472 lines
64 KiB
XML
472 lines
64 KiB
XML
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<mods:title id="C946165687AAC8482EB427331FA0ED0C">An integrative taxonomic review of the South Asian microhylid genus Uperodon</mods:title>
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<mods:namePart id="2DAE6F76B79D57FDA22690B51F2D08E4">Senevirathne, Gayani</mods:namePart>
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<mods:namePart id="AC0B495676F35132B6095F876F72315C">Wijayathilaka, Nayana</mods:namePart>
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<mods:namePart id="8A95EEC79F5AF22C862CAE7CEE9787C4">Phuge, Samadhan</mods:namePart>
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<mods:namePart id="E5E6113BB642F500E2225ED31B38A5D6">Deuti, Kaushik</mods:namePart>
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<mods:namePart id="9E75DA559136E0242536778F7721AF85">Manamendra-Arachchi, Kelum</mods:namePart>
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<mods:title id="B6D51C061AE6C0EF77FAEA3131B11DF4">Zootaxa</mods:title>
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<mods:number id="8625A2903558F9746900BC735AB92566">2018-02-23</mods:number>
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<treatment id="03D4A4166C1D9469FF5ACEBDFEBEFDDF" ID-DOI="http://doi.org/10.5281/zenodo.5587715" ID-GBIF-Taxon="141119104" ID-Zenodo-Dep="5587715" LSID="urn:lsid:plazi:treatment:03D4A4166C1D9469FF5ACEBDFEBEFDDF" httpUri="http://treatment.plazi.org/id/03D4A4166C1D9469FF5ACEBDFEBEFDDF" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
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<paragraph id="8BC215006C1D946BFF5ACEBDFD0FFD4D" blockId="32.[151,751,727,823]" box="[151,687,727,754]" pageId="32" pageNumber="33">
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<taxonomicName id="4C7D6E836C1D946BFF5ACEBDFE23FD4E" ID-CoL="7DM7K" box="[151,387,727,753]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
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<emphasis id="B909C9126C1D946BFF5ACEBDFE23FD4E" bold="true" box="[151,387,727,753]" italics="true" pageId="32" pageNumber="33">Uperodon obscurus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFEC68F16C1D946BFE5ECEB2FDE7FD4D" author="Gunther, A. C. L. G." bookContentInfo="452 pp." box="[403,583,728,754]" journalOrPublisher="Hardwicke, London" pageId="32" pageNumber="33" refId="ref44945" refString="Gunther, A. C. L. G. (1864 " 1863 ") The Reptiles of British India. Ray Society by R. Hardwicke, London, 452 pp." title="1863 ") The Reptiles of British India. Ray Society by R" type="book" year="1864">Günther, 1864</bibRefCitation>
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“1863”)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC215006C1D946BFF5ACE90FE18FCAC" blockId="32.[151,751,727,823]" box="[151,440,762,787]" pageId="32" pageNumber="33">
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<materialsCitation id="3B151F5D6C1D946BFF5ACE90FE18FCAC" ID-GBIF-Occurrence="1821690905" box="[151,440,762,787]" location="Sri" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Sri">
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<collectingRegion id="49B9DBE26C1D946BFF5ACE90FF18FCAC" box="[151,184,762,787]" country="United Kingdom" name="Surrey" pageId="32" pageNumber="33">Sri</collectingRegion>
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Lankan Globular Frog
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</paragraph>
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<subSubSection id="C367468B6C1D946BFF5ACF74FBE1F9DA" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BC215006C1D946BFF5ACF74FD4FFC88" blockId="32.[151,751,727,823]" box="[151,751,798,823]" pageId="32" pageNumber="33">
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(
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<figureCitation id="134609856C1D946BFF52CF74FF4CFC88" box="[159,236,798,823]" captionStart="FIGURE 1" captionStartId="13.[151,250,1632,1654]" captionTargetBox="[154,1433,199,1603]" captionTargetId="figure@13.[151,1436,195,1608]" captionTargetPageId="13" captionText="FIGURE 1. Maximum Likelihood phylogram based on 511 bp of mitochondrial 16S rRNA gene sequences from 73 individuals representing all known members of the genus Uperodon. Three samples representing taxon under synonymy are shown inquotationmarks.Outgroup not shown for graphical purposes.BayesianPosterior Probabilities and bootstrap values of>50 are shown above and below the branches, respectively. Values towards the species labels represent mean uncorrected genetic distances between the species with range values in parentheses." httpUri="https://zenodo.org/record/1183457/files/figure.png" pageId="32" pageNumber="33">Figs. 1</figureCitation>
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,
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<figureCitation id="134609856C1D946BFF37CF75FEEFFC88" box="[250,335,799,823]" captionStart="FIGURE 2" captionStartId="14.[151,250,1617,1639]" captionTargetBox="[152,1434,196,1594]" captionTargetId="figure@14.[151,1436,193,1595]" captionTargetPageId="14" captionText="FIGURE 2. A. Geographic distribution of the genus Uperodon in Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka; B. Distributionof Uperodon globulosus, U. systomaand U.taprobanicus; C. Distribution of four endemic Uperodon species of the Western Ghats (excluding the widely distributed Indian member U. variegatus, shown in Fig.12A); D. Distribution of four endemic Uperdodon species of Sri Lanka. Details of localities reported in the present study are provided in Table 1. Records from literature are listed in supplementary Table S1 and indicated here based onapproximate coordinates." httpUri="https://zenodo.org/record/1183459/files/figure.png" pageId="32" pageNumber="33">2A, 2D</figureCitation>
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,
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<figureCitation id="134609856C1D946BFE96CF75FEDFFC88" box="[347,383,799,823]" captionStart="FIGURE 3" captionStartId="15.[151,250,1749,1771]" captionTargetBox="[151,1436,644,1728]" captionTargetId="figure@15.[151,1436,644,1728]" captionTargetPageId="15" captionText="FIGURE 3. Preserved type specimens of Uperodon species in dorsal and ventral view. A. Uperodon systoma, Syntype, ZMB 3551 (female); B. Uperodon globulosus, Syntype, NHM 62.12.29.4 (ex. BMNH 1947.2.10.97) (female); C. Kaloula pulchra taprobanica, Holotype, NHM 1905.3.25.138 (ex. BMNH 1947.2.10.62) (female); D. Ramanella anamalaiensis, Neotype, ZSI/ WGRC/V/A/956 (male); E. Hylaedactylus montanus, Holotype, NHM 72.4.17.212 (ex. BMNH 1947.2.11.19) (female); F. Ramanella mormorata, Neotype, ZSI/WGRC/V/A/958, (female); G. Ramanella nagaoi, Holotype, WHT 2826 (male); H. Callula obscura, Holotype, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20) (female); I. Ramanella palmata, Holotype, NHM 90.11.8.62 (ex.BMNH 1947.2.11.21); J. Callula triangularis, Lectotype, NHM 74.4.29.897 (ex.BMNH 1947.2.11.15) (male); K.Hylaedactylus carnaticus, Lectotype, ZSI 2761(female); L. Uperodonrohani sp. nov., Holotype, DZ1449 (male).(Images: SD Biju and Sonali Garg)." httpUri="https://zenodo.org/record/1183461/files/figure.png" pageId="32" pageNumber="33">3H</figureCitation>
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,
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<figureCitation id="134609856C1D946BFE46CF74FE0DFC88" box="[395,429,798,823]" captionStart="FIGURE 5" captionStartId="17.[151,250,1589,1611]" captionTargetBox="[155,1432,1093,1561]" captionTargetId="figure@17.[151,1436,1087,1567]" captionTargetPageId="17" captionText="FIGURE 5. Ventral surface of the palate in Uperodon species showing vomerine odontophores separated or fused with dermal projectionsand/or neopalatinalridgeson posterior sides ofthe choanae (indicated witharrows).A. Uperodon systoma, SDBDU 2005.2787 (female) and SDBDU 2005.2806 (male); B. Uperodon globulosus, SDBDU 2014.2673 (female) and SDBDU 2005.4762 (female); C. Uperodon taprobanicus, SDBDU 2007.6364 (female) and SDBDU 2005.4732A (male); D. Uperodon anamalaiensis, SDBDU 2013.2302 (female) and SDBDU 2002.3050 (male); E. Uperodon montanus, SDBDU 2002.24B (female) and SDBDU 2003.40115 (male); F. Uperodon mormorata, SDBDU 2011.507 (male) and SDBDU 2012.43 (male); G. Uperodon obscurus, DZ 1366 (male); H. Uperodon palmatus, DZ 1331 (male); I. Uperodon triangularis, SDBDU 2007.6400 (female) and ZSI/WGRC/V/A/959 (male); J. Uperodon variegatus, SDBDU 2005.4726 (female) and SDBDU 2005.4727 (male); K.Uperodon rohani sp. nov., DZ 1521 (male).(Images:SD Biju)." httpUri="https://zenodo.org/record/1183465/files/figure.png" pageId="32" pageNumber="33">5G</figureCitation>
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,
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<figureCitation id="134609856C1D946BFE75CF75FE5FFC88" box="[440,511,799,823]" captionStart="FIGURE9" captionStartId="30.[151,250,1723,1745]" captionTargetBox="[151,1436,885,1697]" captionTargetId="figure@30.[151,1436,885,1701]" captionTargetPageId="30" captionText="FIGURE9. Uperodon mormorata, U. nagaoi and U.obscurus inlife.A–C. Uperodonmormorata (SDBDU 2016.3404,male): A. Dorsolateral view; B. dorsal view; C. ventral view; D. Uperodon mormorata (SDBDU 2011.835, male), dorsolateral view; E–G. Uperodon nagaoi (DZ 1367, male): E. Dorsolateral view; F. dorsal view; G. frontal view; H–K. Uperodon obscurus: H. Dorsolateral view (DZ 1366, male); I. Dorsal view (DZ 1366, male); J. dorsolateral view (DZ 1408, male); K. dorsolateral view (DZ 1369, male); L. ventral view (DZ 1369, male). (Images: SD Biju and E–G Madhava Meegaskumbura)." httpUri="https://zenodo.org/record/1183473/files/figure.png" pageId="32" pageNumber="33">9H–K</figureCitation>
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,
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||
<figureCitation id="134609856C1D946BFDC7CF75FDC0FC88" box="[522,608,799,823]" captionStart="FIGURE 10" captionStartId="33.[151,250,1229,1251]" captionTargetBox="[157,1426,537,1193]" captionTargetId="figure@33.[151,1436,531,1208]" captionTargetPageId="33" captionText="FIGURE 10. Head, hand, foot and foot webbing in Uperodon obscurus, U. palmatus, U. triangularis and U.variegatus. A–D. Holotype ofUperodon obscurus(NHM58.11.28.42 ex.BMNH 1947.2.11.20, female):A. Lateralview of head; B. ventral view of hand; C. ventral view of foot; D. schematic illustration of foot webbing; E–H. Holotype of Uperodon palmatus (NHM 90.11.8.62 ex. BMNH 1947.2.11.21, female): E. Lateral view of head; F. ventral view of hand; G. ventral view of foot; H. schematic illustration of foot webbing; I–L. Lectotype of Uperodon triangularis (NHM 74.4.29.897 ex. BMNH 1947.2.11.15, male): I. Lateral view of head; J. ventral view of hand; K. ventral view of foot; L. schematic illustration of foot webbing; M. Lectotype of Uperodon variegatus (ZSI 2761, female), lateral view of head; N–P. Referred specimen of Uperodon variegatus (ZSI 4726, female): N. ventral view of hand; O. ventral view of foot; P. schematic illustration of foot webbing. (Illustrations: SD Biju)." httpUri="https://zenodo.org/record/1183475/files/figure.png" pageId="32" pageNumber="33">10A–D</figureCitation>
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;
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||
<tableCitation id="C6FF20BB6C1D946BFDA1CF74FD46FC88" box="[620,742,798,823]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 4" captionStartId-0="4.[151,239,1593,1616]" captionStartId-1="9.[151,239,1084,1107]" captionStartId-2="48.[119,193,547,566]" captionTargetBox-2="[135,1884,616,1625]" captionText-0="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." captionText-1="TABLE 2. Samples used for the molecular study." captionText-2="TABLE 4. Morphometric measurements of Uperodon species.All measurements are in millimetres (mm). The status of each specimen is given after the museum number:Holotype (HT), Paratype (PT), Neotype (NT), Lectotype (LT), Syntype (ST)." httpUri-2="http://table.plazi.org/id/DF0245886C0D947BFFBACE49F8ECFD89" pageId="32" pageNumber="33" tableUuid-2="DF0245886C0D947BFFBACE49F8ECFD89">Tables 1–4</tableCitation>
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)
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</paragraph>
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<paragraph id="8BC215006C1D946BFF5ACF0CFD0BFC54" blockId="32.[151,1437,870,2034]" pageId="32" pageNumber="33">
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<emphasis id="B909C9126C1D946BFF5ACF0CFDB2FCC0" bold="true" box="[151,530,870,895]" pageId="32" pageNumber="33">Original name and description.</emphasis>
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<taxonomicName id="4C7D6E836C1D946BFDD6CF0CFCDEFCC0" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[539,894,870,895]" class="Amphibia" family="Microhylidae" genus="Callula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscura">
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<emphasis id="B909C9126C1D946BFDD6CF0CFD72FCC0" box="[539,722,870,895]" italics="true" pageId="32" pageNumber="33">Callula obscura</emphasis>
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<bibRefCitation id="EFEC68F16C1D946BFD16CF0CFCDEFCC0" author="Gunther, A. C. L. G." bookContentInfo="452 pp." box="[731,894,870,895]" journalOrPublisher="Hardwicke, London" pageId="32" pageNumber="33" refId="ref44945" refString="Gunther, A. C. L. G. (1864 " 1863 ") The Reptiles of British India. Ray Society by R. Hardwicke, London, 452 pp." title="1863 ") The Reptiles of British India. Ray Society by R" type="book" year="1864">Günther, 1864</bibRefCitation>
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</taxonomicName>
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“1863”.
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<emphasis id="B909C9126C1D946BFC25CF0CFA8DFCC0" box="[1000,1325,870,895]" italics="true" pageId="32" pageNumber="33">
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The
|
||
<taxonomicName id="4C7D6E836C1D946BFBD6CF0CFBD5FCC0" authorityName="Linnæus" authorityYear="1758" box="[1051,1141,870,895]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptiles" pageId="32" pageNumber="33" phylum="Chordata" rank="order">Reptiles</taxonomicName>
|
||
of British India
|
||
</emphasis>
|
||
. London: Ray Society by R. Hardwicke: 438.
|
||
<emphasis id="B909C9126C1D946BFDF3CFE0FD12FC1C" bold="true" box="[574,690,906,931]" pageId="32" pageNumber="33">Holotype.</emphasis>
|
||
By monotypy, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20), an adult female, SVL
|
||
<quantity id="4C85B8E56C1D946BFEA2CFC5FE77FC79" box="[367,471,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="32" pageNumber="33" unit="mm" value="32.2">32.2 mm</quantity>
|
||
.
|
||
<emphasis id="B909C9126C1D946BFE2ECFC4FD22FC78" bold="true" box="[483,642,942,967]" pageId="32" pageNumber="33">Type locality.</emphasis>
|
||
“Ceylon” (Sri Lanka).
|
||
<emphasis id="B909C9126C1D946BFC40CFC4FAADFC78" bold="true" box="[909,1293,942,967]" pageId="32" pageNumber="33">Current status of specific name.</emphasis>
|
||
Valid name, as
|
||
<taxonomicName id="4C7D6E836C1D946BFF78CFB8FE32FC54" box="[181,402,978,1003]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1D946BFF78CFB8FE32FC54" box="[181,402,978,1003]" italics="true" pageId="32" pageNumber="33">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEC68F16C1D946BFE6DCFB8FDE3FC54" author="Gunther, A. C. L. G." bookContentInfo="452 pp." box="[416,579,978,1003]" journalOrPublisher="Hardwicke, London" pageId="32" pageNumber="33" refId="ref44945" refString="Gunther, A. C. L. G. (1864 " 1863 ") The Reptiles of British India. Ray Society by R. Hardwicke, London, 452 pp." title="1863 ") The Reptiles of British India. Ray Society by R" type="book" year="1864">Günther, 1864</bibRefCitation>
|
||
“1863”).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C1D946BFF0ACF9CFD1EFA6A" blockId="32.[151,1437,870,2034]" pageId="32" pageNumber="33">
|
||
<emphasis id="B909C9126C1D946BFF0ACF9CFEC1FBB0" bold="true" box="[199,353,1014,1039]" pageId="32" pageNumber="33">Comparison.</emphasis>
|
||
For comparison of
|
||
<taxonomicName id="4C7D6E836C1D946BFD80CF9CFC8DFBB0" box="[589,813,1014,1039]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1D946BFD80CF9CFC8DFBB0" box="[589,813,1014,1039]" italics="true" pageId="32" pageNumber="33">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C7D6E836C1D946BFCB9CF9DFBB3FBB0" box="[884,1043,1014,1039]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="globulosus">
|
||
<emphasis id="B909C9126C1D946BFCB9CF9DFBB3FBB0" box="[884,1043,1014,1039]" italics="true" pageId="32" pageNumber="33">U. globulosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1D946BFBE8CF9DFB05FBB1" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[1061,1189,1015,1038]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C1D946BFBE8CF9DFB05FBB1" box="[1061,1189,1015,1038]" italics="true" pageId="32" pageNumber="33">U. systoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1D946BFB7BCF9DFAD0FBB0" baseAuthorityName="Parker" baseAuthorityYear="1934" box="[1206,1392,1014,1039]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="taprobanicus">
|
||
<emphasis id="B909C9126C1D946BFB7BCF9DFAD0FBB0" box="[1206,1392,1014,1039]" italics="true" pageId="32" pageNumber="33">U. taprobanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1D946BFA4CCF9DFE99FB8C" authorityName="M Parambikulam" baseAuthorityName="Rao" baseAuthorityYear="1937" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="anamalaiensis">
|
||
<emphasis id="B909C9126C1D946BFA4CCF9DFE99FB8C" italics="true" pageId="32" pageNumber="33">U. anamalaiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1D946BFE84C876FE7AFB8C" box="[329,474,1052,1075]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B909C9126C1D946BFE84C876FE7AFB8C" box="[329,474,1052,1075]" italics="true" pageId="32" pageNumber="33">U. montanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1D946BFE27C876FD2CFB8C" baseAuthorityName="Rao" baseAuthorityYear="1937" box="[490,652,1052,1075]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="mormorata">
|
||
<emphasis id="B909C9126C1D946BFE27C876FD2CFB8C" box="[490,652,1052,1075]" italics="true" pageId="32" pageNumber="33">U. mormorata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D6E836C1D946BFD0AC876FC99FB8C" baseAuthorityName="Manamendra-Arachchi and Pethiyagoda" baseAuthorityYear="2001" box="[711,825,1052,1075]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="nagaoi">
|
||
<emphasis id="B909C9126C1D946BFD0AC876FC99FB8C" box="[711,825,1052,1075]" italics="true" pageId="32" pageNumber="33">U. nagaoi</emphasis>
|
||
</taxonomicName>
|
||
see ‘comparison’ section of those species.
|
||
<taxonomicName id="4C7D6E836C1D946BFAE1C870FF5CFBE8" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1D946BFAE1C870FF5CFBE8" italics="true" pageId="32" pageNumber="33">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName id="4C7D6E836C1D946BFE2CC855FDC9FBE8" box="[481,617,1086,1111]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="palmatus">
|
||
<emphasis id="B909C9126C1D946BFE2CC855FDC9FBE8" box="[481,617,1086,1111]" italics="true" pageId="32" pageNumber="33">U. palmatus</emphasis>
|
||
</taxonomicName>
|
||
, both genetically and morphologically. However, it differs from the latter by its snout length longer than horizontal diameter of eye, male: EL/SL ratio 0.6–0.9,
|
||
<emphasis id="B909C9126C1D946BFB92C80EFBD2FBC4" box="[1119,1138,1124,1147]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||
= 17; female: EL/SL ratio 0.6–0.9,
|
||
<emphasis id="B909C9126C1D946BFF36C8EDFEAEFB21" box="[251,270,1159,1182]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||
= 5 (vs. equal, male: EL/SL ratio 1.0,
|
||
<emphasis id="B909C9126C1D946BFD1DC8EDFD43FB21" box="[720,739,1159,1182]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||
= 4, female: EL/SL ratio 1.0,
|
||
<emphasis id="B909C9126C1D946BFB89C8EDFBF7FB21" box="[1092,1111,1159,1182]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||
= 2) (
|
||
<tableCitation id="C6FF20BB6C1D946BFB6DC8ECFB59FB20" box="[1184,1273,1158,1183]" captionStart="TABLE 4" captionStartId="48.[119,193,547,566]" captionTargetBox="[135,1884,616,1625]" captionText="TABLE 4. Morphometric measurements of Uperodon species.All measurements are in millimetres (mm). The status of each specimen is given after the museum number:Holotype (HT), Paratype (PT), Neotype (NT), Lectotype (LT), Syntype (ST)." httpUri="http://table.plazi.org/id/DF0245886C0D947BFFBACE49F8ECFD89" pageId="32" pageNumber="33" tableUuid="DF0245886C0D947BFFBACE49F8ECFD89">Table 4</tableCitation>
|
||
), snout nearly acute in lateral view (vs. vertical), relatively less webbing,
|
||
<date id="FFC333C06C1D946BFCE1C8C4FC20FB79" box="[812,896,1198,1222]" pageId="32" pageNumber="33">I1–2II1</date>
|
||
1/2–3–
|
||
<date id="FFC333C06C1D946BFC77C8C4FBA0FB79" box="[954,1024,1198,1222]" pageId="32" pageNumber="33">III2–3</date>
|
||
+
|
||
<date id="FFC333C06C1D946BFBC4C8C4FB94FB79" box="[1033,1076,1198,1222]" pageId="32" pageNumber="33">IV3</date>
|
||
+–
|
||
<date id="FFC333C06C1D946BFB81C8C4FBCDFB79" box="[1100,1133,1198,1222]" pageId="32" pageNumber="33">2V</date>
|
||
(vs. more,
|
||
<date id="FFC333C06C1D946BFB21C8C4FAFCFB79" box="[1260,1372,1198,1222]" pageId="32" pageNumber="33">I1–2II1–3</date>
|
||
–
|
||
<date id="FFC333C06C1D946BFAA9C8C4FF70FB50" pageId="32" pageNumber="33">III1– 3IV2</date>
|
||
2/3–
|
||
<date id="FFC333C06C1D946BFF35C8BDFEB9FB50" box="[248,281,1239,1263]" pageId="32" pageNumber="33">1V</date>
|
||
) (
|
||
<figureCitation id="134609856C1D946BFEFAC8BCFE05FB50" box="[311,421,1238,1263]" captionStart="FIGURE 10" captionStartId="33.[151,250,1229,1251]" captionTargetBox="[157,1426,537,1193]" captionTargetId="figure@33.[151,1436,531,1208]" captionTargetPageId="33" captionText="FIGURE 10. Head, hand, foot and foot webbing in Uperodon obscurus, U. palmatus, U. triangularis and U.variegatus. A–D. Holotype ofUperodon obscurus(NHM58.11.28.42 ex.BMNH 1947.2.11.20, female):A. Lateralview of head; B. ventral view of hand; C. ventral view of foot; D. schematic illustration of foot webbing; E–H. Holotype of Uperodon palmatus (NHM 90.11.8.62 ex. BMNH 1947.2.11.21, female): E. Lateral view of head; F. ventral view of hand; G. ventral view of foot; H. schematic illustration of foot webbing; I–L. Lectotype of Uperodon triangularis (NHM 74.4.29.897 ex. BMNH 1947.2.11.15, male): I. Lateral view of head; J. ventral view of hand; K. ventral view of foot; L. schematic illustration of foot webbing; M. Lectotype of Uperodon variegatus (ZSI 2761, female), lateral view of head; N–P. Referred specimen of Uperodon variegatus (ZSI 4726, female): N. ventral view of hand; O. ventral view of foot; P. schematic illustration of foot webbing. (Illustrations: SD Biju)." httpUri="https://zenodo.org/record/1183475/files/figure.png" pageId="32" pageNumber="33">Fig. 10D</figureCitation>
|
||
), and dorsal skin shagreened to sparsely granular (vs. distinctly granular with warty projections). It differs from
|
||
<taxonomicName id="4C7D6E836C1D946BFE16C895FD26FAA9" box="[475,646,1277,1302]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="triangularis">
|
||
<emphasis id="B909C9126C1D946BFE16C895FD26FAA9" box="[475,646,1277,1302]" italics="true" pageId="32" pageNumber="33">U. triangularis</emphasis>
|
||
</taxonomicName>
|
||
by its fourth toe webbing extending close up to second subarticular tubercle:
|
||
<date id="FFC333C06C1D946BFECAC94FFEFBFA82" box="[263,347,1317,1341]" pageId="32" pageNumber="33">I1–2II1</date>
|
||
1/2–3–
|
||
<date id="FFC333C06C1D946BFE58C94FFE7CFA82" box="[405,476,1317,1341]" pageId="32" pageNumber="33">III2–3</date>
|
||
+
|
||
<date id="FFC333C06C1D946BFE29C94FFDB0FA82" box="[484,528,1317,1341]" pageId="32" pageNumber="33">IV3</date>
|
||
+–
|
||
<date id="FFC333C06C1D946BFDEAC94FFDE8FA82" box="[551,584,1317,1341]" pageId="32" pageNumber="33">2V</date>
|
||
(vs. toes completely free of webbing).
|
||
<taxonomicName id="4C7D6E836C1D946BFBF8C94EFABAFA82" box="[1077,1306,1316,1341]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1D946BFBF8C94EFABAFA82" box="[1077,1306,1316,1341]" italics="true" pageId="32" pageNumber="33">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
cannot be confused with three other species,
|
||
<taxonomicName id="4C7D6E836C1D946BFDD2C926FD30FADC" baseAuthorityName="Manamendra-Arachchi and Pethiyagoda" baseAuthorityYear="2001" box="[543,656,1356,1379]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="nagaoi">
|
||
<emphasis id="B909C9126C1D946BFDD2C926FD30FADC" box="[543,656,1356,1379]" italics="true" pageId="32" pageNumber="33">U. nagaoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1D946BFD52C926FCABFADB" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[671,779,1355,1380]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C1D946BFD52C926FCABFADB" box="[671,779,1355,1380]" italics="true" pageId="32" pageNumber="33">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C1D946BFCD9C921FCCFFADB" bold="true" box="[788,879,1355,1380]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="A23A74696C1D946BFCD9C921FCCFFADB" box="[788,879,1355,1380]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7D6E836C1D946BFC6AC926FBE0FADC" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[935,1088,1356,1379]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C1D946BFC6AC926FBE0FADC" box="[935,1088,1356,1379]" italics="true" pageId="32" pageNumber="33">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
, due to its relatively extensive foot webbing,
|
||
<date id="FFC333C06C1D946BFEF7C919FE2EFA34" box="[314,398,1395,1419]" pageId="32" pageNumber="33">I1–2II1</date>
|
||
1/2–3–
|
||
<date id="FFC333C06C1D946BFE05C919FDAEFA34" box="[456,526,1395,1419]" pageId="32" pageNumber="33">III2–3</date>
|
||
+
|
||
<date id="FFC333C06C1D946BFDDAC919FDE2FA34" box="[535,578,1395,1419]" pageId="32" pageNumber="33">IV3</date>
|
||
+–
|
||
<date id="FFC333C06C1D946BFD97C919FDDBFA34" box="[602,635,1395,1419]" pageId="32" pageNumber="33">2V</date>
|
||
(vs. rudimentary or absent), and ventral skin being light to dark brown with distinct blotches or spots (vs. brown without prominent mottling in
|
||
<taxonomicName id="4C7D6E836C1D946BFC0CC9F0FB90FA0E" baseAuthorityName="Manamendra-Arachchi and Pethiyagoda" baseAuthorityYear="2001" box="[961,1072,1434,1457]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="nagaoi">
|
||
<emphasis id="B909C9126C1D946BFC0CC9F0FB90FA0E" box="[961,1072,1434,1457]" italics="true" pageId="32" pageNumber="33">U. nagaoi</emphasis>
|
||
</taxonomicName>
|
||
, and white without any blotches or spots in
|
||
<taxonomicName id="4C7D6E836C1D946BFEDFC9D4FEDEFA6A" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[274,382,1468,1493]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C1D946BFEDFC9D4FEDEFA6A" box="[274,382,1468,1493]" italics="true" pageId="32" pageNumber="33">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C1D946BFE48C9D6FE7EFA6A" bold="true" box="[389,478,1468,1493]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="A23A74696C1D946BFE48C9D6FE7EFA6A" box="[389,478,1468,1493]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7D6E836C1D946BFDD8C9D4FD0DFA6A" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[533,685,1470,1493]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C1D946BFDD8C9D4FD0DFA6A" box="[533,685,1470,1493]" italics="true" pageId="32" pageNumber="33">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C1D946BFF0AC98BFBE1F9DA" blockId="32.[151,1437,870,2034]" pageId="32" pageNumber="33">
|
||
<emphasis id="B909C9126C1D946BFF0AC98BFE12FA45" bold="true" box="[199,434,1505,1530]" pageId="32" pageNumber="33">Genetic divergence.</emphasis>
|
||
For 16S mitochondrial gene sequences, intraspecific variation was not observed between the sampled populations of
|
||
<taxonomicName id="4C7D6E836C1D946BFE01CA6FFD09F9A1" box="[460,681,1541,1566]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1D946BFE01CA6FFD09F9A1" box="[460,681,1541,1566]" italics="true" pageId="32" pageNumber="33">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B909C9126C1D946BFD77CA6CFD6DF9A2" box="[698,717,1542,1565]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||
= 3). Genetically,
|
||
<taxonomicName id="4C7D6E836C1D946BFC50CA6CFB84F9A1" box="[925,1060,1541,1566]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1D946BFC50CA6CFB84F9A1" box="[925,1060,1541,1566]" italics="true" pageId="32" pageNumber="33">U. obscurus</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName id="4C7D6E836C1D946BFAC1CA6CFA35F9A1" box="[1292,1429,1541,1566]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="palmatus">
|
||
<emphasis id="B909C9126C1D946BFAC1CA6CFA35F9A1" box="[1292,1429,1541,1566]" italics="true" pageId="32" pageNumber="33">U. palmatus</emphasis>
|
||
</taxonomicName>
|
||
, from which it was found to differ by an average uncorrected genetic distance of 2.4% (range 2.4–2.6%,
|
||
<emphasis id="B909C9126C1D946BFAF1CA40FAEFF9FE" box="[1340,1359,1578,1601]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||
= 12). For interspecific genetic distances with all other members of the genus, see Table 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367468B6C1D946BFF0ACA1BFB53F96D" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8BC215006C1D946BFF0ACA1BFB53F96D" blockId="32.[151,1437,870,2034]" pageId="32" pageNumber="33">
|
||
<emphasis id="B909C9126C1D946BFF0ACA1BFE9BF935" bold="true" box="[199,315,1649,1674]" pageId="32" pageNumber="33">Remarks.</emphasis>
|
||
The original description by
|
||
<bibRefCitation id="EFEC68F16C1D946BFD46CA1BFC93F935" author="Gunther, A. C. L. G." bookContentInfo="452 pp." box="[651,819,1649,1674]" journalOrPublisher="Hardwicke, London" pageId="32" pageNumber="33" refId="ref44945" refString="Gunther, A. C. L. G. (1864 " 1863 ") The Reptiles of British India. Ray Society by R. Hardwicke, London, 452 pp." title="1863 ") The Reptiles of British India. Ray Society by R" type="book" year="1864">Günther (1864</bibRefCitation>
|
||
“1863”) was very brief. Subsequently
|
||
<bibRefCitation id="EFEC68F16C1D946BFB33CA1BFA3CF936" author="Parker, H. W." bookContentInfo="208 pp." box="[1278,1436,1649,1674]" journalOrPublisher="Trustees of the British Museum, London" pageId="32" pageNumber="33" refId="ref38477" refString="Parker, H. W. (1934) A Monograph of the Frogs of the Family Microhylidae. Trustees of the British Museum, London, 208 pp." title="A Monograph of the Frogs of the Family Microhylidae" type="book" year="1934">Parker (1934)</bibRefCitation>
|
||
provided a general description based on the
|
||
<typeStatus id="54C6ABA26C1D946BFD68CAFCFD43F911" box="[677,739,1686,1710]" pageId="32" pageNumber="33">types</typeStatus>
|
||
, without a comparison with other closely related members. Hence, a detailed description and comparison of this species with all its congeners is provided here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367468B6C1D9469FF0ACAB7FEBEFDDF" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BC215006C1D946AFF0ACAB7FB9FFE61" blockId="32.[151,1437,870,2034]" lastBlockId="33.[151,1436,151,478]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<emphasis id="B909C9126C1D946BFF0ACAB7FE5BF949" bold="true" box="[199,507,1757,1782]" pageId="32" pageNumber="33">Redescription of holotype</emphasis>
|
||
(measurements in mm) (
|
||
<figureCitation id="134609856C1D946BFCDBCAB7FCDAF94A" box="[790,890,1757,1782]" captionStart="FIGURE 3" captionStartId="15.[151,250,1749,1771]" captionTargetBox="[151,1436,644,1728]" captionTargetId="figure@15.[151,1436,644,1728]" captionTargetPageId="15" captionText="FIGURE 3. Preserved type specimens of Uperodon species in dorsal and ventral view. A. Uperodon systoma, Syntype, ZMB 3551 (female); B. Uperodon globulosus, Syntype, NHM 62.12.29.4 (ex. BMNH 1947.2.10.97) (female); C. Kaloula pulchra taprobanica, Holotype, NHM 1905.3.25.138 (ex. BMNH 1947.2.10.62) (female); D. Ramanella anamalaiensis, Neotype, ZSI/ WGRC/V/A/956 (male); E. Hylaedactylus montanus, Holotype, NHM 72.4.17.212 (ex. BMNH 1947.2.11.19) (female); F. Ramanella mormorata, Neotype, ZSI/WGRC/V/A/958, (female); G. Ramanella nagaoi, Holotype, WHT 2826 (male); H. Callula obscura, Holotype, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20) (female); I. Ramanella palmata, Holotype, NHM 90.11.8.62 (ex.BMNH 1947.2.11.21); J. Callula triangularis, Lectotype, NHM 74.4.29.897 (ex.BMNH 1947.2.11.15) (male); K.Hylaedactylus carnaticus, Lectotype, ZSI 2761(female); L. Uperodonrohani sp. nov., Holotype, DZ1449 (male).(Images: SD Biju and Sonali Garg)." httpUri="https://zenodo.org/record/1183461/files/figure.png" pageId="32" pageNumber="33">Figs. 3H</figureCitation>
|
||
,
|
||
<figureCitation id="134609856C1D946BFC4BCAB7FC7DF94A" box="[902,989,1757,1781]" captionStart="FIGURE 10" captionStartId="33.[151,250,1229,1251]" captionTargetBox="[157,1426,537,1193]" captionTargetId="figure@33.[151,1436,531,1208]" captionTargetPageId="33" captionText="FIGURE 10. Head, hand, foot and foot webbing in Uperodon obscurus, U. palmatus, U. triangularis and U.variegatus. A–D. Holotype ofUperodon obscurus(NHM58.11.28.42 ex.BMNH 1947.2.11.20, female):A. Lateralview of head; B. ventral view of hand; C. ventral view of foot; D. schematic illustration of foot webbing; E–H. Holotype of Uperodon palmatus (NHM 90.11.8.62 ex. BMNH 1947.2.11.21, female): E. Lateral view of head; F. ventral view of hand; G. ventral view of foot; H. schematic illustration of foot webbing; I–L. Lectotype of Uperodon triangularis (NHM 74.4.29.897 ex. BMNH 1947.2.11.15, male): I. Lateral view of head; J. ventral view of hand; K. ventral view of foot; L. schematic illustration of foot webbing; M. Lectotype of Uperodon variegatus (ZSI 2761, female), lateral view of head; N–P. Referred specimen of Uperodon variegatus (ZSI 4726, female): N. ventral view of hand; O. ventral view of foot; P. schematic illustration of foot webbing. (Illustrations: SD Biju)." httpUri="https://zenodo.org/record/1183475/files/figure.png" pageId="32" pageNumber="33">10A–D</figureCitation>
|
||
). Small-sized (SVL 32.2), slender adult female; head small (HW 9.3, HL 8.8, IFE 4.9, IBE 8.6), less than one-third (27.3%) of body length, wider than long (HW/HL ratio 1.1); snout truncate in dorsal and ventral view, nearly acute in lateral view, its length (SL 4.2) longer than horizontal diameter of eye (EL 3.9); loreal region acute with rounded canthus rostralis; interorbital space more than twice as wide (IUE 3.3) as upper eyelid width (UEW 1.4), and wider than internarial distance (IN 2.7); nostril much closer to tip of snout (NS 0.7) than to eye (EN 2.4); supratympanic fold distinct, extending from posterior corner of upper eyelid to insertion of forelimb at axilla; eye diameter (EL 3.9); vomerine odontophores present on the palate, without teeth; weakly-developed neopalatinal ridges on posterior side of each choana, appear fused with the vomerine odontophores; tongue moderately large, oval, slightly emarginate. Forelimbs (FAL 8.8) shorter than hand length (HAL 9.5); finger length formula I<II<IV<III, tips of all fingers with oval discs, moderately wide compared to finger width (FD I 0.8, FW I 0.4; FD II 1.1, FW II 0.5; FD III 1.0, FW III 0.5; FD IV 0.9, FW IV 0.4); subarticular tubercles prominent, oval, all present; two well-developed palmar tubercles, oval. Hind limbs relatively long and thin, thigh length (TL 13.5) longer than shank (SHL 13.3), and shorter than foot (FOL 14.9); relative digit lengths I<II<V<III<IV, tips of all toes with small truncate to rounded discs, rather wide compared to toe width (TD I 0.7, TW I 0.5; TD II 0.8, TW II 0.5; TD III 0.7, TW III 0.5; TD IV 0.7, TW IV 0.5; TD V 0.5, TW V 0.5); foot webbing present,
|
||
<date id="FFC333C06C1C946AFABBCD1CFF65FE08" pageId="33" pageNumber="34">I1– 2II1</date>
|
||
1/2–3–
|
||
<date id="FFC333C06C1C946AFECDCDF5FEE6FE08" box="[256,326,415,439]" pageId="33" pageNumber="34">III2–3</date>
|
||
+
|
||
<date id="FFC333C06C1C946AFE83CDF5FEDAFE08" box="[334,378,415,439]" pageId="33" pageNumber="34">IV3</date>
|
||
+–
|
||
<date id="FFC333C06C1C946AFE5FCDF5FE14FE08" box="[402,436,415,439]" pageId="33" pageNumber="34">2V</date>
|
||
; well-developed dermal fringes present on all toes; subarticular tubercles prominent, oval; two smooth metatarsal tubercles, oval, outer one slightly larger than the inner.
|
||
</paragraph>
|
||
<caption id="DF0245886C1C946AFF5AC8A7FF59FA67" httpUri="https://zenodo.org/record/1183475/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1229,1251]" targetBox="[157,1426,537,1193]" targetPageId="33">
|
||
<paragraph id="8BC215006C1C946AFF5AC8A7FF59FA67" blockId="33.[151,1437,1229,1496]" pageId="33" pageNumber="34">
|
||
<emphasis id="B909C9126C1C946AFF5AC8A7FEBFFB5C" bold="true" box="[151,287,1229,1251]" pageId="33" pageNumber="34">FIGURE 10.</emphasis>
|
||
Head, hand, foot and foot webbing in
|
||
<taxonomicName id="4C7D6E836C1C946AFD6AC8A7FCCFFB5C" box="[679,879,1229,1251]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1C946AFD6AC8A7FCCFFB5C" box="[679,879,1229,1251]" italics="true" pageId="33" pageNumber="34">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1C946AFCB1C8A4FC58FB5C" box="[892,1016,1229,1251]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="palmatus">
|
||
<emphasis id="B909C9126C1C946AFCB1C8A4FC58FB5C" box="[892,1016,1229,1251]" italics="true" pageId="33" pageNumber="34">U. palmatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C1C946AFBC9C8A4FB3CFB5C" box="[1028,1180,1229,1251]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="triangularis">
|
||
<emphasis id="B909C9126C1C946AFBC9C8A4FB3CFB5C" box="[1028,1180,1229,1251]" italics="true" pageId="33" pageNumber="34">U. triangularis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D6E836C1C946AFB00C8A4FAF7FB5C" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[1229,1367,1230,1251]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C1C946AFB00C8A4FAF7FB5C" box="[1229,1367,1230,1251]" italics="true" pageId="33" pageNumber="34">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B909C9126C1C946AFAA9C8A7FA3CFB5C" bold="true" box="[1380,1436,1229,1251]" pageId="33" pageNumber="34">A–D.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C7D6E836C1C946AFED8C886FE7DFABD" box="[277,477,1260,1282]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1C946AFED8C886FE7DFABD" box="[277,477,1260,1282]" italics="true" pageId="33" pageNumber="34">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
(NHM 58.11.28.42 ex. BMNH 1947.2.11.20, female):
|
||
<emphasis id="B909C9126C1C946AFBC5C886FB80FABD" bold="true" box="[1032,1056,1260,1282]" pageId="33" pageNumber="34">A.</emphasis>
|
||
Lateral view of head;
|
||
<emphasis id="B909C9126C1C946AFACFC886FAB9FABD" bold="true" box="[1282,1305,1260,1282]" pageId="33" pageNumber="34">B.</emphasis>
|
||
ventral view of hand;
|
||
<emphasis id="B909C9126C1C946AFF3AC960FEB0FA9F" bold="true" box="[247,272,1290,1312]" pageId="33" pageNumber="34">C.</emphasis>
|
||
ventral view of foot;
|
||
<emphasis id="B909C9126C1C946AFE36C960FDB4FA9F" bold="true" box="[507,532,1290,1312]" pageId="33" pageNumber="34">D.</emphasis>
|
||
schematic illustration of foot webbing;
|
||
<emphasis id="B909C9126C1C946AFC71C960FC53FA9F" bold="true" box="[956,1011,1290,1312]" pageId="33" pageNumber="34">E–H.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C7D6E836C1C946AFB4EC960FAEFFA9F" box="[1155,1359,1290,1312]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="palmatus">
|
||
<emphasis id="B909C9126C1C946AFB4EC960FAEFFA9F" box="[1155,1359,1290,1312]" italics="true" pageId="33" pageNumber="34">Uperodon palmatus</emphasis>
|
||
</taxonomicName>
|
||
(NHM 90.11.8.62 ex. BMNH 1947.2.11.21, female):
|
||
<emphasis id="B909C9126C1C946AFDBBC943FD2DFA80" bold="true" box="[630,653,1321,1343]" pageId="33" pageNumber="34">E.</emphasis>
|
||
Lateral view of head;
|
||
<emphasis id="B909C9126C1C946AFCB3C943FC2FFA80" bold="true" box="[894,911,1321,1343]" pageId="33" pageNumber="34">F.</emphasis>
|
||
ventral view of hand;
|
||
<emphasis id="B909C9126C1C946AFB4EC943FB39FA80" bold="true" box="[1155,1177,1321,1343]" pageId="33" pageNumber="34">G.</emphasis>
|
||
ventral view of foot;
|
||
<emphasis id="B909C9126C1C946AFA4FC943FA3CFA80" bold="true" box="[1410,1436,1321,1343]" pageId="33" pageNumber="34">H.</emphasis>
|
||
schematic illustration of foot webbing;
|
||
<emphasis id="B909C9126C1C946AFDE5C922FDF6FAE1" bold="true" box="[552,598,1352,1374]" pageId="33" pageNumber="34">I–L.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName id="4C7D6E836C1C946AFD29C922FC6AFAE1" box="[740,970,1352,1374]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="triangularis">
|
||
<emphasis id="B909C9126C1C946AFD29C922FC6AFAE1" box="[740,970,1352,1374]" italics="true" pageId="33" pageNumber="34">Uperodon triangularis</emphasis>
|
||
</taxonomicName>
|
||
(NHM 74.4.29.897 ex. BMNH 1947.2.11.15, male):
|
||
<emphasis id="B909C9126C1C946AFF12C90CFF4EFAC3" bold="true" box="[223,238,1382,1404]" pageId="33" pageNumber="34">I.</emphasis>
|
||
Lateral view of head;
|
||
<emphasis id="B909C9126C1C946AFE1AC90CFE4AFAC3" bold="true" box="[471,490,1382,1404]" pageId="33" pageNumber="34">J.</emphasis>
|
||
ventral view of hand;
|
||
<emphasis id="B909C9126C1C946AFD1EC90CFD4DFAC3" bold="true" box="[723,749,1382,1404]" pageId="33" pageNumber="34">K.</emphasis>
|
||
ventral view of foot;
|
||
<emphasis id="B909C9126C1C946AFC01C90CFC43FAC3" bold="true" box="[972,995,1382,1404]" pageId="33" pageNumber="34">L.</emphasis>
|
||
schematic illustration of foot webbing;
|
||
<emphasis id="B909C9126C1C946AFAB3C90CFA3CFAC3" bold="true" box="[1406,1436,1382,1404]" pageId="33" pageNumber="34">M.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName id="4C7D6E836C1C946AFED2C9EFFE57FA24" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[287,503,1413,1435]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C1C946AFED2C9EFFE57FA24" box="[287,503,1413,1435]" italics="true" pageId="33" pageNumber="34">Uperodon variegatus</emphasis>
|
||
</taxonomicName>
|
||
(ZSI 2761, female), lateral view of head;
|
||
<emphasis id="B909C9126C1C946AFC65C9EFFC7AFA24" bold="true" box="[936,986,1413,1435]" pageId="33" pageNumber="34">N–P.</emphasis>
|
||
Referred specimen of
|
||
<taxonomicName id="4C7D6E836C1C946AFB09C9EFFA3CFA24" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[1220,1436,1413,1435]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C1C946AFB09C9EFFA3CFA24" box="[1220,1436,1413,1435]" italics="true" pageId="33" pageNumber="34">Uperodon variegatus</emphasis>
|
||
</taxonomicName>
|
||
(ZSI 4726, female):
|
||
<emphasis id="B909C9126C1C946AFEA5C9CEFE21FA05" bold="true" box="[360,385,1444,1466]" pageId="33" pageNumber="34">N.</emphasis>
|
||
ventral view of hand;
|
||
<emphasis id="B909C9126C1C946AFDA6C9CEFD24FA05" bold="true" box="[619,644,1444,1466]" pageId="33" pageNumber="34">O.</emphasis>
|
||
ventral view of foot;
|
||
<emphasis id="B909C9126C1C946AFCABC9CEFCD7FA05" bold="true" box="[870,887,1444,1466]" pageId="33" pageNumber="34">P.</emphasis>
|
||
schematic illustration of foot webbing. (Illustrations: SD Biju).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC215006C1C946AFF0ACA4EFDB6F917" blockId="33.[151,1436,1572,2029]" pageId="33" pageNumber="34">
|
||
Skin of snout, between eyes, sides of head, and anterior part of dorsum, shagreened to granular; posterior parts of back, and upper and lower parts of flank, granular; dorsal surfaces of forelimb, thigh, tibia and tarsus, shagreened to sparsely granular; throat, chest and abdomen, shagreened; ventral surfaces of forelimb and hind limb, shagreened (
|
||
<figureCitation id="134609856C1C946AFEABCAE5FE69F917" box="[358,457,1679,1704]" captionStart="FIGURE 3" captionStartId="15.[151,250,1749,1771]" captionTargetBox="[151,1436,644,1728]" captionTargetId="figure@15.[151,1436,644,1728]" captionTargetPageId="15" captionText="FIGURE 3. Preserved type specimens of Uperodon species in dorsal and ventral view. A. Uperodon systoma, Syntype, ZMB 3551 (female); B. Uperodon globulosus, Syntype, NHM 62.12.29.4 (ex. BMNH 1947.2.10.97) (female); C. Kaloula pulchra taprobanica, Holotype, NHM 1905.3.25.138 (ex. BMNH 1947.2.10.62) (female); D. Ramanella anamalaiensis, Neotype, ZSI/ WGRC/V/A/956 (male); E. Hylaedactylus montanus, Holotype, NHM 72.4.17.212 (ex. BMNH 1947.2.11.19) (female); F. Ramanella mormorata, Neotype, ZSI/WGRC/V/A/958, (female); G. Ramanella nagaoi, Holotype, WHT 2826 (male); H. Callula obscura, Holotype, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20) (female); I. Ramanella palmata, Holotype, NHM 90.11.8.62 (ex.BMNH 1947.2.11.21); J. Callula triangularis, Lectotype, NHM 74.4.29.897 (ex.BMNH 1947.2.11.15) (male); K.Hylaedactylus carnaticus, Lectotype, ZSI 2761(female); L. Uperodonrohani sp. nov., Holotype, DZ1449 (male).(Images: SD Biju and Sonali Garg)." httpUri="https://zenodo.org/record/1183461/files/figure.png" pageId="33" pageNumber="34">Figs. 3H</figureCitation>
|
||
,
|
||
<figureCitation id="134609856C1C946AFE18CAFAFDA9F917" box="[469,521,1680,1704]" captionStart="FIGURE 10" captionStartId="33.[151,250,1229,1251]" captionTargetBox="[157,1426,537,1193]" captionTargetId="figure@33.[151,1436,531,1208]" captionTargetPageId="33" captionText="FIGURE 10. Head, hand, foot and foot webbing in Uperodon obscurus, U. palmatus, U. triangularis and U.variegatus. A–D. Holotype ofUperodon obscurus(NHM58.11.28.42 ex.BMNH 1947.2.11.20, female):A. Lateralview of head; B. ventral view of hand; C. ventral view of foot; D. schematic illustration of foot webbing; E–H. Holotype of Uperodon palmatus (NHM 90.11.8.62 ex. BMNH 1947.2.11.21, female): E. Lateral view of head; F. ventral view of hand; G. ventral view of foot; H. schematic illustration of foot webbing; I–L. Lectotype of Uperodon triangularis (NHM 74.4.29.897 ex. BMNH 1947.2.11.15, male): I. Lateral view of head; J. ventral view of hand; K. ventral view of foot; L. schematic illustration of foot webbing; M. Lectotype of Uperodon variegatus (ZSI 2761, female), lateral view of head; N–P. Referred specimen of Uperodon variegatus (ZSI 4726, female): N. ventral view of hand; O. ventral view of foot; P. schematic illustration of foot webbing. (Illustrations: SD Biju)." httpUri="https://zenodo.org/record/1183475/files/figure.png" pageId="33" pageNumber="34">10A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C1C946AFF0ACADEFE02F877" blockId="33.[151,1436,1572,2029]" pageId="33" pageNumber="34">
|
||
<emphasis id="B909C9126C1C946AFF0ACADEFEFFF972" bold="true" box="[199,351,1716,1741]" pageId="33" pageNumber="34">Colouration.</emphasis>
|
||
<emphasis id="B909C9126C1C946AFEA4CADFFDB7F973" box="[361,535,1717,1740]" italics="true" pageId="33" pageNumber="34">In preservation</emphasis>
|
||
: Dorsum greyish-brown with dark brown median patch; flanks light greyishbrown; dorsal surfaces of forelimbs, thigh, shank and foot, light brown; chest and belly, dark grey with faint grey spots; ventral surfaces of thigh, tibia and foot, greyish-brown with faint grey spots (
|
||
<figureCitation id="134609856C1C946AFB95CA96FB14F8AB" box="[1112,1204,1788,1813]" captionStart="FIGURE 3" captionStartId="15.[151,250,1749,1771]" captionTargetBox="[151,1436,644,1728]" captionTargetId="figure@15.[151,1436,644,1728]" captionTargetPageId="15" captionText="FIGURE 3. Preserved type specimens of Uperodon species in dorsal and ventral view. A. Uperodon systoma, Syntype, ZMB 3551 (female); B. Uperodon globulosus, Syntype, NHM 62.12.29.4 (ex. BMNH 1947.2.10.97) (female); C. Kaloula pulchra taprobanica, Holotype, NHM 1905.3.25.138 (ex. BMNH 1947.2.10.62) (female); D. Ramanella anamalaiensis, Neotype, ZSI/ WGRC/V/A/956 (male); E. Hylaedactylus montanus, Holotype, NHM 72.4.17.212 (ex. BMNH 1947.2.11.19) (female); F. Ramanella mormorata, Neotype, ZSI/WGRC/V/A/958, (female); G. Ramanella nagaoi, Holotype, WHT 2826 (male); H. Callula obscura, Holotype, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20) (female); I. Ramanella palmata, Holotype, NHM 90.11.8.62 (ex.BMNH 1947.2.11.21); J. Callula triangularis, Lectotype, NHM 74.4.29.897 (ex.BMNH 1947.2.11.15) (male); K.Hylaedactylus carnaticus, Lectotype, ZSI 2761(female); L. Uperodonrohani sp. nov., Holotype, DZ1449 (male).(Images: SD Biju and Sonali Garg)." httpUri="https://zenodo.org/record/1183461/files/figure.png" pageId="33" pageNumber="34">Fig. 3H</figureCitation>
|
||
).
|
||
<emphasis id="B909C9126C1C946AFB07CA96FAC5F8AA" box="[1226,1381,1788,1813]" italics="true" pageId="33" pageNumber="34">Colour in life</emphasis>
|
||
(DZ 1366): Dorsum light orangish-brown with irregular blackish-brown spots, prominent blackish-brown band extending from behind the eye up to vent where it becomes wider; lower parts of flank brown with light bluish-grey spots; dorsal surfaces of forelimb, thigh, shank and foot, orangish-brown with dark brown cross bands; throat light brown; chest greyish-brown; belly and ventral surfaces of forelimb, thigh, tibia and foot, brown with light bluishgrey spots (
|
||
<figureCitation id="134609856C1C946AFED5CBC5FE36F877" box="[280,406,1967,1992]" captionStart="FIGURE9" captionStartId="30.[151,250,1723,1745]" captionTargetBox="[151,1436,885,1697]" captionTargetId="figure@30.[151,1436,885,1701]" captionTargetPageId="30" captionText="FIGURE9. Uperodon mormorata, U. nagaoi and U.obscurus inlife.A–C. Uperodonmormorata (SDBDU 2016.3404,male): A. Dorsolateral view; B. dorsal view; C. ventral view; D. Uperodon mormorata (SDBDU 2011.835, male), dorsolateral view; E–G. Uperodon nagaoi (DZ 1367, male): E. Dorsolateral view; F. dorsal view; G. frontal view; H–K. Uperodon obscurus: H. Dorsolateral view (DZ 1366, male); I. Dorsal view (DZ 1366, male); J. dorsolateral view (DZ 1408, male); K. dorsolateral view (DZ 1369, male); L. ventral view (DZ 1369, male). (Images: SD Biju and E–G Madhava Meegaskumbura)." httpUri="https://zenodo.org/record/1183473/files/figure.png" pageId="33" pageNumber="34">Figs. 9H–I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C1C9469FF0ACBBEFE64FF47" blockId="33.[151,1436,1572,2029]" lastBlockId="34.[151,1436,151,608]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<emphasis id="B909C9126C1C946AFF0ACBBEFEEAF852" bold="true" box="[199,330,2004,2029]" pageId="33" pageNumber="34">Variations.</emphasis>
|
||
Morphometric measurements of 17 males and five females, including the holotype, are given in
|
||
<tableCitation id="C6FF20BB6C1F9469FF5ACCFDFF50FF0F" box="[151,240,151,176]" captionStart="TABLE 4" captionStartId="48.[119,193,547,566]" captionTargetBox="[135,1884,616,1625]" captionText="TABLE 4. Morphometric measurements of Uperodon species.All measurements are in millimetres (mm). The status of each specimen is given after the museum number:Holotype (HT), Paratype (PT), Neotype (NT), Lectotype (LT), Syntype (ST)." httpUri="http://table.plazi.org/id/DF0245886C0D947BFFBACE49F8ECFD89" pageId="34" pageNumber="35" tableUuid="DF0245886C0D947BFFBACE49F8ECFD89">Table 4</tableCitation>
|
||
. DZ 1408: dorsal skin bright orange and more prominently granular compared to DZ 1366; DZ 1369: dorsum orangish-grey with a dark grey median band, and ventral surfaces coffee brown with light bluish-grey spots, except on the throat.
|
||
</paragraph>
|
||
<paragraph id="8BC215006C1F9469FF0ACD6EFC7DFEFF" blockId="34.[151,1436,151,608]" pageId="34" pageNumber="35">
|
||
<emphasis id="B909C9126C1F9469FF0ACD6EFD84FEA2" bold="true" box="[199,548,260,285]" pageId="34" pageNumber="35">Secondary sexual characters.</emphasis>
|
||
Male (DZ 1366): vocal sac externally visible on the lower jaw; female (NHM 1982.1290): ova white, pigmented on pole (diameter 0.7–1.0 mm,
|
||
<emphasis id="B909C9126C1F9469FC4DCD43FC33FEFF" box="[896,915,297,320]" italics="true" pageId="34" pageNumber="35">N</emphasis>
|
||
= 20).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C1F9469FF0ACD26FEBEFDDF" blockId="34.[151,1436,151,608]" pageId="34" pageNumber="35">
|
||
<emphasis id="B909C9126C1F9469FF0ACD26FD38FEDA" bold="true" box="[199,664,332,357]" pageId="34" pageNumber="35">Geographical distribution and habitat.</emphasis>
|
||
<taxonomicName id="4C7D6E836C1F9469FD6DCD26FCDEFEDA" box="[672,894,332,357]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C1F9469FD6DCD26FCDEFEDA" box="[672,894,332,357]" italics="true" pageId="34" pageNumber="35">Uperodon obscurus</emphasis>
|
||
</taxonomicName>
|
||
is endemic to
|
||
<collectingCountry id="F36A55906C1F9469FBEBCD26FB34FEDA" box="[1062,1172,332,357]" name="Sri Lanka" pageId="34" pageNumber="35">
|
||
<collectingRegion id="49B9DBE26C1F9469FBEBCD26FBE7FEDA" box="[1062,1095,332,357]" country="United Kingdom" name="Surrey" pageId="34" pageNumber="35">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
and known to occur in
|
||
<collectingRegion id="49B9DBE26C1F9469FF5ACD05FF4CFE37" box="[151,236,367,392]" country="Sri Lanka" name="Central" pageId="34" pageNumber="35">Central</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C1F9469FF3ACD05FE3EFE38" box="[247,414,367,392]" country="Sri Lanka" name="North Western" pageId="34" pageNumber="35">
|
||
North
|
||
<collectingRegion id="49B9DBE26C1F9469FE8DCD1AFE3EFE38" box="[320,414,368,391]" country="American Samoa" name="Western" pageId="34" pageNumber="35">Western</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C1F9469FE64CD05FDF7FE37" box="[425,599,367,392]" country="Sri Lanka" name="Sabaragamuwa" pageId="34" pageNumber="35">Sabaragamuwa</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C1F9469FDAECD05FD68FE37" box="[611,712,367,392]" country="Sri Lanka" name="Southern" pageId="34" pageNumber="35">Southern</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B9DBE26C1F9469FD32CD1AFC8EFE38" box="[767,814,368,391]" country="Sri Lanka" name="Uva" pageId="34" pageNumber="35">Uva</collectingRegion>
|
||
Provinces (
|
||
<figureCitation id="134609856C1F9469FC7ECD05FBACFE37" box="[947,1036,367,392]" captionStart="FIGURE 2" captionStartId="14.[151,250,1617,1639]" captionTargetBox="[152,1434,196,1594]" captionTargetId="figure@14.[151,1436,193,1595]" captionTargetPageId="14" captionText="FIGURE 2. A. Geographic distribution of the genus Uperodon in Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka; B. Distributionof Uperodon globulosus, U. systomaand U.taprobanicus; C. Distribution of four endemic Uperodon species of the Western Ghats (excluding the widely distributed Indian member U. variegatus, shown in Fig.12A); D. Distribution of four endemic Uperdodon species of Sri Lanka. Details of localities reported in the present study are provided in Table 1. Records from literature are listed in supplementary Table S1 and indicated here based onapproximate coordinates." httpUri="https://zenodo.org/record/1183459/files/figure.png" pageId="34" pageNumber="35">Fig. 2D</figureCitation>
|
||
;
|
||
<tableCitation id="C6FF20BB6C1F9469FBD4CD05FB04FE37" box="[1049,1188,367,392]" captionStart="TABLE 1" captionStartId="4.[151,239,1593,1616]" captionText="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." pageId="34" pageNumber="35">Tables 1, S1</tableCitation>
|
||
). In the present study, we confirmed the occurrence of this species in Kandy district (Kandy, Peradeniya, Panwila, Pussellawa and Udawatta Kele), Kegalle district (Babilapitiya estate), Kurunegala district (Kurunegala), Matale district (Gammaduwa), Nuwara Eliya district (Bogawantalawa and Laxapana) and Ratnapura district (Sinharaja and Morningside), from elevations of
|
||
<quantity id="4C85B8E56C1F9469FDD4CE6AFDC2FDA8" box="[537,610,512,536]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="m" value="100.0">100 m</quantity>
|
||
up to
|
||
<quantity id="4C85B8E56C1F9469FD63CE6AFCA5FDA8" box="[686,773,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="34" pageNumber="35" unit="m" value="1220.0">1220 m</quantity>
|
||
asl. It was collected either from tree hollows in disturbed forests adjacent to urban areas (Peradeniya and Panwila), or temporary ponds surrounded with vegetation (Laxapana).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |