423 lines
53 KiB
XML
423 lines
53 KiB
XML
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<mods:title id="EF94D6A7C6364498F4192BB719A58DF0">Rhagasostoma (Bryozoa) from the Late Cretaceous of Eurasia: taxonomic revision, stratigraphy and palaeobiogeography</mods:title>
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<mods:namePart id="0635D289BA5A9E8FCA74EE43EF96957C">Koromyslova, Anna V.</mods:namePart>
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<mods:affiliation id="A078F8EF74051D6A6972F9C077C647BC">Borissiak Paleontological Institute of the Russian Academy of Science, Profsoyuznaya st. 123, Moscow, 117997, Russian Federation. Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Senckenberg Forschungsinstitute und Naturmuseun, Sektion Marine Evertebraten III (Bryozoologie), Senckenberganlage 25, 60325 Frankfurt am Main, Germany. The Sedgwick Museum of Earth Sciences, Department of Earth Sciences, University of Cambridge, UK.</mods:affiliation>
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<mods:namePart id="E8F0AF3C79C5828ED045B71783B4AC47">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="CEFAE86A2F3CA1E836B6A65D295CDBFC">Martha, Silviu O.</mods:namePart>
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<mods:namePart id="007387E3001A34750FA6017FB04645BF">Riley, Matthew</mods:namePart>
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<treatment id="03D587D7966DFFB0F36BF9C4FBF9FBD0" ID-DOI="http://doi.org/10.5281/zenodo.3849962" ID-GBIF-Taxon="164165738" ID-Zenodo-Dep="3849962" LSID="urn:lsid:zoobank.org:act:23F08F8D-29DB-4470-B8FA-B176256858F8" httpUri="http://treatment.plazi.org/id/03D587D7966DFFB0F36BF9C4FBF9FBD0" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
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<paragraph id="8BC336C1966DFFB5F36BF9C4FC13F9B5" blockId="27.[373,1214,1559,1656]" box="[576,1012,1559,1586]" pageId="27" pageNumber="28">
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<taxonomicName id="4C7C4D42966DFFB5F36BF9C4FC75F9B5" ID-CoL="86RGM" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[576,914,1559,1586]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="27" pageNumber="28" phylum="Bryozoa" rank="species" species="operculatum" status="sp. nov.">
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<emphasis id="B908EAD3966DFFB5F36BF9C4FC75F9B5" bold="true" box="[576,914,1559,1586]" italics="true" pageId="27" pageNumber="28">Rhagasostoma operculatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B57A8966DFFB5F2B2F9CBFC13F9B5" box="[921,1012,1560,1586]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BC336C1966DFFB5F395F98DFC91F9FF" blockId="27.[373,1214,1559,1656]" box="[702,886,1630,1656]" pageId="27" pageNumber="28">
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<figureCitation id="13472A44966DFFB5F395F98DFCF5F9FF" box="[702,786,1630,1656]" captionStart="Fig" captionStartId="29.[189,232,1852,1878]" captionTargetBox="[189,1398,738,1810]" captionTargetId="figure@29.[189,1399,738,1810]" captionTargetPageId="29" captionText="Fig. 11. Rhagasostoma operculatum sp. nov. A–C. Holotype, TsNIGR Museum 27/12582, western Kopetdag, Turkmenistan, boundary between the early and late Campanian (Cibicidoides temirensis/ Bolivinoides decoratus decoratus Zone, LS13). A. Overview of erect bifoliate colony. B. Autozooids with putative opercula and avicularia. C. Autozooids with putative opercula and avicularium. Scale bars: A = 1 mm; B = 500 µm; C = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3830867" httpUri="https://zenodo.org/record/3830867/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
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,
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<tableCitation id="C6FE037A966DFFB5F235F98DFC91F9FF" box="[798,886,1630,1656]" captionStart="Table 6" captionStartId="28.[189,258,268,294]" captionText="Table 6. Summary of measurements of Rhagasostoma operculatum sp. nov. For each parameter the range is given with the number of measurements in brackets. The arithmetic mean is given ± standard deviation. All measurements in µm." pageId="27" pageNumber="28">Table 6</tableCitation>
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</paragraph>
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<subSubSection id="C366654A966DFFB5F196F94CFBB9F899" pageId="27" pageNumber="28" type="reference_group">
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<paragraph id="8BC336C1966DFFB5F196F94CFD1AF95A" blockId="27.[189,1398,1695,1757]" pageId="27" pageNumber="28">
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<treatmentCitationGroup id="AB6C11EF966DFFB5F196F94CFD1AF95A" pageId="27" pageNumber="28">
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<taxonomicName id="4C7C4D42966DFFB5F196F94CFD71F93D" ID-CoL="8GR5Y" authority="(Lonsdale, 1850)" baseAuthorityName="Lonsdale" baseAuthorityYear="1850" box="[189,662,1695,1722]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="27" pageNumber="28" phylum="Bryozoa" rank="species" species="inelegans">
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<emphasis id="B908EAD3966DFFB5F196F94CFE2FF93E" box="[189,456,1695,1721]" italics="true" pageId="27" pageNumber="28">Onychocella inelegans</emphasis>
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(
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<bibRefCitation id="EFED4B30966DFFB5F0FCF94CFD69F93D" author="Lonsdale W." box="[471,654,1695,1722]" pageId="27" pageNumber="28" pagination="237 - 324" refId="ref31211" refString="Lonsdale W. 1850. Description of the fossils of the Chalk Formation. Note on the corals. In: Dixon F. (ed) The Geology and Fossils of the Tertiary and Cretaceous Formations of Sussex: 237 - 324. Longman, Brighton." type="book chapter" year="1850">Lonsdale, 1850</bibRefCitation>
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)
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||
</taxonomicName>
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||
–
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||
<treatmentCitation id="0ADD10D0966DFFB5F399F94CFC5FF93D" author="Favorskaya T. A." box="[690,952,1695,1722]" page="125" pageId="27" pageNumber="28" year="1992">
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<bibRefCitation id="EFED4B30966DFFB5F399F94CFC5FF93D" author="Favorskaya T. A." box="[690,952,1695,1722]" pageId="27" pageNumber="28" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. & Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya 1992: 125</bibRefCitation>
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</treatmentCitation>
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||
, pl. 64, fig. 7. —
|
||
<bibRefCitation id="EFED4B30966DFFB5F5AAF94CFAA9F93E" author="Favorskaya T. A." box="[1153,1358,1695,1721]" pageId="27" pageNumber="28" refId="ref30528" refString="Favorskaya T. A. 1996. Pрakticiсkoi рukovodсtvo po мakрoфauнi Рoссii i сopрidiльныk tiррitoрiй [A Practical Handbook on the Macrofauna of Russia and Adjacent Territories: Bryozoans of the Mesocenozoic]. Oleynikov A. N. (ed). Ministerstvo prirodnich resursov RF i Vserossiyskiy Nauchno-issledovatelskiy Geologicheskiy Institut, St Petersburg." type="book" year="1996">Favorskaya 1996</bibRefCitation>
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: pl. 3, fig. 4. —
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<bibRefCitation id="EFED4B30966DFFB5F05CF910FD8BF95A" author="Koromyslova A. V." box="[375,620,1731,1757]" pageId="27" pageNumber="28" pagination="587 - 593" refId="ref30881" refString="Koromyslova A. V. 2014 b. The earliest calcified opercula of bryozoans of the order Cheilostomata. Paleontological Journal 48 (6): 587 - 593. https: // doi. org / 10.1134 / S 0031030114060082" type="journal article" year="2014">Koromyslova 2014b</bibRefCitation>
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: pl. 3, fig. 1.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC336C1966DFFB5F196F8D6FBB9F899" blockId="27.[189,1118,1796,1823]" box="[189,1118,1796,1823]" pageId="27" pageNumber="28">
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<treatmentCitationGroup id="AB6C11EF966DFFB5F196F8D6FBB9F899" box="[189,1118,1796,1823]" pageId="27" pageNumber="28">
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non
|
||
<taxonomicName id="4C7C4D42966DFFB5F1DBF8D7FE1BF899" ID-CoL="8GR5Y" baseAuthorityName="Lonsdale" baseAuthorityYear="1850" box="[240,508,1796,1822]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="27" pageNumber="28" phylum="Bryozoa" rank="species" species="inelegans">
|
||
<emphasis id="B908EAD3966DFFB5F1DBF8D7FE1BF899" box="[240,508,1796,1822]" italics="true" pageId="27" pageNumber="28">Onychocella inelegans</emphasis>
|
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</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10D0966DFFB5F331F8D7FCC7F898" author="Favorskaya T. A." box="[538,800,1796,1823]" page="125" pageId="27" pageNumber="28" year="1992">
|
||
<bibRefCitation id="EFED4B30966DFFB5F331F8D7FCC7F898" author="Favorskaya T. A." box="[538,800,1796,1823]" pageId="27" pageNumber="28" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. & Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya 1992: 125</bibRefCitation>
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||
</treatmentCitation>
|
||
, pl. 64, fig. 6, pl. 65, fig. 1.
|
||
</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366654A966DFFB2F196F896FE3FFA65" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="diagnosis">
|
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<paragraph id="8BC336C1966DFFB5F196F896FED1F8D8" blockId="27.[189,310,1861,1887]" box="[189,310,1861,1887]" pageId="27" pageNumber="28">
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<heading id="D08B81AD966DFFB5F196F896FED1F8D8" bold="true" box="[189,310,1861,1887]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B908EAD3966DFFB5F196F896FED1F8D8" bold="true" box="[189,310,1861,1887]" pageId="27" pageNumber="28">Diagnosis</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BC336C1966DFFB2F196F8A7FE3FFA65" blockId="27.[189,1399,1908,2005]" lastBlockId="28.[189,1399,1410,1506]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">Colony erect, bifoliate. Autozooids subrectangular, zooidal boundaries raised; gymnocyst lacking; cryptocyst extensive; opesia terminal, semielliptical; putative calcified opercula rounded trapezoidal. Avicularia interzooidal; rostrum channeled with rounded tip, between conical in outline and spade- shaped, asymmetrical; proximal part rounded, shorter and wider than rostrum; opesia egg-shaped with thin articular ridges bearing two short teeth proximally and a short opesiular indentation between the teeth; opesiules lacking.</paragraph>
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||
</subSubSection>
|
||
<caption id="DF036649966AFFB2F196FEDFFD87FEEB" pageId="28" pageNumber="29">
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<paragraph id="8BC336C1966AFFB2F196FEDFFD87FEEB" blockId="28.[189,1399,267,364]" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F196FEDFFEC4FEA2" bold="true" box="[189,291,267,294]" pageId="28" pageNumber="29">Table 6.</emphasis>
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||
Summary of measurements of
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<taxonomicName id="4C7C4D42966AFFB2F389FED8FC0BFEA2" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[674,1004,267,293]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="28" pageNumber="29" phylum="Bryozoa" rank="species" species="operculatum" status="sp. nov.">
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||
<emphasis id="B908EAD3966AFFB2F389FED8FC0BFEA2" box="[674,1004,267,293]" italics="true" pageId="28" pageNumber="29">Rhagasostoma operculatum</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A23B57A8966AFFB2F2DCFEDFFBB2FEA1" box="[1015,1109,268,294]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
For each parameter the range is given with the number of measurements in brackets. The arithmetic mean is given ± standard deviation. All measurements in µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC336C1966AFFB2F34AFE7DFC3CFAB3" pageId="28" pageNumber="29">
|
||
<table id="F97CC461966A0051F308FE7DFBFFFAB3" box="[547,1048,430,1332]" gridcols="3" gridrows="7" pageId="28" pageNumber="29">
|
||
<tr id="354C3483966A0051F308FE7DFBFFFDB9" box="[547,1048,430,574]" gridrow="0" pageId="28" pageNumber="29">
|
||
<th id="769D5DFF966A0051F308FE7DFD34FDB9" box="[547,723,430,574]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F34AFE7DFD48FDB9" bold="true" pageId="28" pageNumber="29">Basin Stratigraphy Locality</emphasis>
|
||
</th>
|
||
<th id="769D5DFF966A0051F23BFE7DFBFFFDB9" box="[784,1048,430,574]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F259FE63FC14FDBB" bold="true" pageId="28" pageNumber="29">Tethys Boundary early to middle Campanian Western Kopetdag</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="354C3483966A0051F308FD8AFBFFFCE9" box="[547,1048,601,878]" gridrow="1" pageId="28" pageNumber="29">
|
||
<th id="769D5DFF966A0051F308FD8AFDA5FCE9" box="[547,578,601,878]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F308FD41FDA5FCAC" bold="true" box="[547,578,658,811]" pageId="28" pageNumber="29">Autozooids</emphasis>
|
||
</th>
|
||
<td id="769D5DFF966A0051F356FD8AFD34FCE9" box="[637,723,601,878]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F3BBFDB5FD34FCE6" bold="true" pageId="28" pageNumber="29">AzL AzW OperL OperW</emphasis>
|
||
</td>
|
||
<td id="769D5DFF966A0051F23BFD8AFBFFFCE9" box="[784,1048,601,878]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">550–690 (8) 602.50 ± 49.50 300–410 (8) 331.25 ± 39.44 130–180 (8) 155.00 ± 17.73 150–200 (8) 180.00 ± 20.00</td>
|
||
</tr>
|
||
<tr id="354C3483966A0051F308FC54FBFFFC3D" box="[547,1048,903,954]" gridrow="2" pageId="28" pageNumber="29" rowspan-0="1">
|
||
<td id="769D5DFF966A0051F356FC54FD34FC3D" box="[637,723,903,954]" gridcol="1" gridrow="2" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F3BBFC47FD59FC2B" bold="true" box="[656,702,916,940]" pageId="28" pageNumber="29">AvL</emphasis>
|
||
</td>
|
||
<td id="769D5DFF966A0051F23BFC54FBFFFC3D" box="[784,1048,903,954]" gridcol="2" gridrow="2" pageId="28" pageNumber="29">680–870 (8) 771.25 ± 59.63</td>
|
||
</tr>
|
||
<tr id="354C3483966A0051F308FC00FBFFFB82" box="[547,1048,979,1029]" gridrow="3" pageId="28" pageNumber="29" rowspan-0="1">
|
||
<td id="769D5DFF966A0051F356FC00FD34FB82" box="[637,723,979,1029]" gridcol="1" gridrow="3" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F3BDFC33FD5CFC7F" bold="true" box="[662,699,992,1016]" pageId="28" pageNumber="29">RL</emphasis>
|
||
</td>
|
||
<td id="769D5DFF966A0051F23BFC00FBFFFB82" box="[784,1048,979,1029]" gridcol="2" gridrow="3" pageId="28" pageNumber="29">400–520 (8) 461.25 ± 42.91</td>
|
||
</tr>
|
||
<tr id="354C3483966A0051F308FBC1FBFFFB27" box="[547,1048,1042,1184]" gridrow="4" pageId="28" pageNumber="29">
|
||
<th id="769D5DFF966A0051F308FBC1FDA5FB27" box="[547,578,1042,1184]" gridcol="0" gridrow="4" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F308FBC1FDA5FB27" bold="true" box="[547,578,1042,1184]" pageId="28" pageNumber="29">Avicularia</emphasis>
|
||
</th>
|
||
<td id="769D5DFF966A0051F356FBC1FD34FB27" box="[637,723,1042,1184]" gridcol="1" gridrow="4" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F3BAFBFFFD58FB17" bold="true" box="[657,703,1068,1168]" pageId="28" pageNumber="29">RW PrL</emphasis>
|
||
</td>
|
||
<td id="769D5DFF966A0051F23BFBC1FBFFFB27" box="[784,1048,1042,1184]" gridcol="2" gridrow="4" pageId="28" pageNumber="29">100–130 (8) 115.00 ± 11.95 270–380 (8) 318.75 ± 35.23</td>
|
||
</tr>
|
||
<tr id="354C3483966A0051F308FB66FBFFFB6F" box="[547,1048,1205,1256]" gridrow="5" pageId="28" pageNumber="29" rowspan-0="1">
|
||
<td id="769D5DFF966A0051F356FB66FD34FB6F" box="[637,723,1205,1256]" gridcol="1" gridrow="5" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F3A6FB10FD25FB5C" bold="true" box="[653,706,1219,1243]" pageId="28" pageNumber="29">OpL</emphasis>
|
||
</td>
|
||
<td id="769D5DFF966A0051F23BFB66FBFFFB6F" box="[784,1048,1205,1256]" gridcol="2" gridrow="5" pageId="28" pageNumber="29">120–150 (6) 140.00 ± 12.65</td>
|
||
</tr>
|
||
<tr id="354C3483966A0051F308FAD2FBFFFAB3" box="[547,1048,1281,1332]" gridrow="6" pageId="28" pageNumber="29" rowspan-0="1">
|
||
<td id="769D5DFF966A0051F356FAD2FD34FAB3" box="[637,723,1281,1332]" gridcol="1" gridrow="6" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F3A3FADDFD21FAA1" bold="true" box="[648,710,1294,1318]" pageId="28" pageNumber="29">OpW</emphasis>
|
||
</td>
|
||
<td id="769D5DFF966A0051F23BFAD2FBFFFAB3" box="[784,1048,1281,1332]" gridcol="2" gridrow="6" pageId="28" pageNumber="29">100–140 (6) 118.33 ± 11.69</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C366654A966AFFB2F196F9D9FDEBF9F1" pageId="28" pageNumber="29" type="etymology">
|
||
<paragraph id="8BC336C1966AFFB2F196F9D9FEA3F9A3" blockId="28.[189,324,1546,1572]" box="[189,324,1546,1572]" pageId="28" pageNumber="29">
|
||
<heading id="D08B81AD966AFFB2F196F9D9FEA3F9A3" bold="true" box="[189,324,1546,1572]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
|
||
<emphasis id="B908EAD3966AFFB2F196F9D9FEA3F9A3" bold="true" box="[189,324,1546,1572]" pageId="28" pageNumber="29">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336C1966AFFB2F196F9EBFDEBF9F1" blockId="28.[189,1399,1592,1654]" pageId="28" pageNumber="29">
|
||
The species is named from the Latin ‘
|
||
<emphasis id="B908EAD3966AFFB2F3A5F9EBFCEDF9D5" box="[654,778,1592,1618]" italics="true" pageId="28" pageNumber="29">operculum</emphasis>
|
||
’ because the opesiae of autozooids are presumably closed by calcified opercula.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366654A966AFFB2F196F94EFB4DF865" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="8BC336C1966AFFB2F196F94EFE6BF930" blockId="28.[189,396,1693,1719]" box="[189,396,1693,1719]" pageId="28" pageNumber="29">
|
||
<heading id="D08B81AD966AFFB2F196F94EFE6BF930" bold="true" box="[189,396,1693,1719]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
|
||
<emphasis id="B908EAD3966AFFB2F196F94EFE6BF930" bold="true" box="[189,396,1693,1719]" pageId="28" pageNumber="29">Material studied</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B143C9C966AFFB2F1C1F91FFCB1F8F4" ID-GBIF-Occurrence="2620322338" collectionCode="LS, TsNIGR" country="Turkmenistan" location="Kara-Kala" municipality="Western Kopetdag" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BC336C1966AFFB2F1C1F91FFEBBF961" blockId="28.[189,1399,1740,1907]" box="[234,348,1740,1766]" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F1C1F91FFEBBF961" bold="true" box="[234,348,1740,1766]" pageId="28" pageNumber="29">
|
||
<typeStatus id="54C78863966AFFB2F1C1F91FFEBBF961" box="[234,348,1740,1766]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC336C1966AFFB2F196F93CFCB1F8F4" blockId="28.[189,1399,1740,1907]" pageId="28" pageNumber="29">
|
||
<collectingCountry id="F36B7651966AFFB2F196F93CFE45F88E" box="[189,418,1775,1801]" name="Turkmenistan" pageId="28" pageNumber="29">TURKMENISTAN</collectingCountry>
|
||
•
|
||
<collectingMunicipality id="6BA7ACBB966AFFB2F0F9F93CFD5EF88E" box="[466,697,1775,1801]" pageId="28" pageNumber="29">Western Kopetdag</collectingMunicipality>
|
||
,
|
||
<location id="8EA3601A966AFFB2F3FAF93CFCA8F88E" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F3FAF93CFCA8F88E" box="[721,847,1775,1801]" country="Turkmenistan" municipality="Western Kopetdag" name="Kara-Kala" pageId="28" pageNumber="29">Kara-Kala</location>
|
||
; boundary layers of the early
|
||
<location id="8EA3601A966AFFB2F5DAF93CFA90F88E" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F5DAF93CFA90F88E" box="[1265,1399,1775,1801]" country="Turkmenistan" municipality="Western Kopetdag" name="Campanian" pageId="28" pageNumber="29">Campanian</location>
|
||
(
|
||
<location id="8EA3601A966AFFB2F1ECF8C1FC0FF8AA" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F1ECF8C1FC0FF8AA" box="[199,1000,1810,1837]" country="Turkmenistan" municipality="Western Kopetdag" name="Cibicidoides temirensis / Bolivinoides decoratus decoratus Zone" pageId="28" pageNumber="29">
|
||
<taxonomicName id="4C7C4D42966AFFB2F1ECF8C1FE00F8AB" box="[199,487,1810,1836]" class="Globothalamea" family="Anomalinidae" genus="Cibicidoides" kingdom="Chromista" order="Rotaliida" pageId="28" pageNumber="29" phylum="Foraminifera" rank="species" species="temirensis">
|
||
<emphasis id="B908EAD3966AFFB2F1ECF8C1FE00F8AB" box="[199,487,1810,1836]" italics="true" pageId="28" pageNumber="29">Cibicidoides temirensis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7C4D42966AFFB2F0C4F8C1FC0FF8AA" authority="decoratus Zone" authorityName="Zone" box="[495,1000,1810,1837]" class="Globothalamea" family="Bolivinoididae" genus="Bolivinoides" kingdom="Chromista" order="Rotaliida" pageId="28" pageNumber="29" phylum="Foraminifera" rank="subSpecies" species="decoratus" subSpecies="decoratus">
|
||
<emphasis id="B908EAD3966AFFB2F0C4F8C1FC74F8AB" box="[495,915,1810,1836]" italics="true" pageId="28" pageNumber="29">Bolivinoides decoratus decoratus</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
</location>
|
||
;
|
||
<collectionCode id="ED6DAE04966AFFB2F52AF8C1FBC1F8AB" box="[1025,1062,1810,1836]" collectionName="United Kingdom, London, Linnean Society" pageId="28" pageNumber="29">LS</collectionCode>
|
||
13) to middle
|
||
<location id="8EA3601A966AFFB2F5DAF8C1FA90F8AB" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F5DAF8C1FA90F8AB" box="[1265,1399,1810,1836]" country="Turkmenistan" municipality="Western Kopetdag" name="Campanian" pageId="28" pageNumber="29">Campanian</location>
|
||
(
|
||
<taxonomicName id="4C7C4D42966AFFB2F1ECF8E6FDB5F8D7" authority="Zone" authorityName="Zone" box="[199,594,1845,1872]" class="Globothalamea" family="Cancrisidae" genus="Brotzenella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Rotaliida" pageId="28" pageNumber="29" phylum="Foraminifera" rank="species" species="monterelensis">
|
||
<location id="8EA3601A966AFFB2F1ECF8E6FDB5F8D7" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F1ECF8E6FDB5F8D7" box="[199,594,1845,1872]" country="Turkmenistan" municipality="Western Kopetdag" name="Brotzenella monterelensis Zone" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EAD3966AFFB2F1ECF8E6FDE6F8C8" box="[199,513,1845,1871]" italics="true" pageId="28" pageNumber="29">Brotzenella monterelensis</emphasis>
|
||
Zone
|
||
</location>
|
||
</taxonomicName>
|
||
;
|
||
<collectionCode id="ED6DAE04966AFFB2F343F8E6FD6BF8C8" box="[616,652,1845,1871]" collectionName="United Kingdom, London, Linnean Society" pageId="28" pageNumber="29">LS</collectionCode>
|
||
14) of the western
|
||
<location id="8EA3601A966AFFB2F2A4F8E6FBE2F8C8" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F2A4F8E6FBE2F8C8" box="[911,1029,1845,1871]" country="Turkmenistan" municipality="Western Kopetdag" name="Kopetdag" pageId="28" pageNumber="29">Kopetdag</location>
|
||
;
|
||
<collectionCode id="ED6DAE04966AFFB2F530F8E6FB99F8C8" box="[1051,1150,1845,1871]" pageId="28" pageNumber="29">TsNIGR</collectionCode>
|
||
<location id="8EA3601A966AFFB2F5A4F8E5FB11F8D7" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F5A4F8E5FB11F8D7" box="[1167,1270,1846,1872]" country="Turkmenistan" municipality="Western Kopetdag" name="Museum" pageId="28" pageNumber="29">Museum</location>
|
||
27/12582 (
|
||
<figureCitation id="13472A44966AFFB2F1EDF88BFEF0F8F4" box="[198,279,1880,1907]" captionStart="Fig" captionStartId="29.[189,232,1852,1878]" captionTargetBox="[189,1398,738,1810]" captionTargetId="figure@29.[189,1399,738,1810]" captionTargetPageId="29" captionText="Fig. 11. Rhagasostoma operculatum sp. nov. A–C. Holotype, TsNIGR Museum 27/12582, western Kopetdag, Turkmenistan, boundary between the early and late Campanian (Cibicidoides temirensis/ Bolivinoides decoratus decoratus Zone, LS13). A. Overview of erect bifoliate colony. B. Autozooids with putative opercula and avicularia. C. Autozooids with putative opercula and avicularium. Scale bars: A = 1 mm; B = 500 µm; C = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3830867" httpUri="https://zenodo.org/record/3830867/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
|
||
A–C; figured by
|
||
<bibRefCitation id="EFED4B30966AFFB2F0F4F88BFD4BF8F4" author="Favorskaya T. A." box="[479,684,1880,1907]" pageId="28" pageNumber="29" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. & Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">
|
||
<location id="8EA3601A966AFFB2F0F4F88BFD81F8F5" LSID="urn:lsid:plazi:treatment:03D587D7966DFFB0F36BF9C4FBF9FBD0:8EA3601A966AFFB2F0F4F88BFD81F8F5" box="[479,614,1880,1906]" country="Turkmenistan" municipality="Western Kopetdag" name="Favorskaya" pageId="28" pageNumber="29">Favorskaya</location>
|
||
1992
|
||
</bibRefCitation>
|
||
: pl. 64, fig. 7).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BC336C1966AFFB2F196F849FE1CF833" blockId="28.[189,507,1946,1972]" box="[189,507,1946,1972]" pageId="28" pageNumber="29">
|
||
<heading id="D08B81AD966AFFB2F196F849FE1CF833" bold="true" box="[189,507,1946,1972]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
|
||
<emphasis id="B908EAD3966AFFB2F196F849FE1CF833" bold="true" box="[189,507,1946,1972]" pageId="28" pageNumber="29">Type locality and horizon</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336C1966AFFB2F196F81BFB4DF865" blockId="28.[189,1194,1992,2019]" box="[189,1194,1992,2019]" pageId="28" pageNumber="29">
|
||
<collectingCountry id="F36B7651966AFFB2F196F81BFE85F865" box="[189,354,1992,2018]" name="Turkmenistan" pageId="28" pageNumber="29">Turkmenistan</collectingCountry>
|
||
, Western Kopetdag; boundary layers of the early to middle Campanian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366654A966BFFB0F196FED8FB04FECF" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8BC336C1966BFFB3F196FED8FEA9FEA2" blockId="29.[189,334,267,293]" box="[189,334,267,293]" pageId="29" pageNumber="30">
|
||
<heading id="D08B81AD966BFFB3F196FED8FEA9FEA2" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis id="B908EAD3966BFFB3F196FED8FEA9FEA2" bold="true" box="[189,334,267,293]" pageId="29" pageNumber="30">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336C1966BFFB0F196FEEAFB04FECF" blockId="29.[189,1399,313,692]" lastBlockId="30.[189,1399,267,329]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
Colony erect, flattened, bifoliate; fragments 6.0 mm long by
|
||
<quantity id="4C849B24966BFFB3F2AFFEE9FC2FFED3" box="[900,968,314,340]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="2.0">2 mm</quantity>
|
||
wide. Ancestrula and early astogeny not observed. Autozooids variable in shape, often broad and subrectangular with rounded distal ends; zooidal boundaries raised. Gymnocyst lacking. Cryptocyst extensive, finely pustulose, slightly depressed centrally, without peripheral caverns. Opesia terminal, semielliptical, presumably closed by calcified opercula. Putative opercula rounded trapezoidal and slightly concave, tubercular externally (
|
||
<figureCitation id="13472A44966BFFB3F1EDFE3AFED6FD83" box="[198,305,489,516]" captionStart="Fig" captionStartId="29.[189,232,1852,1878]" captionTargetBox="[189,1398,738,1810]" captionTargetId="figure@29.[189,1399,738,1810]" captionTargetPageId="29" captionText="Fig. 11. Rhagasostoma operculatum sp. nov. A–C. Holotype, TsNIGR Museum 27/12582, western Kopetdag, Turkmenistan, boundary between the early and late Campanian (Cibicidoides temirensis/ Bolivinoides decoratus decoratus Zone, LS13). A. Overview of erect bifoliate colony. B. Autozooids with putative opercula and avicularia. C. Autozooids with putative opercula and avicularium. Scale bars: A = 1 mm; B = 500 µm; C = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3830867" httpUri="https://zenodo.org/record/3830867/files/figure.png" pageId="29" pageNumber="30">Fig. 11C</figureCitation>
|
||
), proximal edge slightly arched and with ‘ears’ at the ends, which are
|
||
<quantity id="4C849B24966BFFB3F5ABFE3AFACAFD83" box="[1152,1325,489,516]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="mm" value="0.04" valueMax="0.05" valueMin="0.03">0.03–0.05 mm</quantity>
|
||
wide, ‘ears’ projecting
|
||
<quantity id="4C849B24966BFFB3F0AFFDDEFDC8FDA0" box="[388,559,525,551]" metricMagnitude="-5" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="0.015" valueMax="0.02" valueMin="0.01">0.01–0.02 mm</quantity>
|
||
below the proximal edge. Septula not observed. Ovicells not observed. Avicularia interzooidal, longer than autozooids, elongate. Rostrum channeled with elevated walls and often rounded top, between conical in outline and spade-shaped, asymmetrical, dextral or sinistral. Proximal part rounded, shorter and wider than rostrum, cryptocyst pustulose, concave. Opesia eggshaped with the narrow end pointing upwards and with thin articular ridges bearing two short teeth proximally and short opesiular indentation between the teeth; opesiules lacking. Kenozooids, closure plates, intramural reparative budding autozooids, kenozooids, and avicularia not observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF036649966BFFB3F196F8EFFD53F864" ID-DOI="http://doi.org/10.5281/zenodo.3830867" ID-Zenodo-Dep="3830867" httpUri="https://zenodo.org/record/3830867/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1852,1878]" targetBox="[189,1398,738,1810]" targetPageId="29">
|
||
<paragraph id="8BC336C1966BFFB3F196F8EFFD53F864" blockId="29.[189,1399,1852,2019]" pageId="29" pageNumber="30">
|
||
<emphasis id="B908EAD3966BFFB3F196F8EFFEFBF8D1" bold="true" box="[189,284,1852,1878]" pageId="29" pageNumber="30">Fig. 11.</emphasis>
|
||
<taxonomicName id="4C7C4D42966BFFB3F003F8EFFD93F8D1" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[296,628,1852,1878]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="29" pageNumber="30" phylum="Bryozoa" rank="species" species="operculatum" status="sp. nov.">
|
||
<emphasis id="B908EAD3966BFFB3F003F8EFFD93F8D1" box="[296,628,1852,1878]" italics="true" pageId="29" pageNumber="30">Rhagasostoma operculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57A8966BFFB3F3ABF8EFFD38F8D1" box="[640,735,1852,1878]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B908EAD3966BFFB3F3C0F8EFFCC3F8D1" bold="true" box="[747,804,1852,1878]" pageId="29" pageNumber="30">A–C</emphasis>
|
||
. Holotype, TsNIGR Museum 27/12582, western Kopetdag, Turkmenistan, boundary between the early and late Campanian (
|
||
<taxonomicName id="4C7C4D42966BFFB3F57CF88CFA89F8FE" box="[1111,1390,1887,1913]" class="Globothalamea" family="Anomalinidae" genus="Cibicidoides" kingdom="Chromista" order="Rotaliida" pageId="29" pageNumber="30" phylum="Foraminifera" rank="species" species="temirensis">
|
||
<emphasis id="B908EAD3966BFFB3F57CF88CFA89F8FE" box="[1111,1390,1887,1913]" italics="true" pageId="29" pageNumber="30">Cibicidoides temirensis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7C4D42966BFFB3F196F851FD74F81A" authority="decoratus Zone" authorityName="Zone" box="[189,659,1922,1949]" class="Globothalamea" family="Bolivinoididae" genus="Bolivinoides" kingdom="Chromista" order="Rotaliida" pageId="29" pageNumber="30" phylum="Foraminifera" rank="subSpecies" species="decoratus" subSpecies="decoratus">
|
||
<emphasis id="B908EAD3966BFFB3F196F851FDADF81B" box="[189,586,1922,1948]" italics="true" pageId="29" pageNumber="30">Bolivinoides decoratus decoratus</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
, LS13).
|
||
<emphasis id="B908EAD3966BFFB3F3D0F851FCF7F81B" bold="true" box="[763,784,1922,1948]" pageId="29" pageNumber="30">A</emphasis>
|
||
. Overview of erect bifoliate colony.
|
||
<emphasis id="B908EAD3966BFFB3F5E7F851FB07F81B" bold="true" box="[1228,1248,1922,1948]" pageId="29" pageNumber="30">B</emphasis>
|
||
. Autozooids with putative opercula and avicularia.
|
||
<emphasis id="B908EAD3966BFFB3F3B9F876FD40F838" bold="true" box="[658,679,1957,1983]" pageId="29" pageNumber="30">C</emphasis>
|
||
. Autozooids with putative opercula and avicularium. Scale bars: A = 1 mm; B = 500 µm; C = 200 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C366654A9668FFB0F196FEA3FD88FC6F" pageId="30" pageNumber="31" type="discussion">
|
||
<paragraph id="8BC336C19668FFB0F196FEA3FEC8FE0D" blockId="30.[189,303,368,394]" box="[189,303,368,394]" pageId="30" pageNumber="31">
|
||
<heading id="D08B81AD9668FFB0F196FEA3FEC8FE0D" bold="true" box="[189,303,368,394]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B908EAD39668FFB0F196FEA3FEC8FE0D" bold="true" box="[189,303,368,394]" pageId="30" pageNumber="31">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336C19668FFB0F196FE4CFAEEFDEF" blockId="30.[189,1399,414,617]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="4C7C4D429668FFB0F196FE4CFDE0FE3E" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[189,519,415,441]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="operculatum" status="sp. nov.">
|
||
<emphasis id="B908EAD39668FFB0F196FE4CFDE0FE3E" box="[189,519,415,441]" italics="true" pageId="30" pageNumber="31">Rhagasostoma operculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57A89668FFB0F33AFE4CFD89FE3E" box="[529,622,415,441]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C7C4D429668FFB0F23AFE4CFC7BFE3E" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[785,924,415,441]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="brydonei" status="sp. nov.">
|
||
<emphasis id="B908EAD39668FFB0F23AFE4CFC7BFE3E" box="[785,924,415,441]" italics="true" pageId="30" pageNumber="31">R. brydonei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57A89668FFB0F28DFE4CFBE1FE3E" box="[934,1030,415,441]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, another species with a spadeshaped avicularian rostrum, in the rostrum having almost the same width along the whole of their length and having a pointed or rounded rostral tip instead of the rostrum being narrowed at the base, enlarged centrally and with a pointed tip. The new species differs from the other species described herein in having a spade-shaped avicularian rostrum instead of one that is conical in outline or falciform. Moreover, opesiae of autozooids of
|
||
<taxonomicName id="4C7C4D429668FFB0F0C9FD9CFD71FDEE" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[482,662,591,617]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="operculatum" status="sp. nov.">
|
||
<emphasis id="B908EAD39668FFB0F0C9FD9CFD71FDEE" box="[482,662,591,617]" italics="true" pageId="30" pageNumber="31">R. operculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57A89668FFB0F3B6FD9CFD1FFDEE" box="[669,760,591,617]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are presumably closed by calcified opercula.
|
||
</paragraph>
|
||
<paragraph id="8BC336C19668FFB0F196FD43FD88FC6F" blockId="30.[189,1399,656,1000]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="4C7C4D429668FFB0F196FD43FDE6FD2D" authority="Koromyslova & Taylor & Martha & Riley, 2018" authorityName="Koromyslova & Taylor & Martha & Riley" authorityYear="2018" box="[189,513,656,682]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="operculatum" status="sp. nov.">
|
||
<emphasis id="B908EAD39668FFB0F196FD43FDE6FD2D" box="[189,513,656,682]" italics="true" pageId="30" pageNumber="31">Rhagasostoma operculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57A89668FFB0F32EFD43FDBBFD2D" box="[517,604,656,682]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the earliest known species in cryptocystidean anascan cheilostomes that has putative calcified opercula. Late Cretaceous cheilostomes with calcified opercula from the family
|
||
<taxonomicName id="4C7C4D429668FFB0F039FD05FE2DFD77" authorityName="Jullien" authorityYear="1882" box="[274,458,726,752]" class="Gymnolaemata" family="Onychocellidae" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="family">Onychocellidae</taxonomicName>
|
||
include
|
||
<taxonomicName id="4C7C4D429668FFB0F31BFD04FBCEFD76" authority="(von Hagenow, 1846)" baseAuthorityName="von Hagenow" baseAuthorityYear="1846" box="[560,1065,726,753]" class="Gymnolaemata" family="Onychocellidae" genus="Inversaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="flabellula">
|
||
<emphasis id="B908EAD39668FFB0F31BFD04FCC6FD77" box="[560,801,726,752]" italics="true" pageId="30" pageNumber="31">Inversaria flabellula</emphasis>
|
||
(von Hagenow, 1846)
|
||
</taxonomicName>
|
||
from the late Campanian of
|
||
<collectingRegion id="49B8F8239668FFB0F196FD2AFEE9FC94" box="[189,270,761,787]" country="Sweden" name="Skane" pageId="30" pageNumber="31">Scania</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36B76519668FFB0F037FD2AFE9BFC94" box="[284,380,761,787]" name="Sweden" pageId="30" pageNumber="31">Sweden</collectingCountry>
|
||
,?
|
||
<taxonomicName id="4C7C4D429668FFB0F0B3FD29FDF5FC94" box="[408,530,762,787]" class="Gymnolaemata" family="Onychocellidae" genus="Inversaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B908EAD39668FFB0F0B3FD29FDF5FC94" box="[408,530,762,787]" italics="true" pageId="30" pageNumber="31">Inversaria</emphasis>
|
||
</taxonomicName>
|
||
sp. from the late Campanian to Maastrichtian of
|
||
<collectingCountry id="F36B76519668FFB0F541FD2AFA95FC94" box="[1130,1394,761,787]" name="United Arab Emirates" pageId="30" pageNumber="31">United Arab Emirates</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C7C4D429668FFB0F1D8FCCDFD3BFCBF" authority="(Goldfuss, 1826)" baseAuthorityName="Goldfuss" baseAuthorityYear="1826" box="[243,732,797,824]" class="Gymnolaemata" family="Onychocellidae" genus="Inversaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="tubiporacea">
|
||
<emphasis id="B908EAD39668FFB0F1D8FCCDFDE1FCB0" box="[243,518,797,823]" italics="true" pageId="30" pageNumber="31">Inversaria tubiporacea</emphasis>
|
||
(Goldfuss, 1826)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D429668FFB0F235FCCEFF1FFCDC" authority="Koromyslova & Shcherbinina, 2015" authorityName="Koromyslova & Shcherbinina" authorityYear="2015" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="exilis">
|
||
<emphasis id="B908EAD39668FFB0F235FCCEFC1BFCB0" box="[798,1020,797,823]" italics="true" pageId="30" pageNumber="31">Onychocella exilis</emphasis>
|
||
<bibRefCitation id="EFED4B309668FFB0F52CFCCEFF1FFCDC" author="Koromyslova A. V. & Shcherbinina E. A." pageId="30" pageNumber="31" pagination="474 - 484" refId="ref30922" refString="Koromyslova A. V. & Shcherbinina E. A. 2015. New data on the morphology and age of the bryozoans of the genus Onychocella from the Campanian-Maastrichtian of Turkmenistan and Uzbekistan. Paleontological Journal 49 (5): 474 - 484. https: // doi. org / 10.1134 / S 0031030115050093" type="journal article" year="2015">Koromyslova & Shcherbinina, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Maastrichtian of NW Europe and
|
||
<collectingCountry id="F36B76519668FFB0F3C5FC93FC92FCDD" box="[750,885,832,858]" name="Uzbekistan" pageId="30" pageNumber="31">Uzbekistan</collectingCountry>
|
||
, respectively (
|
||
<bibRefCitation id="EFED4B309668FFB0F530FC93FACAFCDD" author="Voigt E. & Williams A." box="[1051,1325,832,858]" pageId="30" pageNumber="31" pagination="139 - 178" refId="ref33198" refString="Voigt E. & Williams A. 1973. Revision des Genus Inversaria v. Hagenow 1851 (Bryoz. Cheilost.) und seine Beziehungen zu Solen Onychocella n. g. Nachrichten der Akademie der Wissenschaften in G ottingen. II. Mathematisch-Physikalische Klasse 8: 139 - 178." type="journal article" year="1973">Voigt & Williams 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B309668FFB0F41CFC93FF1CFCFA" author="Voigt E." pageId="30" pageNumber="31" pagination="214 - 229" refId="ref32830" refString="Voigt E. 1974. Uber Opercula bei der fossilen Bryozoengattung Inversaria v. Hagenow 1851 (Cheilostomata, Ob. Kreide). Palaontologische Zeitschrift 48 (3 - 4): 214 - 229. https: // doi. org / 10.1007 / BF 02985952" type="journal article" year="1974">Voigt 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B309668FFB0F023FCB0FDA4FCFA" author="Di Martino E. & Taylor P. D." box="[264,579,867,893]" pageId="30" pageNumber="31" pagination="80 - 96" refId="ref30098" refString="Di Martino E. & Taylor P. D. 2013. First bryozoan fauna from a tropical Cretaceous carbonate: Simsima Formation, United Arab Emirates-Oman border region. Cretaceous Research 43: 80 - 96. https: // doi. org / 10.1016 / j. cretres. 2013.02.004" type="journal article" year="2013">Di Martino & Taylor 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B309668FFB0F37BFCB0FCA3FCFA" author="Koromyslova A. V." box="[592,836,867,893]" pageId="30" pageNumber="31" pagination="587 - 593" refId="ref30881" refString="Koromyslova A. V. 2014 b. The earliest calcified opercula of bryozoans of the order Cheilostomata. Paleontological Journal 48 (6): 587 - 593. https: // doi. org / 10.1134 / S 0031030114060082" type="journal article" year="2014">Koromyslova 2014b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B309668FFB0F27BFCB0FB1EFCF9" author="Koromyslova A. V. & Shcherbinina E. A." box="[848,1273,867,894]" pageId="30" pageNumber="31" pagination="474 - 484" refId="ref30922" refString="Koromyslova A. V. & Shcherbinina E. A. 2015. New data on the morphology and age of the bryozoans of the genus Onychocella from the Campanian-Maastrichtian of Turkmenistan and Uzbekistan. Paleontological Journal 49 (5): 474 - 484. https: // doi. org / 10.1134 / S 0031030115050093" type="journal article" year="2015">Koromyslova & Shcherbinina 2015</bibRefCitation>
|
||
). Calcified opercula have also been observed in several species belonging to the family
|
||
<taxonomicName id="4C7C4D429668FFB0F516FC55FA90FC27" authority="Hincks, 1879" authorityName="Hincks" authorityYear="1879" box="[1085,1399,902,928]" class="Gymnolaemata" family="Cribrilinidae" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="family">Cribrilinidae Hincks, 1879</taxonomicName>
|
||
including
|
||
<taxonomicName id="4C7C4D429668FFB0F018FC79FD52FC43" authority="Turner, 1975" authorityName="Turner" authorityYear="1975" box="[307,693,937,964]" class="Gymnolaemata" family="Cribrilinidae" genus="Castanopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="lambi">
|
||
<emphasis id="B908EAD39668FFB0F018FC79FDF0FC43" box="[307,535,938,964]" italics="true" pageId="30" pageNumber="31">Castanopora lambi</emphasis>
|
||
<bibRefCitation id="EFED4B309668FFB0F335FC79FD52FC43" author="Turner R. F." box="[542,693,937,964]" pageId="30" pageNumber="31" pagination="273 - 279" refId="ref32556" refString="Turner R. F. 1975. A new Cretaceous cribrimorph from North America with calcareous opercula. Documents des Laboratoires de Geologie de la Faculte des Sciences de Lyon (hors serie) 3: 273 - 279." type="journal article" year="1975">Turner, 1975</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Maastrichtian of
|
||
<collectingCountry id="F36B76519668FFB0F2DAFC79FBCDFC43" box="[1009,1066,938,964]" name="United States of America" pageId="30" pageNumber="31">USA</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFED4B309668FFB0F511FC79FB37FC43" author="Turner R. F." box="[1082,1232,937,964]" pageId="30" pageNumber="31" pagination="273 - 279" refId="ref32556" refString="Turner R. F. 1975. A new Cretaceous cribrimorph from North America with calcareous opercula. Documents des Laboratoires de Geologie de la Faculte des Sciences de Lyon (hors serie) 3: 273 - 279." type="journal article" year="1975">Turner 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B309668FFB0F5F7FC7AFEC7FC60" author="McKinney F. K. & Taylor P. D. & Lidgard S." pageId="30" pageNumber="31" pagination="239 - 261" refId="ref31510" refString="McKinney F. K. Taylor P. D. & Lidgard S. 2003. Predation on Bryozoans and its reflection in the fossil record. In: Kelley P. H., Kowalewski M. & Hansen T. A. (eds) Predator Prey Interactions in the Fossil Record. Topics in Geobiology 20: 239 - 261. Springer, Boston (MA). https: // doi. org / 10.1007 / 978 - 1 - 4615 - 0161 - 9 _ 10" type="journal article" year="2003">
|
||
McKinney
|
||
<emphasis id="B908EAD39668FFB0F44AFC79FF3CFC60" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B309668FFB0F007FC1EFD84FC6F" author="Taylor P. D. & McKinney F. K." box="[300,611,973,1000]" pageId="30" pageNumber="31" pagination="1 - 346" refId="ref32304" refString="Taylor P. D. & McKinney F. K. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geologica 132: 1 - 346." type="journal article" year="2006">Taylor & McKinney 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366654A9668FFB0F196FBDDFBF9FBD0" box="[189,1054,1038,1111]" pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph id="8BC336C19668FFB0F196FBDDFEB1FBAF" blockId="30.[189,342,1038,1064]" box="[189,342,1038,1064]" pageId="30" pageNumber="31">
|
||
<heading id="D08B81AD9668FFB0F196FBDDFEB1FBAF" bold="true" box="[189,342,1038,1064]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B908EAD39668FFB0F196FBDDFEB1FBAF" bold="true" box="[189,342,1038,1064]" pageId="30" pageNumber="31">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336C19668FFB0F196FBEEFBF9FBD0" blockId="30.[189,1054,1085,1111]" box="[189,1054,1085,1111]" pageId="30" pageNumber="31">
|
||
Early to middle Campanian
|
||
<collectingCountry id="F36B76519668FFB0F32CFBEEFD49FBD0" box="[519,686,1085,1111]" name="Turkmenistan" pageId="30" pageNumber="31">Turkmenistan</collectingCountry>
|
||
: western Kopetdag, Kara-Kala.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |