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<document id="B776A6C7A9068D6F336E586547A18FBD" ID-DOI="10.5252/adansonia2020v42a6" ID-GBIF-Dataset="a270ab31-7e52-4655-aaa4-55d34fcf7fb3" ID-ISSN="1639-4798" ID-Zenodo-Dep="3744689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586354780961" checkinUser="carolina" docAuthor="Florence, Jacques &amp; Butaud, Jean-François" docDate="2020" docId="03D63B06A016FFDCC9F0F8C4FBFDFBD1" docLanguage="fr" docName="adansonia2020v42a6.pdf.imf" docOrigin="Adansonia 42 (6)" docStyle="DocumentStyle:2FE19AC740987067EC2A6C89A85F9648.8:Adansonia.2018-.journal_article" docStyleId="2FE19AC740987067EC2A6C89A85F9648" docStyleName="Adansonia.2018-.journal_article" docStyleVersion="8" docTitle="Psychotria niauensis Florence &amp; Butaud 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="128" masterDocId="FFEF437EA013FFD7C944FF97FFB2FFA0" masterDocTitle="Psychotria niauensis sp. nov. (Rubiaceae), espèce calcicole endémique de la Polynésie française, décrite de l’atoll soulevé de Niau (Tuamotu)" masterLastPageNumber="130" masterPageNumber="119" pageNumber="122" updateTime="1698736476534" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="9B855CF7EA4927ED5F0F12D61B60FC91" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="58A7806484F4684FE4D6D9D2E738D252">
<mods:title id="E7B4821610F62110533CE1284387E88F">Psychotria niauensis sp. nov. (Rubiaceae), espèce calcicole endémique de la Polynésie française, décrite de l’atoll soulevé de Niau (Tuamotu)</mods:title>
</mods:titleInfo>
<mods:name id="B0045320AA7D7F04567852B3212D3A2D" type="personal">
<mods:role id="42DA47F98B6719E5817368EDC381F0AE">
<mods:roleTerm id="158755801C695642647D6C1ED33A026F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA17BF0F7EAF83F081F77D939C9F1010">Florence, Jacques</mods:namePart>
<mods:affiliation id="A14B17302BDE987FB0954AB1BC94A5BF">Herbier national, Muséum national d'Histoire naturalle</mods:affiliation>
<mods:nameIdentifier id="242D46E64FC7216221F677D899674517" type="email">jflo@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="6E8B29A2EC350DB09395D1650571A4B6" type="personal">
<mods:role id="E7182E4BAC943BC22770174B52E362D9">
<mods:roleTerm id="DD1457FE8C6C4B1200D5860AF161008D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="96A0EB5818F50F1409421600673F5B4D">Butaud, Jean-François</mods:namePart>
<mods:affiliation id="56320F86232023F83039C86319790313">Consultant en foresterie et botanique polynésienne</mods:affiliation>
<mods:nameIdentifier id="90D2AC632CF5928810733F58DEC5BB43" type="email">jfbutaud@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5A02D955BA22249823F04CECCB562CE6">text</mods:typeOfResource>
<mods:relatedItem id="FD7F42CEF10407AC97A81AFBFC275121" type="host">
<mods:titleInfo id="DA4B0682A6A19FE05EEE0C3B69F162ED">
<mods:title id="D59EA8C4054BF9205DD47CEEF51B375F">Adansonia</mods:title>
</mods:titleInfo>
<mods:part id="27B9EFAB7AE6194C610E180DED47B41E">
<mods:date id="D7ADA2D0811191D021208BCF5050BBCB">2020</mods:date>
<mods:detail id="F69B68E21EA113C5BDF106F4DDD7507C" type="pubDate">
<mods:number id="8E003E5AB73375B3F6F80E34E5B5DC8D">2020-03-30</mods:number>
</mods:detail>
<mods:detail id="400A4AFEDACC8E7BE392E7AA68F18A64" type="volume">
<mods:number id="CC0A2E0F4CB6850A165B1032BE1253F5">42</mods:number>
</mods:detail>
<mods:detail id="0489334F6629B8CFE581ED036ADFFB19" type="issue">
<mods:number id="479045274E653328095C32014EF96C08">6</mods:number>
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<mods:start id="32DE928D328959BB7ACB30CCD00812D3">119</mods:start>
<mods:end id="47C6A52EE1270B20D524C1C4CCD02291">130</mods:end>
</mods:extent>
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<mods:classification id="FCE2DCEFF3B1B19715247A9FA67FE87F">journal article</mods:classification>
<mods:identifier id="3C944A949F6C4BFAF1426406211A58FD" type="DOI">10.5252/adansonia2020v42a6</mods:identifier>
<mods:identifier id="9C4580B5B1925A0D27A8DE7B242B4031" type="GBIF-Dataset">a270ab31-7e52-4655-aaa4-55d34fcf7fb3</mods:identifier>
<mods:identifier id="17FABA043723670ACDF7C504F1DB157A" type="ISSN">1639-4798</mods:identifier>
<mods:identifier id="0A73F0D384F717BA06A5A99016A74CA2" type="Zenodo-Dep">3744689</mods:identifier>
</mods:mods>
<treatment id="03D63B06A016FFDCC9F0F8C4FBFDFBD1" ID-DOI="http://doi.org/10.5281/zenodo.3809609" ID-GBIF-Taxon="163538784" ID-Zenodo-Dep="3809609" LSID="urn:lsid:plazi:treatment:03D63B06A016FFDCC9F0F8C4FBFDFBD1" httpUri="http://treatment.plazi.org/id/03D63B06A016FFDCC9F0F8C4FBFDFBD1" lastPageId="11" lastPageNumber="128" pageId="5" pageNumber="122">
<subSubSection id="C365D99BA016FFD2C9F0F8C4FD6AF8CE" box="[180,728,1875,1902]" pageId="5" pageNumber="122" type="nomenclature">
<paragraph id="8BC08A10A016FFD2C9F0F8C4FD6AF8CE" blockId="5.[180,728,1875,1902]" box="[180,728,1875,1902]" pageId="5" pageNumber="122">
<heading id="D0883D7CA016FFD2C9F0F8C4FD6AF8CE" box="[180,728,1875,1902]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="122" reason="3">
<taxonomicName id="4C7FF193A016FFD2C9F0F8C4FDC8F8CD" ID-CoL="9V3W9" authority="Butaud &amp; J.Florence" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[180,634,1875,1902]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="122" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A016FFD2C9F0F8C4FE3DF8CD" bold="true" box="[180,399,1875,1902]" italics="true" pageId="5" pageNumber="122">Psychotrianiauensis</emphasis>
Butaud &amp; J.Florence
</taxonomicName>
,
<taxonomicNameLabel id="A238EB79A016FFD2CBC1F8C3FD6AF8CE" box="[645,728,1876,1902]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2C8CBF8E4FE4EF82E" box="[399,508,1907,1934]" pageId="5" pageNumber="122" type="description">
<paragraph id="8BC08A10A016FFD2C8CBF8E4FE4EF82E" blockId="5.[399,508,1907,1934]" box="[399,508,1907,1934]" pageId="5" pageNumber="122">
(
<figureCitation id="13449695A016FFD2C8DCF8E4FE65F82D" box="[408,471,1907,1933]" captionStart="FIG" captionStartId="7.[297,308,1833,1850]" captionTargetBox="[172,1416,215,1792]" captionTargetId="figure@7.[172,1416,212,1795]" captionTargetPageId="7" captionText="FIG. 3. — Inflorescence de Psychotria niauensis Butaud &amp; J.Florence, sp. nov. (Butaud 1540 [PAP015890, PAP015891])." figureDoi="http://doi.org/10.5281/zenodo.3744695" httpUri="https://zenodo.org/record/3744695/files/figure.png" pageId="5" pageNumber="122">Figs 3</figureCitation>
;
<figureCitation id="13449695A016FFD2C8A1F8E4FE41F82D" box="[485,499,1907,1933]" captionStart="FIG" captionStartId="8.[351,362,1870,1887]" captionTargetBox="[172,1416,215,1830]" captionTargetId="figure@8.[172,1416,215,1834]" captionTargetPageId="8" captionText="FIG. 4. — Infrutescence de Psychotria niauensis Butaud &amp; J.Florence, sp. nov. (Butaud 1498 [PAP015764])." figureDoi="http://doi.org/10.5281/zenodo.3744697" httpUri="https://zenodo.org/record/3744697/files/figure.png" pageId="5" pageNumber="122">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2C9C0F82CFA02FE86" pageId="5" pageNumber="122" type="diagnosis">
<paragraph id="8BC08A10A016FFD2C9C0F82CFA02FE86" blockId="5.[132,775,1978,2029]" lastBlockId="5.[813,1456,217,294]" pageId="5" pageNumber="122">
P. leiophyllae
<emphasis id="B90B5602A016FFD2C840F82CFA02FE86" italics="true" pageId="5" pageNumber="122">
Merr. &amp; L.M.Perry affinis, sedsemper munitispedunculo longiori quam
<quantity id="4C8727F5A016FFD2C849F842FEFFF84C" box="[269,333,2005,2029]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="122" unit="cm" value="3.5">3,5 cm</quantity>
inflorescentiis (vs sessilia atquee base ramosa vel pedunculominusquam
<quantity id="4C8727F5A016FFD2CD47FF4EFB9FFF50" box="[1027,1069,217,241]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="122" unit="cm" value="3.0">3 cm</quantity>
inflorescentia), ovoideisminus quam
<quantity id="4C8727F5A016FFD2CC3EFF4EFA1DFF50" box="[1402,1455,217,241]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="122" unit="mm" value="3.0">3 mm</quantity>
crassis pyrenis (vs subellipsoideas vel subglobosas ultra
<quantity id="4C8727F5A016FFD2CC66FF64FADFFEAB" box="[1314,1389,243,267]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="122" unit="mm" value="3.5">3,5 mm</quantity>
crassas pyrenas), minus quam
<quantity id="4C8727F5A016FFD2CABFFE99FB95FE85" box="[1019,1063,270,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="122" unit="cm" value="1.0">1 cm</quantity>
longo petiolo (vs
<quantity id="4C8727F5A016FFD2CD85FE99FAB7FE85" box="[1217,1285,270,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="5" pageNumber="122" unit="cm" value="3.0" valueMax="5.0" valueMin="1.0">1-5 cm</quantity>
), praecipue differt.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2CA69FEAAFB6DFE05" pageId="5" pageNumber="122" type="type_taxon">
<paragraph id="8BC08A10A016FFD2CA69FEAAFB6DFE05" blockId="5.[813,1456,317,421]" pageId="5" pageNumber="122">
<materialsCitation id="3B17804DA016FFD2CA69FEAAFB6DFE05" ID-GBIF-Occurrence="2620271301" collectingDate="2006-10-17" collectionCode="PAP" collectorName="F. Butaud" country="French Polynesia" latitude="-16.15" location="Ouest" longLatPrecision="1282" longitude="-146.4" municipality="Archipeldes Tuamotu. Atollde Niau" pageId="5" pageNumber="122" specimenCode="PAP015889" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C434B2A016FFD2CA69FEAAFCDAFEF2" box="[813,872,317,340]" pageId="5" pageNumber="122">TYPUS</typeStatus>
. —
<emphasis id="B90B5602A016FFD2CAD4FEAAFA93FEF5" bold="true" box="[912,1313,317,341]" pageId="5" pageNumber="122">
<collectingCountry id="F368CA80A016FFD2CAD4FEAAFBF7FEF5" box="[912,1093,317,341]" name="French Polynesia" pageId="5" pageNumber="122">Polynésie française</collectingCountry>
,
<collectingMunicipality id="6BA4106AA016FFD2CD0AFEAAFA93FEF5" box="[1102,1313,317,341]" pageId="5" pageNumber="122">Archipeldes Tuamotu</collectingMunicipality>
</emphasis>
. Atollde Niau,
<location id="8EA0DCCBA016FFD2CA69FECFFCDAFED0" LSID="urn:lsid:plazi:treatment:03D63B06A016FFDCC9F0F8C4FBFDFBD1:8EA0DCCBA016FFD2CA69FECFFCDAFED0" box="[813,872,344,368]" country="French Polynesia" latitude="-16.15" longLatPrecision="1282" longitude="-146.4" municipality="Archipeldes Tuamotu. Atollde Niau" name="Ouest" pageId="5" pageNumber="122">Ouest</location>
de l’aéroport, coté lagon en arrière des zones d’extraction,
<quantity id="4C8727F5A016FFD2CA69FEE5FCE5FE2A" box="[813,855,370,394]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="122" unit="m" value="8.0">8 m</quantity>
,
<geoCoordinate id="EE4BECD7A016FFD2CA27FEE5FC00FE2A" box="[867,946,370,394]" degrees="16" direction="south" minutes="09" orientation="latitude" pageId="5" pageNumber="122" precision="925" value="-16.15">16°09’S</geoCoordinate>
,
<geoCoordinate id="EE4BECD7A016FFD2CAFAFEE5FB91FE2A" box="[958,1059,370,394]" degrees="146" direction="west" minutes="24" orientation="longitude" pageId="5" pageNumber="122" precision="925" value="-146.4">146°24’W</geoCoordinate>
,
<date id="FFC1ACD0A016FFD2CD6AFEE5FB2BFE2A" box="[1070,1177,370,394]" pageId="5" pageNumber="122" value="2006-10-17">
<collectingDate id="EF855538A016FFD2CD6AFEE5FB2BFE2A" box="[1070,1177,370,394]" pageId="5" pageNumber="122" value="2006-10-17">17.X.2006</collectingDate>
</date>
, fl. &amp; fr.,
<emphasis id="B90B5602A016FFD2CC44FEE4FA1DFE2A" box="[1280,1455,370,394]" italics="true" pageId="5" pageNumber="122">
J.-
<collectorName id="268AEFC6A016FFD2CC50FEE4FAC7FE2A" box="[1300,1397,370,394]" pageId="5" pageNumber="122">F. Butaud</collectorName>
1519
</emphasis>
(holo-, P!; iso-,
<collectionCode id="ED6E12D5A016FFD2CA88FE1AFC4BFE05" box="[972,1017,397,421]" collectionName="PAP" pageId="5" pageNumber="122">PAP</collectionCode>
[
<specimenCode id="DBD9226BA016FFD2CABAFE1BFBC7FE04" box="[1022,1141,396,420]" pageId="5" pageNumber="122">PAP015889</specimenCode>
!]) (
<figureCitation id="13449695A016FFD2CDDDFE1AFB63FE04" box="[1177,1233,396,421]" captionStart="FIG" captionStartId="10.[331,342,2008,2025]" captionTargetBox="[241,1346,215,1954]" captionTargetId="figure@10.[241,1346,215,1954]" captionTargetPageId="10" captionText="FIG. 5. — Image de l’isotype de Psychotria niauensis Butaud &amp; J.Florence, sp. nov. (Butaud 1519 [PAP015889])." figureDoi="http://doi.org/10.5281/zenodo.3744699" httpUri="https://zenodo.org/record/3744699/files/figure.png" pageId="5" pageNumber="122">Fig. 5</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2CA69FE2BFB16FDD3" pageId="5" pageNumber="122" type="materials_examined">
<paragraph id="8BC08A10A016FFD2CA69FE2BFB16FDD3" blockId="5.[813,1456,444,627]" pageId="5" pageNumber="122">
<materialsCitation id="3B17804DA016FFD2CA69FE2BFAF8FDA9" ID-GBIF-Occurrence="2597224108" collectingDate="2006-10-13" collectionCode="PAP" collectorName="F. Butaud" country="French Polynesia" elevation="8" latitude="-16.133333" location="Tupana" longLatPrecision="1283" longitude="-146.33333" pageId="5" pageNumber="122" specimenCode="PAP015764" specimenCount="1" stateProvince="Tupana" typeStatus="paratype">
<typeStatus id="54C434B2A016FFD2CA69FE2BFC3BFE71" box="[813,905,444,467]" pageId="5" pageNumber="122" type="paratype">PARATYPI</typeStatus>
. —
<emphasis id="B90B5602A016FFD2CAFDFE2BFADDFE74" bold="true" box="[953,1391,444,468]" pageId="5" pageNumber="122">
<collectingCountry id="F368CA80A016FFD2CAFDFE2BFBCFFE74" box="[953,1149,444,468]" name="French Polynesia" pageId="5" pageNumber="122" value="French Polynesia">Polynésie française</collectingCountry>
, Archipeldes Tuamotu
</emphasis>
. Atoll de Niau,
<collectingRegion id="49BB44F2A016FFD2CACEFE40FC64FE4E" box="[906,982,471,494]" pageId="5" pageNumber="122">Tupana</collectingRegion>
, derrière le cimetière,
<quantity id="4C8727F5A016FFD2CDF0FE40FB6DFE4E" box="[1204,1247,471,495]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="122" unit="m" value="8.0">
<elevation id="00526D23A016FFD2CDF0FE40FB6DFE4E" box="[1204,1247,471,495]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="122" unit="m" value="8.0">8 m</elevation>
</quantity>
,
<geoCoordinate id="EE4BECD7A016FFD2CDAFFE40FA88FE4F" box="[1259,1338,471,495]" degrees="16" direction="south" minutes="08" orientation="latitude" pageId="5" pageNumber="122" precision="925" value="-16.133333">16°08’S</geoCoordinate>
,
<geoCoordinate id="EE4BECD7A016FFD2CC02FE40FA1EFE4F" box="[1350,1452,471,495]" degrees="146" direction="west" minutes="20" orientation="longitude" pageId="5" pageNumber="122" precision="925" value="-146.33333">146°20’W</geoCoordinate>
,
<date id="FFC1ACD0A016FFD2CA69FE66FC2BFDA9" box="[813,921,497,521]" pageId="5" pageNumber="122" value="2006-10-13">
<collectingDate id="EF855538A016FFD2CA69FE66FC2BFDA9" box="[813,921,497,521]" pageId="5" pageNumber="122" value="2006-10-13">13.X.2006</collectingDate>
</date>
, fr.,
<emphasis id="B90B5602A016FFD2CA8CFE65FBCBFDA9" box="[968,1145,497,521]" italics="true" pageId="5" pageNumber="122">
J.-
<collectorName id="268AEFC6A016FFD2CA99FE65FB8CFDA9" box="[989,1086,497,521]" pageId="5" pageNumber="122">F. Butaud</collectorName>
1498
</emphasis>
(
<collectionCode id="ED6E12D5A016FFD2CDCCFE66FB05FDA9" box="[1160,1207,497,521]" collectionName="PAP" pageId="5" pageNumber="122">PAP</collectionCode>
[
<specimenCode id="DBD9226BA016FFD2CDF9FE66FA86FDA9" box="[1213,1332,497,521]" pageId="5" pageNumber="122">PAP015764</specimenCode>
!])
</materialsCitation>
;
<materialsCitation id="3B17804DA016FFD2CC1BFE65FBC1FD9E" ID-GBIF-Occurrence="2597224106" collectingDate="2006-10-13" collectionCode="PAP" collectorName="F. Butaud" country="French Polynesia" latitude="-16.133333" location="Tupana" longLatPrecision="1283" longitude="-146.33333" pageId="5" pageNumber="122" specimenCode="PAP015763" specimenCount="8" specimenCount-male="8" stateProvince="Tupana">
<collectingRegion id="49BB44F2A016FFD2CC1BFE65FA19FDA9" box="[1375,1451,498,521]" pageId="5" pageNumber="122">Tupana</collectingRegion>
, derrière le cimetière,
<quantity id="4C8727F5A016FFD2CABDFD9BFB93FD83" box="[1017,1057,524,548]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="122" unit="m" value="8.0">
<specimenCount id="9D794199A016FFD2CABDFD9BFB93FD83" box="[1017,1057,524,548]" pageId="5" pageNumber="122" type="male">8 m</specimenCount>
</quantity>
,
<geoCoordinate id="EE4BECD7A016FFD2CD6FFD9BFBC8FD84" box="[1067,1146,524,548]" degrees="16" direction="south" minutes="08" orientation="latitude" pageId="5" pageNumber="122" precision="925" value="-16.133333">16°08’S</geoCoordinate>
,
<geoCoordinate id="EE4BECD7A016FFD2CDC0FD9BFB5BFD84" box="[1156,1257,524,548]" degrees="146" direction="west" minutes="20" orientation="longitude" pageId="5" pageNumber="122" precision="925" value="-146.33333">146°20’W</geoCoordinate>
,
<date id="FFC1ACD0A016FFD2CDB7FD9BFAECFD84" box="[1267,1374,524,548]" pageId="5" pageNumber="122" value="2006-10-13">
<collectingDate id="EF855538A016FFD2CDB7FD9BFAECFD84" box="[1267,1374,524,548]" pageId="5" pageNumber="122" value="2006-10-13">13.X.2006</collectingDate>
</date>
, fl.,
<emphasis id="B90B5602A016FFD2CCCCFD9BFC1BFD9E" italics="true" pageId="5" pageNumber="122">
J.-
<collectorName id="268AEFC6A016FFD2CCD9FD9BFCC0FD9E" pageId="5" pageNumber="122">F. Butaud</collectorName>
1499
</emphasis>
(
<collectionCode id="ED6E12D5A016FFD2CAF0FDB1FC50FD9E" box="[948,994,550,574]" collectionName="PAP" pageId="5" pageNumber="122">PAP</collectionCode>
[
<specimenCode id="DBD9226BA016FFD2CAACFDB1FBEFFD9E" box="[1000,1117,550,574]" pageId="5" pageNumber="122">PAP015763</specimenCode>
!])
</materialsCitation>
;
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<collectingRegion id="49BB44F2A016FFD2CDC5FDB0FB7EFD9E" box="[1153,1228,551,574]" pageId="5" pageNumber="122">Tupana</collectingRegion>
, enarrièreducimetière,
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<specimenCount id="9D794199A016FFD2CA69FDD6FCEBFDF8" box="[813,857,577,601]" pageId="5" pageNumber="122" type="male">8 m</specimenCount>
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,
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,
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,
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<collectingDate id="EF855538A016FFD2CD73FDD7FB15FDF8" box="[1079,1191,576,600]" pageId="5" pageNumber="122" value="2006-10-24">24.X.2006</collectingDate>
</date>
, fl., fr.,
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J.-
<collectorName id="268AEFC6A016FFD2CC4BFDD6FAC1FDF8" box="[1295,1395,576,600]" pageId="5" pageNumber="122">F. Butaud</collectorName>
1540
</emphasis>
(
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[
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,
<specimenCode id="DBD9226BA016FFD2CAAFFDCCFBD0FDD3" box="[1003,1122,603,627]" pageId="5" pageNumber="122">PAP015891</specimenCode>
]!
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;
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<collectionCode id="ED6E12D5A016FFD2CDC6FDCCFB26FDD3" box="[1154,1172,603,627]" pageId="5" pageNumber="122">P</collectionCode>
!)
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.
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2CA69FD1DFBA6FD77" pageId="5" pageNumber="122" type="biology_ecology">
<paragraph id="8BC08A10A016FFD2CA69FD1DFBA6FD77" blockId="5.[813,1455,650,727]" pageId="5" pageNumber="122">PHÉNOLOGIE. — En fleurs et enfruits en février, mars, juin, octobre et novembre, d’après les échantillons d’herbier et les observations de différents botanistes.</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2CA69FD7BFBA8FC82" pageId="5" pageNumber="122" type="etymology">
<paragraph id="8BC08A10A016FFD2CA69FD7BFBA8FC82" blockId="5.[813,1455,748,802]" pageId="5" pageNumber="122">
ÉTYMOLOGIE.
<emphasis id="B90B5602A016FFD2CAFCFD67FC63FCA7" box="[952,977,752,775]" italics="true" pageId="5" pageNumber="122">—</emphasis>
Le nom d’espèce se réfère à l’atoll de Niau duquel la plante est endémique.
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFD2CA69FCAFFB71FCCC" pageId="5" pageNumber="122" type="vernacular_names">
<paragraph id="8BC08A10A016FFD2CA69FCAFFB71FCCC" blockId="5.[813,1455,824,876]" pageId="5" pageNumber="122">NOM LOCAL ET USAGES. — Aucun nom polynésien ni aucun usage n’ont été relevés pour cette espèce à Niau.</paragraph>
</subSubSection>
<subSubSection id="C365D99BA016FFDECA69FC01FE21FA8F" lastPageId="9" lastPageNumber="126" pageId="5" pageNumber="122" type="description">
<paragraph id="8BC08A10A016FFD2CA69FC01FC0FFC0E" blockId="5.[813,1456,918,1137]" box="[813,957,918,944]" pageId="5" pageNumber="122">DESCRIPTION</paragraph>
<paragraph id="8BC08A10A016FFD2CA69FC20FAF4FBD0" blockId="5.[813,1456,918,1137]" pageId="5" pageNumber="122">
Arbuste atteignant
<quantity id="4C8727F5A016FFD2CABCFC21FBEFFC71" box="[1016,1117,950,977]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="122" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1,5 à 2 m</quantity>
de hauteur et
<quantity id="4C8727F5A016FFD2CDB2FC21FA99FC71" box="[1270,1323,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="122" unit="cm" value="5.0">5 cm</quantity>
de diamètre à la base, entièrement glabre, au port parfois retombant, aux rameaux longtemps chlorophylliens de
<quantity id="4C8727F5A016FFD2CD82FC60FA9AFBB1" box="[1222,1320,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="122" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2 à 3 mm</quantity>
de diamètre, souvent aplatis, puis à écorce brunâtre et lisse. Rameaux, face inférieure desfeuilles et ramilles de l’inflorescenceparsemés de faisceaux de raphides, de
<quantity id="4C8727F5A016FFD2CD7DFBC1FB08FBD1" box="[1081,1210,1110,1137]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="5" pageNumber="122" unit="mm" value="0.44999999999999996" valueMax="0.7" valueMin="0.2">0,2-0,7 mm</quantity>
de longueur.
</paragraph>
<paragraph id="8BC08A10A016FFD2CA69FB01FCCAFB10" blockId="5.[813,1456,1174,1487]" box="[813,888,1174,1200]" pageId="5" pageNumber="122">
<emphasis id="B90B5602A016FFD2CA69FB01FCCAFB10" box="[813,888,1174,1200]" italics="true" pageId="5" pageNumber="122">Feuilles</emphasis>
</paragraph>
<paragraph id="8BC08A10A016FFD2CA69FB21FA19FA6F" blockId="5.[813,1456,1174,1487]" pageId="5" pageNumber="122">
Opposées-décussées, limbede 7,5-15 ×
<quantity id="4C8727F5A016FFD2CD8BFB21FAF6FB70" box="[1231,1348,1206,1233]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.5" metricValueMin="2.5" pageId="5" pageNumber="122" unit="cm" value="4.0" valueMax="5.5" valueMin="2.5">2,5-5,5 cm</quantity>
(1,8 &lt;L/l &lt;3,2), elliptique à obovale, charnu, souple, brillant sur la face supérieure àl’état frais, base cunéiforme longuement décurrente sur le pétiole, apex aigu à obtus, (4)5 à 7 paires de nervures secondaires, eucamptodromesà faiblementbrochidodromes; pétiolede
<quantity id="4C8727F5A016FFD2CADEFAC2FC4CFAD0" box="[922,1022,1365,1392]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="5" pageNumber="122" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4-10 mm</quantity>
; stipules interpétiolaires, sefragmentant rapidement par l’apex, largement triangulaires à engainantes, 1,5 ×
<quantity id="4C8727F5A016FFD2CA23FA02FC15FA10" box="[871,935,1429,1456]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="122" unit="mm" value="2.0">2 mm</quantity>
, parfois bifidessur moins de
<quantity id="4C8727F5A016FFD2CD8BFA02FA90FA10" box="[1231,1314,1429,1456]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="122" unit="mm" value="0.5">0,5 mm</quantity>
en se fendant longitudinalement, face interne à nombreux colleters bruns.
</paragraph>
<paragraph id="8BC08A10A016FFD2CA69FA62FC03F9AF" blockId="5.[813,1456,1525,1774]" box="[813,945,1525,1551]" pageId="5" pageNumber="122">
<emphasis id="B90B5602A016FFD2CA69FA62FC03F9AF" box="[813,945,1525,1551]" italics="true" pageId="5" pageNumber="122">Inflorescences</emphasis>
</paragraph>
<paragraph id="8BC08A10A016FFD2CA69F983FBF7F94E" blockId="5.[813,1456,1525,1774]" pageId="5" pageNumber="122">
Terminales, cymeuses, non calyptrées, de (5-)6,7-15,5 × (2,2-)
<quantity id="4C8727F5A016FFD2CA2EF9A3FC6DF9EF" box="[874,991,1588,1615]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="10.0" metricValueMin="4.5" pageId="5" pageNumber="122" unit="cm" value="7.25" valueMax="10.0" valueMin="4.5">4,5-10 cm</quantity>
, avec 50 à 65 fleurs, pédoncule unique de
<quantity id="4C8727F5A016FFD2CA69F9C3FC28F9CF" box="[813,922,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="10.0" metricValueMin="3.5" pageId="5" pageNumber="122" unit="cm" value="6.75" valueMax="10.0" valueMin="3.5">3,5-10 cm</quantity>
, se divisant en 4 à 5 cymesdichasiales ou verticillées, parfoiscontractées jusqu’à devenir subcapitées, ramifiées 3 à 4 fois, les ramifications ultimes portant 3 (ou 4) fleurs; paires de bractées séteuses, 0,8 ×
<quantity id="4C8727F5A016FFD2CD06F923FB2BF96F" box="[1090,1177,1716,1743]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="122" unit="mm" value="0.3">0,3 mm</quantity>
, au sommet du pédoncule et des divisons successives.
</paragraph>
<paragraph id="8BC08A10A016FFD2CA69F883FCD8F88E" blockId="5.[813,1455,1812,2030]" box="[813,874,1812,1838]" pageId="5" pageNumber="122">
<emphasis id="B90B5602A016FFD2CA69F883FCD8F88E" box="[813,874,1812,1838]" italics="true" pageId="5" pageNumber="122">Fleurs</emphasis>
</paragraph>
<paragraph id="8BC08A10A016FFD0CA69F8A4FE49F84E" blockId="5.[813,1455,1812,2030]" lastBlockId="7.[132,775,1907,2030]" lastPageId="7" lastPageNumber="124" pageId="5" pageNumber="122">
Hermaphrodites, pédicellede
<quantity id="4C8727F5A016FFD2CD24F8A4FB79F8EE" box="[1120,1227,1843,1870]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="122" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1,5 mm</quantity>
. Hypantheobconique de
<quantity id="4C8727F5A016FFD2CA08F8C3FC05F8CE" box="[844,951,1876,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="122" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1-1,2 mm</quantity>
de longueur. Calice verdâtre, tube de
<quantity id="4C8727F5A016FFD2CC00F8C3FA1DF8CE" box="[1348,1455,1876,1902]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="122" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0,8-1 mm</quantity>
de longueur, (4) 5 (6) lobes dressés largement triangulaires de
<quantity id="4C8727F5A016FFD2CA69F804FC33F80E" box="[813,897,1939,1966]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="122" unit="mm" value="0.5">0,5 mm</quantity>
de longueur. Corolle blanche, de
<quantity id="4C8727F5A016FFD2CDA5F804FA92F80E" box="[1249,1312,1939,1966]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="122" unit="mm" value="3.0">3 mm</quantity>
de hauteur et
<quantity id="4C8727F5A016FFD2CA69F823FCDCF86E" box="[813,878,1972,1998]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="122" unit="mm" value="7.0">7 mm</quantity>
de diamètre, hypocratériforme, tube barbu sur la face interne,
<quantity id="4C8727F5A016FFD2CAC1F844FC43F84E" box="[901,1009,2003,2030]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="5" pageNumber="122" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2-2,5 mm</quantity>
de longueur et (4) 5 (6) lobes étroitement ovales, 2,5-3 ×
<quantity id="4C8727F5A014FFD0C865F8E4FE3CF82E" box="[289,398,1907,1934]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="7" pageNumber="124" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1,5-2 mm</quantity>
, révolutés; étaminesblanchesàjaunâtres, exsertes, (4) 5 (6) anthères longues de
<quantity id="4C8727F5A014FFD0CB2BF804FD7AF80E" box="[623,712,1939,1966]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="124" unit="mm" value="1.3">1,3 mm</quantity>
. Style long de
<quantity id="4C8727F5A014FFD0C99BF824FE85F86E" box="[223,311,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="124" unit="mm" value="3.5">3,5 mm</quantity>
et stigmate divisé en deux lobes papilleux ovoïdes; hétérostylie non observée.
</paragraph>
<caption id="DF00DA98A015FFD1C9C0F856FE62F849" ID-DOI="http://doi.org/10.5281/zenodo.3744693" ID-Zenodo-Dep="3744693" httpUri="https://zenodo.org/record/3744693/files/figure.png" pageId="6" pageNumber="123" startId="6.[132,143,1985,2002]" targetBox="[194,1393,215,1947]" targetPageId="6">
<paragraph id="8BC08A10A015FFD1C9C0F856FE62F849" blockId="6.[132,1457,1985,2025]" pageId="6" pageNumber="123">
<subScript id="17FB8855A015FFD1C9C0F856FF2FF872" attach="left" box="[132,157,1985,2002]" fontSize="5" pageId="6" pageNumber="123">FIG</subScript>
. 2. — Imagedel’holotypede
<taxonomicName id="4C7FF193A015FFD1C8D4F856FD79F872" authority="Merr. &amp; L. M. Perry" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[400,715,1985,2002]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="123" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A015FFD1C8D4F856FD85F872" box="[400,567,1985,2002]" italics="true" pageId="6" pageNumber="123">Psychotrialeiophylla</emphasis>
Merr.&amp; L.M.Perry
</taxonomicName>
(
<emphasis id="B90B5602A015FFD1CB90F856FC9DF872" box="[724,815,1985,2002]" italics="true" pageId="6" pageNumber="123">Brass 3133</emphasis>
[A00095390]); crédit: Herbariumof the Arnold Arboretum,Harvard University, Cambridge, Massachusetts, États-Unis.
</paragraph>
</caption>
<caption id="DF00DA98A014FFD0C86DF8BEFAB9F89A" ID-DOI="http://doi.org/10.5281/zenodo.3744695" ID-Zenodo-Dep="3744695" box="[297,1291,1833,1850]" httpUri="https://zenodo.org/record/3744695/files/figure.png" pageId="7" pageNumber="124" startId="7.[297,308,1833,1850]" targetBox="[172,1416,215,1792]" targetPageId="7">
<paragraph id="8BC08A10A014FFD0C86DF8BEFAB9F89A" blockId="7.[297,1291,1833,1850]" box="[297,1291,1833,1850]" pageId="7" pageNumber="124">
<subScript id="17FB8855A014FFD0C86DF8BEFEF0F89A" attach="left" box="[297,322,1833,1850]" fontSize="5" pageId="7" pageNumber="124">FIG</subScript>
. 3. — Inflorescence de
<taxonomicName id="4C7FF193A014FFD0CB4CF8BEFCD1F89A" authority="Butaud &amp; J.Florence" authorityName="Butaud &amp; J. Florence" authorityYear="2020" box="[520,867,1833,1850]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="124" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A014FFD0CB4CF8BEFD03F89A" box="[520,689,1833,1850]" italics="true" pageId="7" pageNumber="124">Psychotria niauensis</emphasis>
Butaud &amp; J.Florence
</taxonomicName>
,
<taxonomicNameLabel id="A238EB79A014FFD0CA2FF8BEFC1DF89A" box="[875,943,1833,1850]" pageId="7" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B90B5602A014FFD0CAF3F8BEFB90F89A" box="[951,1058,1833,1850]" italics="true" pageId="7" pageNumber="124">Butaud 1540</emphasis>
[PAP015890, PAP015891]).
</paragraph>
</caption>
<paragraph id="8BC08A10A014FFD0CA69F8E3FCD1F82D" blockId="7.[813,1455,1908,2030]" box="[813,867,1908,1933]" pageId="7" pageNumber="124">
<emphasis id="B90B5602A014FFD0CA69F8E3FCD1F82D" box="[813,867,1908,1933]" italics="true" pageId="7" pageNumber="124">Fruit</emphasis>
</paragraph>
<paragraph id="8BC08A10A014FFDFCA69F804FC5BF84D" blockId="7.[813,1455,1908,2030]" lastBlockId="8.[813,1455,1939,2030]" lastPageId="8" lastPageNumber="125" pageId="7" pageNumber="124">
Drupacé, tronconique à l’état immature à cylindrique à maturité, de 7 ×
<quantity id="4C8727F5A014FFD0CAA1F824FBF0F86E" box="[997,1090,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="7" pageNumber="124" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6-7 mm</quantity>
, rose à violet noirâtre à maturité, couronné par les dents du calice verdâtres, comportant deuxpyrènes; pédicellede
<quantity id="4C8727F5A01BFFDFC8EAF804FE5DF80E" box="[430,495,1939,1966]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="125" unit="mm" value="2.0">2 mm</quantity>
; pyrènes ovoïdes, hémisphériques en coupe, 4,5-5 × 2,5-3 ×
<quantity id="4C8727F5A01BFFDFCB5EF824FDEDF86E" box="[538,607,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="125" unit="mm" value="2.0">2 mm</quantity>
, à face ventrale plane, traversée longitudinalement par un sillon bisulqué central, largede
<quantity id="4C8727F5A01BFFDFCAA5F804FB89F80E" box="[993,1083,1939,1966]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="125" unit="mm" value="0.5">0,5 mm</quantity>
, etàface dorsalearrondieetlisse; absence de fente de germination préformée, mais présence d’unporeapical.
</paragraph>
<caption id="DF00DA98A01BFFDFC81BF8D9FB67F8FF" ID-DOI="http://doi.org/10.5281/zenodo.3744697" ID-Zenodo-Dep="3744697" box="[351,1237,1870,1887]" httpUri="https://zenodo.org/record/3744697/files/figure.png" pageId="8" pageNumber="125" startId="8.[351,362,1870,1887]" targetBox="[172,1416,215,1830]" targetPageId="8">
<paragraph id="8BC08A10A01BFFDFC81BF8D9FB67F8FF" blockId="8.[351,1237,1870,1887]" box="[351,1237,1870,1887]" pageId="8" pageNumber="125">
<subScript id="17FB8855A01BFFDFC81BF8D9FECAF8FF" attach="left" box="[351,376,1870,1887]" fontSize="5" pageId="8" pageNumber="125">FIG</subScript>
. 4. — Infrutescence de
<taxonomicName id="4C7FF193A01BFFDFCB7BF8D9FC28F8FF" authority="Butaud &amp; J.Florence" authorityName="Butaud &amp; J. Florence" authorityYear="2020" box="[575,922,1870,1887]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="125" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01BFFDFCB7BF8D9FD5BF8FF" box="[575,745,1870,1887]" italics="true" pageId="8" pageNumber="125">Psychotria niauensis</emphasis>
Butaud &amp; J.Florence
</taxonomicName>
,
<taxonomicNameLabel id="A238EB79A01BFFDFCAE6F8D9FC54F8FF" box="[930,998,1870,1887]" pageId="8" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B90B5602A01BFFDFCAAAF8D9FBEBF8FF" box="[1006,1113,1870,1887]" italics="true" pageId="8" pageNumber="125">Butaud 1498</emphasis>
[PAP015764]).
</paragraph>
</caption>
<caption id="DF00DA98A01AFFDEC9C0FF4CFE98FEA3" ID-Table-UUID="DF00DA98A01AFFDEC9C0FF4CFE98FEA3" httpUri="http://table.plazi.org/id/DF00DA98A01AFFDEC9C0FF4CFE98FEA3" pageId="9" pageNumber="126" targetBox="[134,1456,317,1159]" targetIsTable="true" targetPageId="9">
<paragraph id="8BC08A10A01AFFDEC9C0FF4CFE98FEA3" blockId="9.[132,1455,219,259]" pageId="9" pageNumber="126">
<subScript id="17FB8855A01AFFDEC9C0FF4CFF77FF4C" attach="left" box="[132,197,219,236]" fontSize="5" pageId="9" pageNumber="126">TABLEAU</subScript>
1. — Comparaison des caractères morphologiques du genre
<taxonomicName id="4C7FF193A01AFFDECBA4FF4CFC86FF4C" box="[736,820,219,236]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="genus">
<emphasis id="B90B5602A01AFFDECBA4FF4CFC86FF4C" box="[736,820,219,236]" italics="true" pageId="9" pageNumber="126">Eumachia</emphasis>
</taxonomicName>
DC., de
<taxonomicName id="4C7FF193A01AFFDECAC7FF4CFB61FF4C" authority="Merr. &amp; L. M. Perry" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[899,1235,219,236]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECAC7FF4CFB81FF4C" box="[899,1075,219,236]" italics="true" pageId="9" pageNumber="126">Psychotria leiophylla</emphasis>
Merr. &amp; L.M.Perry
</taxonomicName>
et
<taxonomicName id="4C7FF193A01AFFDECDB4FF4CFF6CFEA3" authority="Butaud &amp; J.Florence" authorityName="Butaud &amp; J. Florence" authorityYear="2020" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDECDB4FF4CFAEAFF4C" box="[1264,1368,219,236]" italics="true" pageId="9" pageNumber="126">P. niauensis</emphasis>
Butaud &amp; J.Florence
</taxonomicName>
,
<taxonomicNameLabel id="A238EB79A01AFFDEC9A2FF65FE98FEA3" box="[230,298,242,259]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<table id="F97F78B0A01A0028C9C2FEAAFA02FB27" box="[134,1456,317,1159]" gridcols="3" gridrows="24" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028CABCFEAAFA02FECB" box="[1016,1456,317,363]" gridcol="2" gridrow="0" pageId="9" pageNumber="126">
<paragraph id="8BC08A10A01AFFDECABCFEAAFA27FEF1" box="[1016,1429,317,337]" pageId="9" pageNumber="126">
<taxonomicName id="4C7FF193A01AFFDECABCFEAAFA27FEF1" authority="Merr. &amp; L. M. Perry &amp;" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[1016,1429,317,337]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECABCFEAAFA27FEF1" bold="true" box="[1016,1429,317,337]" pageId="9" pageNumber="126">
<emphasis id="B90B5602A01AFFDECABCFEAAFB7BFEF1" bold="true" box="[1016,1225,317,337]" italics="true" pageId="9" pageNumber="126">Psychotrialeiophylla</emphasis>
Merr. &amp; L.M.Perry &amp;
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FEC0FA02FECB" box="[134,1456,343,363]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FEAAFA02FECB" box="[134,1456,317,363]" gridrow="0" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FEAAFDAFFECB" box="[134,541,317,363]" gridcol="0" gridrow="0" pageId="9" pageNumber="126">Caractère</th>
<emphasis id="B90B5602A01AFFDECB7BFEC0FA02FECB" bold="true" box="[575,1456,343,363]" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028CB7BFEAAFC5BFECB" box="[575,1001,317,363]" gridcol="1" gridrow="0" pageId="9" pageNumber="126">
<taxonomicName id="4C7FF193A01AFFDECB7BFEC0FD17FECB" box="[575,677,343,363]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="genus">
<emphasis id="B90B5602A01AFFDECB7BFEC0FD17FECB" bold="true" box="[575,677,343,363]" italics="true" pageId="9" pageNumber="126">Eumachia</emphasis>
</taxonomicName>
DC.
</th>
<taxonomicName id="4C7FF193A01AFFDECD4AFEC0FAE9FECB" authority="Butaud &amp; J.Florence" authorityName="Butaud &amp; J. Florence" authorityYear="2020" box="[1038,1371,343,363]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDECD4AFEC0FB3AFECB" bold="true" box="[1038,1160,343,363]" italics="true" pageId="9" pageNumber="126">P. niauensis</emphasis>
Butaud &amp; J.Florence
</taxonomicName>
<taxonomicNameLabel id="A238EB79A01AFFDECC25FEC0FA02FECB" box="[1377,1456,343,363]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</tr>
</paragraph>
</th>
<paragraph id="8BC08A10A01AFFDEC9C2FEECFBF0FE2F" box="[134,1090,379,399]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FEECFBF0FE2F" box="[134,1090,379,399]" gridrow="1" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FEECFDAFFE2F" box="[134,541,379,399]" gridcol="0" gridrow="1" pageId="9" pageNumber="126">Type biologique</th>
<td id="769EE12EA01A0028CB7BFEECFA02FE2F" box="[575,1456,379,399]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="9" pageNumber="126">Arbuste ou petit arbre Arbuste</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FE04FBE3FE07" box="[134,1105,403,423]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FE04FBE3FE07" box="[134,1105,403,423]" gridrow="2" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FE04FDAFFE07" box="[134,541,403,423]" gridcol="0" gridrow="2" pageId="9" pageNumber="126">Raphides</th>
<td id="769EE12EA01A0028CB7BFE04FA02FE07" box="[575,1456,403,423]" colspan="2" colspanRight="1" gridcol="1" gridrow="2" pageId="9" pageNumber="126">Présence Présence</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FE3CFABFFE1F" box="[134,1293,427,447]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FE3CFABFFE1F" box="[134,1293,427,447]" gridrow="3" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FE3CFDAFFE1F" box="[134,541,427,447]" gridcol="0" gridrow="3" pageId="9" pageNumber="126">Couleur des parties végétatives</th>
<td id="769EE12EA01A0028CB7BFE3CFA02FE1F" box="[575,1456,427,447]" colspan="2" colspanRight="1" gridcol="1" gridrow="3" pageId="9" pageNumber="126">Vert jaunâtre pâle à l’état sec Vert jaunâtre pâle à l’état sec</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FE54FB72FE77" box="[134,1216,451,471]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FE54FB72FE77" box="[134,1216,451,471]" gridrow="4" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FE54FDAFFE77" box="[134,541,451,471]" gridcol="0" gridrow="4" pageId="9" pageNumber="126">Texture des rameaux</th>
<td id="769EE12EA01A0028CB7BFE54FA02FE77" box="[575,1456,451,471]" colspan="2" colspanRight="1" gridcol="1" gridrow="4" pageId="9" pageNumber="126">Lisses et non liégeux Lisses et non liégeux</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FE4CFB38FE4F" box="[134,1162,475,495]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FE4CFB38FE4F" box="[134,1162,475,495]" gridrow="5" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FE4CFDAFFE4F" box="[134,541,475,495]" gridcol="0" gridrow="5" pageId="9" pageNumber="126">Forme des jeunes rameaux et entrenoeuds</th>
<td id="769EE12EA01A0028CB7BFE4CFA02FE4F" box="[575,1456,475,495]" colspan="2" colspanRight="1" gridcol="1" gridrow="5" pageId="9" pageNumber="126">Souvent aplatis Souvent aplatis</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FE64FC75FD81" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FE64FA1CFD9A" box="[134,1454,499,570]" gridrow="6" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FE64FDAFFD9A" box="[134,541,499,570]" gridcol="0" gridrow="6" pageId="9" pageNumber="126">Position et forme des stipules</th>
<td id="769EE12EA01A0028CB7BFE64FC5BFD9A" box="[575,1001,499,570]" gridcol="1" gridrow="6" pageId="9" pageNumber="126">
Interpétiolaires, entières, généralement unies
<td id="769EE12EA01A0028CABCFE64FA02FD9A" box="[1016,1456,499,570]" gridcol="2" gridrow="6" pageId="9" pageNumber="126">Interpétiolaires, parfois bifides, non calyptrées</td>
à la base ou calyptrées sur une grande
<paragraph id="8BC08A10A01AFFDECB11FDB1FC9EFD9A" box="[597,812,550,570]" pageId="9" pageNumber="126">partie de leur longueur</paragraph>
</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FDA9FAF4FDF2" box="[134,1350,574,594]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FDA9FAF4FD25" box="[134,1350,574,645]" gridrow="7" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FDA9FDAFFD25" box="[134,541,574,645]" gridcol="0" gridrow="7" pageId="9" pageNumber="126">Persistance des stipules</th>
<td id="769EE12EA01A0028CB7BFDA9FC5BFD25" box="[575,1001,574,645]" gridcol="1" gridrow="7" pageId="9" pageNumber="126">
Semi-décidues, devenant indurées et
<td id="769EE12EA01A0028CABCFDA9FA02FD25" box="[1016,1456,574,645]" gridcol="2" gridrow="7" pageId="9" pageNumber="126">
Semi-décidues, leurs extrémités se
<paragraph id="8BC08A10A01AFFDECB11FDCFFD46FD25" pageId="9" pageNumber="126">leurs extrémités se désagrégeant désagrégeant progressivement progressivement</paragraph>
</td>
</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FD1DFAD4FD3E" box="[134,1382,650,670]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FD1DFAD4FD3E" box="[134,1382,650,670]" gridrow="8" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FD1DFDAFFD3E" box="[134,541,650,670]" gridcol="0" gridrow="8" pageId="9" pageNumber="126">Inflorescences</th>
<td id="769EE12EA01A0028CB7BFD1DFA02FD3E" box="[575,1456,650,670]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="9" pageNumber="126">Cymeuses aux ramilles vertes à blanchies Cymes composées aux ramilles vertes</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FD35FD4AFD6F" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FD35FAC1FD6F" box="[134,1395,674,719]" gridrow="9" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FD35FDAFFD6F" box="[134,541,674,719]" gridcol="0" gridrow="9" pageId="9" pageNumber="126">Forme du calice et de la corolle</th>
<td id="769EE12EA01A0028CB7BFD35FC5BFD6F" box="[575,1001,674,719]" gridcol="1" gridrow="9" pageId="9" pageNumber="126">
Actinomorphe, corolle infundibuliforme à
<td id="769EE12EA01A0028CABCFD35FA02FD6F" box="[1016,1456,674,719]" gridcol="2" gridrow="9" pageId="9" pageNumber="126">Actinomorphe, corolle hypocratériforme</td>
hypocratériforme
</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FD44FB9CFD47" box="[134,1070,723,743]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FD44FB9CFD47" box="[134,1070,723,743]" gridrow="10" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FD44FDAFFD47" box="[134,541,723,743]" gridcol="0" gridrow="10" pageId="9" pageNumber="126">Couleur de la corolle</th>
<td id="769EE12EA01A0028CB7BFD44FA02FD47" box="[575,1456,723,743]" colspan="2" colspanRight="1" gridcol="1" gridrow="10" pageId="9" pageNumber="126">Blanc, crème, vert-jaune Blanc</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FD7CFBF1FD5F" box="[134,1091,747,767]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FD7CFBF1FD5F" box="[134,1091,747,767]" gridrow="11" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FD7CFDAFFD5F" box="[134,541,747,767]" gridcol="0" gridrow="11" pageId="9" pageNumber="126">Préfloraison</th>
<td id="769EE12EA01A0028CB7BFD7CFA02FD5F" box="[575,1456,747,767]" colspan="2" colspanRight="1" gridcol="1" gridrow="11" pageId="9" pageNumber="126">Valvaire Valvaire</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC94FB8FFCB7" box="[134,1085,771,791]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC94FB8FFCB7" box="[134,1085,771,791]" gridrow="12" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC94FDAFFCB7" box="[134,541,771,791]" gridcol="0" gridrow="12" pageId="9" pageNumber="126">Pilosité de la gorge de la corolle</th>
<td id="769EE12EA01A0028CB7BFC94FA02FCB7" box="[575,1456,771,791]" colspan="2" colspanRight="1" gridcol="1" gridrow="12" pageId="9" pageNumber="126">Barbue Barbue</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC8BFBECFC90" box="[134,1118,796,816]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC8BFBECFC90" box="[134,1118,796,816]" gridrow="13" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC8BFDAFFC90" box="[134,541,796,816]" gridcol="0" gridrow="13" pageId="9" pageNumber="126">Style</th>
<td id="769EE12EA01A0028CB7BFC8BFA02FC90" box="[575,1456,796,816]" colspan="2" colspanRight="1" gridcol="1" gridrow="13" pageId="9" pageNumber="126">Hétérostyle Homostyle</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FCA3FB8EFCE8" box="[134,1084,820,840]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FCA3FB8EFCE8" box="[134,1084,820,840]" gridrow="14" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FCA3FDAFFCE8" box="[134,541,820,840]" gridcol="0" gridrow="14" pageId="9" pageNumber="126">Polymérie</th>
<td id="769EE12EA01A0028CB7BFCA3FA02FCE8" box="[575,1456,820,840]" colspan="2" colspanRight="1" gridcol="1" gridrow="14" pageId="9" pageNumber="126">4 ou 5-mère 5-mère</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FCDBFB0BFCD9" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FCDBFA36FCD9" box="[134,1412,844,889]" gridrow="15" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FCDBFDAFFCD9" box="[134,541,844,889]" gridcol="0" gridrow="15" pageId="9" pageNumber="126">Ovaire</th>
<td id="769EE12EA01A0028CB7BFCDBFC5BFCD9" box="[575,1001,844,889]" gridcol="1" gridrow="15" pageId="9" pageNumber="126">
Biloculaire aux ovules solitaires et basaux
<td id="769EE12EA01A0028CABCFCDBFA02FCD9" box="[1016,1456,844,889]" gridcol="2" gridrow="15" pageId="9" pageNumber="126">Biloculaire aux ovules solitaires et basaux dans chaque loge dans chaque loge</td>
</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FCEAFB75FC31" box="[134,1223,893,913]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FCEAFB75FC31" box="[134,1223,893,913]" gridrow="16" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FCEAFDAFFC31" box="[134,541,893,913]" gridcol="0" gridrow="16" pageId="9" pageNumber="126">Type et forme de fruit</th>
<td id="769EE12EA01A0028CB7BFCEAFA02FC31" box="[575,1456,893,913]" colspan="2" colspanRight="1" gridcol="1" gridrow="16" pageId="9" pageNumber="126">Drupacé, ellipsoïde à subglobuleux Drupacé, tronconique</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC01FB78FC0A" box="[134,1226,918,938]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC01FB78FC0A" box="[134,1226,918,938]" gridrow="17" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC01FDAFFC0A" box="[134,541,918,938]" gridcol="0" gridrow="17" pageId="9" pageNumber="126">Couleur du fruit</th>
<td id="769EE12EA01A0028CB7BFC01FA02FC0A" box="[575,1456,918,938]" colspan="2" colspanRight="1" gridcol="1" gridrow="17" pageId="9" pageNumber="126">Orange à rouge Rose, pourpre à violet</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC39FB55FC62" box="[134,1255,942,962]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC39FB55FC62" box="[134,1255,942,962]" gridrow="18" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC39FDAFFC62" box="[134,541,942,962]" gridcol="0" gridrow="18" pageId="9" pageNumber="126">Forme des pyrènes</th>
<td id="769EE12EA01A0028CB7BFC39FA02FC62" box="[575,1456,942,962]" colspan="2" colspanRight="1" gridcol="1" gridrow="18" pageId="9" pageNumber="126">Hémisphérique en coupe Hémisphérique en coupe</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC51FB9BFC7A" box="[134,1065,966,986]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC51FB9BFC7A" box="[134,1065,966,986]" gridrow="19" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC51FDAFFC7A" box="[134,541,966,986]" gridcol="0" gridrow="19" pageId="9" pageNumber="126">Face dorsale du pyrène</th>
<td id="769EE12EA01A0028CB7BFC51FA02FC7A" box="[575,1456,966,986]" colspan="2" colspanRight="1" gridcol="1" gridrow="19" pageId="9" pageNumber="126">Lisse à côtelée Lisse</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC49FAD0FC52" box="[134,1378,990,1010]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC49FAD0FC52" box="[134,1378,990,1010]" gridrow="20" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC49FDAFFC52" box="[134,541,990,1010]" gridcol="0" gridrow="20" pageId="9" pageNumber="126">Face ventrale du pyrène</th>
<td id="769EE12EA01A0028CB7BFC49FA02FC52" box="[575,1456,990,1010]" colspan="2" colspanRight="1" gridcol="1" gridrow="20" pageId="9" pageNumber="126">Planeàconcaveetsanssillonlongitudinal Planeavecsillonlongitudinalbisulqué</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FC61FCC3FB83" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FC61FA14FB83" box="[134,1446,1014,1059]" gridrow="21" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FC61FDAFFB83" box="[134,541,1014,1059]" gridcol="0" gridrow="21" pageId="9" pageNumber="126">Fente de germination préformée</th>
<td id="769EE12EA01A0028CB7BFC61FC5BFB83" box="[575,1001,1014,1059]" gridcol="1" gridrow="21" pageId="9" pageNumber="126">
Présencede 2 fentesenpositionmarginale
<td id="769EE12EA01A0028CABCFC61FA02FB83" box="[1016,1456,1014,1059]" gridcol="2" gridrow="21" pageId="9" pageNumber="126">Aucunefentemaisprésenced’unporeapical</td>
sur la base de la face ventrale
</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C2FBBFFB3AFBF5" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FBBFFA19FBF5" box="[134,1451,1064,1109]" gridrow="22" pageId="9" pageNumber="126">
<th id="769EE12EA01A0028C9C2FBBFFDAFFBF5" box="[134,541,1064,1109]" gridcol="0" gridrow="22" pageId="9" pageNumber="126">Albumen</th>
<td id="769EE12EA01A0028CB7BFBBFFC5BFBF5" box="[575,1001,1064,1109]" gridcol="1" gridrow="22" pageId="9" pageNumber="126">
Non ruminé, souvent avec une petite
<td id="769EE12EA01A0028CABCFBBFFA02FBF5" box="[1016,1456,1064,1109]" gridcol="2" gridrow="22" pageId="9" pageNumber="126">Non ruminé, sans invagination au centre de la invagination au centre de la face ventrale face ventrale</td>
</td>
</tr>
</paragraph>
<th id="769EE12EA01A0028C9C2FBCEFDAFFB27" box="[134,541,1113,1159]" gridcol="0" gridrow="23" pageId="9" pageNumber="126">
<paragraph id="8BC08A10A01AFFDEC9C2FBCEFBABFBCD" box="[134,1049,1113,1133]" pageId="9" pageNumber="126">
<tr id="354F8852A01A0028C9C2FBCEFBABFB27" box="[134,1049,1113,1159]" gridrow="23" pageId="9" pageNumber="126">
Présence dans le tégument d’un pigment
<td id="769EE12EA01A0028CB7BFBCEFC5BFB27" box="[575,1001,1113,1159]" gridcol="1" gridrow="23" pageId="9" pageNumber="126">Non</td>
<td id="769EE12EA01A0028CABCFBCEFA02FB27" box="[1016,1456,1113,1159]" gridcol="2" gridrow="23" pageId="9" pageNumber="126">Oui</td>
</tr>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9D8FBE4FE34FB27" box="[156,390,1139,1159]" pageId="9" pageNumber="126">rouge soluble à l’éthanol</paragraph>
</th>
</table>
<paragraph id="8BC08A10A01AFFDEC9C0FB41FF7EFB50" blockId="9.[132,775,1238,1328]" box="[132,204,1238,1264]" pageId="9" pageNumber="126">
<emphasis id="B90B5602A01AFFDEC9C0FB41FF7EFB50" box="[132,204,1238,1264]" italics="true" pageId="9" pageNumber="126">Graine</emphasis>
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C0FB62FE21FA8F" blockId="9.[132,775,1238,1328]" pageId="9" pageNumber="126">Àalbumen non ruminé, doté d’un pigment tégumentaire rouge soluble à l’éthanol.</paragraph>
</subSubSection>
<subSubSection id="C365D99BA01AFFDEC9C0FAC3FA96F84D" pageId="9" pageNumber="126" type="discussion">
<paragraph id="8BC08A10A01AFFDEC9C0FAC3FF42FACC" blockId="9.[132,775,1364,2030]" box="[132,240,1364,1390]" pageId="9" pageNumber="126">AFFINITÉS</paragraph>
<paragraph id="8BC08A10A01AFFDEC9C0FAE1FEF3F80E" blockId="9.[132,775,1364,2030]" pageId="9" pageNumber="126">
Les affinités de
<taxonomicName id="4C7FF193A01AFFDEC805FAE1FE7AFA2F" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[321,456,1398,1423]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDEC805FAE1FE7AFA2F" box="[321,456,1398,1423]" italics="true" pageId="9" pageNumber="126">P. niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A01AFFDEC895FAE1FD9DFA30" box="[465,559,1398,1424]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
sont clairement à rechercher avec
<taxonomicName id="4C7FF193A01AFFDEC801FA01FE62FA0F" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[325,464,1429,1455]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDEC801FA01FE62FA0F" box="[325,464,1429,1455]" italics="true" pageId="9" pageNumber="126">P. leiophylla</emphasis>
</taxonomicName>
des Îles Salomon et de la Polynésie occidentale (Fidji, Samoa et Tonga), puisque seuls des caractères relativement mineurs, mais notables et récurrents permettent de l’en distinguer (voir la diagnose) et d’autres l’en rapprochent (dont les fruits de couleur rose, violet ou pourpre, le pore apical du pyrène lisse, le pigment tégumentaire soluble à l’éthanol). Plus précisément,
<taxonomicName id="4C7FF193A01AFFDEC9C0F9E2FF4CF92E" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[132,254,1653,1678]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDEC9C0F9E2FF4CF92E" box="[132,254,1653,1678]" italics="true" pageId="9" pageNumber="126">P. niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A01AFFDEC840F9E2FEEAF92F" box="[260,344,1653,1679]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
se distingue de
<taxonomicName id="4C7FF193A01AFFDECB41F9E3FD6CF92E" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[517,734,1652,1678]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECB41F9E3FD6CF92E" box="[517,734,1652,1678]" italics="true" pageId="9" pageNumber="126">Psychotria leiophylla</emphasis>
</taxonomicName>
par ses inflorescences toujours pédonculées avec un pédoncule de plus de
<quantity id="4C8727F5A01AFFDEC9BDF923FEFAF96F" box="[249,328,1716,1743]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="126" unit="cm" value="3.5">3,5 cm</quantity>
, tandis qu’elles sont ramifiées dès la base ou pédonculées sur moins de
<quantity id="4C8727F5A01AFFDEC891F943FDBCF94E" box="[469,526,1748,1774]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="126" unit="cm" value="3.0">3 cm</quantity>
chez
<taxonomicName id="4C7FF193A01AFFDECB14F942FD60F94E" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[592,722,1748,1774]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECB14F942FD60F94E" box="[592,722,1748,1774]" italics="true" pageId="9" pageNumber="126">P. leiophylla</emphasis>
</taxonomicName>
. Les pyrènes de cette dernière sont subellipsoïdaux à subglobuleux et mesurent plus de
<quantity id="4C8727F5A01AFFDEC896F884FD9DF88E" box="[466,559,1811,1838]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="126" unit="mm" value="3.5">3,5 mm</quantity>
de largeur, tandis que ceux de
<taxonomicName id="4C7FF193A01AFFDEC854F8A3FE3FF8ED" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[272,397,1844,1869]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDEC854F8A3FE3FF8ED" box="[272,397,1844,1869]" italics="true" pageId="9" pageNumber="126">P. niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A01AFFDEC8D0F8A3FE58F8EE" box="[404,490,1844,1870]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
sont ovoïdes et mesurent moins de
<quantity id="4C8727F5A01AFFDEC9ABF8C3FE83F8CE" box="[239,305,1876,1902]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="126" unit="mm" value="3.0">3 mm</quantity>
de largeur. Enfin, le pétiole de
<taxonomicName id="4C7FF193A01AFFDECBCCF8C3FCB5F8CD" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[648,775,1875,1901]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECBCCF8C3FCB5F8CD" box="[648,775,1875,1901]" italics="true" pageId="9" pageNumber="126">P. leiophylla</emphasis>
</taxonomicName>
atteint
<quantity id="4C8727F5A01AFFDEC991F8E4FE95F82E" box="[213,295,1907,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="9" pageNumber="126" unit="cm" value="3.0" valueMax="5.0" valueMin="1.0">1-5 cm</quantity>
, tandis que celui de
<taxonomicName id="4C7FF193A01AFFDECB4EF8E3FD34F82D" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[522,646,1908,1933]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDECB4EF8E3FD34F82D" box="[522,646,1908,1933]" italics="true" pageId="9" pageNumber="126">P. niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A01AFFDECBC9F8E3FD51F82E" box="[653,739,1908,1934]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
est inférieur à
<quantity id="4C8727F5A01AFFDEC844F804FE8EF80E" box="[256,316,1939,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="126" unit="cm" value="1.0">1 cm</quantity>
.
</paragraph>
<paragraph id="8BC08A10A01AFFDEC9DFF824FB8DF9AF" blockId="9.[132,775,1364,2030]" lastBlockId="9.[813,1456,1238,2030]" pageId="9" pageNumber="126">
Ces deux espèces appartiennent très probablement au sous-clade PNG du clade IV de
<bibRefCitation id="EFEEF7E1A01AFFDEC8A2F844FD3DF84E" author="BARRABE L." bookContentInfo="367 p." box="[486,655,2003,2030]" journalOrPublisher="Universite de Nouvelle Caledonie, Nouvelle-Caledonie" pageId="9" pageNumber="126" publicationUrl="https://tel.archives-ouvertes.fr/tel-00967108" refId="ref6151" refString="BARRABE L. 2013. - Systematique et Evolution du genre Psychotria (Rubiaceae) en Nouvelle-Caledonie. These de doctorat. Universite de Nouvelle Caledonie, Nouvelle-Caledonie, 367 p. https: // tel. archives-ouvertes. fr / tel- 00967108" title="Systematique et Evolution du genre Psychotria (Rubiaceae) en Nouvelle-Caledonie. These de doctorat" type="book" year="2013">Barrabé (2013)</bibRefCitation>
et
<bibRefCitation id="EFEEF7E1A01AFFDECBF5F844FC72FB50" author="BARRABE L. &amp; MAGGIA L. &amp; PILLON Y. &amp; RIGAULT F. &amp; MOULY A. &amp; DAVIS A. P. &amp; BUERKI S." box="[689,960,1238,2029]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="9" pageNumber="126" pagination="15 - 35" part="71" refId="ref6269" refString="BARRABE L., MAGGIA L., PILLON Y., RIGAULT F., MOULY A., DAVIS A. P. &amp; BUERKI S. 2014. - New Caledonian lineages of Psychotria (Rubiaceae) reveal different evolutionary histories and the largest documented plant radiation for the archipelago. Molecular Phylogenetics and Evolution 71: 15 - 35. https: // doi. org / 10.1016 / j. ympev. 2013.10.020" title="New Caledonian lineages of Psychotria (Rubiaceae) reveal different evolutionary histories and the largest documented plant radiation for the archipelago" type="journal article" year="2014">
Barrabé
<emphasis id="B90B5602A01AFFDECA69FB41FCD5FB50" box="[813,871,1238,1264]" italics="true" pageId="9" pageNumber="126">et al.</emphasis>
(2014)
</bibRefCitation>
présent en Nouvelle-Guinée, Micronésie, Mélanésie et Polynésie occidentale, jusqu’alors absent de la Polynésie orientale.
<taxonomicName id="4C7FF193A01AFFDECD6DFA82FAB3FA8F" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[1065,1281,1301,1327]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A01AFFDECD6DFA82FAB3FA8F" box="[1065,1281,1301,1327]" italics="true" pageId="9" pageNumber="126">Psychotrianiauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A01AFFDECC4CFA81FAEDFA90" box="[1288,1375,1302,1328]" pageId="9" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
résulte ainsi d’un événement de dispersion distinct de ceux de tous les autres
<taxonomicName id="4C7FF193A01AFFDECADEFAC2FBB5FACF" box="[922,1031,1365,1391]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="genus">
<emphasis id="B90B5602A01AFFDECADEFAC2FBB5FACF" box="[922,1031,1365,1391]" italics="true" pageId="9" pageNumber="126">Psychotria</emphasis>
</taxonomicName>
de la région, qui appartiennent apparemment uniquement au clade Pacifique. Le seul
<taxonomicName id="4C7FF193A01AFFDECC00FAE2FA1DFA2F" box="[1348,1455,1397,1423]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="genus">
<emphasis id="B90B5602A01AFFDECC00FAE2FA1DFA2F" box="[1348,1455,1397,1423]" italics="true" pageId="9" pageNumber="126">Eumachia</emphasis>
</taxonomicName>
de Polynésie orientale,
<taxonomicName id="4C7FF193A01AFFDECD75FA01FB04FA0F" authorityName="Barrabe, C. M. Taylor &amp; Razafim." baseAuthorityName="Baill. ex Drake" box="[1073,1206,1429,1455]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="lepiniana">
<emphasis id="B90B5602A01AFFDECD75FA01FB04FA0F" box="[1073,1206,1429,1455]" italics="true" pageId="9" pageNumber="126">E. lepiniana</emphasis>
</taxonomicName>
endémique de l’île de Tahiti, ainsi qu’un taxon proche non décrit propre à l’île de Raiatea, font partie, quant à eux, du sous-clade
<emphasis id="B90B5602A01AFFDECC2DFA42FA02FA4E" box="[1385,1456,1493,1518]" italics="true" pageId="9" pageNumber="126">carnea</emphasis>
de
<bibRefCitation id="EFEEF7E1A01AFFDECA0BFA62FB88F9AF" author="BARRABE L. &amp; BUERKI S. &amp; MOULY A. &amp; DAVIS A. P. &amp; MUNZINGER J. &amp; MAGGIA L." box="[847,1082,1525,1551]" journalOrPublisher="Taxon" pageId="9" pageNumber="126" pagination="1251 - 1268" part="61" refId="ref6194" refString="BARRABE L., BUERKI S., MOULY A., DAVIS A. P., MUNZINGER J. &amp; MAGGIA L. 2012. - Delimitation of the genus Margaritopsis (Rubiaceae) in the Asian, Australasian and Pacific region, based on molecular phylogenetic inference and morphology. Taxon 61 (6): 1251 - 1268. https: // doi. org / 10.1002 / tax. 616007" title="Delimitation of the genus Margaritopsis (Rubiaceae) in the Asian, Australasian and Pacific region, based on molecular phylogenetic inference and morphology" type="journal article" year="2012">
Barrabé
<emphasis id="B90B5602A01AFFDECAE9FA62FC51F9AF" box="[941,995,1525,1551]" italics="true" pageId="9" pageNumber="126">et al.</emphasis>
(2012)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BC08A10A01AFFDECA07F982FA96F84D" blockId="9.[813,1456,1238,2030]" pageId="9" pageNumber="126">
Les affinités avec
<taxonomicName id="4C7FF193A01AFFDECAB9F982FBCBF98E" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[1021,1145,1556,1582]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECAB9F982FBCBF98E" box="[1021,1145,1556,1582]" italics="true" pageId="9" pageNumber="126">P. leiophylla</emphasis>
</taxonomicName>
sont également écologiques, puisque ces deux espèces se cantonnent le plus souvent aux stations littorales, souvent sur calcaire soulevé, aux Salomon, Fidji, Tonga et Tuamotu avec, néanmoins, des stations de
<taxonomicName id="4C7FF193A01AFFDECA69F902FC1CF90E" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[813,942,1684,1710]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECA69F902FC1CF90E" box="[813,942,1684,1710]" italics="true" pageId="9" pageNumber="126">P. leiophylla</emphasis>
</taxonomicName>
plus à l’intérieur des terres aux Samoa, entre 400 et
<quantity id="4C8727F5A01AFFDECA3CF923FB82F96E" box="[888,1072,1716,1743]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="126" unit="m" value="850.0">850 md’altitude</quantity>
(
<bibRefCitation id="EFEEF7E1A01AFFDECD04F923FA8EF96E" author="SMITH A. C. &amp; DARWIN S. P." box="[1088,1340,1716,1742]" editor="SMITH A. C." journalOrPublisher="National Tropical Botanical Garden, Hawai'i" pageId="9" pageNumber="126" pagination="143 - 376" refId="ref7141" refString="SMITH A. C. &amp; DARWIN S. P. 1988. - Rubiaceae, in SMITH A. C. (ed.), Flora Vitiensis Nova 4. National Tropical Botanical Garden, Hawai'i: 143 - 376." title="Rubiaceae" type="book chapter" volumeTitle="Flora Vitiensis Nova 4" year="1988">Smith &amp; Darwin 1988</bibRefCitation>
;
<bibRefCitation id="EFEEF7E1A01AFFDECC0AF923FCD5F94E" author="WHISTLER W. A." bookContentInfo="173 p." journalOrPublisher="Report Prepared for the Tonga Trust Ltd. Funded by Conservation International and Critical Ecosystem Partnership Fund. Isle Botanica, Honolulu" pageId="9" pageNumber="126" refId="ref7323" refString="WHISTLER W. A. 2011. - The Rare Plants of Tonga. Report Prepared for the Tonga Trust Ltd. Funded by Conservation International and Critical Ecosystem Partnership Fund. Isle Botanica, Honolulu, 173 p." title="The Rare Plants of Tonga" type="book" year="2011">Whistler 2011</bibRefCitation>
).
<bibRefCitation id="EFEEF7E1A01AFFDECA3AF943FB86F94E" author="WHISTLER W. A." bookContentInfo="173 p." box="[894,1076,1748,1774]" journalOrPublisher="Report Prepared for the Tonga Trust Ltd. Funded by Conservation International and Critical Ecosystem Partnership Fund. Isle Botanica, Honolulu" pageId="9" pageNumber="126" refId="ref7323" refString="WHISTLER W. A. 2011. - The Rare Plants of Tonga. Report Prepared for the Tonga Trust Ltd. Funded by Conservation International and Critical Ecosystem Partnership Fund. Isle Botanica, Honolulu, 173 p." title="The Rare Plants of Tonga" type="book" year="2011">Whistler (2011)</bibRefCitation>
met en doute la présence de cette dernière aux Samoa, notamment en raison de la couleur rouge des fruits mûrs de ce taxon, rappelant donc plutôt ceux d’
<taxonomicName id="4C7FF193A01AFFDECA3DF8A4FBEEF8EE" baseAuthorityName="Barrabe, C. M. Taylor &amp; Razafim. Gray" box="[889,1116,1843,1870]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="forsteriana">
<emphasis id="B90B5602A01AFFDECA3DF8A4FBEEF8EE" box="[889,1116,1843,1870]" italics="true" pageId="9" pageNumber="126">Eumachia forsteriana</emphasis>
</taxonomicName>
, tandis que ceux de Tonga sont pourpres. Des études semblent encore nécessaires afin de clarifier la situation aux Samoa, mais l’examen des images de certains spécimens d’herbier (notamment
<emphasis id="B90B5602A01AFFDECC66F804FA02F80D" box="[1314,1456,1939,1965]" italics="true" pageId="9" pageNumber="126">Whistler 899</emphasis>
[BISH417668] collectéà Savai’i) nousfait penchersurune présence effective de
<taxonomicName id="4C7FF193A01AFFDECD5DF843FB28F84D" authorityName="Merr. &amp; L. M. Perry de la Melanesie" authorityYear="1946" box="[1049,1178,2003,2029]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="126" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B90B5602A01AFFDECD5DF843FB28F84D" box="[1049,1178,2003,2029]" italics="true" pageId="9" pageNumber="126">P. leiophylla</emphasis>
</taxonomicName>
aux Samoa.
</paragraph>
</subSubSection>
<caption id="DF00DA98A019FFDDC80FF84FFB5BF849" ID-DOI="http://doi.org/10.5281/zenodo.3744699" ID-Zenodo-Dep="3744699" box="[331,1257,2008,2025]" httpUri="https://zenodo.org/record/3744699/files/figure.png" pageId="10" pageNumber="127" startId="10.[331,342,2008,2025]" targetBox="[241,1346,215,1954]" targetPageId="10">
<paragraph id="8BC08A10A019FFDDC80FF84FFB5BF849" blockId="10.[331,1257,2008,2025]" box="[331,1257,2008,2025]" pageId="10" pageNumber="127">
<subScript id="17FB8855A019FFDDC80FF84FFED6F849" attach="left" box="[331,356,2008,2025]" fontSize="5" pageId="10" pageNumber="127">FIG</subScript>
. 5. — Image de l’isotype de
<taxonomicName id="4C7FF193A019FFDDCB17F84FFC1CF849" authority="Butaud &amp; J.Florence" authorityName="Butaud &amp; J. Florence" authorityYear="2020" box="[595,942,2008,2025]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="127" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A019FFDDCB17F84FFD4EF849" box="[595,764,2008,2025]" italics="true" pageId="10" pageNumber="127">Psychotria niauensis</emphasis>
Butaud &amp; J.Florence
</taxonomicName>
,
<taxonomicNameLabel id="A238EB79A019FFDDCAF1F84FFC4BF849" box="[949,1017,2008,2025]" pageId="10" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B90B5602A019FFDDCD46F84FFBDFF849" box="[1026,1133,2008,2025]" italics="true" pageId="10" pageNumber="127">Butaud 1519</emphasis>
[PAP015889]).
</paragraph>
</caption>
<subSubSection id="C365D99BA018FFDCC9C0FF4FFE52F9EE" pageId="11" pageNumber="128" type="biology_ecology">
<paragraph id="8BC08A10A018FFDCC9C0FF4FFDBEFF50" blockId="11.[132,775,216,1614]" box="[132,524,216,242]" pageId="11" pageNumber="128">RÉPARTITION, HABITAT &amp; ÉCOLOGIE</paragraph>
<paragraph id="8BC08A10A018FFDCC9C0FF6EFE8CFED3" blockId="11.[132,775,216,1614]" pageId="11" pageNumber="128">
<taxonomicName id="4C7FF193A018FFDCC9C0FF6EFEF8FEB3" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[132,330,249,275]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A018FFDCC9C0FF6EFEF8FEB3" box="[132,330,249,275]" italics="true" pageId="11" pageNumber="128">Psychotria niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A018FFDCC80BFF6DFE2FFEB4" box="[335,413,250,276]" pageId="11" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
n’est connue en Polynésie française que de l’atoll soulevé de Niau (
<quantity id="4C8727F5A018FFDCC893FE8EFDB6FE94" box="[471,516,281,308]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="128" unit="m" value="6.0">6 m</quantity>
d’altitude maximale) et constitue le seul représentant de son genre dans tout l’archipel des Tuamotu.
</paragraph>
<paragraph id="8BC08A10A018FFDCC9DFFEEEFEE0FDF3" blockId="11.[132,775,216,1614]" pageId="11" pageNumber="128">
Sur l’atoll de Niau,
<taxonomicName id="4C7FF193A018FFDCC8C2FEEFFDD6FE32" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[390,612,376,402]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A018FFDCC8C2FEEFFDD6FE32" box="[390,612,376,402]" italics="true" pageId="11" pageNumber="128">Psychotria niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A018FFDCCB2AFEEEFD7AFE33" box="[622,712,377,403]" pageId="11" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
a été identifiée après des prospections intensives dans des sites qu’ilestpossiblederegrouperentroissous-populations: à l’ouest du village de Tupana dans le nord-est, à Vaiava dans l’est et à Tapae à l’ouest (
<figureCitation id="13449695A018FFDCC8D1FE6FFE66FDB2" box="[405,468,504,530]" captionStart="FIG" captionStartId="12.[494,505,1088,1105]" captionTargetBox="[144,1423,214,1049]" captionTargetId="figure@12.[132,1455,158,1094]" captionTargetPageId="12" captionText="FIG. 6. — Répartition de Psychotria niauensis sp. nov. sur l’atoll de Niau." figureDoi="http://doi.org/10.5281/zenodo.3744701" httpUri="https://zenodo.org/record/3744701/files/figure.png" pageId="11" pageNumber="128">Fig. 6</figureCitation>
). Un total de 78 pieds a été décompté lors de ces prospections, dont la moitié dans la localité de Tupana.
</paragraph>
<paragraph id="8BC08A10A018FFDCC9DFFDCFFD28FB50" blockId="11.[132,775,216,1614]" pageId="11" pageNumber="128">
Il s’agit d’un arbuste de sous-bois des forêts peu perturbées sur corail soulevé dominées par les arbres indigènes
<taxonomicName id="4C7FF193A018FFDCCBFEFDEFFD39FD13" authority="J. Florence" authorityName="J. Florence" authorityYear="1997" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="tuamotuense">
<emphasis id="B90B5602A018FFDCCBFEFDEFFEF7FD11" italics="true" pageId="11" pageNumber="128">Glochidion tuamotuense</emphasis>
J.Florence (Phyllanthaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCCBDCFD0FFF60FD71" class="Magnoliopsida" family="Rubiaceae" genus="Guettarda" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B90B5602A018FFDCCBDCFD0FFF60FD71" italics="true" pageId="11" pageNumber="128">Guettarda speciosa</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C7FF193A018FFDCC844FD20FEC6FD72" authorityName="de Jussieu" authorityYear="1789" box="[256,372,695,722]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
),
<taxonomicName id="4C7FF193A018FFDCC8C3FD20FEE7FD52" authority="Parkinson ex Du Roi" authorityName="Parkinson ex Du Roi" authorityYear="1773" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="tectorius">
<emphasis id="B90B5602A018FFDCC8C3FD20FDFBFD71" box="[391,585,695,721]" italics="true" pageId="11" pageNumber="128">Pandanus tectorius</emphasis>
Parkinson ex Du Roi (Pandanaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCC826FD4FFCB1FD52" authority="R. Br." authorityName="R. Br." box="[354,771,727,754]" class="Magnoliopsida" family="Nyctaginaceae" genus="Pisonia" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="B90B5602A018FFDCC826FD4FFDB7FD51" box="[354,517,727,753]" italics="true" pageId="11" pageNumber="128">Pisonia grandis</emphasis>
R.Br. (Nyctaginaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCC9C0FD60FF63FC92" authority="(Nadeaud) Pierre ex Dubard" authorityName="Pierre ex Dubard" authorityYear="1912" baseAuthorityName="Nadeaud" class="Magnoliopsida" family="Sapotaceae" genus="Planchonella" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="tahitensis">
<emphasis id="B90B5602A018FFDCC9C0FD60FEC6FCB1" box="[132,372,759,785]" italics="true" pageId="11" pageNumber="128">Planchonella tahitensis</emphasis>
(Nadeaud) Pierre ex Dubard (Sapotaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCC999FC80FEB4FCF2" authority="(J. R. Forst. &amp; G. Forst.) G. Forst." authorityName="G. Forst." baseAuthorityName="J. R. Forst. &amp; G. Forst." class="Magnoliopsida" family="Salicaceae" genus="Xylosma" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="suaveolens">
<emphasis id="B90B5602A018FFDCC999FC80FE10FC91" box="[221,418,791,817]" italics="true" pageId="11" pageNumber="128">Xylosma suaveolens</emphasis>
(J.R.Forst. &amp; G.Forst.) G.Forst. (Salicaceae)
</taxonomicName>
, les arbustes indigènes
<taxonomicName id="4C7FF193A018FFDCCB4DFCA0FE47FCD2" authority="(Nadeaud) Radlk." authorityName="Radlk." baseAuthorityName="Nadeaud" class="Magnoliopsida" family="Sapindaceae" genus="Allophylus" kingdom="Animalia" order="Sapindales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="rhomboidalis">
<emphasis id="B90B5602A018FFDCCB4DFCA0FCB5FCF1" box="[521,775,823,849]" italics="true" pageId="11" pageNumber="128">Allophylus rhomboidalis</emphasis>
(Nadeaud) Radlk. (Sapindaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCCB47FCC0FDE7FC32" authority="(G. Forst.) N. Halle &amp; J. Florence" authorityName="N. Halle &amp; J. Florence" baseAuthorityName="G. Forst." class="Magnoliopsida" family="Rubiaceae" genus="Cyclophyllum" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="barbatum">
<emphasis id="B90B5602A018FFDCCB47FCC0FCB5FCD1" box="[515,775,855,881]" italics="true" pageId="11" pageNumber="128">Cyclophyllum barbatum</emphasis>
(G.Forst.) N.Hallé &amp; J.Florence (Rubiaceae)
</taxonomicName>
,
<emphasis id="B90B5602A018FFDCCB1AFCEFFD00FC31" box="[606,690,888,913]" italics="true" pageId="11" pageNumber="128">Ixora sp.</emphasis>
(
<taxonomicName id="4C7FF193A018FFDCCBF9FCE0FF04FC11" authorityName="de Jussieu" authorityYear="1789" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
), «
<emphasis id="B90B5602A018FFDCC99DFC00FE96FC10" box="[217,292,919,944]" italics="true" pageId="11" pageNumber="128">Meryta</emphasis>
sp. nov.» (
<taxonomicName id="4C7FF193A018FFDCC8DBFC00FDA6FC12" authorityName="Durande" authorityYear="1782" box="[415,532,919,946]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
),
<taxonomicName id="4C7FF193A018FFDCCB6CFC00FD55FC11" box="[552,743,919,945]" class="Insecta" family="Cicadellidae" genus="Morinda" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="citrifolia">
<emphasis id="B90B5602A018FFDCCB6CFC00FD55FC11" box="[552,743,919,945]" italics="true" pageId="11" pageNumber="128">Morinda citrifolia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C7FF193A018FFDCC9C8FC20FEB1FC72" authorityName="de Jussieu" authorityYear="1789" box="[140,259,951,978]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
),
<taxonomicName id="4C7FF193A018FFDCC853FC20FF62FC51" authority="Fosberg &amp; Sachet" authorityName="Fosberg &amp; Sachet" authorityYear="1975" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis">
<emphasis id="B90B5602A018FFDCC853FC20FE64FC70" box="[279,470,951,976]" italics="true" pageId="11" pageNumber="128">Myrsine niauensis</emphasis>
Fosberg &amp; Sachet (Primulaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCC99AFC40FEBAFBB1" authority="(G. Forst.) T. Durand ex Drake" authorityName="T. Durand ex Drake" authorityYear="1890" baseAuthorityName="G. Forst." class="Magnoliopsida" family="Rubiaceae" genus="Tarenna" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="sambucina">
<emphasis id="B90B5602A018FFDCC99AFC40FE1DFC50" box="[222,431,982,1008]" italics="true" pageId="11" pageNumber="128">Tarenna sambucina</emphasis>
(G.Forst.) T.Durand ex Drake (Rubiaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCC851FC60FE6CFBB0" authorityName="Moldenke" authorityYear="1977" baseAuthorityName="L." box="[277,478,1014,1040]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="serratifolia">
<emphasis id="B90B5602A018FFDCC851FC60FE6CFBB0" box="[277,478,1014,1040]" italics="true" pageId="11" pageNumber="128">Premna serratifolia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C7FF193A018FFDCCB4AFC61FD34FBB1" box="[526,646,1014,1041]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
), l’herbacée indigène
<taxonomicName id="4C7FF193A018FFDCC9A0FB80FD51FB91" authority="(G. Forst.) A. Diet." authorityName="A. Diet." baseAuthorityName="G. Forst." box="[228,739,1046,1073]" class="Magnoliopsida" family="Piperaceae" genus="Peperomia" kingdom="Plantae" order="Piperales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="pallida">
<emphasis id="B90B5602A018FFDCC9A0FB80FE28FB90" box="[228,410,1046,1072]" italics="true" pageId="11" pageNumber="128">Peperomia pallida</emphasis>
(G.Forst.) A.Diet. (Piperaceae)
</taxonomicName>
, les lianes indigènes
<taxonomicName id="4C7FF193A018FFDCC875FBA0FE83FBD1" authority="(Lam.) Fici" authorityName="Fici" baseAuthorityName="Lam." pageId="11" pageNumber="128" rank="subSpecies" subSpecies="cordifolia">
<emphasis id="B90B5602A018FFDCC875FBA0FE6EFBF0" box="[305,476,1079,1105]" italics="true" pageId="11" pageNumber="128">Capparis spinosa</emphasis>
subsp.
<emphasis id="B90B5602A018FFDCCB6EFBA1FD39FBF0" box="[554,651,1078,1104]" italics="true" pageId="11" pageNumber="128">cordifolia</emphasis>
(Lam.) Fici (Capparidaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCC87DFBC1FEDDFB31" authority="(A. Gray) Razafim. &amp; B. Bremer" authorityName="Razafim. &amp; B. Bremer" baseAuthorityName="A. Gray" class="Magnoliopsida" family="Rubiaceae" genus="Gynochthodes" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="myrtifolia">
<emphasis id="B90B5602A018FFDCC87DFBC1FD9AFBD0" box="[313,552,1110,1136]" italics="true" pageId="11" pageNumber="128">Gynochthodes myrtifolia</emphasis>
(A.Gray) Razafim. &amp; B.Bremer (Rubiaceae)
</taxonomicName>
, lesfougères indigènes
<taxonomicName id="4C7FF193A018FFDCCB1AFBE1FCB5FB30" box="[606,775,1142,1168]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="nidus">
<emphasis id="B90B5602A018FFDCCB1AFBE1FCB5FB30" box="[606,775,1142,1168]" italics="true" pageId="11" pageNumber="128">Asplenium nidus</emphasis>
</taxonomicName>
L. &amp;
<taxonomicName id="4C7FF193A018FFDCC9FFFB01FD31FB11" authority="G. Forst." authorityName="G. Forst." box="[187,643,1174,1201]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="polyodon">
<emphasis id="B90B5602A018FFDCC9FFFB01FE37FB10" box="[187,389,1174,1200]" italics="true" pageId="11" pageNumber="128">Asplenium polyodon</emphasis>
G.Forst. (Aspleniaceae)
</taxonomicName>
,
<taxonomicName id="4C7FF193A018FFDCCBC8FB00FD5EFB71" authority="(Langsd. &amp; Fisch.) S. B. Andrews" authorityName="S. B. Andrews" authorityYear="1990" baseAuthorityName="Langsd. &amp; Fisch." class="Polypodiopsida" family="Polypodiaceae" genus="Microsorum" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="grossum">
<emphasis id="B90B5602A018FFDCCBC8FB00FF66FB6F" italics="true" pageId="11" pageNumber="128">Microsorum grossum</emphasis>
(Langsd. &amp; Fisch.) S.B.Andrews (Polypodiaceae)
</taxonomicName>
et le palmier introduit
<taxonomicName id="4C7FF193A018FFDCC81BFB41FE41FB50" authorityName="Linneo" authorityYear="1753" box="[351,499,1238,1264]" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="nucifera">
<emphasis id="B90B5602A018FFDCC81BFB41FE41FB50" box="[351,499,1238,1264]" italics="true" pageId="11" pageNumber="128">Cocos nucifera</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C7FF193A018FFDCCB5AFB41FD3DFB50" authorityName="Bercht. &amp; J.Presl" authorityYear="1820" box="[542,655,1238,1264]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
).
</paragraph>
<paragraph id="8BC08A10A018FFDCC9DFFB61FE52F9EE" blockId="11.[132,775,216,1614]" pageId="11" pageNumber="128">
Avec ses fruits charnus violets à pourpres à maturité, cet arbuste est très probablement disséminé par des oiseauxfrugivores, au premier rang desquels figure le Ptilope des Tuamotu (
<emphasis id="B90B5602A018FFDCC9C9FAC2FEE8FACF" box="[141,346,1365,1391]" italics="true" pageId="11" pageNumber="128">Ptilinopus coralensis</emphasis>
Peale, 1848), endémique de l’archipel et abondant dans les forêts naturelles de l’île. La pollinisation est très probablement assurée par des insectes, mais aucune observation probante n’a été réalisée. Seule la fourmi d’introduction moderne
<emphasis id="B90B5602A018FFDCC814FA42FDF0FA4F" box="[336,578,1493,1519]" italics="true" pageId="11" pageNumber="128">Monomorium floricola</emphasis>
(Jerdon, 1851) a été observée sur les fleurs de
<taxonomicName id="4C7FF193A018FFDCC8FCFA62FD81F9AE" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[440,563,1525,1551]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A018FFDCC8FCFA62FE75F9AE" box="[440,455,1525,1550]" italics="true" pageId="11" pageNumber="128">P</emphasis>
.
<emphasis id="B90B5602A018FFDCC891FA62FD81F9AE" box="[469,563,1525,1550]" italics="true" pageId="11" pageNumber="128">niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A018FFDCCB7EFA62FD3FF9AF" box="[570,653,1525,1551]" pageId="11" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
en train de prospecter pour du nectar, mais sa très petite taille exclut tout rôle de dissémination du pollen.
</paragraph>
</subSubSection>
<subSubSection id="C365D99BA018FFDCC9C0F9E4FBFDFBD1" pageId="11" pageNumber="128" type="conservation">
<paragraph id="8BC08A10A018FFDCC9C0F9E4FE1AF92B" blockId="11.[132,775,1651,2030]" box="[132,424,1651,1677]" pageId="11" pageNumber="128">STATUT DE CONSERVATION</paragraph>
<paragraph id="8BC08A10A018FFDCC9C0F902FABCFDF2" blockId="11.[132,775,1651,2030]" lastBlockId="11.[813,1456,217,1137]" pageId="11" pageNumber="128">
<taxonomicName id="4C7FF193A018FFDCC9C0F902FF48F90E" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[132,250,1685,1710]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A018FFDCC9C0F902FF48F90E" box="[132,250,1685,1710]" italics="true" pageId="11" pageNumber="128">P. niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A018FFDCC845F902FEE1F90F" box="[257,339,1685,1711]" pageId="11" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
n’est connue que de l’atoll de Niau dont la superficie atteint
<quantity id="4C8727F5A018FFDCC812F923FE07F96F" box="[342,437,1715,1743]" metricMagnitude="4" metricUnit="m" metricValue="2.15" pageId="11" pageNumber="128" unit="km" value="21.5">21,5 km</quantity>
². Elle est présente sous la forme de 3 sous-populations distantes les unes des autres de moins de
<quantity id="4C8727F5A018FFDCC9E2F963FF53F8AE" box="[166,225,1780,1807]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="128" unit="km" value="5.0">5 km</quantity>
(
<figureCitation id="13449695A018FFDCC9B6F963FE83F8AE" box="[242,305,1780,1806]" captionStart="FIG" captionStartId="12.[494,505,1088,1105]" captionTargetBox="[144,1423,214,1049]" captionTargetId="figure@12.[132,1455,158,1094]" captionTargetPageId="12" captionText="FIG. 6. — Répartition de Psychotria niauensis sp. nov. sur l’atoll de Niau." figureDoi="http://doi.org/10.5281/zenodo.3744701" httpUri="https://zenodo.org/record/3744701/files/figure.png" pageId="11" pageNumber="128">Fig. 6</figureCitation>
) et au sein desquelles près de 80 pieds ont étédénombrés; cenombre estprobablementsous-estiméen raison de sa discrétion à l’état stérile. Elle est essentiellement menacée par la dégradation de sonhabitat liéeà l’urbanisation (la sous-population principale située à proximité du village de Tupana étant incluse dans la zone urbaine – UB du Plan Général d’Aménagement [PGA] de la Commune de Fakarava dont fait partie l’atoll de Niau), aux incendies consécutifs à la culture du coprah et débordant sur les forêts naturelles et à l’extension de plantes envahissantes, en particulier l’arbre
<emphasis id="B90B5602A018FFDCCA69FE8EFC6CFE93" box="[813,990,281,307]" italics="true" pageId="11" pageNumber="128">Syzygium cumini</emphasis>
(L.) Skeels (
<taxonomicName id="4C7FF193A018FFDCCD26FE8EFB67FE93" authorityName="de Jussieu" authorityYear="1789" box="[1122,1237,281,307]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) déjà présent sous la forme de pieds-mères isolés et de plantules à proximité de ses stations, et faisant officiellement partie des espèces menaçant la biodiversité en Polynésie française (arrêté 466 CM du
<date id="FFC1ACD0A018FFDCCCD7FEEEFC13FE13" pageId="11" pageNumber="128" value="2018-03-22">22 mars 2018</date>
). La diversité des menaces pesant sur lesdifférentes sous-populationsconduit à retenir deux localités, la première de Tupana étant très menacée par l’urbanisation, car non prise en compte par le PGA, tandis que la seconde regroupant les deux autres sous-populations est intégrée à la zone de site protégé de la zone tampon – NDzt du PGA.
</paragraph>
<paragraph id="8BC08A10A018FFDCCA07FDCFFBC5FC51" blockId="11.[813,1456,217,1137]" pageId="11" pageNumber="128">
L’aire d’occupation de l’espèce, en se fondant sur des carrés d’un kilomètre de côté, atteint
<quantity id="4C8727F5A018FFDCCDCDFDEFFB75FD33" box="[1161,1223,631,659]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="128" unit="km" value="6.0">6 km</quantity>
², tandis que son aire d’occurrence est inférieure à
<quantity id="4C8727F5A018FFDCCD2CFD00FB07FD13" box="[1128,1205,663,691]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="128" unit="km" value="50.0">50 km</quantity>
². Cette aire de répartition n’est pas considérée comme sévèrement fragmentée, en raison de la proximité des trois sous-populations ou des deux localitéset de la présence d’un oiseau frugivore disséminateur des semences. En raison des incendies récurrents, de l’agrandissement progressif du village de Tupana et du développement de l’envahissant
<emphasis id="B90B5602A018FFDCCD58FCC0FB7CFCD1" box="[1052,1230,855,881]" italics="true" pageId="11" pageNumber="128">Syzygium cumini</emphasis>
, un déclin continu a été estimé pour l’étendue et la qualité de son habitat. Ainsi, en utilisant le critère Bde la liste rouge de l’UICN (
<bibRefCitation id="EFEEF7E1A018FFDCCC23FC00FCD6FC71" author="IUCN" journalOrPublisher="IUCN, Cambridge" pageId="11" pageNumber="128" publicationUrl="http://www.iucnredlist.org" refId="ref6681" refString="IUCN 2019. - The IUCN Red List of Threatened Species. Version 2018 - 2. IUCN, Cambridge. http: // www. iucnredlist. org" title="The IUCN Red List of Threatened Species. Version 2018 - 2" type="url" year="2019">IUCN 2019</bibRefCitation>
),
<taxonomicName id="4C7FF193A018FFDCCA3CFC20FC58FC70" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[888,1002,951,976]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A018FFDCCA3CFC20FC58FC70" box="[888,1002,951,976]" italics="true" pageId="11" pageNumber="128">P.niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A018FFDCCAB4FC20FB8DFC71" box="[1008,1087,951,977]" pageId="11" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
peut être évaluée comme en danger d’extinction EN B1, B2ab(iii).
</paragraph>
<paragraph id="8BC08A10A018FFDCCA07FC60FBFDFBD1" blockId="11.[813,1456,217,1137]" pageId="11" pageNumber="128">
En utilisant le critère Det avec 80 individus recensés (dont des juvéniles) et moins de 250 individus matures estimés,
<taxonomicName id="4C7FF193A018FFDCCA69FBA0FC16FBF0" authority="Florence &amp; Butaud, 2020" authorityName="Florence &amp; Butaud" authorityYear="2020" box="[813,932,1079,1104]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="niauensis" status="sp. nov.">
<emphasis id="B90B5602A018FFDCCA69FBA0FC16FBF0" box="[813,932,1079,1104]" italics="true" pageId="11" pageNumber="128">P. niauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238EB79A018FFDCCAE9FBA0FBB3FBF1" box="[941,1025,1079,1105]" pageId="11" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
peut également être évaluée comme en danger d’extinction EN D.
</paragraph>
</subSubSection>
</treatment>
</document>
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