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<document id="F1C079ACE4E121FD844029812DB261A4" ID-DOI="10.11646/zootaxa.3630.1.2" ID-GBIF-Dataset="28703b77-1579-49e7-885f-5c2edb895f83" ID-ISSN="1175-5326" ID-Zenodo-Dep="283467" ID-ZooBank="A338EE37-E8AC-4AC3-8D6B-D0177FB680BF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460582983698" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="03D687E5B0429A6DBDD147F01110FC85" docLanguage="en" docName="zt03630p079.pdf" docOrigin="Zootaxa 3630 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanichnus Jałoszyński, 2013, gen. nov." docType="treatment" docUuid="530A7704-1C3E-4518-ABEE-AE4A40C7F08D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="61" masterDocId="FFEFFF9DB0509A7BBD46427C1313FF91" masterDocTitle="Revision of subgenera of Stenichnus Thomson, with review of Australo-Pacific species (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="79" masterPageNumber="39" pageNumber="57" updateTime="1698324670604" updateUser="plazi">
<mods:mods id="FF4820280F6486A35A52324D3944A8D7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3C5A3960F5DC57B2259EF6963D76A3AE">Revision of subgenera of Stenichnus Thomson, with review of Australo-Pacific species (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="0A1C8C463E9557EF4EB3E5F403D08B43" type="personal">
<mods:role id="3618B4DE751A61D9397060DC47F08773">
<mods:roleTerm id="E981DA98EBA8CCD98EB2BD93AB4307BE">Author</mods:roleTerm>
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<mods:namePart id="122DE20D0C16C498A791BAD435CD286F">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="4DB701645BD6705F1D5EBEF052D1B357">Zootaxa</mods:title>
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<mods:part id="8F96B7B69F508D6244095A4922208243">
<mods:date id="C69860D5E39D951CDE719B2CCD9C2BC8">2013</mods:date>
<mods:detail id="CBEFC3BB46F685A3FD10B431DA724C00" type="volume">
<mods:number id="378882A7D64E1F0C3BB9B218A00240AE">3630</mods:number>
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<mods:classification id="D745F3C3C093BD8F6F365F8F427C9DB3">journal article</mods:classification>
<mods:identifier id="F8CA793B7080AA862CB375F2D6D7BE17" type="DOI">10.11646/zootaxa.3630.1.2</mods:identifier>
<mods:identifier id="70D963F72E10CDEAC0BE8A5B96C1790E" type="GBIF-Dataset">28703b77-1579-49e7-885f-5c2edb895f83</mods:identifier>
<mods:identifier id="D9349C745FE79478BD170B9F23B91B2C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6E13925E6E80B5C1D7DA4FFF4EFD81CF" type="Zenodo-Dep">283467</mods:identifier>
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<treatment id="03D687E5B0429A6DBDD147F01110FC85" ID-DOI="http://doi.org/10.5281/zenodo.5696222" ID-GBIF-Taxon="127663494" ID-Zenodo-Dep="5696222" LSID="urn:lsid:zoobank.org:act:530A7704-1C3E-4518-ABEE-AE4A40C7F08D" httpUri="http://treatment.plazi.org/id/03D687E5B0429A6DBDD147F01110FC85" lastPageId="22" lastPageNumber="61" pageId="18" pageNumber="57">
<subSubSection id="C3656578B0429A69BDD147F01029FA78" pageId="18" pageNumber="57" type="nomenclature">
<paragraph id="8BC036F3B0429A69BDD147F012E1FA37" blockId="18.[151,498,1420,1447]" box="[151,498,1420,1447]" pageId="18" pageNumber="57">
<heading id="D088819FB0429A69BDD147F012E1FA37" bold="true" box="[151,498,1420,1447]" fontSize="11" level="1" pageId="18" pageNumber="57" reason="1">
<emphasis id="B90BEAE1B0429A69BDD147F012E1FA37" bold="true" box="[151,498,1420,1447]" pageId="18" pageNumber="57">
Genus
<taxonomicName id="4C7F4D70B0429A69BDA847F1126DFA36" ID-CoL="8XQSL" box="[238,382,1421,1447]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0429A69BDA847F1126DFA36" bold="true" box="[238,382,1421,1447]" italics="true" pageId="18" pageNumber="57">Zeanichnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238579AB0429A69BCC247F012E1FA37" box="[388,498,1420,1446]" pageId="18" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036F3B0429A69BDD147A81029FA78" blockId="18.[151,826,1491,1513]" box="[151,826,1491,1513]" pageId="18" pageNumber="57">
<typeStatus id="54C48851B0429A69BDD147A813DBFA78" box="[151,200,1492,1513]" pageId="18" pageNumber="57">Type</typeStatus>
species:
<taxonomicName id="4C7F4D70B0429A69BC6047AF116EFA78" authority="Broun, 1893" authorityName="Broun" authorityYear="1893" box="[294,637,1491,1513]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0429A69BC6047AF12EBFA78" box="[294,504,1491,1513]" italics="true" pageId="18" pageNumber="57">Scydmaenus insignis</emphasis>
Broun, 1893
</taxonomicName>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0429A69BDD1446910ADF84F" pageId="18" pageNumber="57" type="diagnosis">
<paragraph id="8BC036F3B0429A69BDD1446910ADF84F" blockId="18.[151,1436,1557,2014]" pageId="18" pageNumber="57">
<emphasis id="B90BEAE1B0429A69BDD144691202F9BF" bold="true" box="[151,273,1557,1582]" pageId="18" pageNumber="57">Diagnosis.</emphasis>
Eyes located posteriorly; occipital constriction nearly as broad as vertex or only slightly narrower; fronto-clypeal groove absent; mandibles subtriangular, with subapical tooth on mesal margin; submentum subtriangular, with very short posterior margin and sinuate lateral margins, completely demarcated from hypostomae by lateral sutures; hypostomae elongate and demarcated from genae only in short anterior parts by incomplete hypostomal ridges; posterior tentorial pits hidden in deep groove between submentum and gular plate; antennae with 4–segmented club or gradually thickened distally; pronotum with one pair of lateral ante-basal pits connected by transverse groove, without lateral marginal carinae or edges; prosternum with narrow basisternal part not demarcated from procoxal cavities, procoxal cavities separated in middle by fine carina; hypomera not demarcated from pronotum; mesoventrite without asetose impressions and with lateral setose impressions narrowly separated in middle and broadly separated from mesocoxal cavities; mesoventral intercoxal process moderately narrow and moderately projecting ventrally, with distinct posterior tip separated from metaventrite; mesocoxal projections with posterior lobes; metaventrite with metaventral anterior process, with short and broad metaventral intercoxal process bearing median notch; aedeagus with free parameres.
</paragraph>
</subSubSection>
<caption id="DF00667BB0439A68BDD145D3171BF879" httpUri="https://zenodo.org/record/283479/files/figure.png" pageId="19" pageNumber="58" targetBox="[160,1426,189,1935]" targetPageId="19">
<paragraph id="8BC036F3B0439A68BDD145D3171BF879" blockId="19.[151,1436,1967,2025]" pageId="19" pageNumber="58">
<emphasis id="B90BEAE1B0439A68BDD145D31246F854" bold="true" box="[151,341,1967,1990]" pageId="19" pageNumber="58">FIGURES 62–66.</emphasis>
<taxonomicName id="4C7F4D70B0439A68BC1A45CC1192F857" authority="Broun" authorityName="Broun" box="[348,641,1968,1990]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="58" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0439A68BC1A45CC113BF857" box="[348,552,1968,1990]" italics="true" pageId="19" pageNumber="58">Zeanichnus insignis</emphasis>
(Broun)
</taxonomicName>
, holotype male (62, 65) and
<taxonomicName id="4C7F4D70B0439A68BEED45CC17ECF857" authority="Franz" authorityName="Franz" box="[939,1279,1968,1990]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="58" phylum="Arthropoda" rank="species" species="kuschelianus">
<emphasis id="B90BEAE1B0439A68BEED45CC17B8F857" box="[939,1195,1968,1990]" italics="true" pageId="19" pageNumber="58">Zeanichnus kuschelianus</emphasis>
(Franz)
</taxonomicName>
, holotype male (63–64, 66). Dorsal (62–63) and ventral (64) habitus and original set of labels (65–66).
</paragraph>
</caption>
<subSubSection id="C3656578B0449A6FBD8142EB1694F9BD" pageId="20" pageNumber="59" type="description">
<paragraph id="8BC036F3B0449A6FBD8142EB12B5FF45" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
<emphasis id="B90BEAE1B0449A6FBD8142EB124AFF21" bold="true" box="[199,345,151,176]" pageId="20" pageNumber="59">Description.</emphasis>
Body (
<figureCitation id="13442A76B0449A6FBCEC42EB113CFF3E" box="[426,559,151,175]" captionStart="FIGURES 62 – 66" captionStartId="19.[151,264,1967,1990]" captionTargetBox="[160,1426,189,1935]" captionTargetId="figure@19.[151,1436,167,1946]" captionTargetPageId="19" captionText="FIGURES 62 – 66. Zeanichnus insignis (Broun), holotype male (62, 65) and Zeanichnus kuschelianus (Franz), holotype male (63 – 64, 66). Dorsal (62 – 63) and ventral (64) habitus and original set of labels (65 – 66)." httpUri="https://zenodo.org/record/283479/files/figure.png" pageId="20" pageNumber="59">Figs. 62–63</figureCitation>
) strongly convex, strongly elongate and slender, with long appendages, cuticle glossy, distinctly setose.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD8142A31228FEF5" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Head (
<figureCitation id="13442A76B0449A6FBC5542A31275FF66" box="[275,358,223,247]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
) subtriangular, broadest at posteriorly located eyes; neck region short and subcylindrical, only slightly broadened in middle; occipital constriction (
<figureCitation id="13442A76B0449A6FBFAE4378102EFE8D" box="[744,829,260,284]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBE0F43791063FE8D" box="[841,880,261,284]" italics="true" pageId="20" pageNumber="59">occ</emphasis>
) in the narrowest place nearly as wide as vertex; tempora (
<figureCitation id="13442A76B0449A6FBC44435B1245FEAE" box="[258,342,295,319]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBC244355126DFED1" box="[354,382,297,320]" italics="true" pageId="20" pageNumber="59">tm</emphasis>
) much shorter than eyes; vertex and frons between eyes confluent and convex, fronto-clypeal groove absent.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD814313104EFE16" blockId="20.[151,1436,151,2012]" box="[199,861,367,391]" pageId="20" pageNumber="59">Labrum transverse with rounded sides and anterior margin.</paragraph>
<paragraph id="8BC036F3B0449A6FBD8143E81221FE5E" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">Mandibles symmetrical, each with broad basal part and moderately slender distal part, mesal margin with median tooth.</paragraph>
<paragraph id="8BC036F3B0449A6FBD8143A017E6FDAD" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Each maxilla with elongate galea and lacinia (
<figureCitation id="13442A76B0449A6FBFF943A01003FE65" box="[703,784,476,500]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBE5A43A0102CFE65" box="[796,831,476,500]" italics="true" pageId="20" pageNumber="59">gal</emphasis>
,
<emphasis id="B90BEAE1B0449A6FBE0B43A0107DFE65" box="[845,878,476,500]" italics="true" pageId="20" pageNumber="59">lac</emphasis>
) and long maxillary palp (
<figureCitation id="13442A76B0449A6FB9D443A017F0FE65" box="[1170,1251,476,500]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FB9A943A1160EFE65" box="[1263,1309,477,500]" italics="true" pageId="20" pageNumber="59">mxp</emphasis>
) composed of minute palpomere I, strongly elongate and broadening distally palpomere II, long and pedunculate palpomere III nearly equally broad from middle to apex, and slender, elongate subconical and pointed palpomere IV.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD81403B1659FD7E" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Labium with subtriangular submentum (
<figureCitation id="13442A76B0449A6FBFD1403B11FDFDCE" box="[663,750,583,607]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBFBB40351039FDF1" box="[765,810,585,608]" italics="true" pageId="20" pageNumber="59">smn</emphasis>
) with sinuate lateral margins and very short posterior margin demarcated from gular plate (
<figureCitation id="13442A76B0449A6FBF054010118BFD15" box="[579,664,620,644]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBFE0401111D1FD15" box="[678,706,621,644]" italics="true" pageId="20" pageNumber="59">gp</emphasis>
) by deep transverse groove in which posterior tentorial pits are hidden, laterally demarcated from hypostomae (
<figureCitation id="13442A76B0449A6FBFF440F31016FD36" box="[690,773,655,679]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBE5740EC100CFD39" box="[785,799,656,680]" italics="true" pageId="20" pageNumber="59">h</emphasis>
) by lateral sutures (
<figureCitation id="13442A76B0449A6FB94640F31740FD36" box="[1024,1107,655,679]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FB91940EC176EFD39" box="[1119,1149,656,680]" italics="true" pageId="20" pageNumber="59">lss</emphasis>
); mentum subtrapezoidal (
<figureCitation id="13442A76B0449A6FBDD940C813E5FD5D" box="[159,246,692,716]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBC4340C91234FD5D" box="[261,295,693,716]" italics="true" pageId="20" pageNumber="59">mn</emphasis>
); prementum short with broadly separated, small 3-segmented labial palps (
<figureCitation id="13442A76B0449A6FB9E640C817E4FD5D" box="[1184,1271,692,716]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FB84040C8160FFD5D" box="[1286,1308,692,716]" italics="true" pageId="20" pageNumber="59">lp</emphasis>
). Laterally mouthparts are demarcated from genae only anteriorly by short, incomplete hypostomal ridges (
<figureCitation id="13442A76B0449A6FB98F40AB1606FD7E" box="[1225,1301,727,751]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig.69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FB86740A41629FD61" box="[1313,1338,728,752]" italics="true" pageId="20" pageNumber="59">hr</emphasis>
).
</paragraph>
<paragraph id="8BC036F3B0449A6FBD81408017E7FC85" blockId="20.[151,1436,151,2012]" box="[199,1268,764,788]" pageId="20" pageNumber="59">
Gular plate (
<figureCitation id="13442A76B0449A6FBC15408012B4FC85" box="[339,423,764,788]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBCF4408112DDFC85" box="[434,462,765,788]" italics="true" pageId="20" pageNumber="59">gp</emphasis>
) large and trapezoidal in shape, with distinct gular sutures (
<figureCitation id="13442A76B0449A6FB92A408017ACFC85" box="[1132,1215,764,788]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Fig. 69</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FB98D408117F7FC85" box="[1227,1252,765,788]" italics="true" pageId="20" pageNumber="59">gs</emphasis>
).
</paragraph>
<paragraph id="8BC036F3B0449A6FBD81415C124CFCCD" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Antennae (
<figureCitation id="13442A76B0449A6FBC04416312DBFCA6" box="[322,456,799,823]" captionStart="FIGURES 62 – 66" captionStartId="19.[151,264,1967,1990]" captionTargetBox="[160,1426,189,1935]" captionTargetId="figure@19.[151,1436,167,1946]" captionTargetPageId="19" captionText="FIGURES 62 – 66. Zeanichnus insignis (Broun), holotype male (62, 65) and Zeanichnus kuschelianus (Franz), holotype male (63 – 64, 66). Dorsal (62 – 63) and ventral (64) habitus and original set of labels (65 – 66)." httpUri="https://zenodo.org/record/283479/files/figure.png" pageId="20" pageNumber="59">Figs. 62–63</figureCitation>
) with club composed of four antennomeres (
<taxonomicName id="4C7F4D70B0449A6FBE87415D1726FCA9" box="[961,1077,801,824]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0449A6FBE87415D1726FCA9" box="[961,1077,801,824]" italics="true" pageId="20" pageNumber="59">Z. insignis</emphasis>
</taxonomicName>
) or gradually thickened distally (
<taxonomicName id="4C7F4D70B0449A6FBDE64139125CFCCD" box="[160,335,836,860]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="kuschelianus">
<emphasis id="B90BEAE1B0449A6FBDE64139125CFCCD" box="[160,335,836,860]" italics="true" pageId="20" pageNumber="59">Z. kuschelianus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC036F3B0449A6FBD81411B113FFBA5" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Prothorax (
<figureCitation id="13442A76B0449A6FBC01411B12C3FCEE" box="[327,464,871,895]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
) in dorsal view with rounded anterior and lateral margins of pronotum, at base with transverse groove connecting one pair of lateral ante-basal foveae, without lateral marginal carinae or edges; hypomera (
<figureCitation id="13442A76B0449A6FBC5041D3128DFC56" box="[278,414,943,967]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBCEF41CC12D0FC59" box="[425,451,944,968]" italics="true" pageId="20" pageNumber="59">hy</emphasis>
) not demarcated from notum. Prosternum (
<figureCitation id="13442A76B0449A6FBEE141D3173EFC56" box="[935,1069,943,967]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
) with very short basisternal part; procoxal cavities (
<figureCitation id="13442A76B0449A6FBC3641A812EDFC7D" box="[368,510,980,1004]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBF4941A91125FC7D" box="[527,566,981,1004]" italics="true" pageId="20" pageNumber="59">pcc</emphasis>
) without anterior marginal carina, separated in middle by fine prosternal intercoxal carina; procoxal sockets (
<figureCitation id="13442A76B0449A6FBF72418B11ACFB9E" box="[564,703,1015,1039]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBF8B418511E1FB81" box="[717,754,1017,1040]" italics="true" pageId="20" pageNumber="59">pcs</emphasis>
) nearly closed by lateral expansion of sternum; notosternal sutures (
<figureCitation id="13442A76B0449A6FBDB04660126CFBA5" box="[246,383,1052,1076]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBCCF466112BEFBA5" box="[393,429,1053,1076]" italics="true" pageId="20" pageNumber="59">nss</emphasis>
) complete.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD8146431737FB76" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Mesoventrite (
<figureCitation id="13442A76B0449A6FBC3246431116FBC6" box="[372,517,1087,1111]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
) with narrow anterior ridge, without lateral asetose impressions and with moderately large setose impressions (
<figureCitation id="13442A76B0449A6FBF01461811C0FBED" box="[583,723,1124,1148]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBFA6461911E0FBED" box="[736,755,1125,1148]" italics="true" pageId="20" pageNumber="59">si</emphasis>
) narrowly separated in middle and broadly separated from mesocoxal cavities (
<figureCitation id="13442A76B0449A6FBC3F46FB1111FB0E" box="[377,514,1159,1183]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBF4A46F51157FB31" box="[524,580,1161,1184]" italics="true" pageId="20" pageNumber="59">mscc</emphasis>
); mesocoxal projection (
<figureCitation id="13442A76B0449A6FBE1C46FB10F1FB0E" box="[858,994,1159,1183]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBEAA46F51709FB31" box="[1004,1050,1161,1184]" italics="true" pageId="20" pageNumber="59">mcp</emphasis>
) without carina and with posterior lobe (
<figureCitation id="13442A76B0449A6FBD9F46D01276FB55" box="[217,357,1196,1220]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBC3546D0129AFB55" box="[371,393,1196,1220]" italics="true" pageId="20" pageNumber="59">pl</emphasis>
) bearing dense setae along posterior margin; mesoventral intercoxal process (
<figureCitation id="13442A76B0449A6FB84A46D0168BFB55" box="[1292,1432,1196,1220]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBDD146AD13C2FB79" box="[151,209,1233,1256]" italics="true" pageId="20" pageNumber="59">msvp</emphasis>
) moderately narrow, with distinct posterior tip separated from metaventrite.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD81468817D7FABE" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Metaventrite (
<figureCitation id="13442A76B0449A6FBC28468812EDFA9D" box="[366,510,1268,1292]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
) broader than long, with metaventral anterior process (
<figureCitation id="13442A76B0449A6FB9EB4688162CFA9D" box="[1197,1343,1268,1292]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FB81546891698FA9D" box="[1363,1419,1269,1292]" italics="true" pageId="20" pageNumber="59">mtap</emphasis>
); metaventral intercoxal process (
<figureCitation id="13442A76B0449A6FBCBA476B1196FABE" box="[508,645,1303,1327]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 67–68</figureCitation>
;
<emphasis id="B90BEAE1B0449A6FBFD6476511D5FAA1" box="[656,710,1305,1328]" italics="true" pageId="20" pageNumber="59">mtvp</emphasis>
) broad and short, with distinct median notch.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD81474010C1FAE6" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Elytra (
<figureCitation id="13442A76B0449A6FBC58474012BAFAC5" box="[286,425,1340,1364]" captionStart="FIGURES 62 – 66" captionStartId="19.[151,264,1967,1990]" captionTargetBox="[160,1426,189,1935]" captionTargetId="figure@19.[151,1436,167,1946]" captionTargetPageId="19" captionText="FIGURES 62 – 66. Zeanichnus insignis (Broun), holotype male (62, 65) and Zeanichnus kuschelianus (Franz), holotype male (63 – 64, 66). Dorsal (62 – 63) and ventral (64) habitus and original set of labels (65 – 66)." httpUri="https://zenodo.org/record/283479/files/figure.png" pageId="20" pageNumber="59">Figs. 62–63</figureCitation>
) oval, with well-marked humeral calli; each elytron with one rudimentary basal fovea barely noticeable in dry-mounted specimens.
<collectingCountry id="F3687663B0449A6FBFD3472311DEFAE6" box="[661,717,1375,1399]" name="India" pageId="20" pageNumber="59">Hind</collectingCountry>
wings well-developed.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD8147F91601FA0D" blockId="20.[151,1436,151,2012]" box="[199,1298,1412,1437]" pageId="20" pageNumber="59">
Legs (
<figureCitation id="13442A76B0449A6FBC4A47F81280FA0D" box="[268,403,1412,1436]" captionStart="FIGURES 62 – 66" captionStartId="19.[151,264,1967,1990]" captionTargetBox="[160,1426,189,1935]" captionTargetId="figure@19.[151,1436,167,1946]" captionTargetPageId="19" captionText="FIGURES 62 – 66. Zeanichnus insignis (Broun), holotype male (62, 65) and Zeanichnus kuschelianus (Franz), holotype male (63 – 64, 66). Dorsal (62 – 63) and ventral (64) habitus and original set of labels (65 – 66)." httpUri="https://zenodo.org/record/283479/files/figure.png" pageId="20" pageNumber="59">Figs. 62–63</figureCitation>
) long and slender; femora clavate, tibiae nearly straight, tarsi strongly elongate.
</paragraph>
<paragraph id="8BC036F3B0449A6FBD8147DB1080FA2E" blockId="20.[151,1436,151,2012]" box="[199,915,1447,1471]" pageId="20" pageNumber="59">Abdominal sternites VII and VIII separated by indistinct suture.</paragraph>
<paragraph id="8BC036F3B0449A6FBD8147B01694F9BD" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
Aedeagus (
<figureCitation id="13442A76B0449A6FBC0E47B012C0FA75" box="[328,467,1484,1508]" captionStart="FIGURES 67 – 73" captionStartId="21.[151,264,1769,1792]" captionTargetBox="[167,1417,350,1731]" captionTargetId="figure@21.[151,1436,337,1748]" captionTargetPageId="21" captionText="FIGURES 67 – 73. Zeanichnus insignis (Broun), holotype male (67, 70 – 71) and Zeanichnus kuschelianus (Franz), holotype male (68 – 69, 72 – 73). Venter (67 – 68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest 2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v 3, metaventrite." httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="20" pageNumber="59">Figs. 70–73</figureCitation>
) with complex and symmetrical internal armature composed of sub-basally or submedially located voluminous central complex and sub-apical lateral groups of needle-like sclerites; apical part of dorsal wall forming strongly projecting distally, narrow projection; parameres free and slender, with apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0449A6FBD81444B1624F9DE" box="[199,1335,1591,1616]" pageId="20" pageNumber="59" type="distribution">
<paragraph id="8BC036F3B0449A6FBD81444B1624F9DE" blockId="20.[151,1436,151,2012]" box="[199,1335,1591,1616]" pageId="20" pageNumber="59">
<emphasis id="B90BEAE1B0449A6FBD81444B113FF9C1" bold="true" box="[199,556,1591,1616]" pageId="20" pageNumber="59">Distribution and composition.</emphasis>
Two species are known to occur in the North Island of
<collectingCountry id="F3687663B0449A6FB9DF44441620F9DE" box="[1177,1331,1591,1615]" name="New Zealand" pageId="20" pageNumber="59">New Zealand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0449A6FBD8144201240F906" pageId="20" pageNumber="59" type="etymology">
<paragraph id="8BC036F3B0449A6FBD8144201240F906" blockId="20.[151,1436,151,2012]" pageId="20" pageNumber="59">
<emphasis id="B90BEAE1B0449A6FBD814420125DF9E5" bold="true" box="[199,334,1628,1652]" pageId="20" pageNumber="59">Etymology.</emphasis>
The genus name is a combination of &quot;Zea-&quot; derived from
<collectingCountry id="F3687663B0449A6FBEB24421179FF9E5" box="[1012,1164,1628,1652]" name="New Zealand" pageId="20" pageNumber="59">New Zealand</collectingCountry>
and &quot;-nichnus&quot; derived from
<taxonomicName id="4C7F4D70B0449A6FBD9244FC125FF909" box="[212,332,1664,1688]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1B0449A6FBD9244FC125FF909" box="[212,332,1664,1688]" italics="true" pageId="20" pageNumber="59">Stenichnus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0449A6DBD8144D81110FC85" lastPageId="22" lastPageNumber="61" pageId="20" pageNumber="59" type="discussion">
<paragraph id="8BC036F3B0449A6EBD8144D811DFFE8D" blockId="20.[151,1436,151,2012]" lastBlockId="21.[151,1436,151,284]" lastPageId="21" lastPageNumber="60" pageId="20" pageNumber="59">
<emphasis id="B90BEAE1B0449A6FBD8144D81228F92D" bold="true" box="[199,315,1700,1724]" pageId="20" pageNumber="59">Remarks.</emphasis>
Broun (1893) in the description of
<taxonomicName id="4C7F4D70B0449A6FBFAD44D810CBF92D" box="[747,984,1700,1724]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0449A6FBFAD44D810CBF92D" box="[747,984,1700,1724]" italics="true" pageId="20" pageNumber="59">Scydmaenus insignis</emphasis>
</taxonomicName>
stated that &quot;I have little doubt it will become the
<typeStatus id="54C48851B0449A6FBC6544B41240F971" box="[291,339,1736,1760]" pageId="20" pageNumber="59">type</typeStatus>
of a new genus&quot;. This opinion is confirmed here and supported by a number of characters that define the new genus,
<taxonomicName id="4C7F4D70B0449A6FBCD244901106F895" box="[404,533,1772,1796]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0449A6FBCD244901106F895" box="[404,533,1772,1796]" italics="true" pageId="20" pageNumber="59">Zeanichnus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C7F4D70B0449A6FBF65449011B7F895" box="[547,676,1772,1796]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0449A6FBF65449011B7F895" box="[547,676,1772,1796]" italics="true" pageId="20" pageNumber="59">Zeanichnus</emphasis>
</taxonomicName>
is unique among
<taxonomicName id="4C7F4D70B0449A6FBE2B44901700F895" box="[877,1043,1772,1796]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
in having the submentum strongly narrowing caudad, with sinuate lateral margins and very short posterior margin, as short as about half length of the anterior margin of gular plate, and in having incomplete hypostomal ridges, well-developed only in their anterior parts. The lack of asetose impressions of the mesoventrite differentiates
<taxonomicName id="4C7F4D70B0449A6FBEA045241774F8E1" box="[998,1127,1880,1904]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0449A6FBEA045241774F8E1" box="[998,1127,1880,1904]" italics="true" pageId="20" pageNumber="59">Zeanichnus</emphasis>
</taxonomicName>
from all other genera and subgenera revised in the present paper.
<taxonomicName id="4C7F4D70B0449A6FBF1745001002F805" box="[593,785,1916,1940]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1B0449A6FBF1745001002F805" box="[593,785,1916,1940]" italics="true" pageId="20" pageNumber="59">Austrostenichnus</emphasis>
</taxonomicName>
, in which
<taxonomicName id="4C7F4D70B0449A6FBEC1450110EFF805" box="[903,1020,1917,1940]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0449A6FBEC1450110EFF805" box="[903,1020,1917,1940]" italics="true" pageId="20" pageNumber="59">Z. insignis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D70B0449A6FB975450117F2F805" box="[1075,1249,1916,1940]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="kuschelianus">
<emphasis id="B90BEAE1B0449A6FB975450117F2F805" box="[1075,1249,1916,1940]" italics="true" pageId="20" pageNumber="59">Z. kuschelianus</emphasis>
</taxonomicName>
were previously placed, differs from
<taxonomicName id="4C7F4D70B0449A6FBCC645DC1112F829" box="[384,513,1952,1976]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0449A6FBCC645DC1112F829" box="[384,513,1952,1976]" italics="true" pageId="20" pageNumber="59">Zeanichnus</emphasis>
</taxonomicName>
in a large number of other characters, e.g., in having confluent procoxal cavities (separated by carina in
<taxonomicName id="4C7F4D70B0449A6FBCFD45B8112FF84D" box="[443,572,1988,2012]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0449A6FBCFD45B8112FF84D" box="[443,572,1988,2012]" italics="true" pageId="20" pageNumber="59">Zeanichnus</emphasis>
</taxonomicName>
) and carina demarcating procoxal cavities from the basisternal part of prosternum (absent in
<taxonomicName id="4C7F4D70B0459A6EBCD342E41105FF21" box="[405,534,152,176]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0459A6EBCD342E41105FF21" box="[405,534,152,176]" italics="true" pageId="21" pageNumber="60">Zeanichnus</emphasis>
</taxonomicName>
); the setose impressions of mesoventrite broadly separated in middle (narrowly in
<taxonomicName id="4C7F4D70B0459A6EBDFC42C01228FF45" box="[186,315,188,212]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0459A6EBDFC42C01228FF45" box="[186,315,188,212]" italics="true" pageId="21" pageNumber="60">Zeanichnus</emphasis>
</taxonomicName>
); a broad mesoventral intercoxal process posteriorly fused with the metaventrite (narrow and posteriorly separated from metaventrite in
<taxonomicName id="4C7F4D70B0459A6EBFC5429C1017FF69" box="[643,772,224,248]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0459A6EBFC5429C1017FF69" box="[643,772,224,248]" italics="true" pageId="21" pageNumber="60">Zeanichnus</emphasis>
</taxonomicName>
); and the lack of metaventral anterior process (present in
<taxonomicName id="4C7F4D70B0459A6EBDD14378120BFE8D" box="[151,280,260,284]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0459A6EBDD14378120BFE8D" box="[151,280,260,284]" italics="true" pageId="21" pageNumber="60">Zeanichnus</emphasis>
</taxonomicName>
) (differences are compiled in
<tableCitation id="C6FD0348B0459A6EBF2F437811A8FE8D" box="[617,699,260,284]" captionStart="TABLE 1" captionStartId="14.[151,239,295,319]" captionTargetBox="[165,1418,351,1998]" captionTargetPageId="14" captionText="TABLE 1. Major morphological differences between genera and subgenera." httpUri="http://table.plazi.org/id/DF00667BB05E9A75BDD1435B10D6FEAE" pageId="21" pageNumber="60" tableUuid="DF00667BB05E9A75BDD1435B10D6FEAE">Table 1</tableCitation>
).
</paragraph>
<caption id="DF00667BB0459A6EBDD14495108BF87A" httpUri="https://zenodo.org/record/283480/files/figure.png" pageId="21" pageNumber="60" targetBox="[167,1417,350,1731]" targetPageId="21">
<paragraph id="8BC036F3B0459A6EBDD14495108BF87A" blockId="21.[151,1437,1769,2027]" pageId="21" pageNumber="60">
<emphasis id="B90BEAE1B0459A6EBDD144951244F96E" bold="true" box="[151,343,1769,1792]" pageId="21" pageNumber="60">FIGURES 67–73.</emphasis>
<taxonomicName id="4C7F4D70B0459A6EBC1944961194F891" authority="Broun" authorityName="Broun" box="[351,647,1770,1792]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0459A6EBC194496113FF891" box="[351,556,1770,1792]" italics="true" pageId="21" pageNumber="60">Zeanichnus insignis</emphasis>
(Broun)
</taxonomicName>
, holotype male (67, 70–71) and
<taxonomicName id="4C7F4D70B0459A6EBEA644961625F891" authority="Franz" authorityName="Franz" box="[992,1334,1770,1792]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="kuschelianus">
<emphasis id="B90BEAE1B0459A6EBEA6449617F3F891" box="[992,1248,1770,1792]" italics="true" pageId="21" pageNumber="60">Zeanichnus kuschelianus</emphasis>
(Franz)
</taxonomicName>
, holotype male (68–69, 72–73). Venter (67–68), head in ventral view (69), aedeagus in ventral (70, 72) and lateral (71, 73) views. Abbreviations: aest2, mesoanepisternum; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; h, hypostoma; hr, hypostomal ridge; hy, hypomeron; lac, lacinia; lb, labrum; lp, labial palp; lss, lateral suture of submentum; mcp, mesocoxal projection; mn, mentum; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pl, posterior lobe of mesocoxal projection; pre, prepectus; si, setose impression of mesoventrite; smn, submentum; tm, temple; v3, metaventrite.
</paragraph>
</caption>
<paragraph id="8BC036F3B0469A6DBD8142E41110FE5E" blockId="22.[151,1436,151,788]" pageId="22" pageNumber="61">
It was not possible to examine details of the mandibles (especially their proximal parts) and prementum in the fragile
<typeStatus id="54C48851B0469A6DBDAE42C01246FF45" box="[232,341,188,212]" pageId="22" pageNumber="61" type="holotype">holotypes</typeStatus>
available for the study. The general shape of the mandible in
<taxonomicName id="4C7F4D70B0469A6DB95F42C01789FF45" box="[1049,1178,188,212]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEAE1B0469A6DB95F42C01789FF45" box="[1049,1178,188,212]" italics="true" pageId="22" pageNumber="61">Zeanichnus</emphasis>
</taxonomicName>
seems to be similar to that of
<taxonomicName id="4C7F4D70B0469A6DBDA9429C12BCFF69" box="[239,431,224,248]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1B0469A6DBDA9429C12BCFF69" box="[239,431,224,248]" italics="true" pageId="22" pageNumber="61">Austrostenichnus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D70B0469A6DBCA8429C1196FF69" box="[494,645,224,248]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1B0469A6DBCA8429C1196FF69" box="[494,645,224,248]" italics="true" pageId="22" pageNumber="61">Scydmaenilla</emphasis>
</taxonomicName>
, i.e., a broad base, abruptly demarcated from a moderately slender apical part bearing a subapical tooth on the mesal margin. The subapical mandibular tooth was well-visible, but the basal part of mandibles, concealed by maxillae, were not visible well enough to describe their shape. Labial discs on the prementum were also not found, but the broadly separated bases of labial palps and lack of a median pair of long bristle-like sensilla (typically located between the palps in many
<taxonomicName id="4C7F4D70B0469A6DBEE843131745FE16" box="[942,1110,367,391]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
) suggest that the prementum may be modified in a similar way as in
<taxonomicName id="4C7F4D70B0469A6DBF2643E811CBFE3D" box="[608,728,404,428]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1B0469A6DBF2643E811CBFE3D" box="[608,728,404,428]" italics="true" pageId="22" pageNumber="61">Stenichnus</emphasis>
</taxonomicName>
. However, additional specimens are necessary to examine the mouthpart morphology in detail.
</paragraph>
<paragraph id="8BC036F3B0469A6DBD8143A01110FC85" blockId="22.[151,1436,151,788]" pageId="22" pageNumber="61">
<taxonomicName id="4C7F4D70B0469A6DBD8143A012B4FE65" box="[199,423,476,500]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B90BEAE1B0469A6DBD8143A012B4FE65" box="[199,423,476,500]" italics="true" pageId="22" pageNumber="61">Zeanichnus insignis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D70B0469A6DBCA743A11183FE65" box="[481,656,476,500]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="species" species="kuschelianus">
<emphasis id="B90BEAE1B0469A6DBCA743A11183FE65" box="[481,656,476,500]" italics="true" pageId="22" pageNumber="61">Z. kuschelianus</emphasis>
</taxonomicName>
differ strikingly in the structure of the antennae. The former species has slender antennae with 4-antennomere club, while
<taxonomicName id="4C7F4D70B0469A6DBE6E407D10CDFD89" box="[808,990,512,536]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="species" species="kuschelianus">
<emphasis id="B90BEAE1B0469A6DBE6E407D10CDFD89" box="[808,990,512,536]" italics="true" pageId="22" pageNumber="61">Z. kuschelianus</emphasis>
</taxonomicName>
has massive antennae with elongate antennomeres I–III, and nearly spherical or transverse antennomeres IV–X. Such big differences in the shape and composition of the antennal club are rare within genera of
<taxonomicName id="4C7F4D70B0469A6DBE67403B10DBFDCE" box="[801,968,583,607]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="61" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
and if they occur, then usually are used as subgeneric diagnostic characters. However, both species from
<collectingCountry id="F3687663B0469A6DBE14401110FBFD15" box="[850,1000,620,644]" name="New Zealand" pageId="22" pageNumber="61">New Zealand</collectingCountry>
share many character states associated with the head capsule, prothorax, pterothorax, elytra and aedeagus. Especially their aedeagi are highly similar, making it possible to interpret various parts (e.g., the strongly narrowing apex of dorsal wall, sub-apical lateral groups of needle-like sclerites, various components of the central complex) as homologous and clearly suggesting close evolutionary relationships.
</paragraph>
</subSubSection>
</treatment>
</document>
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