treatments-xml/data/03/D4/A7/03D4A739FFEE686725FFFD789073FDFA.xml

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<mods:title id="A5D2F2C9399F038B830FA355243039CF">A new Afrotropical genus Monocentroptilum gen. n. (Ephemeroptera: Baetidae: Protopatellata)</mods:title>
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<treatment id="03D4A739FFEE686725FFFD789073FDFA" ID-DOI="http://doi.org/10.5281/zenodo.5966870" ID-GBIF-Taxon="148403022" ID-Zenodo-Dep="5966870" LSID="urn:lsid:plazi:treatment:03D4A739FFEE686725FFFD789073FDFA" httpUri="http://treatment.plazi.org/id/03D4A739FFEE686725FFFD789073FDFA" lastPageId="3" lastPageNumber="118" pageId="1" pageNumber="116">
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<paragraph id="8BC2162FFFEE686525FFFD7895F1FD0B" blockId="1.[151,467,690,750]" box="[151,467,690,716]" pageId="1" pageNumber="116">
<heading id="D08AA143FFEE686525FFFD7895F1FD0B" bold="true" box="[151,467,690,716]" fontSize="11" level="1" pageId="1" pageNumber="116" reason="1">
<emphasis id="B909CA3DFFEE686525FFFD7895F1FD0B" bold="true" box="[151,467,690,716]" pageId="1" pageNumber="116">
<taxonomicName id="4C7D6DACFFEE686525FFFD789558FD0B" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[151,378,690,716]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE686525FFFD789558FD0B" bold="true" box="[151,378,690,716]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFEE686524E8FD7895F1FD0B" box="[384,467,690,716]" pageId="1" pageNumber="116" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC2162FFFEE686525FFFD1F953BFD29" blockId="1.[151,467,690,750]" box="[151,281,725,750]" pageId="1" pageNumber="116">
(
<figureCitation id="13460AAAFFEE686525F7FD1F9533FD29" box="[159,273,725,750]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–9" captionStart-2="FIGURES 10–27" captionStart-3="FIGURES 28–31" captionStart-4="FIGURES 32–39" captionStart-5="FIGURES 40–43" captionStart-6="FIGURES 44–47" captionStart-7="FIGURES 48–52" captionStart-8="FIGURES 53–58" captionStartId-0="2.[151,264,1932,1955]" captionStartId-1="3.[151,264,1969,1992]" captionStartId-2="4.[151,264,1871,1894]" captionStartId-3="5.[151,264,1493,1516]" captionStartId-4="6.[151,264,1926,1949]" captionStartId-5="7.[151,264,1072,1095]" captionStartId-6="9.[151,264,1870,1893]" captionStartId-7="10.[151,264,1655,1678]" captionStartId-8="11.[151,264,1784,1807]" captionTargetBox-0="[151,1436,879,1896]" captionTargetBox-1="[181,1415,663,1951]" captionTargetBox-2="[174,1415,202,1856]" captionTargetBox-3="[151,1435,193,1485]" captionTargetBox-4="[151,1436,193,1903]" captionTargetBox-5="[151,1423,193,1034]" captionTargetBox-6="[183,1400,201,1847]" captionTargetBox-7="[150,1436,193,1634]" captionTargetBox-8="[151,1435,303,1763]" captionTargetId-0="figure@2.[151,1436,879,1908]" captionTargetId-1="figure@3.[174,1423,628,1959]" captionTargetId-2="figure@4.[170,1417,195,1857]" captionTargetId-3="figure@5.[151,1435,193,1485]" captionTargetId-4="figure@6.[151,1436,193,1906]" captionTargetId-5="figure@7.[151,1436,193,1052]" captionTargetId-6="figure@9.[183,1406,193,1854]" captionTargetId-7="figure@10.[150,1436,193,1634]" captionTargetId-8="figure@11.[151,1435,303,1763]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionText-0="FIGURES 1–4. Larva Of Monocentroptilum badium. 1, Lateral view (arrOws shOw prOtuberances On mesOnOtum; 2, exuviae Of right half Of prOnOtum and mesOnOtum; 3, exuviae Of abdOminal sterna and terga; 4, subimaginal sterna and terga extracted frOm mature larva (tO shOw hypOdermal cOlOratiOn)." captionText-1="FIGURES 5–9. Larva Of Monocentroptilum badium. 5, labium; 6, maxilla (dOrsal view); 7, labium (at left ventral view, at right dOrsal view); 8–9, left and right mandible. AbbreviatiOns: ds1–ds3, dentisetae." captionText-2="FIGURES 10–27. Larva Of Monocentroptilum badium. 10–16, tergalii Of I–VII pairs; 17, margin Of tergalius; 18, fOre leg; 19, claw (view frOm anteriOr side); 20, hind leg; 21, setae On apex Of its tibia; 22, claw (view frOm inner side); 23, exuviae Of half Of prOnOtum and mesOnOtum (thick line shOws area enlarged On Fig. 28); 24, setae On mesOnOtum; 25, seta On paired prOtuberances; 26, hind prOtOpterOm (the same scale as Fig. 23); 27, part Of head with clypeus, bases Of antennae, median Ocellus and frOntal suture." captionText-3="FIGURES 28–31. Larval exuviae Of Monocentroptilum badium. 28, fragment Of mesOnOtum (Outlined On Fig. 23); 29, fragment Of cercus; 30, fragment Of tergum VIII; 31, paraprOct (all at the same scale)." captionText-4="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." captionText-5="FIGURES 40–43. Monocentroptilum badium. 40–41, fOre and hind wings; 42, head and mesOnOtum Of male imagO; 43, subimaginal exuviae Of left pOstsubalar sclerite and laterOpOstnOtal crest." captionText-6="FIGURES 44–47. Male genitals Of Monocentroptilum badium. 44, develOping subimaginal genitals, fOlded under larval cuticle; 45, subimaginal exuviae Of gOnOstyli; 46, imaginal genitals Of the same specimen (gOnOduct shOwn by dOtted lines, muscles shOwn by interrupted lines, internal parts Of penis shOwn by interrupted lines and dOtted); 47, its left unistyliger, tO shOw muscles. AbbreviatiOns: 1–3, segments Of gOnOstylus; gv, gOnOvectis; m.sg, median sternO-styligeral muscle; pgs, larval prOtOgOnOstylus; usg, unistyliger." captionText-7="FIGURES 48–52. Male imaginal genitals Of Monocentroptilum badium. 48–49, specimen with prOtracted gOnOvectes: 48, with fOcus On internal parts, including gOnOvectes; 49, with fOcus On ventral surface, including unistyligers and median styligeral sclerite; 50–51, specimen with retracted gOnOvectes: 50, with fOcus On gOnOvectes and penial bridge; 51, with fOcus On median styligeral sclerite and median sternO-styligeral muscle; 52, gOnOstyli." captionText-8="FIGURES 53–58. Eggs Of Monocentroptilum badium." httpUri-0="https://zenodo.org/record/1436672/files/figure.png" httpUri-1="https://zenodo.org/record/1436674/files/figure.png" httpUri-2="https://zenodo.org/record/1436676/files/figure.png" httpUri-3="https://zenodo.org/record/1436678/files/figure.png" httpUri-4="https://zenodo.org/record/1436680/files/figure.png" httpUri-5="https://zenodo.org/record/1436682/files/figure.png" httpUri-6="https://zenodo.org/record/1436684/files/figure.png" httpUri-7="https://zenodo.org/record/1436686/files/figure.png" httpUri-8="https://zenodo.org/record/1436688/files/figure.png" pageId="1" pageNumber="116">Figs 1–58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36745A4FFEE686525FFFCD496B8FCF4" box="[151,666,797,819]" pageId="1" pageNumber="116" type="materials_examined">
<paragraph id="8BC2162FFFEE686525FFFCD496B8FCF4" blockId="1.[151,666,797,819]" box="[151,666,797,819]" pageId="1" pageNumber="116">
Type species. 
<taxonomicName id="4C7D6DACFFEE6865244DFCD796B4FCF4" authority="KOpelke 1980" authorityName="Kopelke" authorityYear="1980" box="[293,662,797,819]" class="Insecta" family="Baetidae" genus="Centroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="B909CA3DFFEE6865244DFCD795DCFCF4" box="[293,510,797,819]" italics="true" pageId="1" pageNumber="116">Centroptilum badium</emphasis>
KOpelke 1980
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36745A4FFEE686525FFFC959522FB27" pageId="1" pageNumber="116" type="etymology">
<paragraph id="8BC2162FFFEE686525FFFC959798FC5A" blockId="1.[151,1437,863,1429]" pageId="1" pageNumber="116">
<emphasis id="B909CA3DFFEE686525FFFC95953CFCBF" bold="true" box="[151,286,863,888]" pageId="1" pageNumber="116">Etymology.</emphasis>
Allusion to the single point of the costal process of hind wing; antonym to the generic name « 
<taxonomicName id="4C7D6DACFFEE686525CDFC4E956DFC5A" authorityName="Wuillot &amp; Gillies" authorityYear="1994" box="[165,335,900,925]" class="Insecta" family="Baetidae" genus="Dicentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE686525CDFC4E956DFC5A" box="[165,335,900,925]" italics="true" pageId="1" pageNumber="116">Dicentroptilum</emphasis>
</taxonomicName>
», whose costal process has two points. Neutral gender.
</paragraph>
<paragraph id="8BC2162FFFEE686525AFFC6D9522FB27" blockId="1.[151,1437,863,1429]" pageId="1" pageNumber="116">
<emphasis id="B909CA3DFFEE686525AFFC6D9599FC07" bold="true" box="[199,443,935,960]" pageId="1" pageNumber="116">Systematic position.</emphasis>
<taxonomicName id="4C7D6DACFFEE686524A0FC6D96B5FC07" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[456,663,935,960]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE686524A0FC6D96B5FC07" box="[456,663,935,960]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
belongs to the plesiomorphon Protopatellata 
<bibRefCitation id="EFEC6BDEFFEE686521D1FC6D94F3FC23" pageId="1" pageNumber="116" refId="ref4813" refString="Kluge, N. J. &amp; NOvikOva, E. A. (2011) Systematics Of the mayfly taxOn Acentrella (EphemerOptera, Baetidae), with descriptiOn Of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article">Kluge &amp; Novikova 2011</bibRefCitation>
; its leg structure is typical for Protopatellata: in larva, patella-tibial suture is present on middle and hind legs (
<figureCitation id="13460AAAFFEE686525F7FC2594D1FBCF" box="[159,243,1007,1032]" captionStart="FIGURES 10–27" captionStartId="4.[151,264,1871,1894]" captionTargetBox="[174,1415,202,1856]" captionTargetId="figure@4.[170,1417,195,1857]" captionTargetPageId="4" captionText="FIGURES 10–27. Larva Of Monocentroptilum badium. 10–16, tergalii Of I–VII pairs; 17, margin Of tergalius; 18, fOre leg; 19, claw (view frOm anteriOr side); 20, hind leg; 21, setae On apex Of its tibia; 22, claw (view frOm inner side); 23, exuviae Of half Of prOnOtum and mesOnOtum (thick line shOws area enlarged On Fig. 28); 24, setae On mesOnOtum; 25, seta On paired prOtuberances; 26, hind prOtOpterOm (the same scale as Fig. 23); 27, part Of head with clypeus, bases Of antennae, median Ocellus and frOntal suture." httpUri="https://zenodo.org/record/1436676/files/figure.png" pageId="1" pageNumber="116">Fig. 20</figureCitation>
), but absent on fore leg (
<figureCitation id="13460AAAFFEE68652779FC259647FBCF" box="[529,613,1007,1032]" captionStart="FIGURES 10–27" captionStartId="4.[151,264,1871,1894]" captionTargetBox="[174,1415,202,1856]" captionTargetId="figure@4.[170,1417,195,1857]" captionTargetPageId="4" captionText="FIGURES 10–27. Larva Of Monocentroptilum badium. 10–16, tergalii Of I–VII pairs; 17, margin Of tergalius; 18, fOre leg; 19, claw (view frOm anteriOr side); 20, hind leg; 21, setae On apex Of its tibia; 22, claw (view frOm inner side); 23, exuviae Of half Of prOnOtum and mesOnOtum (thick line shOws area enlarged On Fig. 28); 24, setae On mesOnOtum; 25, seta On paired prOtuberances; 26, hind prOtOpterOm (the same scale as Fig. 23); 27, part Of head with clypeus, bases Of antennae, median Ocellus and frOntal suture." httpUri="https://zenodo.org/record/1436676/files/figure.png" pageId="1" pageNumber="116">Fig. 18</figureCitation>
); in female subimago and imago it is also expressed on middle and hind legs (
<figureCitation id="13460AAAFFEE686525BDFBDE9508FBEB" box="[213,298,1044,1069]" captionStart="FIGURES 32–39" captionStartId="6.[151,264,1926,1949]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." httpUri="https://zenodo.org/record/1436680/files/figure.png" pageId="1" pageNumber="116">Fig. 38</figureCitation>
), but absent on fore leg (
<figureCitation id="13460AAAFFEE68652721FBDE96BEFBEB" box="[585,668,1044,1069]" captionStart="FIGURES 32–39" captionStartId="6.[151,264,1926,1949]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." httpUri="https://zenodo.org/record/1436680/files/figure.png" pageId="1" pageNumber="116">Fig. 37</figureCitation>
) (similarly to male subimago and imago of this taxon and all other 
<taxonomicName id="4C7D6DACFFEE686525FFFBFD94DCFB97" box="[151,254,1079,1104]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
). As well as other Protopatellata, 
<taxonomicName id="4C7D6DACFFEE68652711FBFD976AFB97" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[633,840,1079,1104]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE68652711FBFD976AFB97" box="[633,840,1079,1104]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
has following plesiomorphies: fore wing with single marginal intercalary per space (
<figureCitation id="13460AAAFFEE68652491FB969669FBB3" box="[505,587,1116,1141]" captionStart="FIGURES 40–43" captionStartId="7.[151,264,1072,1095]" captionTargetBox="[151,1423,193,1034]" captionTargetId="figure@7.[151,1436,193,1052]" captionTargetPageId="7" captionText="FIGURES 40–43. Monocentroptilum badium. 40–41, fOre and hind wings; 42, head and mesOnOtum Of male imagO; 43, subimaginal exuviae Of left pOstsubalar sclerite and laterOpOstnOtal crest." httpUri="https://zenodo.org/record/1436682/files/figure.png" pageId="1" pageNumber="116">Fig. 40</figureCitation>
) (in contrast to Baetovectata); larval femur without villopore (in contrast to Baetofemorata); subimaginal tarsi are covered with pointed microlepides (
<figureCitation id="13460AAAFFEE6865269AFBB5906BFB5F" box="[1010,1097,1151,1176]" captionStart="FIGURES 32–39" captionStartId="6.[151,264,1926,1949]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." httpUri="https://zenodo.org/record/1436680/files/figure.png" pageId="1" pageNumber="116">Fig. 39</figureCitation>
); subimaginal gonostyli buds under cuticle of mature larva are curved by the « 
<taxonomicName id="4C7D6DACFFEE686527DAFB6E9721FB7A" authorityName="Leach" authorityYear="1815" box="[690,771,1188,1213]" class="Insecta" family="Baetidae" genus="Cloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE686527DAFB6E9721FB7A" box="[690,771,1188,1213]" italics="true" pageId="1" pageNumber="116">Cloeon</emphasis>
</taxonomicName>
- 
<typeStatus id="54C6A88DFFEE68652663FB6F9762FB7A" box="[779,832,1189,1213]" pageId="1" pageNumber="116">type</typeStatus>
», i.e. with 2nd (the longest) segment directed laterally (
<figureCitation id="13460AAAFFEE686525F7FB0D94D6FB27" box="[159,244,1223,1248]" captionStart="FIGURES 44–47" captionStartId="9.[151,264,1870,1893]" captionTargetBox="[183,1400,201,1847]" captionTargetId="figure@9.[183,1406,193,1854]" captionTargetPageId="9" captionText="FIGURES 44–47. Male genitals Of Monocentroptilum badium. 44, develOping subimaginal genitals, fOlded under larval cuticle; 45, subimaginal exuviae Of gOnOstyli; 46, imaginal genitals Of the same specimen (gOnOduct shOwn by dOtted lines, muscles shOwn by interrupted lines, internal parts Of penis shOwn by interrupted lines and dOtted); 47, its left unistyliger, tO shOw muscles. AbbreviatiOns: 1–3, segments Of gOnOstylus; gv, gOnOvectis; m.sg, median sternO-styligeral muscle; pgs, larval prOtOgOnOstylus; usg, unistyliger." httpUri="https://zenodo.org/record/1436684/files/figure.png" pageId="1" pageNumber="116">Fig. 44</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36745A4FFEE686525AFFB2696DCF84A" pageId="1" pageNumber="116" type="diagnosis">
<paragraph id="8BC2162FFFEE686525AFFB269594FAEF" blockId="1.[151,1437,863,1429]" pageId="1" pageNumber="116">
<emphasis id="B909CA3DFFEE686525AFFB269563FAC2" bold="true" box="[199,321,1260,1285]" pageId="1" pageNumber="116">Diagnosis.</emphasis>
<taxonomicName id="4C7D6DACFFEE68652423FB269638FAC2" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[331,538,1260,1285]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE68652423FB269638FAC2" box="[331,538,1260,1285]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
can be distinguished from other taxa of Protopatellata by combination of the following two characters:
</paragraph>
<paragraph id="8BC2162FFFEE686525D4FAFE9128FAB7" blockId="1.[151,1437,863,1429]" pageId="1" pageNumber="116">
(1)Labial palp of the « 
<taxonomicName id="4C7D6DACFFEE686524A4FAFE9666FA8A" authorityName="Gillies" authorityYear="1990" box="[460,580,1332,1357]" class="Insecta" family="Baetidae" genus="Afroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE686524A4FAFE9666FA8A" box="[460,580,1332,1357]" italics="true" pageId="1" pageNumber="116">Afroptilum</emphasis>
</taxonomicName>
-type»: its 2nd segment is long and widened distally, forming a rounded convexity on median side close to base of 3rd segment; 3rd segment small, narrowed distally (
<figureCitation id="13460AAAFFEE686521D0FA9D90DEFAB7" box="[1208,1276,1367,1392]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[181,1415,663,1951]" captionTargetId="figure@3.[174,1423,628,1959]" captionTargetPageId="3" captionText="FIGURES 5–9. Larva Of Monocentroptilum badium. 5, labium; 6, maxilla (dOrsal view); 7, labium (at left ventral view, at right dOrsal view); 8–9, left and right mandible. AbbreviatiOns: ds1–ds3, dentisetae." httpUri="https://zenodo.org/record/1436674/files/figure.png" pageId="1" pageNumber="116">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BC2162FFFEE686525D4FAB6901FFA53" blockId="1.[151,1437,863,1429]" box="[188,1085,1404,1429]" pageId="1" pageNumber="116">
(2)Costal projection of hind wing is hooked and pointed, not bifurcate (
<figureCitation id="13460AAAFFEE686526B5FAB69012FA53" box="[989,1072,1404,1429]" captionStart="FIGURES 40–43" captionStartId="7.[151,264,1072,1095]" captionTargetBox="[151,1423,193,1034]" captionTargetId="figure@7.[151,1436,193,1052]" captionTargetPageId="7" captionText="FIGURES 40–43. Monocentroptilum badium. 40–41, fOre and hind wings; 42, head and mesOnOtum Of male imagO; 43, subimaginal exuviae Of left pOstsubalar sclerite and laterOpOstnOtal crest." httpUri="https://zenodo.org/record/1436682/files/figure.png" pageId="1" pageNumber="116">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="8BC2162FFFEE686525D4FA0E913FFA1A" blockId="1.[151,1436,1476,1573]" box="[188,1309,1476,1501]" pageId="1" pageNumber="116">
The single species of 
<taxonomicName id="4C7D6DACFFEE686524C7FA0E965CFA1A" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[431,638,1476,1501]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE686524C7FA0E965CFA1A" box="[431,638,1476,1501]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
has unique autapomorphies, among which is the following:
</paragraph>
<paragraph id="8BC2162FFFEE686525D4FA22950CF9E3" blockId="1.[151,1436,1476,1573]" pageId="1" pageNumber="116">
(3)On femur of each larval leg, most distal setae of the row on outer margin form 2 groups, with 2 setae in each (
<figureCitation id="13460AAAFFEE686525F7F9C69503F9E3" box="[159,289,1548,1573]" captionStart="FIGURES 10–27" captionStartId="4.[151,264,1871,1894]" captionTargetBox="[174,1415,202,1856]" captionTargetId="figure@4.[170,1417,195,1857]" captionTargetPageId="4" captionText="FIGURES 10–27. Larva Of Monocentroptilum badium. 10–16, tergalii Of I–VII pairs; 17, margin Of tergalius; 18, fOre leg; 19, claw (view frOm anteriOr side); 20, hind leg; 21, setae On apex Of its tibia; 22, claw (view frOm inner side); 23, exuviae Of half Of prOnOtum and mesOnOtum (thick line shOws area enlarged On Fig. 28); 24, setae On mesOnOtum; 25, seta On paired prOtuberances; 26, hind prOtOpterOm (the same scale as Fig. 23); 27, part Of head with clypeus, bases Of antennae, median Ocellus and frOntal suture." httpUri="https://zenodo.org/record/1436676/files/figure.png" pageId="1" pageNumber="116">Figs 18, 20</figureCitation>
).
</paragraph>
<paragraph id="8BC2162FFFEE686525AFF99E94FFF973" blockId="1.[151,1437,1620,2005]" pageId="1" pageNumber="116">
Other characters are described bellow, in the species description. As only one species of 
<taxonomicName id="4C7D6DACFFEE686521A4F99E91B9F9AA" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[1228,1435,1620,1645]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE686521A4F99E91B9F9AA" box="[1228,1435,1620,1645]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFEE686525FFF9BD94CAF957" box="[151,232,1655,1680]" pageId="1" pageNumber="116" rank="genus">
<emphasis id="B909CA3DFFEE686525FFF9BD94CAF957" bold="true" box="[151,232,1655,1680]" pageId="1" pageNumber="116">gen. n.</emphasis>
</taxonomicNameLabel>
is known, it is unclear, which of its characters are species-specific, and which belong to a supra-species taxon.
</paragraph>
<paragraph id="8BC2162FFFEE686525AFF97596DCF84A" blockId="1.[151,1437,1620,2005]" pageId="1" pageNumber="116">
<emphasis id="B909CA3DFFEE686525AFF9759543F91F" bold="true" box="[199,353,1727,1752]" pageId="1" pageNumber="116">Comparison.</emphasis>
Labial palp of the « 
<taxonomicName id="4C7D6DACFFEE6865271DF97596CFF91F" authorityName="Gillies" authorityYear="1990" box="[629,749,1727,1752]" class="Insecta" family="Baetidae" genus="Afroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE6865271DF97596CFF91F" box="[629,749,1727,1752]" italics="true" pageId="1" pageNumber="116">Afroptilum</emphasis>
</taxonomicName>
- 
<typeStatus id="54C6A88DFFEE6865279DF90B9708F91E" box="[757,810,1729,1753]" pageId="1" pageNumber="116">type</typeStatus>
», besides 
<taxonomicName id="4C7D6DACFFEE686526D8F975905DF91F" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[944,1151,1727,1752]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEE686526D8F975905DF91F" box="[944,1151,1727,1752]" italics="true" pageId="1" pageNumber="116">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFEE686521F8F97590CBF91F" box="[1168,1257,1727,1752]" pageId="1" pageNumber="116" rank="genus">
<emphasis id="B909CA3DFFEE686521F8F97590CBF91F" bold="true" box="[1168,1257,1727,1752]" pageId="1" pageNumber="116">gen. n.</emphasis>
</taxonomicNameLabel>
, is present in 
<taxonomicName id="4C7D6DACFFEE686525FFF92E952DF93A" authorityName="Gillies" authorityYear="1990" box="[151,271,1764,1789]" class="Insecta" family="Baetidae" genus="Afroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE686525FFF92E952DF93A" box="[151,271,1764,1789]" italics="true" pageId="1" pageNumber="116">Afroptilum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFEE68652477F92E96F9F93B" authority="Wuillot &amp; Gillies 1994" authorityName="Wuillot &amp; Gillies" authorityYear="1994" box="[287,731,1764,1789]" class="Insecta" family="Baetidae" genus="Dicentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE68652477F92E95EBF93A" box="[287,457,1764,1789]" italics="true" pageId="1" pageNumber="116">Dicentroptilum</emphasis>
Wuillot &amp; Gillies 1994
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFEE68652780F92E90DCF93A" authority="Lugo-Ortiz &amp; McCafferty 1997 b" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" box="[744,1278,1764,1789]" class="Insecta" family="Baetidae" genus="Xyrodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE68652780F92E975FF93A" box="[744,893,1764,1789]" italics="true" pageId="1" pageNumber="116">Xyrodromeus</emphasis>
Lugo-Ortiz &amp; McCafferty 1997b
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFEE68652063F92E9511F8E7" authority="Gillies 1990 a" authorityName="Gillies" authorityYear="1990" class="Insecta" family="Baetidae" genus="Afroptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE68652063F92E91BEF93A" box="[1291,1436,1764,1789]" italics="true" pageId="1" pageNumber="116">Afroptiloides</emphasis>
<bibRefCitation id="EFEC6BDEFFEE686525FFF8CD9511F8E7" box="[151,307,1799,1824]" pageId="1" pageNumber="116" refId="ref4206" refString="Gillies, M. T. (1990 a) A revisiOn Of the African species Of Centroptilum EatOn (Baetidae, EphemerOptera). Aquatic Insects, 12 (2), 97 - 128. https: // dOi. Org / 10.1080 / 01650429009361395" type="journal article">Gillies 1990a</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFEE6865242AF8CD96C3F8E7" authority="Gillies &amp; Wuillot 1997" authorityName="Gillies &amp; Wuillot" authorityYear="1997" box="[322,737,1799,1824]" class="Insecta" family="Baetidae" genus="Platycloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE6865242AF8CD95EAF8E7" box="[322,456,1799,1824]" italics="true" pageId="1" pageNumber="116">Platycloeon</emphasis>
<bibRefCitation id="EFEC6BDEFFEE686524BAF8CD96C3F8E7" box="[466,737,1799,1824]" pageId="1" pageNumber="116" refId="ref4291" refString="Gillies, M. T. &amp; WuillOt, J. (1997) Platycloeon, a new genus Of Baetidae (EphemerOptera) frOm East Africa. Aquatic Insects, 19, 185 - 189. https: // dOi. Org / 10.1080 / 01650429709361654" type="journal article">Gillies &amp; Wuillot 1997</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFEE68652798F8CD90FDF8E7" authority="Lugo-Ortiz &amp; McCafferty 1998 a" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[752,1247,1799,1824]" class="Insecta" family="Baetidae" genus="Peuhlella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEE68652798F8CD9779F8E7" box="[752,859,1799,1824]" italics="true" pageId="1" pageNumber="116">Peuhlella</emphasis>
Lugo-Ortiz &amp; McCafferty 1998a
</taxonomicName>
and some other taxa; probably, such shape of labial palp is initial for 
<taxonomicName id="4C7D6DACFFEE68652781F8E6976DF882" box="[745,847,1836,1861]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
. It differs both from the «chelate» 
<typeStatus id="54C6A88DFFEE686521B9F8E79123F882" box="[1233,1281,1837,1861]" pageId="1" pageNumber="116">type</typeStatus>
of labial palp (i.e. with prominent medio-apical projection of 2nd segment), and from labial palp with widened 3rd segment, which repeatedly appeared in various taxa of 
<taxonomicName id="4C7D6DACFFEE686527FCF8BE96D8F84A" box="[660,762,1908,1933]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36745A4FFEE686625AFF853905AFDA7" lastPageId="2" lastPageNumber="117" pageId="1" pageNumber="116" type="reference_group">
<paragraph id="8BC2162FFFEE686625AFF8539159FEDA" blockId="1.[151,1437,1620,2005]" lastBlockId="2.[151,1436,151,824]" lastPageId="2" lastPageNumber="117" pageId="1" pageNumber="116">
In 
<taxonomicName id="4C7D6DACFFEE6865258DF8529568F876" ID-CoL="6ZG" box="[229,330,1944,1969]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
, costal projection of hind wing either has two points, or has one point, or is blunt without points, or is absent; in many baetid taxa hind wings are absent at all. Among Anteropatellata, two-pointed costal projection is not found, and one-pointed is usual. Among Protopatellata, two-pointed costal projection occurs in 
<taxonomicName id="4C7D6DACFFED68662074FF5D91B6FF77" ID-CoL="QCK" authorityName="Gillies" authorityYear="1990" box="[1308,1428,151,176]" class="Insecta" family="Baetidae" genus="Afroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662074FF5D91B6FF77" box="[1308,1428,151,176]" italics="true" pageId="2" pageNumber="117">Afroptilum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED686625FFFF769563FF12" ID-CoL="44HT" authorityName="Wuillot &amp; Gillies" authorityYear="1994" box="[151,321,188,213]" class="Insecta" family="Baetidae" genus="Dicentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686625FFFF769563FF12" box="[151,321,188,213]" italics="true" pageId="2" pageNumber="117">Dicentroptilum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED6866243EFF7695C9FF12" ID-CoL="89MX" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" box="[342,491,188,213]" class="Insecta" family="Baetidae" genus="Xyrodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866243EFF7695C9FF12" box="[342,491,188,213]" italics="true" pageId="2" pageNumber="117">Xyrodromeus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED68662769FF7696B0FF12" authorityName="Gillies" authorityYear="1990" box="[513,658,188,213]" class="Insecta" family="Baetidae" genus="Afroptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662769FF7696B0FF12" box="[513,658,188,213]" italics="true" pageId="2" pageNumber="117">Afroptiloides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED686627CFFF76970FFF12" authorityName="Gillies &amp; Wuillot" authorityYear="1997" box="[679,813,188,213]" class="Insecta" family="Baetidae" genus="Platycloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686627CFFF76970FFF12" box="[679,813,188,213]" italics="true" pageId="2" pageNumber="117">Platycloeon</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED6866262BFF7690FEFF13" ID-CoL="62ZZJ" authority="McCafferty et al. 1997" authorityName="McCafferty et al." authorityYear="1997" box="[835,1244,188,213]" class="Insecta" family="Baetidae" genus="Micksiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866262BFF769797FF12" box="[835,949,188,213]" italics="true" pageId="2" pageNumber="117">Micksiops</emphasis>
McCafferty 
<emphasis id="B909CA3DFFED6866213BFF7790AEFF12" box="[1107,1164,188,213]" italics="true" pageId="2" pageNumber="117">et al</emphasis>
. 1997
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED68662187FF76950DFF3F" ID-CoL="62JV6" authority="Lestage 1918" authorityName="Lestage" authorityYear="1918" class="Insecta" family="Baetidae" genus="Centroptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662187FF7691BEFF12" box="[1263,1436,188,213]" italics="true" pageId="2" pageNumber="117">Centroptiloides</emphasis>
<bibRefCitation id="EFEC6BDEFFED686625FFFF2A950DFF3F" box="[151,303,224,248]" pageId="2" pageNumber="117" refId="ref5015" refString="Lestage, J. - A. (1918) Les Ephemeres d'Afrique (NOtes critiques sur les especes cOnnues). Revue Zoologique Africaine, 6, 65 - 114. https: // dOi. Org / 10.5962 / bhl. part. 19452" type="journal article">Lestage 1918</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7D6DACFFED68662403FF1590F1FF3F" ID-CoL="4VRP" authority="McCafferty &amp; Lugo-Ortiz (in Lugo-Ortiz &amp; McCafferty)" authorityName="McCafferty &amp; Lugo-Ortiz (in Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[363,1235,223,248]" class="Insecta" family="Baetidae" genus="Herbrossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662403FF1595C9FF3F" box="[363,491,223,248]" italics="true" pageId="2" pageNumber="117">Herbrossus</emphasis>
McCafferty &amp; Lugo-Ortiz (in Lugo-Ortiz &amp; McCafferty) 1998a
</taxonomicName>
; according to my unpublished data, costal projection of 
<taxonomicName id="4C7D6DACFFED6866272AFECE968FFEDA" ID-CoL="6M7T" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[578,685,260,285]" class="Insecta" family="Baetidae" genus="Peuhlella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866272AFECE968FFEDA" box="[578,685,260,285]" italics="true" pageId="2" pageNumber="117">Peuhlella</emphasis>
</taxonomicName>
is also two-pointed, while it was described as having one point.
</paragraph>
<paragraph id="8BC2162FFFED686625AFFEE2905AFDA7" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="117">
Among Protopatellata, one-pointed costal projection is present in 
<taxonomicName id="4C7D6DACFFED686626DBFEED9013FE87" ID-CoL="3V95" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[947,1073,295,320]" class="Insecta" family="Baetidae" genus="Crassabwa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">Crassabwa</taxonomicName>
/g1 (incl. 
<taxonomicName id="4C7D6DACFFED686621F3FEE290FCFEF8" ID-CoL="7QGR" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[1179,1246,296,319]" class="Insecta" family="Baetidae" genus="Susua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686621F3FEE290FCFEF8" box="[1179,1246,296,319]" italics="true" pageId="2" pageNumber="117">Susua</emphasis>
</taxonomicName>
), 
<taxonomicName id="4C7D6DACFFED6866219FFEED9600FEA2" ID-CoL="3Z86" authority="Lugo-Ortiz &amp; McCafferty 1996 a" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" class="Insecta" family="Baetidae" genus="Dabulamanzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866219FFEED91BEFE87" box="[1271,1436,295,320]" italics="true" pageId="2" pageNumber="117">Dabulamanzia</emphasis>
Lugo-Ortiz &amp; McCafferty 1996a
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED6866275EFE869758FEA3" ID-CoL="78J5" authority="Gattolliat 2001" authorityName="Gattolliat" authorityYear="2001" box="[566,890,332,357]" class="Insecta" family="Baetidae" genus="Rheoptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866275EFE869695FEA2" box="[566,695,332,357]" italics="true" pageId="2" pageNumber="117">Rheoptilum</emphasis>
<bibRefCitation id="EFEC6BDEFFED686627AEFE869758FEA3" box="[710,890,332,357]" pageId="2" pageNumber="117" refId="ref4055" refString="GattOlliat, J. - L. (2001) Rheoptilum: A new genus Of twO-tailed baetidae (EphemerOptera) frOm Madagascar. Aquatic Insects, 23, 67 - 81. https: // dOi. Org / 10.1076 / aqin. 23.1.67.4932" type="journal article">Gattolliat 2001</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED686626E6FE8690D2FEA3" ID-CoL="63KP" authority="Demoulin 1973" authorityName="Demoulin" authorityYear="1973" box="[910,1264,332,357]" class="Insecta" family="Baetidae" genus="Nesoptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686626E6FE869004FEA2" box="[910,1062,332,357]" italics="true" pageId="2" pageNumber="117">Nesoptiloides</emphasis>
<bibRefCitation id="EFEC6BDEFFED6866215EFE8690D2FEA3" box="[1078,1264,332,357]" pageId="2" pageNumber="117" refId="ref3994" refString="DemOulin, G. (1973) EphemerOpteres de Madagascar. III. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 49 (7), 1 - 20." type="journal article">Demoulin 1973</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED6866206CFE8695E0FE4F" ID-CoL="4RK2" authority="Gattolliat &amp; Sartori 2000" authorityName="Gattolliat &amp; Sartori" authorityYear="2000" class="Insecta" family="Baetidae" genus="Guloptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866206CFE8691BEFEA2" box="[1284,1436,332,357]" italics="true" pageId="2" pageNumber="117">Guloptiloides</emphasis>
<bibRefCitation id="EFEC6BDEFFED686625FFFEA595E0FE4F" box="[151,450,367,392]" pageId="2" pageNumber="117" refId="ref4144" refString="GattOlliat, J. - L. &amp; SartOri, M. (2000) Guloptiloides: an extraOrdinary new carnivOrOus genus Of Baetidae (EphemerOptera). Aquatic Insects, 22, 148 - 159." type="journal article">Gattolliat &amp; Sartori 2000</bibRefCitation>
</taxonomicName>
(if association of its larvae and imagoes is correct) and in that representatives of 
<taxonomicName id="4C7D6DACFFED686625FFFE5E95E7FE6B" ID-CoL="62BNC" authority="Kluge 2012 a" authorityName="Kluge" authorityYear="2012" box="[151,453,404,429]" class="Insecta" family="Baetidae" genus="Anafroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686625FFFE5E9509FE6A" box="[151,299,404,429]" italics="true" pageId="2" pageNumber="117">Anafroptilum</emphasis>
<bibRefCitation id="EFEC6BDEFFED6866245CFE5E95E7FE6B" box="[308,453,404,429]" pageId="2" pageNumber="117" refId="ref4569" refString="Kluge, N. J. (2012 a) NOn-African representatives Of the plesiOmOrphOn PrOtOpatellata (EphemerOptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article">Kluge 2012a</bibRefCitation>
</taxonomicName>
and Rhithrocloeoninae 
<bibRefCitation id="EFEC6BDEFFED686627BFFE5E974FFE6A" box="[727,877,404,429]" pageId="2" pageNumber="117" refId="ref4604" refString="Kluge, N. J. (2012 b) Systematics Of RhithrOclOeOninae with new species frOm Uganda. Russian Entomological Journal, 21 (1), 1 - 13." type="journal article">Kluge 2012b</bibRefCitation>
, which retain hind wings. These taxa differ from the new genus 
<taxonomicName id="4C7D6DACFFED68662429FE7D9632FE17" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[321,528,439,464]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFED68662429FE7D9632FE17" box="[321,528,439,464]" italics="true" pageId="2" pageNumber="117">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFED68662771FE7D964BFE17" box="[537,617,439,464]" pageId="2" pageNumber="117" rank="genus">
<emphasis id="B909CA3DFFED68662771FE7D964BFE17" bold="true" box="[537,617,439,464]" pageId="2" pageNumber="117">gen. n.</emphasis>
</taxonomicNameLabel>
at least by the following characters: 
<taxonomicName id="4C7D6DACFFED68662165FE7D90A8FE17" ID-CoL="3V95" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[1037,1162,439,464]" class="Insecta" family="Baetidae" genus="Crassabwa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">Crassabwa</taxonomicName>
/fg1 differs by «chelate» labial palp and single subapical seta on larval femur (
<bibRefCitation id="EFEC6BDEFFED68662782FE169791FE33" box="[746,947,476,501]" pageId="2" pageNumber="117" refId="ref4689" refString="Kluge, N. J., GattOlliat, J. - L. &amp; Salles, F. F. (2017) RedescriptiOn Of the AfrOtrOpical genus Crassabwa LugO-OrtiZ &amp; McCafferty 1996 (EphemerOptera: Baetidae: PrOtOpatellata). Zootaxa, 4350 (3), 401 - 435. https: // dOi. Org / 10.11646 / ZOOtaxa. 4350.3.1" type="journal article">
Kluge 
<emphasis id="B909CA3DFFED6866265FFE17974AFE32" box="[823,872,476,501]" italics="true" pageId="2" pageNumber="117">et al</emphasis>
. 2017
</bibRefCitation>
). 
<taxonomicName id="4C7D6DACFFED686626ADFE169048FE32" ID-CoL="3Z86" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[965,1130,476,501]" class="Insecta" family="Baetidae" genus="Dabulamanzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686626ADFE169048FE32" box="[965,1130,476,501]" italics="true" pageId="2" pageNumber="117">Dabulamanzia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7D6DACFFED686621C8FE169117FE32" ID-CoL="62BNC" authorityName="Kluge" authorityYear="2012" box="[1184,1333,476,501]" class="Insecta" family="Baetidae" genus="Anafroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686621C8FE169117FE32" box="[1184,1333,476,501]" italics="true" pageId="2" pageNumber="117">Anafroptilum</emphasis>
</taxonomicName>
differ by widened 3rd segment of labial palp. Rhithrocloeoninae differ from all other taxa by peculiar structure of gonostyli and mode of their folding in larva (
<bibRefCitation id="EFEC6BDEFFED6866275BFDEE96E2FDFA" box="[563,704,548,573]" pageId="2" pageNumber="117" refId="ref4636" refString="Kluge, N. J. (2015) First descriptiOn Of winged stages Of Thraulobaetodes ElOuard &amp; Hideux 1991 and reclassificatiOn Of RhithrOclOeOninae (EphemerOptera, Baetidae). Zootaxa, 3949 (4), 491 - 514. https: // dOi. Org / 10.11646 / ZOOtaxa. 3949.4.2" type="journal article">Kluge 2015</bibRefCitation>
). Larvae of 
<taxonomicName id="4C7D6DACFFED68662625FDEE97ECFDFA" ID-CoL="78J5" authorityName="Gattolliat" authorityYear="2001" box="[845,974,548,573]" class="Insecta" family="Baetidae" genus="Rheoptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662625FDEE97ECFDFA" box="[845,974,548,573]" italics="true" pageId="2" pageNumber="117">Rheoptilum</emphasis>
</taxonomicName>
are highly modified as the « 
<taxonomicName id="4C7D6DACFFED68662077FDEE91B1FDFA" ID-CoL="MKW" authorityName="Bengtsson" authorityYear="1912" box="[1311,1427,548,573]" class="Insecta" family="Baetidae" genus="Acentrella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662077FDEE91B1FDFA" box="[1311,1427,548,573]" italics="true" pageId="2" pageNumber="117">Acentrella</emphasis>
</taxonomicName>
- 
<typeStatus id="54C6A88DFFED686625FFFD8294E9FDA7" box="[151,203,584,608]" pageId="2" pageNumber="117">type</typeStatus>
», and larvae of 
<taxonomicName id="4C7D6DACFFED68662414FD8D9636FDA7" ID-CoL="63KP" authorityName="Demoulin" authorityYear="1973" box="[380,532,583,608]" class="Insecta" family="Baetidae" genus="Nesoptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662414FD8D9636FDA7" box="[380,532,583,608]" italics="true" pageId="2" pageNumber="117">Nesoptiloides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7D6DACFFED68662723FD8D96C6FDA7" ID-CoL="4RK2" authorityName="Gattolliat &amp; Sartori" authorityYear="2000" box="[587,740,583,608]" class="Insecta" family="Baetidae" genus="Guloptiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662723FD8D96C6FDA7" box="[587,740,583,608]" italics="true" pageId="2" pageNumber="117">Guloptiloides</emphasis>
</taxonomicName>
are highly modified as carnivorous.
</paragraph>
</subSubSection>
<subSubSection id="C36745A4FFED686725AFFDA6908FFE32" lastPageId="3" lastPageNumber="118" pageId="2" pageNumber="117" type="description">
<paragraph id="8BC2162FFFED686625AFFDA696F7FCFF" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="117">
Some taxa have no hind wings, and initial structure of their hind wings is unknown; among Protopatellata these are 
<taxonomicName id="4C7D6DACFFED686625AAFD45962AFD6F" authority="Gillies 1990 b" authorityName="Gillies" authorityYear="1990" box="[194,520,655,680]" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686625AAFD459546FD6F" box="[194,356,655,680]" italics="true" pageId="2" pageNumber="117">Potamocloeon</emphasis>
<bibRefCitation id="EFEC6BDEFFED68662406FD45962AFD6F" box="[366,520,655,680]" pageId="2" pageNumber="117" refId="ref4253" refString="Gillies, M. T. (1990 b) A new genus fOr the AfrOtrOpical mayfly, Cloeon dentatum Kimmins (Ephem., Baetidae). Entomologist's Monthly Magazine, 126, 207 - 208." type="journal article">Gillies 1990b</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="4C7D6DACFFED6866275CFD459036FD6F" authority="Lugo-Ortiz &amp; McCafferty 1997 c" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" box="[564,1044,655,680]" class="Insecta" family="Baetidae" genus="Maliqua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866275CFD4596B1FD6F" box="[564,659,655,680]" italics="true" pageId="2" pageNumber="117">Maliqua</emphasis>
Lugo-Ortiz &amp; McCafferty 1997c
</taxonomicName>
) and 
<taxonomicName id="4C7D6DACFFED6866213FFD4594F1FD0B" authority="Muller-Liebenau 1982" authorityName="Muller-Liebenau" authorityYear="1982" class="Insecta" family="Baetidae" genus="Indocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866213FFD4590F7FD6F" box="[1111,1237,655,680]" italics="true" pageId="2" pageNumber="117">Indocloeon</emphasis>
Müller-Liebenau 1982
</taxonomicName>
; the same in the genera of unknown systematic position 
<taxonomicName id="4C7D6DACFFED68662631FD7E905FFD0B" authority="Gillies 1990 a" authorityName="Gillies" authorityYear="1990" box="[857,1149,692,717]" class="Insecta" family="Baetidae" genus="Demoulinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662631FD7E97FFFD0A" box="[857,989,692,717]" italics="true" pageId="2" pageNumber="117">Demoulinia</emphasis>
<bibRefCitation id="EFEC6BDEFFED6866268EFD7E905FFD0B" box="[998,1149,692,717]" pageId="2" pageNumber="117" refId="ref4206" refString="Gillies, M. T. (1990 a) A revisiOn Of the African species Of Centroptilum EatOn (Baetidae, EphemerOptera). Aquatic Insects, 12 (2), 97 - 128. https: // dOi. Org / 10.1080 / 01650429009361395" type="journal article">Gillies 1990a</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7D6DACFFED686621DEFD7E916CFD0A" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" box="[1206,1358,692,717]" class="Insecta" family="Baetidae" genus="Edmulmeatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686621DEFD7E916CFD0A" box="[1206,1358,692,717]" italics="true" pageId="2" pageNumber="117">Edmulmeatus</emphasis>
</taxonomicName>
Lugo- Ortiz &amp; McCafferty 1997a. Among them, 
<taxonomicName id="4C7D6DACFFED68662714FD1D973CFD37" authorityName="Gillies" authorityYear="1990" box="[636,798,727,752]" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662714FD1D973CFD37" box="[636,798,727,752]" italics="true" pageId="2" pageNumber="117">Potamocloeon</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7D6DACFFED686626D0FD1D90A5FD37" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[952,1159,727,752]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFED686626D0FD1D90A5FD37" box="[952,1159,727,752]" italics="true" pageId="2" pageNumber="117">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFED686621F8FD1D90C3FD37" box="[1168,1249,727,752]" pageId="2" pageNumber="117" rank="genus">
<emphasis id="B909CA3DFFED686621F8FD1D90C3FD37" bold="true" box="[1168,1249,727,752]" pageId="2" pageNumber="117">gen. n.</emphasis>
</taxonomicNameLabel>
by widened 3rd segment of labial palp and other characters; 
<taxonomicName id="4C7D6DACFFED68662787FD36974FFCD2" authorityName="Muller-Liebenau" authorityYear="1982" box="[751,877,764,789]" class="Insecta" family="Baetidae" genus="Indocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED68662787FD36974FFCD2" box="[751,877,764,789]" italics="true" pageId="2" pageNumber="117">Indocloeon</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFED686626E3FD36902DFCD2" authorityName="Gillies" authorityYear="1990" box="[907,1039,764,789]" class="Insecta" family="Baetidae" genus="Demoulinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED686626E3FD36902DFCD2" box="[907,1039,764,789]" italics="true" pageId="2" pageNumber="117">Demoulinia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7D6DACFFED6866210BFD3690D9FCD2" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" box="[1123,1275,764,789]" class="Insecta" family="Baetidae" genus="Edmulmeatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFED6866210BFD3690D9FCD2" box="[1123,1275,764,789]" italics="true" pageId="2" pageNumber="117">Edmulmeatus</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="4C7D6DACFFED686625FFFCD59544FCFF" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[151,358,799,824]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFED686625FFFCD59544FCFF" box="[151,358,799,824]" italics="true" pageId="2" pageNumber="117">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFED68662405FCD5959EFCFF" box="[365,444,799,824]" pageId="2" pageNumber="117" rank="genus">
<emphasis id="B909CA3DFFED68662405FCD5959EFCFF" bold="true" box="[365,444,799,824]" pageId="2" pageNumber="117">gen. n.</emphasis>
</taxonomicNameLabel>
by «chelate» labial palp.
</paragraph>
<caption id="DF0246A7FFED686625FFF846968AF827" httpUri="https://zenodo.org/record/1436672/files/figure.png" pageId="2" pageNumber="117" startId="2.[151,264,1932,1955]" targetBox="[151,1436,879,1896]" targetPageId="2">
<paragraph id="8BC2162FFFED686625FFF846968AF827" blockId="2.[151,1436,1932,2016]" pageId="2" pageNumber="117">
<emphasis id="B909CA3DFFED686625FFF846951BF865" bold="true" box="[151,313,1932,1955]" pageId="2" pageNumber="117">FIGURES 1–4.</emphasis>
Larva 
<taxonomicName id="4C7D6DACFFED686624E9F847968BF864" authorityName="Kluge" authorityYear="2018" baseAuthorityName="Kopelke" baseAuthorityYear="1980" box="[385,681,1933,1955]" pageId="2" pageNumber="117" rank="subSpecies" subSpecies="badium">
Of 
<emphasis id="B909CA3DFFED686624F4F847968BF864" box="[412,681,1933,1955]" italics="true" pageId="2" pageNumber="117">Monocentroptilum badium</emphasis>
</taxonomicName>
. 1, Lateral view (arrOws shOw prOtuberances On mesOnOtum; 2, exuviae Of right half Of prOnOtum and mesOnOtum; 3, exuviae Of abdOminal sterna and terga; 4, subimaginal sterna and terga extracted frOm mature larva (tO shOw hypOdermal cOlOratiOn).
</paragraph>
</caption>
<paragraph id="8BC2162FFFEC686725AFFF52908FFE32" blockId="3.[151,1436,151,573]" pageId="3" pageNumber="118">
Some taxa of Protopatellata have hind wings, but are known only as larvae, so structure of their costal projection is unknown. These are 
<taxonomicName id="4C7D6DACFFEC68672777FF7697F4FF13" authority="Waltz &amp; McCafferty 1987" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[543,982,188,213]" class="Insecta" family="Baetidae" genus="Acanthiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC68672777FF7696BFFF12" box="[543,669,188,213]" italics="true" pageId="3" pageNumber="118">Acanthiops</emphasis>
Waltz &amp; McCafferty 1987
</taxonomicName>
, 
<taxonomicName id="4C7D6DACFFEC6867268CFF7694C2FF3F" authority="Lugo-Ortiz &amp; McCafferty 1998 b" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" class="Insecta" family="Baetidae" genus="Nesydemius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC6867268CFF769048FF12" box="[996,1130,188,213]" italics="true" pageId="3" pageNumber="118">Nesydemius</emphasis>
Lugo-Ortiz &amp; McCafferty 1998b
</taxonomicName>
, 
<emphasis id="B909CA3DFFEC68672583FF15955AFF3F" box="[235,376,223,248]" italics="true" pageId="3" pageNumber="118">Scutoptilium</emphasis>
<bibRefCitation id="EFEC6BDEFFEC686724E8FF15960BFF3F" box="[384,553,223,248]" pageId="3" pageNumber="118" refId="ref4100" refString="GattOlliat, J. - L. (2002) TwO new genera Of Baetidae (EphemerOptera; Insecta) frOm Madagascar. Aquatic Insects, 24, 143 - 159. https: // dOi. Org / 10.1076 / aqin. 24.2.143.4903" type="journal article">Gattolliat 2002</bibRefCitation>
and 
<taxonomicName id="4C7D6DACFFEC68672708FF1597A8FF3F" authority="Gattolliat 2002" authorityName="Gattolliat" authorityYear="2002" box="[608,906,223,248]" class="Insecta" family="Baetidae" genus="Echinopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC68672708FF1596F4FF3F" box="[608,726,223,248]" italics="true" pageId="3" pageNumber="118">Echinopus</emphasis>
<bibRefCitation id="EFEC6BDEFFEC686727B5FF1597A8FF3F" box="[733,906,223,248]" pageId="3" pageNumber="118" refId="ref4100" refString="GattOlliat, J. - L. (2002) TwO new genera Of Baetidae (EphemerOptera; Insecta) frOm Madagascar. Aquatic Insects, 24, 143 - 159. https: // dOi. Org / 10.1076 / aqin. 24.2.143.4903" type="journal article">Gattolliat 2002</bibRefCitation>
</taxonomicName>
. Among them, 
<taxonomicName id="4C7D6DACFFEC6867215EFF159096FF3F" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[1078,1204,223,248]" class="Insecta" family="Baetidae" genus="Acanthiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC6867215EFF159096FF3F" box="[1078,1204,223,248]" italics="true" pageId="3" pageNumber="118">Acanthiops</emphasis>
</taxonomicName>
has highly modified larval structure. 
<taxonomicName id="4C7D6DACFFEC68672439FECE95F4FEDA" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[337,470,260,285]" class="Insecta" family="Baetidae" genus="Nesydemius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC68672439FECE95F4FEDA" box="[337,470,260,285]" italics="true" pageId="3" pageNumber="118">Nesydemius</emphasis>
</taxonomicName>
seems to be closely related to 
<taxonomicName id="4C7D6DACFFEC68672651FECE97FCFEDA" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[825,990,260,285]" class="Insecta" family="Baetidae" genus="Dabulamanzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC68672651FECE97FCFEDA" box="[825,990,260,285]" italics="true" pageId="3" pageNumber="118">Dabulamanzia</emphasis>
</taxonomicName>
and differs from 
<taxonomicName id="4C7D6DACFFEC686721C0FECE9155FEDA" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[1192,1399,260,285]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEC686721C0FECE9155FEDA" box="[1192,1399,260,285]" italics="true" pageId="3" pageNumber="118">Monocentroptilum</emphasis>
</taxonomicName>
by widened 3rd segment of labial palp. The genera 
<emphasis id="B909CA3DFFEC686727BCFEED9740FE87" box="[724,866,295,320]" italics="true" pageId="3" pageNumber="118">Scutoptilium</emphasis>
and 
<taxonomicName id="4C7D6DACFFEC686726C9FEED9035FE87" authorityName="Gattolliat" authorityYear="2002" box="[929,1047,295,320]" class="Insecta" family="Baetidae" genus="Echinopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC686726C9FEED9035FE87" box="[929,1047,295,320]" italics="true" pageId="3" pageNumber="118">Echinopus</emphasis>
</taxonomicName>
, described from 
<collectingCountry id="F36A56BFFFEC68672189FEED9148FE87" box="[1249,1386,295,320]" name="Madagascar" pageId="3" pageNumber="118">Madagascar</collectingCountry>
, are regarded to be related one to another, based on presence of transverse subapical row of setae on their femora (
<bibRefCitation id="EFEC6BDEFFEC686725F6FEA59572FE4F" box="[158,336,367,392]" pageId="3" pageNumber="118" refId="ref4100" refString="GattOlliat, J. - L. (2002) TwO new genera Of Baetidae (EphemerOptera; Insecta) frOm Madagascar. Aquatic Insects, 24, 143 - 159. https: // dOi. Org / 10.1076 / aqin. 24.2.143.4903" type="journal article">Gattolliat 2002</bibRefCitation>
); in this respect they both differ from 
<taxonomicName id="4C7D6DACFFEC68672662FEA597FBFE4F" authority="Kluge, 2018" authorityName="Kluge" authorityYear="2018" box="[778,985,367,392]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909CA3DFFEC68672662FEA597FBFE4F" box="[778,985,367,392]" italics="true" pageId="3" pageNumber="118">Monocentroptilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFEC6867268BFEA59011FE4F" box="[995,1075,367,392]" pageId="3" pageNumber="118" rank="genus">
<emphasis id="B909CA3DFFEC6867268BFEA59011FE4F" bold="true" box="[995,1075,367,392]" pageId="3" pageNumber="118">gen. n.</emphasis>
</taxonomicNameLabel>
However, 
<taxonomicName id="4C7D6DACFFEC686721DBFEA5910BFE4F" authorityName="Gattolliat" authorityYear="2002" box="[1203,1321,367,392]" class="Insecta" family="Baetidae" genus="Echinopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B909CA3DFFEC686721DBFEA5910BFE4F" box="[1203,1321,367,392]" italics="true" pageId="3" pageNumber="118">Echinopus</emphasis>
</taxonomicName>
has some similarities with 
<taxonomicName id="4C7D6DACFFEC6867243DFE5E96A3FE6A" authorityName="Kluge" authorityYear="2018" baseAuthorityName="Kopelke" baseAuthorityYear="1980" box="[341,641,404,429]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="B909CA3DFFEC6867243DFE5E96A3FE6A" box="[341,641,404,429]" italics="true" pageId="3" pageNumber="118">Monocentroptilum badium</emphasis>
</taxonomicName>
: similar structure of mouth parts with elongated mandibles (
<bibRefCitation id="EFEC6BDEFFEC68672058FE5E9544FE17" pageId="3" pageNumber="118" refId="ref4100" refString="GattOlliat, J. - L. (2002) TwO new genera Of Baetidae (EphemerOptera; Insecta) frOm Madagascar. Aquatic Insects, 24, 143 - 159. https: // dOi. Org / 10.1076 / aqin. 24.2.143.4903" type="journal article">
Gattolliat 2002: 
<figureCitation id="13460AAAFFEC6867258AFE7D9544FE17" box="[226,358,439,464]" captionStart-0="FIGURES 28–31" captionStart-1="FIGURES 32–39" captionStartId-0="5.[151,264,1493,1516]" captionStartId-1="6.[151,264,1926,1949]" captionTargetBox-0="[151,1435,193,1485]" captionTargetBox-1="[151,1436,193,1903]" captionTargetId-0="figure@5.[151,1435,193,1485]" captionTargetId-1="figure@6.[151,1436,193,1906]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 28–31. Larval exuviae Of Monocentroptilum badium. 28, fragment Of mesOnOtum (Outlined On Fig. 23); 29, fragment Of cercus; 30, fragment Of tergum VIII; 31, paraprOct (all at the same scale)." captionText-1="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." httpUri-0="https://zenodo.org/record/1436678/files/figure.png" httpUri-1="https://zenodo.org/record/1436680/files/figure.png" pageId="3" pageNumber="118">Figs 28–32</figureCitation>
</bibRefCitation>
), claws with reduced posterior row of denticles and two subapical setae (ibid., 
<figureCitation id="13460AAAFFEC68672064FE7D9142FE17" box="[1292,1376,439,464]" captionStart="FIGURES 32–39" captionStartId="6.[151,264,1926,1949]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." httpUri="https://zenodo.org/record/1436680/files/figure.png" pageId="3" pageNumber="118">Fig. 34</figureCitation>
) and tergalii of the unusual «inverted» shape (ibid., 
<figureCitation id="13460AAAFFEC686727CAFE1696D0FE32" box="[674,754,476,501]" captionStart="FIGURES 32–39" captionStartId="6.[151,264,1926,1949]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 32–39. Monocentroptilum badium. 32, anteriOr part Of male imagO with subimaginal exuviae; 33, head and thOrax Of male imagO; 34, abdOmen Of male imagO (spread On slide); 35, had and thOrax Of female imagO; 36, female imagO; 37–38, exuviae Of fOre and hind legs Of female subimagO; 39, exuviae Of hind tarsus Of female subimagO." httpUri="https://zenodo.org/record/1436680/files/figure.png" pageId="3" pageNumber="118">Fig. 35</figureCitation>
) (see description of 
<taxonomicName id="4C7D6DACFFEC686726BFFE179072FE32" authorityName="Kluge" authorityYear="2018" baseAuthorityName="Kopelke" baseAuthorityYear="1980" box="[983,1104,476,501]" class="Insecta" family="Baetidae" genus="Monocentroptilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="B909CA3DFFEC686726BFFE179072FE32" box="[983,1104,476,501]" italics="true" pageId="3" pageNumber="118">M. badium</emphasis>
</taxonomicName>
below).
</paragraph>
</subSubSection>
<subSubSection id="C36745A4FFEC686725AFFE359073FDFA" pageId="3" pageNumber="118" type="distribution">
<paragraph id="8BC2162FFFEC686725AFFE35975EFDDF" blockId="3.[151,1436,151,573]" box="[199,892,511,536]" pageId="3" pageNumber="118">
<emphasis id="B909CA3DFFEC686725AFFE359542FDDF" bold="true" box="[199,352,511,536]" pageId="3" pageNumber="118">Distribution.</emphasis>
Afrotropical (known from 
<collectingRegion id="49B9D8CDFFEC686727FAFDCA96FFFDDF" box="[658,733,512,536]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="3" pageNumber="118">
<collectingCountry id="F36A56BFFFEC686727FAFDCA96FFFDDF" box="[658,733,512,536]" name="Democratic Republic of the Congo" pageId="3" pageNumber="118">Congo</collectingCountry>
</collectingRegion>
and 
<collectingCountry id="F36A56BFFFEC6867267CFE359752FDDF" box="[788,880,511,536]" name="Uganda" pageId="3" pageNumber="118">Uganda</collectingCountry>
).
</paragraph>
<paragraph id="8BC2162FFFEC686725AFFDEE9073FDFA" blockId="3.[151,1436,151,573]" box="[199,1105,548,573]" pageId="3" pageNumber="118">
<emphasis id="B909CA3DFFEC686725AFFDEE9545FDFA" bold="true" box="[199,359,548,573]" pageId="3" pageNumber="118">Composition.</emphasis>
One species
<taxonomicName id="4C7D6DACFFEC6867249EFDEE97C6FDFB" authority="(Kopelke 1980)" authorityName="Kluge" authorityYear="2018" baseAuthorityName="Kopelke" baseAuthorityYear="1980" box="[502,996,548,573]" pageId="3" pageNumber="118" rank="subSpecies" status="comb. nov." subSpecies="badium">
, 
<emphasis id="B909CA3DFFEC6867276AFDEE970EFDFA" box="[514,812,548,573]" italics="true" pageId="3" pageNumber="118">Monocentroptilum badium</emphasis>
(Kopelke 1980)
</taxonomicName>
<taxonomicNameLabel id="A23A7746FFEC68672683FDEE9073FDFA" box="[1003,1105,548,573]" pageId="3" pageNumber="118">
<emphasis id="B909CA3DFFEC68672683FDEE9073FDFA" bold="true" box="[1003,1105,548,573]" pageId="3" pageNumber="118">comb. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>