355 lines
38 KiB
XML
355 lines
38 KiB
XML
<document id="C559A73F59AA97264F86072D128B1C07" ID-CLB-Dataset="55717" ID-DOI="10.5281/zenodo.5341014" ID-GBIF-Dataset="b9741f08-928d-4423-9e6f-478cebcd02cc" ID-ISSN="0253-116X" ID-Zenodo-Dep="5341014" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630189109691" checkinUser="marcus" docAuthor="Assing, V., Schülke, M., M. R & A. S" docDate="2013" docId="03D574268E52FFBDFF1AFE57D98CFEDF" docLanguage="en" docName="LinzerbiolBeitr.2013.45.1.141-154.pdf" docOrigin="Linzer biologische Beiträge 45 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Tachyporus saudicus SCHULKE 2013, nov.sp." docType="treatment" docVersion="9" lastPageNumber="152" masterDocId="FFEC0C5E8E5BFFB6FFB9FFFBD97DFFE6" masterDocTitle="On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera)" masterLastPageNumber="154" masterPageNumber="141" pageNumber="150" updateTime="1699181010772" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="DA87FA20C36FBEA0A15CD7F95FB711D1">
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<mods:title id="31BB298F5ED62FE3D6231EB911C13372">On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="E15980B90FE081A789B302049DE708AA" type="personal">
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<mods:namePart id="B567B88C94D43582B8F2035430A334D0">Assing, V.</mods:namePart>
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<mods:namePart id="4A03093356B430EC16C6313B35CDB979">Schülke, M.</mods:namePart>
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<mods:name id="A928848B55E8A3DE1C314353EF1EBE3F" type="personal">
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<mods:namePart id="BC18ACA85D37903CBC1E62BFDCA48DF6">M. R</mods:namePart>
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</mods:name>
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<mods:name id="898212643F1DEF7E081409B512CAC266" type="personal">
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<mods:roleTerm id="A1592276CEF4C828774B2677A39A0616">Author</mods:roleTerm>
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<mods:namePart id="DE7C994E90F43BF5C0E5061AB783CEBA">A. S</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7B353149E86FBCC0EFC3E2409E2C48C2">text</mods:typeOfResource>
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<mods:title id="A8902579B41C5E041056F13E6C22DD40">Linzer biologische Beiträge</mods:title>
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<mods:part id="00435B031E33BD0033A8F41923DA743B">
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<mods:date id="B1E1C8A27C278EB0923AC36BE4715BE3">2013</mods:date>
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<mods:number id="F9784320B944D1BDE5CE66A96D048E31">2013-07-31</mods:number>
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<mods:detail id="F0068512B0DC086CA4D14FD24819AE5B" type="volume">
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<mods:number id="90ABA5986115AD08A3D185F39E3AE84B">45</mods:number>
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<mods:identifier id="1DBB41784292782E8DF61B8A8CA12707" type="CLB-Dataset">55717</mods:identifier>
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<mods:identifier id="47D5FF29FAAF70E79565E63C73DB7DBF" type="DOI">10.5281/zenodo.5341014</mods:identifier>
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<mods:identifier id="A735F0A5F0022E192E69A5207B3416F7" type="GBIF-Dataset">b9741f08-928d-4423-9e6f-478cebcd02cc</mods:identifier>
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<mods:identifier id="70C298BF6F1EB9A898013AD293BF8289" type="ISSN">0253-116X</mods:identifier>
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<treatment id="03D574268E52FFBDFF1AFE57D98CFEDF" ID-DOI="http://doi.org/10.5281/zenodo.6521792" ID-GBIF-Taxon="195191928" ID-Zenodo-Dep="6521792" LSID="urn:lsid:plazi:treatment:03D574268E52FFBDFF1AFE57D98CFEDF" httpUri="http://treatment.plazi.org/id/03D574268E52FFBDFF1AFE57D98CFEDF" lastPageId="11" lastPageNumber="152" pageId="9" pageNumber="150">
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<subSubSection id="C36696BB8E52FFBFFF1AFE57DB43FE24" box="[163,574,428,450]" pageId="9" pageNumber="150" type="nomenclature">
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<paragraph id="8BC3C5308E52FFBFFF1AFE57DB43FE24" blockId="9.[163,693,428,450]" box="[163,574,428,450]" pageId="9" pageNumber="150">
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<heading id="D08B725C8E52FFBFFF1AFE57DB43FE24" box="[163,574,428,450]" fontSize="8" level="2" pageId="9" pageNumber="150" reason="2">
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<emphasis id="B90819228E52FFBFFF1AFE57DB43FE24" bold="true" box="[163,574,428,450]" pageId="9" pageNumber="150">
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<taxonomicName id="4C7CBEB38E52FFBFFF1AFE57D897FE24" ID-CoL="54D95" authority="SCHULKE" authorityName="SCHULKE" authorityYear="2013" box="[163,490,428,450]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="saudicus" status="sp. nov.">
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<emphasis id="B90819228E52FFBFFF1AFE57D8FDFE24" bold="true" box="[163,384,428,450]" italics="true" pageId="9" pageNumber="150">Tachyporus saudicus</emphasis>
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SCHÜLKE
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</taxonomicName>
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<taxonomicNameLabel id="A23BA4598E52FFBFFE48FE57DB43FE24" box="[497,574,428,450]" pageId="9" pageNumber="150" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36696BB8E52FFBFFDFDFE57DBC8FE24" box="[580,693,428,450]" pageId="9" pageNumber="150" type="description">
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<paragraph id="8BC3C5308E52FFBFFDFDFE57DBC8FE24" blockId="9.[163,693,428,450]" box="[580,693,428,450]" pageId="9" pageNumber="150">
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<heading id="D08B725C8E52FFBFFDFDFE57DBC8FE24" box="[580,693,428,450]" fontSize="8" level="2" pageId="9" pageNumber="150" reason="2">
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(
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<figureCitation id="1347D9B58E52FFBFFDF2FE57DBD0FE24" box="[587,685,428,450]" captionStart-0="Figs 9-11" captionStart-1="Fig" captionStartId-0="9.[163,203,1550,1570]" captionStartId-1="11.[163,196,1079,1099]" captionTargetBox-0="[163,1070,945,1538]" captionTargetBox-1="[163,1077,554,1068]" captionTargetId-0="figure-330@9.[163,1070,945,1538]" captionTargetId-1="figure-122@11.[163,1077,554,1068]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="Figs 9-11: Tachyporus saudicus nov.sp.: (9) habitus; (10) aedeagus in ventral view; (11) sternite VIII. Scale bars: 9: 2.0 mm; 10-11: 0.2 mm." captionText-1="Fig. 12: Type locality of Tachyporus saudicus nov.sp." figureDoi-0="http://doi.org/10.5281/zenodo.5341020" figureDoi-1="http://doi.org/10.5281/zenodo.5341022" httpUri-0="https://zenodo.org/record/5341020/files/figure.png" httpUri-1="https://zenodo.org/record/5341022/files/figure.png" pageId="9" pageNumber="150">Figs 9-12</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36696BB8E52FFBFFF1AFE29D8CBFDCC" pageId="9" pageNumber="150" type="materials_examined">
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<paragraph id="8BC3C5308E52FFBFFF1AFE29D8CBFDCC" blockId="9.[163,1070,465,554]" pageId="9" pageNumber="150">
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Type material:
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<materialsCitation id="3B14CF6D8E52FFBFFE26FE29DB1DFDF2" collectingDate="2011-05-10" collectionCode="KSMA" collectorName="Sharaf" country="Saudi Arabia" determinerName="M. Schulke" elevation="1790" latitude="20.211111" location="Al Mandaq" longLatPrecision="21" longitude="41.288334" municipality="Turabah" pageId="9" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahah" typeStatus="holotype">
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<typeStatus id="54C77B928E52FFBFFE26FE29D88EFE00" box="[415,499,466,486]" pageId="9" pageNumber="150" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D7A0EB98E52FFBFFE46FE2ADB6FFE00" box="[511,530,465,486]" pageId="9" pageNumber="150" type="male">♂</specimenCount>
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: “
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<collectingCountry id="F36B85A08E52FFBFFD92FE29DBA9FE00" box="[555,724,466,486]" name="Saudi Arabia" pageId="9" pageNumber="150">SAUDI ARABIA</collectingCountry>
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740 m,
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<collectingRegion id="49B80BD28E52FFBFFC97FE29DAF1FE00" box="[814,908,466,486]" country="Saudi Arabia" name="Al Bahah" pageId="9" pageNumber="150">Al Bahah</collectingRegion>
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, W
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<collectingMunicipality id="6BA75F4A8E52FFBFFC02FE29DD74FE00" box="[955,1033,466,486]" pageId="9" pageNumber="150">Turabah</collectingMunicipality>
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,
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<location id="8EA393EB8E52FFBFFBAEFE29D98DFE1B" LSID="urn:lsid:plazi:treatment:03D574268E52FFBDFF1AFE57D98CFEDF:8EA393EB8E52FFBFFBAEFE29D98DFE1B" country="Saudi Arabia" latitude="20.211111" longLatPrecision="21" longitude="41.288334" municipality="Turabah" name="Al Mandaq" pageId="9" pageNumber="150" stateProvince="Al Bahah">Al Mandaq</location>
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,
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<geoCoordinate id="EE48A3F78E52FFBFFEB9FE12D811FE1B" box="[256,364,489,509]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="9" pageNumber="150" precision="15" seconds="40" value="20.211111">20°12'40''N</geoCoordinate>
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,
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<geoCoordinate id="EE48A3F78E52FFBFFEC4FE12D89BFE1B" box="[381,486,489,509]" degrees="41" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="150" precision="15" seconds="18" value="41.288334">41°17'18''E</geoCoordinate>
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,
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<quantity id="4C8468D58E52FFBFFE41FE12DB3AFE1B" box="[504,583,489,509]" metricMagnitude="3" metricUnit="m" metricValue="1.7899999999999998" pageId="9" pageNumber="150" unit="m" value="1790.0">
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<elevation id="005122038E52FFBFFE41FE12DB3AFE1B" box="[504,583,489,509]" metricMagnitude="3" metricUnit="m" metricValue="1.7899999999999998" pageId="9" pageNumber="150" unit="m" value="1790.0">1790 m</elevation>
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</quantity>
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,
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<date id="FFC2E3F08E52FFBFFDE1FE12DBC4FE1B" box="[600,697,489,509]" pageId="9" pageNumber="150" value="2011-05-10">
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<collectingDate id="EF861A188E52FFBFFDE1FE12DBC4FE1B" box="[600,697,489,509]" pageId="9" pageNumber="150" value="2011-05-10">10.V.2011</collectingDate>
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</date>
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, leg.
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<collectorName id="2689A0E68E52FFBFFD40FE12DA49FE1B" box="[761,820,489,509]" pageId="9" pageNumber="150">Sharaf</collectorName>
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/ Holotypus
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<taxonomicName id="4C7CBEB38E52FFBFFC7AFE13D98FFDF5" authority="Assing & Schülke & M.R & A.S, 2013" authorityName="Assing & Schülke & M.R & A.S" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="saudicus" status="sp. nov.">
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<emphasis id="B90819228E52FFBFFC7AFE13D98FFDF5" italics="true" pageId="9" pageNumber="150">Tachyporus saudicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23BA4598E52FFBFFF40FDFBD851FDF2" box="[249,300,512,532]" pageId="9" pageNumber="150" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<determinerName id="7E1F29F48E52FFBFFEE7FDFBD8B5FDF2" box="[350,456,512,532]" pageId="9" pageNumber="150">M. Schülke</determinerName>
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2012” (
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<collectionCode id="ED6D5DF58E52FFBFFDAAFDFBDB27FDF2" box="[531,602,512,532]" collectionName="KSMA" pageId="9" pageNumber="150">KSMA</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B14CF6D8E52FFBFFDD3FDFBD8CBFDCC" collectionCode="cSch, cAss" pageId="9" pageNumber="150" specimenCount="10" specimenCount-female="6" specimenCount-male="4" typeStatus="paratype">
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<typeStatus id="54C77B928E52FFBFFDD3FDFBDBBBFDF2" box="[618,710,512,532]" pageId="9" pageNumber="150" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D7A0EB98E52FFBFFD76FDFBDA7DFDF5" box="[719,768,510,532]" pageId="9" pageNumber="150" type="male">3♂♂</specimenCount>
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,
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<specimenCount id="9D7A0EB98E52FFBFFCB5FDFBDA40FDFF" box="[780,829,512,537]" pageId="9" pageNumber="150" type="female">3♀♀</specimenCount>
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[
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<specimenCount id="9D7A0EB98E52FFBFFCF5FDFBDA17FDF5" box="[844,874,510,532]" pageId="9" pageNumber="150" type="male">1♂</specimenCount>
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,
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<specimenCount id="9D7A0EB98E52FFBFFCCFFDFBDADAFDFF" box="[886,935,512,537]" pageId="9" pageNumber="150" type="female">3♀♀</specimenCount>
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teneral]: same data as holotype (
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<collectionCode id="ED6D5DF58E52FFBFFEFAFDEDD80FFDCC" box="[323,370,534,554]" pageId="9" pageNumber="150">cSch</collectionCode>
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,
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<collectionCode id="ED6D5DF58E52FFBFFEC5FDEDD8D1FDCC" box="[380,428,534,554]" pageId="9" pageNumber="150">cAss</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36696BB8E52FFBFFF1AFDC2D883FD8A" pageId="9" pageNumber="150" type="etymology">
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<paragraph id="8BC3C5308E52FFBFFF1AFDC2D883FD8A" blockId="9.[163,1070,569,620]" pageId="9" pageNumber="150">
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Etymology: The specific epithet (Latin, adjective) is derived from
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<collectingCountry id="F36B85A08E52FFBFFC1FFDC2DD53FDA9" box="[934,1070,569,591]" name="Saudi Arabia" pageId="9" pageNumber="150">Saudi Arabia</collectingCountry>
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where the species was discovered.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36696BB8E52FFBCFF1AFD80D8C1FC46" lastPageId="10" lastPageNumber="151" pageId="9" pageNumber="150" type="description">
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<paragraph id="8BC3C5308E52FFBFFF1AFD80DB89FCC2" blockId="9.[163,1071,635,804]" pageId="9" pageNumber="150">Description: Measurements (in mm) and indices: body length 2.7-3.4; length of forebody 1.7-1.9; head width 0.59-0.60; pronotal width 0.96-1.00; pronotal length 0.64- 0.66; elytral sutural length (apex of scutellum to sutural angle) 0.77-0.79; elytral humeral length (anterior to posterior margin) 0.96-1.01; elytral width 1.03-1.10; aedeagus length 0.69-0.72. Ratios: head width / pronotal width 0.60-0.62; pronotal length / elytral length 0.77-0.83; elytral humeral length / elytral width 0.90-0.97.</paragraph>
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<paragraph id="8BC3C5308E52FFBFFF1AFCC8DAA5FC47" blockId="9.[163,1070,819,929]" pageId="9" pageNumber="150">
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Habitus as in
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<figureCitation id="1347D9B58E52FFBFFE88FCC8D80DFCAF" box="[305,368,819,841]" captionStart="Figs 9-11" captionStartId="9.[163,203,1550,1570]" captionTargetBox="[163,1070,945,1538]" captionTargetId="figure-330@9.[163,1070,945,1538]" captionTargetPageId="9" captionText="Figs 9-11: Tachyporus saudicus nov.sp.: (9) habitus; (10) aedeagus in ventral view; (11) sternite VIII. Scale bars: 9: 2.0 mm; 10-11: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341020" httpUri="https://zenodo.org/record/5341020/files/figure.png" pageId="9" pageNumber="150">Fig. 9</figureCitation>
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. Coloration: head black, pronotum and elytra yellowish-brown, disc of pronotum dark-brown, only
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<specimenCount id="9D7A0EB98E52FFBFFE50FCAADB0BFC80" box="[489,630,848,870]" pageId="9" pageNumber="150" type="generic">one specimen</specimenCount>
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with oblique brown elytral markings, abdomen dark-brown to black, posterior margins of segments yellowish-brown, legs and basal antennomeres yellow, antennomeres IV-XI and maxillary palpus brownish.
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</paragraph>
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<caption id="DF0395B88E52FFBFFF1AF9F5DB49F9DF" ID-DOI="http://doi.org/10.5281/zenodo.5341020" ID-Zenodo-Dep="5341020" httpUri="https://zenodo.org/record/5341020/files/figure.png" pageId="9" pageNumber="150" startId="9.[163,203,1550,1570]" targetBox="[163,1070,945,1538]" targetPageId="9">
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<paragraph id="8BC3C5308E52FFBFFF1AF9F5DB49F9DF" blockId="9.[163,1070,1549,1593]" pageId="9" pageNumber="150">
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<emphasis id="B90819228E52FFBFFF1AF9F5D980F9C4" bold="true" box="[163,253,1550,1570]" pageId="9" pageNumber="150">Figs 9-11</emphasis>
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:
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<taxonomicName id="4C7CBEB38E52FFBFFEB2F9F6D8B0F9C4" authority="Assing & Schülke & M.R & A.S, 2013" authorityName="Assing & Schülke & M.R & A.S" authorityYear="2013" box="[267,461,1549,1570]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="saudicus" status="sp. nov.">
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<emphasis id="B90819228E52FFBFFEB2F9F6D8B0F9C4" box="[267,461,1549,1570]" italics="true" pageId="9" pageNumber="150">Tachyporus saudicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23BA4598E52FFBFFE6FF9F5DB61F9C4" box="[470,540,1550,1570]" pageId="9" pageNumber="150" rank="species">nov.sp.</taxonomicNameLabel>
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|
: (
|
|
<emphasis id="B90819228E52FFBFFD97F9F5DB44F9C4" bold="true" box="[558,569,1550,1570]" pageId="9" pageNumber="150">9</emphasis>
|
|
) habitus; (
|
|
<emphasis id="B90819228E52FFBFFD19F9F5DBCAF9C4" bold="true" box="[672,695,1550,1570]" pageId="9" pageNumber="150">10</emphasis>
|
|
) aedeagus in ventral view; (
|
|
<emphasis id="B90819228E52FFBFFC7CF9F5DAA1F9C4" bold="true" box="[965,988,1550,1570]" pageId="9" pageNumber="150">11</emphasis>
|
|
) sternite VIII. Scale bars: 9: 2.0 mm; 10-11: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFF31D862FE42" blockId="10.[163,1071,202,420]" pageId="10" pageNumber="151">
|
|
Head transverse; posterior part concealed under pronotal anterior margin; punctation of frons and vertex fine and sparse, distance between punctures approximately 30 µm; without microsculpture. Eyes weakly convex and rather large, distinctly longer than postocular region in dorsal view. Antenna approximately
|
|
<quantity id="4C8468D58E51FFBCFD4FFEE5DA38FED2" box="[758,837,286,308]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="151" unit="mm" value="1.1">1.1 mm</quantity>
|
|
long; antennomeres I-III more than twice as long as broad, antennomeres IV-X of decreasing length, antennomere X as long as broad, antennomere XI slightly elongated, 1.7 times as long as broad. Antennomeres I-III only with sensory bristles, antennomeres IV-XI with dense pubescence.
|
|
</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFE48D9AAFDDE" blockId="10.[163,1070,434,568]" pageId="10" pageNumber="151">Pronotum distinctly transverse, 1.6-1.7 times as wide as head; anterior angles bent lateroventral, posterior angles broadly rounded; punctation similar to that of head, pubescence almost invisible, only 1-2 µm long; microsculpture absent. Anterior, posterior, and lateral margins each with four distinct bristle-bearing punctures, bristles approximately 100 µm long.</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFDBDDD7AFD0E" blockId="10.[163,1071,582,744]" pageId="10" pageNumber="151">Elytra moderately long, 1.2-1.3 times as long as pronotum, slightly widened posteriad, 0.90-0.97 times as long as wide; punctures as sparse as those of pronotum, but distinctly coarser with pubescence of approximately 50 µm length. Elytra covered with rows of sensory bristles of approximately 100 µm length forming distinct rows: humeral 0-1, sublateral 1-2, inner discal 2, middle discal 3, outer discal 2, sutural 3-4, apical 3 and lateral 5-6; surface of elytra without any microsculpture. Hind wings fully developed.</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFD0DDA00FCA2" blockId="10.[163,1070,758,836]" pageId="10" pageNumber="151">Abdomen with punctation and pubescence of tergites similar to those of pronotum; tergites with indistinct microsculpture composed of transverse striae or meshes (2 meshes per 10 µm); posterior margin of tergite VII with distinct palisade fringe.</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFCA8D8C1FC46" blockId="10.[163,1070,850,928]" pageId="10" pageNumber="151">
|
|
♂: protarsomeres I-IV distinctly dilated; sternite VIII with distinct posterior emargination (
|
|
<figureCitation id="1347D9B58E51FFBCFF60FC95D859FC62" box="[217,292,878,900]" captionStart="Figs 9-11" captionStartId="9.[163,203,1550,1570]" captionTargetBox="[163,1070,945,1538]" captionTargetId="figure-330@9.[163,1070,945,1538]" captionTargetPageId="9" captionText="Figs 9-11: Tachyporus saudicus nov.sp.: (9) habitus; (10) aedeagus in ventral view; (11) sternite VIII. Scale bars: 9: 2.0 mm; 10-11: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341020" httpUri="https://zenodo.org/record/5341020/files/figure.png" pageId="10" pageNumber="151">Fig. 11</figureCitation>
|
|
); aedeagus approximately
|
|
<quantity id="4C8468D58E51FFBCFD8EFC95DBFBFC62" box="[567,646,878,900]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="151" unit="mm" value="0.7">0.7 mm</quantity>
|
|
long, without large sclerotized structures in the internal sac (
|
|
<figureCitation id="1347D9B58E51FFBCFEDDFC71D8CDFC46" box="[356,432,906,928]" captionStart="Figs 9-11" captionStartId="9.[163,203,1550,1570]" captionTargetBox="[163,1070,945,1538]" captionTargetId="figure-330@9.[163,1070,945,1538]" captionTargetPageId="9" captionText="Figs 9-11: Tachyporus saudicus nov.sp.: (9) habitus; (10) aedeagus in ventral view; (11) sternite VIII. Scale bars: 9: 2.0 mm; 10-11: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341020" httpUri="https://zenodo.org/record/5341020/files/figure.png" pageId="10" pageNumber="151">Fig. 10</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C36696BB8E51FFBCFF1AFC55DB62F9C3" pageId="10" pageNumber="151" type="discussion">
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFC55DA3AFB6E" blockId="10.[163,1070,942,1160]" pageId="10" pageNumber="151">
|
|
Comparative notes No records of
|
|
<taxonomicName id="4C7CBEB38E51FFBCFD13FC55DA5CFC25" authority="GRAVENHORST 1802" authorityName="GRAVENHORST" authorityYear="1802" box="[682,801,942,963]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B90819228E51FFBCFD13FC55DA5CFC25" box="[682,801,942,963]" italics="true" pageId="10" pageNumber="151">Tachyporus</emphasis>
|
|
</taxonomicName>
|
|
species were previously known from the Arabian Peninsula. Only few species have been recorded from the adjacent regions, such as
|
|
<taxonomicName id="4C7CBEB38E51FFBCFE3EFC1CDBC2FC1A" authority="(FABRICIUS 1781)" baseAuthorityName="FABRICIUS" baseAuthorityYear="1781" box="[391,703,998,1020]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="nitidulus">
|
|
<emphasis id="B90819228E51FFBCFE3EFC1CD883FC1D" box="[391,510,998,1020]" italics="true" pageId="10" pageNumber="151">T. nitidulus</emphasis>
|
|
(FABRICIUS 1781)
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F36B85A08E51FFBCFCBDFC1CDA3EFC1A" box="[772,835,999,1020]" name="Egypt" pageId="10" pageNumber="151">Egypt</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFCE9FC1DDAD6FC1A" box="[848,939,998,1020]" name="Lebanon" pageId="10" pageNumber="151">Lebanon</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFC00FC1DDA8CFC1A" box="[953,1009,998,1020]" name="Syria" pageId="10" pageNumber="151">Syria</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFC46FC1CDD57FC1A" box="[1023,1066,999,1020]" name="Iran" pageId="10" pageNumber="151">Iran</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFF1AFBF9D9A6FBFE" box="[163,219,1026,1048]" name="Israel" pageId="10" pageNumber="151">Israel</collectingCountry>
|
|
and
|
|
<collectingCountry id="F36B85A08E51FFBCFEA1FBF8D83EFBFE" box="[280,323,1027,1048]" name="Iraq" pageId="10" pageNumber="151">Iraq</collectingCountry>
|
|
,
|
|
<taxonomicName id="4C7CBEB38E51FFBCFEEDFBF8DB33FBFE" authority="SAULCY 1865" authorityName="SAULCY" authorityYear="1865" box="[340,590,1026,1048]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="abner">
|
|
<emphasis id="B90819228E51FFBCFEEDFBF8D8D2FBF1" box="[340,431,1026,1048]" italics="true" pageId="10" pageNumber="151">T. abner</emphasis>
|
|
SAULCY 1865
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F36B85A08E51FFBCFD2EFBF9DBAFFBFE" box="[663,722,1026,1048]" name="Israel" pageId="10" pageNumber="151">Israel</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFD5BFBF9DA58FBFE" box="[738,805,1026,1048]" name="Jordan" pageId="10" pageNumber="151">Jordan</collectingCountry>
|
|
and
|
|
<collectingCountry id="F36B85A08E51FFBCFCDAFBF8DAF3FBFE" box="[867,910,1027,1048]" name="Iran" pageId="10" pageNumber="151">Iran</collectingCountry>
|
|
,
|
|
<taxonomicName id="4C7CBEB38E51FFBCFC27FBF8D838FBD2" authority="KOLENATI 1846" authorityName="KOLENATI" authorityYear="1846" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="caucasicus">
|
|
<emphasis id="B90819228E51FFBCFC27FBF8DD53FBFE" box="[926,1070,1027,1048]" italics="true" pageId="10" pageNumber="151">T. caucasicus</emphasis>
|
|
KOLENATI 1846
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F36B85A08E51FFBCFE3DFBE5D8C2FBD2" box="[388,447,1054,1076]" name="Israel" pageId="10" pageNumber="151">Israel</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFE70FBE5DB72FBD2" box="[457,527,1054,1076]" name="Jordan" pageId="10" pageNumber="151">Jordan</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFDA3FBE5DB2DFBD2" box="[538,592,1054,1076]" name="Syria" pageId="10" pageNumber="151">Syria</collectingCountry>
|
|
and
|
|
<collectingCountry id="F36B85A08E51FFBCFD3BFBE4DBD0FBD2" box="[642,685,1055,1076]" name="Iran" pageId="10" pageNumber="151">Iran</collectingCountry>
|
|
,
|
|
<taxonomicName id="4C7CBEB38E51FFBCFD01FBE4DA8BFBD2" authority="(FABRICIUS 1775)" baseAuthorityName="FABRICIUS" baseAuthorityYear="1775" box="[696,1014,1054,1076]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="hypnorum">
|
|
<emphasis id="B90819228E51FFBCFD01FBE4DA47FBD5" box="[696,826,1054,1076]" italics="true" pageId="10" pageNumber="151">T. hypnorum</emphasis>
|
|
(FABRICIUS 1775)
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F36B85A08E51FFBCFF1AFBC1D981FBB6" box="[163,252,1082,1104]" name="Lebanon" pageId="10" pageNumber="151">Lebanon</collectingCountry>
|
|
and
|
|
<collectingCountry id="F36B85A08E51FFBCFE8CFBC0D81DFBB6" box="[309,352,1083,1104]" name="Iran" pageId="10" pageNumber="151">Iran</collectingCountry>
|
|
, and
|
|
<taxonomicName id="4C7CBEB38E51FFBCFE27FBC0DB96FBB6" authority="GRAVENHORST 1806" authorityName="GRAVENHORST" authorityYear="1806" box="[414,747,1082,1104]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pusillus">
|
|
<emphasis id="B90819228E51FFBCFE27FBC0DB70FBA9" box="[414,525,1082,1104]" italics="true" pageId="10" pageNumber="151">T. pusillus</emphasis>
|
|
GRAVENHORST 1806
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F36B85A08E51FFBCFC88FBC0DA0DFBB6" box="[817,880,1083,1104]" name="Egypt" pageId="10" pageNumber="151">Egypt</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFCC6FBC1DAC7FBB6" box="[895,954,1082,1104]" name="Israel" pageId="10" pageNumber="151">Israel</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36B85A08E51FFBCFC70FBC1DA82FBB6" box="[969,1023,1082,1104]" name="Syria" pageId="10" pageNumber="151">Syria</collectingCountry>
|
|
and
|
|
<collectingCountry id="F36B85A08E51FFBCFF1AFBACD9B2FB8A" box="[163,207,1111,1132]" name="Iran" pageId="10" pageNumber="151">Iran</collectingCountry>
|
|
. The record of
|
|
<taxonomicName id="4C7CBEB38E51FFBCFEC4FBACDBEAFB8A" authority="ERICHSON 1839" authorityName="ERICHSON" authorityYear="1839" box="[381,663,1110,1132]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="solutus">
|
|
<emphasis id="B90819228E51FFBCFEC4FBACD89BFB8D" box="[381,486,1110,1132]" italics="true" pageId="10" pageNumber="151">T. solutus</emphasis>
|
|
ERICHSON 1839
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F36B85A08E51FFBCFD66FBADDA68FB8A" box="[735,789,1110,1132]" name="Syria" pageId="10" pageNumber="151">Syria</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EFEDB8C18E51FFBCFC91FBADDAB6FB8A" author="SMETANA A" box="[808,971,1110,1132]" pageId="10" pageNumber="151" pagination="505 - 698" refId="ref7869" refString="SMETANA A. (2004): Staphylinidae, subfamilies Omaliinae - Dasycerinae, Phloeocharinae - Apaticinae, Piestinae - Staphylininae, pp. 237 - 272, 329 - 495, 505 - 698. - In: LOBL I. & A. SMETANA (eds), Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea - Histeroidea - Staphylinoidea. - Stenstrup, Apollo Books: 942 pp." type="book chapter" year="2004">SMETANA 2004</bibRefCitation>
|
|
) remains doubtful, most likely it is based on a confusion with
|
|
<taxonomicName id="4C7CBEB38E51FFBCFD0EFB88DA3CFB6E" authorityName="KOLENATI" authorityYear="1846" box="[695,833,1139,1160]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="caucasicus">
|
|
<emphasis id="B90819228E51FFBCFD0EFB88DA3CFB6E" box="[695,833,1139,1160]" italics="true" pageId="10" pageNumber="151">T. caucasicus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFB63DB1AFB2D" blockId="10.[163,1071,1175,1227]" pageId="10" pageNumber="151">
|
|
<taxonomicName id="4C7CBEB38E51FFBCFF1AFB63D804FB4B" authorityName="Assing & Schülke & M.R & A.S" authorityYear="2013" box="[163,377,1176,1197]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="saudicus">
|
|
<emphasis id="B90819228E51FFBCFF1AFB63D804FB4B" box="[163,377,1176,1197]" italics="true" pageId="10" pageNumber="151">Tachyporus saudicus</emphasis>
|
|
</taxonomicName>
|
|
is highly similar in size, habitus, and elytral chaetotaxy to
|
|
<taxonomicName id="4C7CBEB38E51FFBCFC61FB63DD52FB4B" authorityName="SAULCY" authorityYear="1865" box="[984,1071,1176,1197]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="abner">
|
|
<emphasis id="B90819228E51FFBCFC61FB63DD52FB4B" box="[984,1071,1176,1197]" italics="true" pageId="10" pageNumber="151">T. abner</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C7CBEB38E51FFBCFF77FB4DD845FB2C" authorityName="GRAVENHORST" authorityYear="1806" box="[206,312,1205,1227]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pusillus">
|
|
<emphasis id="B90819228E51FFBCFF77FB4DD845FB2C" box="[206,312,1205,1227]" italics="true" pageId="10" pageNumber="151">T. pusillus</emphasis>
|
|
</taxonomicName>
|
|
, but distinguished as follows:
|
|
</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFB21D83CFAA1" blockId="10.[163,1070,1242,1352]" pageId="10" pageNumber="151">
|
|
from
|
|
<taxonomicName id="4C7CBEB38E51FFBCFF64FB20D848FB09" authorityName="SAULCY" authorityYear="1865" box="[221,309,1242,1264]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="abner">
|
|
<emphasis id="B90819228E51FFBCFF64FB20D848FB09" box="[221,309,1242,1264]" italics="true" pageId="10" pageNumber="151">T. abner</emphasis>
|
|
</taxonomicName>
|
|
by the much paler coloration of the whole body, especially of the elytra (with characteristic black markings in
|
|
<taxonomicName id="4C7CBEB38E51FFBCFD8DFB03DBF1FAEA" authorityName="SAULCY" authorityYear="1865" box="[564,652,1271,1293]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="abner">
|
|
<emphasis id="B90819228E51FFBCFD8DFB03DBF1FAEA" box="[564,652,1271,1293]" italics="true" pageId="10" pageNumber="151">T. abner</emphasis>
|
|
</taxonomicName>
|
|
), variability of the presence of humeral bristle-bearing punctures on the elytra, and the larger size of the aedeagus (
|
|
<taxonomicName id="4C7CBEB38E51FFBCFC74FAEEDD5AFACC" authorityName="SAULCY" authorityYear="1865" box="[973,1063,1301,1322]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="abner">
|
|
<emphasis id="B90819228E51FFBCFC74FAEEDD5AFACC" box="[973,1063,1301,1322]" italics="true" pageId="10" pageNumber="151">T. abner</emphasis>
|
|
</taxonomicName>
|
|
:
|
|
<quantity id="4C8468D58E51FFBCFF1AFAC9D848FAA1" box="[163,309,1330,1352]" metricMagnitude="-4" metricUnit="m" metricValue="6.05" metricValueMax="6.3" metricValueMin="5.8" pageId="10" pageNumber="151" unit="mm" value="0.605" valueMax="0.63" valueMin="0.58">0.58-0.63 mm</quantity>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BC3C5308E51FFBCFF1AFAACDAD2FA41" blockId="10.[163,1070,1367,1448]" pageId="10" pageNumber="151">
|
|
from
|
|
<taxonomicName id="4C7CBEB38E51FFBCFF65FAA3D83AFA8A" authorityName="GRAVENHORST" authorityYear="1806" box="[220,327,1367,1389]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pusillus">
|
|
<emphasis id="B90819228E51FFBCFF65FAA3D83AFA8A" box="[220,327,1367,1389]" italics="true" pageId="10" pageNumber="151">T. pusillus</emphasis>
|
|
</taxonomicName>
|
|
by usually paler coloration (
|
|
<taxonomicName id="4C7CBEB38E51FFBCFDC8FAA3DBA1FA8A" authorityName="GRAVENHORST" authorityYear="1806" box="[625,732,1367,1389]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pusillus">
|
|
<emphasis id="B90819228E51FFBCFDC8FAA3DBA1FA8A" box="[625,732,1367,1389]" italics="true" pageId="10" pageNumber="151">T. pusillus</emphasis>
|
|
</taxonomicName>
|
|
: body dark-brown), the variability of the presence of humeral bristle-bearing punctures on the elytra, the absence of elytral microsculpture, and the smaller aedeagus (
|
|
<taxonomicName id="4C7CBEB38E51FFBCFD23FA68DA79FA41" authorityName="GRAVENHORST" authorityYear="1806" box="[666,772,1426,1448]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pusillus">
|
|
<emphasis id="B90819228E51FFBCFD23FA68DA79FA41" box="[666,772,1426,1448]" italics="true" pageId="10" pageNumber="151">T. pusillus</emphasis>
|
|
</taxonomicName>
|
|
:
|
|
<quantity id="4C8468D58E51FFBCFCA8FA69DAD9FA41" box="[785,932,1426,1448]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" metricValueMax="8.5" metricValueMin="7.9" pageId="10" pageNumber="151" unit="mm" value="0.82" valueMax="0.85" valueMin="0.79">0.79-0.85 mm</quantity>
|
|
).
|
|
</paragraph>
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<paragraph id="8BC3C5308E51FFBCFF1AFA4CDB62F9C3" blockId="10.[163,1070,1463,1573]" pageId="10" pageNumber="151">
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Other pale-coloured species with similar oblique elytral markings (
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<taxonomicName id="4C7CBEB38E51FFBCFC29FA43D81FFA0C" authority="BERNHAUER 1920" authorityName="BERNHAUER" authorityYear="1920" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="himalayicus">
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|
<emphasis id="B90819228E51FFBCFC29FA43DD53FA2A" box="[912,1070,1463,1485]" italics="true" pageId="10" pageNumber="151">T. himalayicus</emphasis>
|
|
BERNHAUER 1920
|
|
</taxonomicName>
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|
,
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|
<taxonomicName id="4C7CBEB38E51FFBCFECAFA2EDBB4FA0C" authority="BERNHAUER, 1915" authorityName="BERNHAUER" authorityYear="1915" box="[371,713,1492,1514]" class="Insecta" family="Staphylinidae" genus="Tachyporus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="montanus">
|
|
<emphasis id="B90819228E51FFBCFECAFA2ED88BFA0C" box="[371,502,1493,1514]" italics="true" pageId="10" pageNumber="151">T. montanus</emphasis>
|
|
BERNHAUER, 1915
|
|
</taxonomicName>
|
|
, and other probably undescribed species) are distinguished by the reduced number of elytral sensory bristles, especially the reduced number of discal bristles.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C36696BB8E50FFBDFF1AFF30D98CFEDF" pageId="11" pageNumber="152" type="distribution">
|
|
<paragraph id="8BC3C5308E50FFBDFF1AFF30D98CFEDF" blockId="11.[163,1070,203,313]" pageId="11" pageNumber="152">
|
|
Distribution and natural history The
|
|
<typeStatus id="54C77B928E50FFBDFCF0FF37DA10FF07" box="[841,877,204,225]" pageId="11" pageNumber="152">type</typeStatus>
|
|
specimens were collected in a pasture, probably from under stones, in Wadi Turabah,
|
|
<collectingRegion id="49B80BD28E50FFBDFCC4FF12DA9FFF19" box="[893,994,233,255]" country="Saudi Arabia" name="Al Bahah" pageId="11" pageNumber="152">Al Bahah</collectingRegion>
|
|
, southwestern
|
|
<collectingCountry id="F36B85A08E50FFBDFF40FEFDD8FCFEFA" box="[249,385,262,284]" name="Saudi Arabia" pageId="11" pageNumber="152">Saudi Arabia</collectingCountry>
|
|
, at an altitude of
|
|
<quantity id="4C8468D58E50FFBDFD8CFEFDDBFCFEFA" box="[565,641,262,284]" metricMagnitude="3" metricUnit="m" metricValue="1.7899999999999998" pageId="11" pageNumber="152" unit="m" value="1790.0">1790 m</quantity>
|
|
in May. The
|
|
<typeStatus id="54C77B928E50FFBDFCB6FEFCDA46FEFA" box="[783,827,263,284]" pageId="11" pageNumber="152">type</typeStatus>
|
|
locality is illustrated in
|
|
<figureCitation id="1347D9B58E50FFBDFF1AFED8D990FEDF" box="[163,237,291,313]" captionStart="Fig" captionStartId="11.[163,196,1079,1099]" captionTargetBox="[163,1077,554,1068]" captionTargetId="figure-122@11.[163,1077,554,1068]" captionTargetPageId="11" captionText="Fig. 12: Type locality of Tachyporus saudicus nov.sp." figureDoi="http://doi.org/10.5281/zenodo.5341022" httpUri="https://zenodo.org/record/5341022/files/figure.png" pageId="11" pageNumber="152">Fig. 12</figureCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |