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<document id="79BC90CDBC9EC0E896BCC02E0A665CEC" ID-DOI="10.15298/rusentj.32.4.06" ID-ISSN="0132-8069" ID-Zenodo-Dep="13177446" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722592154512" checkinUser="felipe" docAuthor="Manukyan, A. R." docDate="2023" docId="03D687E5FFAEFFD1FE8B0CC2FD14F8D2" docLanguage="en" docName="RussEntomolJ.32.4.403-409.pdf" docOrigin="Russian Entomological Journal 32 (4)" docSource="http://dx.doi.org/10.15298/rusentj.32.4.06" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Vorstadt Manukyan 2023, gen.n." docType="treatment" docVersion="3" lastPageNumber="406" masterDocId="FFEFFF9DFFACFFD2FFBB0A52FF9AFFB3" masterDocTitle="New data on the subfamily Hybrizoninae (Hymenoptera: Ichneumonidae) in Baltic Amber" masterLastPageNumber="409" masterPageNumber="403" pageNumber="405" updateTime="1722631926422" updateUser="ExternalLinkService">
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<mods:title id="40E8EC3283D2EDBC8EE9F16E0865964C">New data on the subfamily Hybrizoninae (Hymenoptera: Ichneumonidae) in Baltic Amber</mods:title>
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<mods:title id="0CB04D1236055C6BA89BE3651A11B980">Russian Entomological Journal</mods:title>
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<mods:date id="273353D0388F24A64CA69FF0205EECE3">2023</mods:date>
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<treatment id="03D687E5FFAEFFD1FE8B0CC2FD14F8D2" ID-DOI="http://doi.org/10.5281/zenodo.13177668" ID-Zenodo-Dep="13177668" LSID="urn:lsid:plazi:treatment:03D687E5FFAEFFD1FE8B0CC2FD14F8D2" httpUri="http://treatment.plazi.org/id/03D687E5FFAEFFD1FE8B0CC2FD14F8D2" lastPageId="3" lastPageNumber="406" pageId="2" pageNumber="405">
<subSubSection id="C3656578FFAEFFD0FE8B0CC2FD07F914" box="[304,669,1680,1703]" pageId="2" pageNumber="405" type="nomenclature">
<paragraph id="8BC036F3FFAEFFD0FE8B0CC2FD07F914" blockId="2.[304,669,1680,1703]" box="[304,669,1680,1703]" pageId="2" pageNumber="405">
<heading id="D088819FFFAEFFD0FE8B0CC2FD07F914" box="[304,669,1680,1703]" fontSize="10" level="2" pageId="2" pageNumber="405" reason="2">
Genus
<taxonomicName id="4C7F4D70FFAEFFD0FEC10CC2FDD5F914" authority="Manukyan" authorityName="Manukyan" authorityYear="2023" box="[378,591,1680,1703]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="genus" status="gen.n.">
<emphasis id="B90BEAE1FFAEFFD0FEC10CC2FE49F914" box="[378,467,1680,1703]" italics="true" pageId="2" pageNumber="405">Vorstadt</emphasis>
Manukyan
</taxonomicName>
,
<emphasis id="B90BEAE1FFAEFFD0FDE20CC2FD07F914" bold="true" box="[601,669,1680,1703]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="A238579AFFAEFFD0FDE20CC2FD07F914" box="[601,669,1680,1703]" pageId="2" pageNumber="405" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656578FFAEFFD0FF580C9AFD6FF96E" box="[227,757,1736,1757]" pageId="2" pageNumber="405" type="materials_examined">
<paragraph id="8BC036F3FFAEFFD0FF580C9AFD6FF96E" blockId="2.[189,785,1736,1894]" box="[227,757,1736,1757]" pageId="2" pageNumber="405">
<typeStatus id="54C48851FFAEFFD0FF580C9AFE89F96E" box="[227,275,1736,1757]" pageId="2" pageNumber="405">Type</typeStatus>
species:
<taxonomicName id="4C7F4D70FFAEFFD0FED10C9AFD21F96E" authority="Manukyan" authorityName="Manukyan" authorityYear="2023" box="[362,699,1736,1757]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAEFFD0FED10C9AFDD6F96E" box="[362,588,1736,1757]" italics="true" pageId="2" pageNumber="405">Vorstadt ambermuseum</emphasis>
Manukyan
</taxonomicName>
,
<emphasis id="B90BEAE1FFAEFFD0FD7E0C9AFD6FF96E" bold="true" box="[709,757,1736,1757]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="A238579AFFAEFFD0FD7E0C9AFD6FF96E" box="[709,757,1736,1757]" pageId="2" pageNumber="405" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3656578FFAEFFD0FF580CB6FC8BF883" pageId="2" pageNumber="405" type="etymology">
<paragraph id="8BC036F3FFAEFFD0FF580CB6FC8BF883" blockId="2.[189,785,1736,1894]" pageId="2" pageNumber="405">
ETYMOLOGY. The genus name comes from the German word meaning “suburb”. This was the name of the
<collectingRegion id="49BBF811FFAEFFD0FD340CADFC8AF8A7" box="[655,784,1791,1812]" country="Russia" name="Yevrey" pageId="2" pageNumber="405">Jewish district</collectingRegion>
in Königsberg where the famous Amber Manufactory was located.
</paragraph>
</subSubSection>
<subSubSection id="C3656578FFAEFFD1FF580D64FD15FA59" lastPageId="3" lastPageNumber="406" pageId="2" pageNumber="405" type="diagnosis">
<paragraph id="8BC036F3FFAEFFD1FF580D64FEF8FACD" blockId="2.[189,785,1736,1894]" lastBlockId="3.[189,787,1251,1889]" lastPageId="3" lastPageNumber="406" pageId="2" pageNumber="405">
DIAGNOSIS. It is distinguished from other genera of recent and fossil
<taxonomicName id="4C7F4D70FFAEFFD0FEE10D03FE47F8D5" authorityName="Blanchard" authorityYear="1845" box="[346,477,1873,1894]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="subFamily" subFamily="Hybrizoninae">Hybrizoninae</taxonomicName>
by flagellomeres with distinct constriction in the middle, and also by three-segmented maxillary palps. The genus
<taxonomicName id="4C7F4D70FFAFFFD1FE360EACFD83FAA0" authorityName="Khalaim" authorityYear="2014" box="[397,537,1278,1299]" class="Insecta" family="Ichneumonidae" genus="Paxylobembra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1FFAFFFD1FE360EACFD83FAA0" box="[397,537,1278,1299]" italics="true" pageId="3" pageNumber="406">Paxylobembra</emphasis>
</taxonomicName>
with unknown morphology of antennae and palpi resembles
<taxonomicName id="4C7F4D70FFAFFFD1FDAA0F4BFDFBFA9D" authorityName="Manukyan" authorityYear="2023" box="[529,609,1305,1326]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1FFAFFFD1FDAA0F4BFDFBFA9D" box="[529,609,1305,1326]" italics="true" pageId="3" pageNumber="406">Vorstadt</emphasis>
</taxonomicName>
<emphasis id="B90BEAE1FFAFFFD1FDDD0F4BFD39FA9D" bold="true" box="[614,675,1305,1326]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="A238579AFFAFFFD1FDDD0F4BFD39FA9D" box="[614,675,1305,1326]" pageId="3" pageNumber="406" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
by absence of notauli and S-curved vein 4Rs. However, the latter genus differs from
<taxonomicName id="4C7F4D70FFAFFFD1FE8F0F1DFE5AFAD7" authorityName="Khalaim" authorityYear="2014" box="[308,448,1359,1380]" class="Insecta" family="Ichneumonidae" genus="Paxylobembra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1FFAFFFD1FE8F0F1DFE5AFAD7" box="[308,448,1359,1380]" italics="true" pageId="3" pageNumber="406">Paxylobembra</emphasis>
</taxonomicName>
by much larger body size and presence of vein r-m.
</paragraph>
<caption id="DF00667BFFAEFFD0FF000DDAFDEEF7A0" ID-DOI="http://doi.org/10.5281/zenodo.13177452" ID-Zenodo-Dep="13177452" httpUri="https://zenodo.org/record/13177452/files/figure.png" pageId="2" pageNumber="405" startId="2.[187,225,1928,1947]" targetBox="[822,1419,1115,2041]" targetPageId="2" targetType="figure">
<paragraph id="8BC036F3FFAEFFD0FF000DDAFC8AF800" blockId="2.[187,784,1928,2067]" pageId="2" pageNumber="405">
<emphasis id="B90BEAE1FFAEFFD0FF000DDAFE91F828" bold="true" box="[187,267,1928,1947]" pageId="2" pageNumber="405">Figs 45.</emphasis>
<taxonomicName id="4C7F4D70FFAEFFD0FEAA0DDAFE43F828" authorityName="Manukyan" authorityYear="2023" box="[273,473,1928,1947]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAEFFD0FEAA0DDAFE43F828" box="[273,473,1928,1947]" italics="true" pageId="2" pageNumber="405">Vorstadt ambermuseum</emphasis>
</taxonomicName>
<emphasis id="B90BEAE1FFAEFFD0FE640DDAFD93F828" bold="true" box="[479,521,1928,1947]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="A238579AFFAEFFD0FE640DDAFD93F828" box="[479,521,1928,1947]" pageId="2" pageNumber="405" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, holotype, female: 4 — flagellomeres of left flagellum, arrows indicate constrictions; 5 — 4th to
</paragraph>
<paragraph id="8BC036F3FFAEFFD0FF000DEAFD7CF878" blockId="2.[187,784,1928,2067]" box="[187,742,1976,1995]" pageId="2" pageNumber="405">11th flagellomere of left flagellum, arrows indicate constrictions.</paragraph>
<paragraph id="8BC036F3FFAEFFD0FF000D82FDEEF7A0" blockId="2.[187,784,1928,2067]" pageId="2" pageNumber="405">
<emphasis id="B90BEAE1FFAEFFD0FF000D82FE94F850" bold="true" box="[187,270,2000,2019]" pageId="2" pageNumber="405">Рис. 45.</emphasis>
<taxonomicName id="4C7F4D70FFAEFFD0FEAF0D82FE47F850" authorityName="Manukyan" authorityYear="2023" box="[276,477,2000,2019]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAEFFD0FEAF0D82FE47F850" box="[276,477,2000,2019]" italics="true" pageId="2" pageNumber="405">Vorstadt ambermuseum</emphasis>
</taxonomicName>
<emphasis id="B90BEAE1FFAEFFD0FE580D82FD94F850" bold="true" box="[483,526,2000,2019]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="A238579AFFAEFFD0FE580D82FD94F850" box="[483,526,2000,2019]" pageId="2" pageNumber="405" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, голотип, самка: 4 — левый жгутик, стрелками покаЗаны перетЯжки; 5 — 411-й членики левого жгутика, стрелками покаЗаны перетЯжки.
</paragraph>
</caption>
<caption id="DF00667BFFAFFFD1FF060ED2FC48FB18" ID-DOI="http://doi.org/10.5281/zenodo.13177454" ID-Zenodo-Dep="13177454" httpUri="https://zenodo.org/record/13177454/files/figure.png" pageId="3" pageNumber="406" startId="3.[189,220,1152,1171]" targetBox="[313,1333,313,1100]" targetPageId="3" targetType="figure">
<paragraph id="8BC036F3FFAFFFD1FF060ED2FC52FB20" blockId="3.[189,978,1152,1195]" box="[189,968,1152,1171]" pageId="3" pageNumber="406">
<emphasis id="B90BEAE1FFAFFFD1FF060ED2FF6FFB20" bold="true" box="[189,245,1152,1171]" pageId="3" pageNumber="406">Fig. 6.</emphasis>
<taxonomicName id="4C7F4D70FFAFFFD1FF400ED2FE59FB20" authorityName="Manukyan" authorityYear="2023" box="[251,451,1152,1171]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAFFFD1FF400ED2FE59FB20" box="[251,451,1152,1171]" italics="true" pageId="3" pageNumber="406">Vorstadt ambermuseum</emphasis>
</taxonomicName>
<emphasis id="B90BEAE1FFAFFFD1FE720ED2FE6EFB20" bold="true" box="[457,500,1152,1171]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="A238579AFFAFFFD1FE720ED2FE6EFB20" box="[457,500,1152,1171]" pageId="3" pageNumber="406" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, holotype, female, head and mesoscutum, dorsal view.
</paragraph>
<paragraph id="8BC036F3FFAFFFD1FF060ECAFC48FB18" blockId="3.[189,978,1152,1195]" box="[189,978,1176,1195]" pageId="3" pageNumber="406">
<emphasis id="B90BEAE1FFAFFFD1FF060ECAFF60FB18" bold="true" box="[189,250,1176,1195]" pageId="3" pageNumber="406">Рис. 6.</emphasis>
<taxonomicName id="4C7F4D70FFAFFFD1FEBB0ECAFE52FB18" authorityName="Manukyan" authorityYear="2023" box="[256,456,1176,1195]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAFFFD1FEBB0ECAFE52FB18" box="[256,456,1176,1195]" italics="true" pageId="3" pageNumber="406">Vorstadt ambermuseum</emphasis>
</taxonomicName>
<emphasis id="B90BEAE1FFAFFFD1FE750ECAFE65FB18" bold="true" box="[462,511,1176,1195]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="A238579AFFAFFFD1FE750ECAFE63FB18" box="[462,505,1176,1195]" pageId="3" pageNumber="406" rank="species">sp.n.</taxonomicNameLabel>
,
</emphasis>
голотип, самка, голова и среднеспинка, вид сверху.
</paragraph>
</caption>
<paragraph id="8BC036F3FFAFFFD1FF580FD6FEE4FA07" blockId="3.[189,787,1251,1889]" pageId="3" pageNumber="406">
<typeStatus id="54C48851FFAFFFD1FF580FD6FE89FA2A" box="[227,275,1412,1433]" pageId="3" pageNumber="406">Type</typeStatus>
species.
<taxonomicName id="4C7F4D70FFAFFFD1FED10FD6FD23FA2A" authority="Manukyan" authorityName="Manukyan" authorityYear="2023" box="[362,697,1412,1433]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAFFFD1FED10FD6FDD6FA2A" box="[362,588,1412,1433]" italics="true" pageId="3" pageNumber="406">Vorstadt ambermuseum</emphasis>
Manukyan
</taxonomicName>
<emphasis id="B90BEAE1FFAFFFD1FD050FD6FD75FA2A" bold="true" box="[702,751,1412,1433]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="A238579AFFAFFFD1FD050FD6FD75FA2A" box="[702,751,1412,1433]" pageId="3" pageNumber="406" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
<paragraph id="8BC036F3FFAFFFD1FF580FE8FD15FA59" blockId="3.[189,787,1251,1889]" pageId="3" pageNumber="406">
Species composition.
<taxonomicName id="4C7F4D70FFAFFFD1FE070FE8FC88FA7C" authority="Manukyan" authorityName="Manukyan" authorityYear="2023" box="[444,786,1466,1487]" class="Insecta" family="Formicidae" genus="Vorstadt" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="ambermuseum" status="sp. nov.">
<emphasis id="B90BEAE1FFAFFFD1FE070FE8FD38FA7C" box="[444,674,1466,1487]" italics="true" pageId="3" pageNumber="406">Vorstadt ambermuseum</emphasis>
Manukyan
</taxonomicName>
<emphasis id="B90BEAE1FFAFFFD1FF060F87FF77FA59" bold="true" box="[189,237,1493,1514]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="A238579AFFAFFFD1FF060F87FF77FA59" box="[189,237,1493,1514]" pageId="3" pageNumber="406" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B90BEAE1FFAFFFD1FF410F87FEC1FA59" box="[250,347,1493,1514]" italics="true" pageId="3" pageNumber="406">V. groehni</emphasis>
(
<bibRefCitation id="EFEE4B02FFAFFFD1FEDC0F87FD1EFA59" author="Tolkanitz V. I. &amp; Perkovsky E. E." box="[359,644,1493,1514]" pageId="3" pageNumber="406" pagination="76 - 85" refId="ref3913" refString="Tolkanitz V. I., Perkovsky E. E. 2015. A new species of the genus Paxylommites (Hymenoptera, Ichneumonidae, Hybrizoninae) from Baltic Amber // Paleontological Journal. Vol. 49. No. 4. P. 391 - 293. [Original Russian text published in Paleontologicheskii Zhurnal, 2009, No. 1. P. 76 - 85]. https: // doi. org / 10.1134 / S 0031030115040164" type="book chapter" year="2015">Tolkanitz et Perkovsky, 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3656578FFAFFFD1FF580FBDFD14F8D2" pageId="3" pageNumber="406" type="description">
<paragraph id="8BC036F3FFAFFFD1FF580FBDFE69F8A3" blockId="3.[189,787,1251,1889]" pageId="3" pageNumber="406">
DESCRIPTION. Flagellum 1113-segmented, all flagellomeres including first one with distinct medial constriction (
<figureCitation id="13442A76FFAFFFD1FF7F0C77FE87F989" box="[196,285,1573,1594]" captionStart="Figs 45" captionStartId="2.[187,225,1928,1947]" captionTargetBox="[822,1419,1115,2041]" captionTargetId="figure-293@2.[822,1419,1115,2047]" captionTargetPageId="2" captionText="Figs 45. Vorstadt ambermuseum sp.n., holotype, female: 4 — flagel- lomeres of left flagellum, arrows indicate constrictions; 5 — 4th to 11th flagellomere of left flagellum, arrows indicate constrictions. Рис. 45. Vorstadt ambermuseum sp.n., голотип, самка: 4 — левый жгутик, стрелками покаЗаны перетЯжки; 5 — 411-й членики левого жгутика, стрелками покаЗаны перетЯжки." figureDoi="http://doi.org/10.5281/zenodo.13177452" httpUri="https://zenodo.org/record/13177452/files/figure.png" pageId="3" pageNumber="406">Figs 45</figureCitation>
). Occipital carina obliterated. Small obscure area at level of stemmaticum (area bounded by the ocelli) probably preserved. Maxillary palps 3-segmented, long, not reduced; segment 2 not widened at apex; labial palps reduced, 2-segmented. Forewings with vein 2m-cu (
<figureCitation id="13442A76FFAFFFD1FD860CC2FD03F916" box="[573,665,1680,1701]" captionStart="Figs 12" captionStartId="1.[188,226,2025,2044]" captionTargetBox="[313,1333,452,1985]" captionTargetId="figure-91@1.[313,1333,452,1986]" captionTargetPageId="1" captionText="Figs 12. Vorstadt ambermuseum sp.n., holotype, female: 1 — dorsal view; 2 — fragment of front wing. Рис. 12. Vorstadt ambermuseum sp.n., голотип, самка: 1 — вид сверху; 2 — фрагмент переднего крыла." figureDoi="http://doi.org/10.5281/zenodo.13177448" httpUri="https://zenodo.org/record/13177448/files/figure.png" pageId="3" pageNumber="406">Figs 12</figureCitation>
). Length of vein 4Rs 4-5.2 times as long as vein r-rs. Vein 1cu-a (nervulus) interstitial. Fore tarsi as long as middle tarsi. Base of 1T distinctly above base of hind coxae. 1T and basal half of 2T probably longitudinally striated.
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<paragraph id="8BC036F3FFAFFFD1FF580D44FD14F8D2" blockId="3.[189,787,1251,1889]" pageId="3" pageNumber="406">NOTE. Accurate determination of the number of apical flagellomeres is problematic because the difference between constrictions and segment boundaries is unclear.</paragraph>
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