,259,283]" pageId="7" pageNumber="6">Nosy Be</collectingCounty>
(
<specimenCount id="9D7569A0751CB13A4ADCFEA1880A6F4D" box="[478,513,258,282]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
,
<specimenCount id="9D7569A0751CB13A490EFEA1883B6F4D" box="[524,560,258,282]" pageId="7" pageNumber="6" type="female">3♀</specimenCount>
;
<collectionCode id="ED623AEC751CB13A493EFEA188976F4D" box="[572,668,258,282]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="6">MNHN</collectionCode>
).
<collectingRegion id="49B76CCB751CB13A49AFFEA1896C6F4D" box="[685,871,258,282]" country="Madagascar" name="Mahajanga" pageId="7" pageNumber="6">MAHAJANGA</collectingRegion>
:
<collectingMunicipality id="6BA83853751CB13A4872FEA18E586F4D" box="[880,1107,258,282]" pageId="7" pageNumber="6">Forêt d’Anobohazo</collectingMunicipality>
(
<specimenCount id="9D7569A0751CB13A4F61FEA18E8F6F4D" box="[1123,1156,258,282]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
<collectionCode id="ED623AEC751CB13A4F89FEA18ED76F4D" box="[1163,1244,258,282]" pageId="7" pageNumber="6">MJPC</collectionCode>
).
<collectingRegion id="49B76CCB751CB13A4FECFEA18F966F4D" box="[1262,1437,258,282]" country="Madagascar" name="Toamasina" pageId="7" pageNumber="6">TOAMASINA</collectingRegion>
:
<location id="8EACF4F2751CB13A4BC2FE808B626F6D" LSID="urn:lsid:plazi:treatment:03DA133F751EB13A4BC2FA0288EE6892:8EACF4F2751CB13A4BC2FE808B626F6D" box="[192,361,291,314]" county="Nosy Be" municipality="Foret d'Anobohazo" name="Maroantsetra" pageId="7" pageNumber="6" stateProvince="Mahajanga">Maroantsetra</location>
(
<specimenCount id="9D7569A0751CB13A4A79FE818B956F6D" box="[379,414,290,314]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
,
<specimenCount id="9D7569A0751CB13A4AA9FE818BDB6F6D" box="[427,464,290,314]" pageId="7" pageNumber="6" type="female">2♀</specimenCount>
;
<collectionCode id="ED623AEC751CB13A4ADFFE8188326F6D" box="[477,569,290,314]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="7" pageNumber="6">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369F1A2751CB13A4BC2FEC189F56C6B" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BCCA229751CB13A4BC2FEC189F56C6B" blockId="7.[191,1441,354,572]" pageId="7" pageNumber="6">
<emphasis id="B9077E3B751CB13A4BC2FEC18B5B6F2D" bold="true" box="[192,336,354,378]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
In males of
<taxonomicName id="4C73D9AA751CB13A4AE0FEC088976F2C" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[482,668,355,379]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A4AE0FEC088976F2C" box="[482,668,355,379]" italics="true" pageId="7" pageNumber="6">G. oberndorferi</emphasis>
</taxonomicName>
the dorso-basal tooth on the vertical ramus is weakly produced and obtuse and much lower than the interior median denticle of the ramus; in
<taxonomicName id="4C73D9AA751CB13A4F70FE208F7A6FCC" box="[1138,1393,387,411]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751CB13A4F70FE208F7A6FCC" box="[1138,1393,387,411]" italics="true" pageId="7" pageNumber="6">G. madagascariensis</emphasis>
</taxonomicName>
the external dorso-basal tooth is strongly produced and near to the interior median denticle of the ramus (see
<figureCitation id="1348BEAC751CB13A4BF5FE608B706F8C" box="[247,379,451,475]" captionStart="Figures 26–27" captionStartId="16.[192,298,837,861]" captionTargetBox="[540,1123,396,805]" captionTargetId="figure-47@16.[509,1123,386,805]" captionTargetPageId="16" captionText="Figures 26–27. Comparison of male mandibles. Arrows indicate relative position of dorsal basal teeth and median denticles. 26) Ganelius madagascariensis. 27) G. oberndorferi." figureDoi="http://doi.org/10.5281/zenodo.5169537" httpUri="https://zenodo.org/record/5169537/files/figure.png" pageId="7" pageNumber="6">Fig. 26–27</figureCitation>
). In females of
<taxonomicName id="4C73D9AA751CB13A492DFE6788E26F8C" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[559,745,451,475]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A492DFE6788E26F8C" box="[559,745,451,475]" italics="true" pageId="7" pageNumber="6">G. oberndorferi</emphasis>
</taxonomicName>
the canthus is sloping and concave (
<figureCitation id="1348BEAC751CB13A4FA1FE608EF16F8B" box="[1187,1274,451,476]" captionStart="Figures 28–32" captionStartId="16.[192,302,1675,1699]" captionTargetBox="[230,1426,1030,1622]" captionTargetId="figure-48@16.[192,1440,1020,1634]" captionTargetPageId="16" captionText="Figures 28–32. Comparison of ocular canthi, females. 28) Ganelius madagascariensis. 29) G. oberndorferi. 30) G. gnamptus. 31) G. zombi. 32) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169539" httpUri="https://zenodo.org/record/5169539/files/figure.png" pageId="7" pageNumber="6">Fig. 29</figureCitation>
), not straight or almost rounded as with female
<emphasis id="B9077E3B751CB13A4969FE47897F6FAB" box="[619,884,484,508]" italics="true" pageId="7" pageNumber="6">
<taxonomicName id="4C73D9AA751CB13A4969FE47897B6FAB" box="[619,880,484,508]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="madagascariensis">G. madagascariensis</taxonomicName>
.
</emphasis>
In addition, the tubercle above the antennal insertion is usually much more elevated in females of
<taxonomicName id="4C73D9AA751CB13A484AFDA78E0A6C4B" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[840,1025,516,540]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A484AFDA78E0A6C4B" box="[840,1025,516,540]" italics="true" pageId="7" pageNumber="6">G. oberndorferi</emphasis>
</taxonomicName>
(
<figureCitation id="1348BEAC751CB13A4F0DFDA78E5F6C4B" box="[1039,1108,516,540]" captionStart="Figures 5–8" captionStartId="6.[192,288,1875,1897]" captionTargetBox="[208,1419,355,1726]" captionTargetId="figure-48@6.[191,1439,311,1741]" captionTargetPageId="6" captionText="Figures 5–8. Ganelius oberndorferi (Nonfried, 1892). 5) Male neotype, dorsal habitus. 6) Female, dorsal habitus. 7) Neotype labels. 8) Male genitalia, flagellum, 41 mm." figureDoi="http://doi.org/10.5281/zenodo.5169529" httpUri="https://zenodo.org/record/5169529/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
). The flagellum of the male genitalia (
<figureCitation id="1348BEAC751CB13A4A3FFD878B886C6B" box="[317,387,548,572]" captionStart="Figures 5–8" captionStartId="6.[192,288,1875,1897]" captionTargetBox="[208,1419,355,1726]" captionTargetId="figure-48@6.[191,1439,311,1741]" captionTargetPageId="6" captionText="Figures 5–8. Ganelius oberndorferi (Nonfried, 1892). 5) Male neotype, dorsal habitus. 6) Female, dorsal habitus. 7) Neotype labels. 8) Male genitalia, flagellum, 41 mm." figureDoi="http://doi.org/10.5281/zenodo.5169529" httpUri="https://zenodo.org/record/5169529/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
) is somewhat shorter than in
<emphasis id="B9077E3B751CB13A49F4FD8689F56C6B" box="[758,1022,548,572]" italics="true" pageId="7" pageNumber="6">
<taxonomicName id="4C73D9AA751CB13A49F4FD8689F16C6B" box="[758,1018,548,572]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="madagascariensis">G. madagascariensis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C369F1A2751CB13A4BC2FDC688EE6892" pageId="7" pageNumber="6" type="discussion">
<paragraph id="8BCCA229751CB13A4BC2FDC68FAB6DA8" blockId="7.[190,1442,613,1733]" pageId="7" pageNumber="6">
<emphasis id="B9077E3B751CB13A4BC2FDC68B486C2A" bold="true" box="[192,323,613,637]" pageId="7" pageNumber="6">Remarks.</emphasis>
The number of specimens examined by
<bibRefCitation id="EFE2DFD8751CB13A4823FDC689EB6C2A" author="Nonfried, A. F." box="[801,992,613,637]" pageId="7" pageNumber="6" pagination="105 - 111" refId="ref7558" refString="Nonfried, A. F. 1892. Beitrage zur Coleopterenfauna von Africa und Madagascar. Entomologische Nachrichten 18: 105 - 111." type="journal article" year="1892">Nonfried (1892)</bibRefCitation>
is not stated, but may have consisted of only a single specimen. The following information was contributed by S. Boucher, MNHN, concerning the whereabouts of Nonfried’s type (pers. comm.). No probable type specimens of
<taxonomicName id="4C73D9AA751CB13A4FE7FD068B266C8A" authority="Nonfried" authorityName="Nonfried" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A4FE7FD068EF26CEB" box="[1253,1273,677,700]" italics="true" pageId="7" pageNumber="6">G</emphasis>
.
<emphasis id="B9077E3B751CB13A4E0BFD068FAB6CEA" box="[1289,1440,677,701]" italics="true" pageId="7" pageNumber="6">oberndorferi</emphasis>
Nonfried
</taxonomicName>
have been located in MNHN, where Nonfried specimens received a special label printed by Oberthür and thus are readily identifiable. This part of Nonfried’s collection arrived in Paris from Oberthür in 1952, but another large part of Nonfried’s scarabaeoids were deposited at ZMHB, Berlin. All
<taxonomicName id="4C73D9AA751CB13A4BECFC858B576D69" authorityName="Benesh" authorityYear="1955" box="[238,348,806,830]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751CB13A4BECFC858B576D69" box="[238,348,806,830]" italics="true" pageId="7" pageNumber="6">Ganelius</emphasis>
</taxonomicName>
material from Berlin was examined and does not appear to have originated with Nonfried. Therefore a
<typeStatus id="54C81C8B751CB13A4AA2FCE4880B6D08" box="[416,512,839,863]" pageId="7" pageNumber="6" type="neotype">neotype</typeStatus>
specimen is chosen from the original locality to stabilize the nomenclature and tie the name to the appropriate species. The identity of
<taxonomicName id="4C73D9AA751CB13A4897FCC48E446D29" box="[917,1103,870,894]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A4897FCC48E446D29" box="[917,1103,870,894]" italics="true" pageId="7" pageNumber="6">G. oberndorferi</emphasis>
</taxonomicName>
was confused with
<taxonomicName id="4C73D9AA751CB13A4E3EFCC48AD96DC8" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B751CB13A4E3EFCC48AD96DC8" italics="true" pageId="7" pageNumber="6">A. nageli</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE2DFD8751CB13A4BFFFC248BB56DC8" author="Benesh, B." box="[253,446,903,927]" pageId="7" pageNumber="6" pagination="59 - 76" refId="ref7381" refString="Benesh, B. 1955 b. Some further notes on the stagbeetles, with especial reference to Figulinae (Coleoptera: Lucanidae). Transactions of the American Entomological Society 81 (2): 59 - 76." type="journal article" year="1955">Benesh (1955b)</bibRefCitation>
, in part. His second figure of
<taxonomicName id="4C73D9AA751CB13A4826FC2489D46DC8" box="[804,991,903,927]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A4826FC2489D46DC8" box="[804,991,903,927]" italics="true" pageId="7" pageNumber="6">G. oberndorferi</emphasis>
</taxonomicName>
(Benesh’s fig. 15) is a specimen of
<taxonomicName id="4C73D9AA751CB13A4E84FC248B076DE8" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B751CB13A4E84FC248B076DE8" italics="true" pageId="7" pageNumber="6">A. nageli</emphasis>
</taxonomicName>
from ‘Sandramora’ forest, near Vohilava in eastern Madagascar. Because
<bibRefCitation id="EFE2DFD8751CB13A4F90FC048F456DE8" author="Benesh, B." box="[1170,1358,935,959]" pageId="7" pageNumber="6" pagination="59 - 76" refId="ref7381" refString="Benesh, B. 1955 b. Some further notes on the stagbeetles, with especial reference to Figulinae (Coleoptera: Lucanidae). Transactions of the American Entomological Society 81 (2): 59 - 76." type="journal article" year="1955">Benesh (1955b)</bibRefCitation>
mixed two different species under the name, and
<bibRefCitation id="EFE2DFD8751CB13A49D2FC6489786D88" author="Didier, R." box="[720,883,967,991]" pageId="7" pageNumber="6" pagination="1 - 32" refId="ref7421" refString="Didier, R. 1928. Etude sur les Coleopteres Lucanides du globe. I. La vie et l &quot; oeuvre scientifique de H. Boileau 1866 - 1924. Catalogue de sa collection Librairie speciale Agricole, Paris Fascicule 1: 1 - 32." type="journal article" year="1928">Didier (1928)</bibRefCitation>
erroneously considered the species to belong to the genus
<taxonomicName id="4C73D9AA751CB13A4A59FC44883F6A57" authority="MacLeay" authorityName="MacLeay" box="[347,564,999,1024]" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751CB13A4A59FC448BB26DA8" box="[347,441,999,1023]" italics="true" pageId="7" pageNumber="6">Figulus</emphasis>
MacLeay
</taxonomicName>
, a
<typeStatus id="54C81C8B751CB13A4957FC4B88BE6A57" box="[597,693,1000,1024]" pageId="7" pageNumber="6" type="neotype">neotype</typeStatus>
is clearly necessary to resolve the identity of
<taxonomicName id="4C73D9AA751CB13A4FDCFC4B8F936DA8" box="[1246,1432,999,1023]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751CB13A4FDCFC4B8F936DA8" box="[1246,1432,999,1023]" italics="true" pageId="7" pageNumber="6">G. oberndorferi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCCA229751CB13A4BF2FBAB8B996B95" blockId="7.[190,1442,613,1733]" pageId="7" pageNumber="6">
Nonfried (1891) clearly described the mandibles and ocular canthus of this species. The mandibles are described as long, bent nearly vertically and reinforced at the bottom of the base with a large tooth, while the tip is made of two bifurcated small teeth. This description confuses the actual apex with the dorsal ramus, but it nevertheless clearly refers to a male
<taxonomicName id="4C73D9AA751CB13A486FFBCB89D06AD7" authorityName="Benesh" authorityYear="1955" box="[877,987,1128,1152]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751CB13A486FFBCB89D06AD7" box="[877,987,1128,1152]" italics="true" pageId="7" pageNumber="6">Ganelius</emphasis>
</taxonomicName>
. In the original description,
<bibRefCitation id="EFE2DFD8751CB13A4E31FBCB8B026AF6" author="Nonfried, A. F." pageId="7" pageNumber="6" pagination="105 - 111" refId="ref7558" refString="Nonfried, A. F. 1892. Beitrage zur Coleopterenfauna von Africa und Madagascar. Entomologische Nachrichten 18: 105 - 111." type="journal article" year="1892">Nonfried (1892)</bibRefCitation>
noted that it differed by its slimmer construction and differences in pronotal shape and mandibular form. The first two characters are variable based on body size. However, based on the male mandibles of material examined from the
<typeStatus id="54C81C8B751CB13A493DFB6A887F6AB6" box="[575,628,1225,1249]" pageId="7" pageNumber="6">type</typeStatus>
locality as described in the diagnosis, this species is distinct from
<emphasis id="B9077E3B751CB13A4BC2FB498BC16B56" box="[192,458,1257,1281]" italics="true" pageId="7" pageNumber="6">
<taxonomicName id="4C73D9AA751CB13A4BC2FB498BCD6B56" box="[192,454,1257,1281]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="madagascariensis">G. madagascariensis</taxonomicName>
.
</emphasis>
Based on the relatively few specimens available for study, the species may be restricted to extreme northern Madagascar near the
<typeStatus id="54C81C8B751CB13A4848FAA989746B75" box="[842,895,1290,1314]" pageId="7" pageNumber="6">type</typeStatus>
locality (
<figureCitation id="1348BEAC751CB13A48F2FAAA8E436B75" box="[1008,1096,1289,1314]" captionStart="Figures 33–37" captionStartId="17.[192,298,1697,1721]" captionTargetBox="[233,1386,399,1638]" captionTargetId="figure-36@17.[192,1440,364,1666]" captionTargetPageId="17" captionText="Figures 33–37. Species distributions. 33) Ganelius madagascariensis. 34) G. oberndorferi. 35) G. gnamptus. 36) G. zombi. 37) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169541" httpUri="https://zenodo.org/record/5169541/files/figure.png" pageId="7" pageNumber="6">Fig. 34</figureCitation>
). A series of specimens from the Lemoult collection in IRSNB are labeled as being from Maroantsetra. If correct these specimens are geographically separated from all others and possibly sympatric with
<taxonomicName id="4C73D9AA751CB13A4F4AFAE98F436B35" box="[1096,1352,1354,1378]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751CB13A4F4AFAE98F436B35" box="[1096,1352,1354,1378]" italics="true" pageId="7" pageNumber="6">G. madagascariensis</emphasis>
</taxonomicName>
, which seems unlikely. However, I do not have any information about whether these might be mislabeled with the locality actually referring to the port from which they were shipped, or if this is a common distributional pattern.
</paragraph>
<paragraph id="8BCCA229751CB13A4BF2FA6888EE6892" blockId="7.[190,1442,613,1733]" pageId="7" pageNumber="6">
<bibRefCitation id="EFE2DFD8751CB13A4BF2FA688B946BB4" author="Benesh, B." box="[240,415,1483,1507]" pageId="7" pageNumber="6" pagination="37 - 49" refId="ref7324" refString="Benesh, B. 1943. Descriptions of some new exotic species of stag beetles. Psyche 50: 37 - 49." type="journal article" year="1943">Benesh (1943)</bibRefCitation>
described
<taxonomicName id="4C73D9AA751CB13A4926FA6888E56BB4" authorityName="Benesh" authorityYear="1943" box="[548,750,1483,1507]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="passaliformis">
<emphasis id="B9077E3B751CB13A4926FA6888E56BB4" box="[548,750,1483,1507]" italics="true" pageId="7" pageNumber="6">G. passaliformis</emphasis>
</taxonomicName>
from ‘West Africa’, which
<bibRefCitation id="EFE2DFD8751CB13A4F31FA688FAB6BB4" author="Bartolozzi, L. &amp; K. Werner" box="[1075,1440,1483,1507]" pageId="7" pageNumber="6" refId="ref7286" refString="Bartolozzi, L., and K. Werner. 2004. Illustrated catalogue of the Lucanidae from Africa and Madagascar (Coleoptera: Lucanidae). Taita Publishers; Hradec Kralove, Czechia. 191 p." type="book" year="2004">Bartolozzi and Werner (2004)</bibRefCitation>
proposed might be referring to Madagascar since the genus is not known from mainland Africa. Benesh discusses the specimen as male, however it is a female, which he confirmed with a subsequent dissection (
<bibRefCitation id="EFE2DFD8751CB13A4A00F9888BBE6814" author="Benesh, B." box="[258,437,1579,1603]" pageId="7" pageNumber="6" pagination="57 - 63" refId="ref7347" refString="Benesh, B. 1955 a. On the genus Macrocrates, with notes on other stagbeetles (Coleoptera: Lucanidae). Entomological News 66 (3): 57 - 63." type="journal article" year="1955">Benesh 1955a</bibRefCitation>
). The
<typeStatus id="54C81C8B751CB13A4906F98888666814" box="[516,621,1579,1603]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
is entirely consistent with females of
<emphasis id="B9077E3B751CB13A4F53F98F8F1D6814" box="[1105,1302,1579,1603]" italics="true" pageId="7" pageNumber="6">
<taxonomicName id="4C73D9AA751CB13A4F53F98F8F196814" box="[1105,1298,1579,1603]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="oberndorferi">G. oberndorferi</taxonomicName>
.
</emphasis>
Specimens of that species on hand are indistinguishable from images of Benesh’s type, especially in the concave, sloping anterior margin of the head, elevated suprantennal tubercles, and vaulted appearance of the pronotum anteriorly in lateral view. Thus a name based on a single, abraded female specimen rather unsurprisingly is reduced here to synonymy.
</paragraph>
</subSubSection>
</treatment>
</document>
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