233 lines
31 KiB
XML
233 lines
31 KiB
XML
<document id="0547B6F91885A0F1205FEAF05CFC25F3" ID-CLB-Dataset="30247" ID-DOI="10.11646/zootaxa.4410.2.2" ID-GBIF-Dataset="1e81ce7c-8f17-41b1-add2-a19ea5f54d60" ID-ISSN="1175-5326" ID-Zenodo-Dep="1221512" ID-ZooBank="96DFDE73-A410-4D9B-9124-A5AC1070D112" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524196034089" checkinUser="plazi" docAuthor="Li, Xin-Ran, Wang, Li-Li & Wang, Zong-Qing" docDate="2018" docId="03D46C14C869010641CBFF1AFCB7FA65" docLanguage="en" docName="zootaxa.4410.2.2.pdf" docOrigin="Zootaxa 4410 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perisphaerinae Brunner von Wattenwyl 1865" docType="treatment" docVersion="7" lastPageNumber="254" masterDocId="FFED146CC86A0105415CFFC4FFD8FF87" masterDocTitle="Rediscovered and new perisphaerine cockroaches from SW China with a review of subfamilial diagnosis (Blattodea: Blaberidae)" masterLastPageNumber="290" masterPageNumber="251" pageNumber="254" updateTime="1698538287180" updateUser="ExternalLinkService">
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<mods:mods id="FA2F2273F0D39907BAD252BB0246C89E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CB0C6F4EC4DDD15898D68BF1B66F7542">
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<mods:title id="3B55B670265EF29EFCFF19DDA428509E">Rediscovered and new perisphaerine cockroaches from SW China with a review of subfamilial diagnosis (Blattodea: Blaberidae)</mods:title>
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<mods:name id="4D081161BA3FF7F595F8A84A239158CA" type="personal">
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<mods:namePart id="10C2E5936A3E759B2B1EC2E18F2153B5">Li, Xin-Ran</mods:namePart>
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<mods:namePart id="02106ACC12D3C5850B179A2A82262FAD">Wang, Li-Li</mods:namePart>
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<mods:roleTerm id="E38BBCC3837D2AFB0047EFE64C6B3C2A">Author</mods:roleTerm>
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<mods:namePart id="B860C250E91850061159BAC2974EEAED">Wang, Zong-Qing</mods:namePart>
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<mods:title id="01BD6DDA51497EFB5361656EF888DCD1">Zootaxa</mods:title>
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<mods:part id="333725D8838FE07346834B912AF6842E">
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<mods:date id="40AAB5254F682A18B330880C0D23ABAB">2018</mods:date>
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<mods:number id="D9FDFC4E16DA2DEEF542A0703138C843">2018-04-17</mods:number>
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<mods:number id="80F1DDFB0F9813FF639BFD907AE0E9C9">4410</mods:number>
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<mods:identifier id="F82E8584FCFD7C0772CECD09216EF4E1" type="DOI">10.11646/zootaxa.4410.2.2</mods:identifier>
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<treatment id="03D46C14C869010641CBFF1AFCB7FA65" ID-DOI="http://doi.org/10.5281/zenodo.5986546" ID-GBIF-Taxon="143618728" ID-Zenodo-Dep="5986546" LSID="urn:lsid:plazi:treatment:03D46C14C869010641CBFF1AFCB7FA65" httpUri="http://treatment.plazi.org/id/03D46C14C869010641CBFF1AFCB7FA65" lastPageNumber="254" pageId="3" pageNumber="254">
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<subSubSection id="C3678E89C869010641CBFF1AFDF2FE9D" pageId="3" pageNumber="254" type="nomenclature">
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<paragraph id="8BC2DD02C869010641CBFF1AFE04FF7F" blockId="3.[151,554,222,282]" box="[151,476,222,248]" pageId="3" pageNumber="254">
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<heading id="D08A6A6EC869010641CBFF1AFE04FF7F" bold="true" box="[151,476,222,248]" fontSize="11" level="1" pageId="3" pageNumber="254" reason="1">
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<emphasis id="B9090110C869010641CBFF1AFE04FF7F" bold="true" box="[151,476,222,248]" pageId="3" pageNumber="254">
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Subfamily
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<taxonomicName id="4C7DA681C86901064043FF1AFE04FF7F" ID-CoL="8N8H2" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[287,476,222,248]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC2DD02C869010641CBFEC5FDF2FE9D" blockId="3.[151,554,222,282]" box="[151,554,257,282]" pageId="3" pageNumber="254">
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See
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<bibRefCitation id="EFECA0F3C8690106419AFEC5FE4DFE9D" box="[198,405,257,282]" pageId="3" pageNumber="254" refId="ref11777" refString="Li, X. - R. & Wang, Z. - Q. (2016) Perisphaerinae Brunner von Wattenwyl and Hyposphaeria Lucas are valid names concealed by the unavailable names Perisphaeriinae and Perisphaeria Burmeister (Blattodea, Blaberidae). ZooKeys, 574, 75 - 80. https: // doi. org / 10.3897 / zookeys. 574.7063" type="journal article">Li & Wang (2016)</bibRefCitation>
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for citations.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3678E89C869010641CBFE8DFA53FC95" pageId="3" pageNumber="254" type="diagnosis">
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<paragraph id="8BC2DD02C869010641CBFE8DFA53FC95" blockId="3.[151,1437,329,1506]" pageId="3" pageNumber="254">
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<emphasis id="B9090110C869010641CBFE8DFE5BFEE5" bold="true" box="[151,387,329,354]" pageId="3" pageNumber="254">Tentative diagnosis.</emphasis>
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Female neoteny (
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<figureCitation id="1346C187C86901064311FE8DFD49FEE6" box="[589,657,329,354]" captionStart="FIGURE 9" captionStartId="5.[151,250,1681,1703]" captionTargetBox="[151,1436,193,1660]" captionTargetId="figure@5.[151,1436,193,1660]" captionTargetPageId="5" captionText="FIGURE 9. Pseudoglomeris magnifica, a famous species and also an example of perisphaerines. A–C. Female, arrowheads indicate pits.D–G. Male, the white line contours the typical carina.Scale bar:5mm (A, B, D, E) or 1mm (C, F, G)." httpUri="https://zenodo.org/record/1221516/files/figure.png" pageId="3" pageNumber="254">Fig. 9</figureCitation>
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), integument hardened (in male adults, limited to pronotum and base of tegmen); ventral pronotum with typical carinae, which end with a lobe or a process, more evident in males (
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<figureCitation id="1346C187C86901064433FEA9FF64FE2E" captionStart="FIGURE 9" captionStartId="5.[151,250,1681,1703]" captionTargetBox="[151,1436,193,1660]" captionTargetId="figure@5.[151,1436,193,1660]" captionTargetPageId="5" captionText="FIGURE 9. Pseudoglomeris magnifica, a famous species and also an example of perisphaerines. A–C. Female, arrowheads indicate pits.D–G. Male, the white line contours the typical carina.Scale bar:5mm (A, B, D, E) or 1mm (C, F, G)." httpUri="https://zenodo.org/record/1221516/files/figure.png" pageId="3" pageNumber="254">Fig. 9G</figureCitation>
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). Front femur type C or D. In females of many species, the abdominal tergites bear one or more pits at sides, along the tergal furrow (
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<figureCitation id="1346C187C869010640FBFE71FE27FE49" box="[423,511,437,462]" captionStart="FIGURE 9" captionStartId="5.[151,250,1681,1703]" captionTargetBox="[151,1436,193,1660]" captionTargetId="figure@5.[151,1436,193,1660]" captionTargetPageId="5" captionText="FIGURE 9. Pseudoglomeris magnifica, a famous species and also an example of perisphaerines. A–C. Female, arrowheads indicate pits.D–G. Male, the white line contours the typical carina.Scale bar:5mm (A, B, D, E) or 1mm (C, F, G)." httpUri="https://zenodo.org/record/1221516/files/figure.png" pageId="3" pageNumber="254">Fig. 9C</figureCitation>
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); an orifice usually present in the pit. The number of pits on either side of the third through seventh tergum is a diagnostic character of species, and it can be stated as ‘
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<emphasis id="B9090110C8690106451FFE1DFB13FE75" bold="true" box="[1091,1227,473,498]" pageId="3" pageNumber="254">pit formula</emphasis>
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’. For example, pit formula 1-1-2-2-2 means that third tergum through seventh tergum typically has one, one, two, two, two pits respectively; pit formula [3] means that either side of third tergum through seventh tergum each has three pits typically. Cerci of nymphs and females very short, coniform, coalesced into one segment, sometimes the cercomeres are observable. The male phallic complex is generalized as
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<figureCitation id="1346C187C86901064550FDADFBA3FD05" box="[1036,1147,617,642]" captionStart="FIGURE 10" captionStartId="6.[151,250,1158,1180]" captionTargetBox="[153,1432,413,1131]" captionTargetId="figure@6.[151,1435,409,1137]" captionTargetPageId="6" captionText="FIGURE 10. Generalized diagram of male and female genitalia. A. Male phallic complex, with membrane depicted as transparent dotted area. B. Female valvulae and surrounding components, with membrane depicted as opaque white area.Note that either the valvula I or the valvulaeII and III are artificially reversed.Dashed line, cutting line; circled line, folding line." httpUri="https://zenodo.org/record/1221518/files/figure.png" pageId="3" pageNumber="254">Fig. 10A</figureCitation>
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. Sclerotizations of cleft phallomere are R1T’, R2’, R3’ and R5’; virga well developed (except for
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<taxonomicName id="4C7DA681C869010642B1FD49FDEBFD4D" authority="(Wang & Che))" authorityName="Li & Wang & Wang" authorityYear="2018" baseAuthorityName="Li & Wang & Wang" baseAuthorityYear="2018" class="Insecta" family="Blaberidae" genus="Pseudoglomeris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="variety" species="semisulcata" variety="angustifolia">
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<emphasis id="B9090110C869010642B1FD49FAF7FD21" box="[1005,1327,653,678]" italics="true" pageId="3" pageNumber="254">Pseudoglomeris semisulcata</emphasis>
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Hanitsch and
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<emphasis id="B9090110C8690106419BFD76FEA1FD4D" box="[199,377,689,714]" italics="true" pageId="3" pageNumber="254">Ps. angustifolia</emphasis>
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(Wang & Che))
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</taxonomicName>
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, the virga sclerite occupies the dorsal wall and sometimes part of the ventral wall of the virga; basolateral sclerite large, approximately as wide as the hook-like phallomere; the hook-base sclerite and the hook at the same side. Female valvulae and surrounding components are generalized as
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<figureCitation id="1346C187C8690106447EFD3DFA5FFC95" box="[1314,1415,761,786]" captionStart="FIGURE 10" captionStartId="6.[151,250,1158,1180]" captionTargetBox="[153,1432,413,1131]" captionTargetId="figure@6.[151,1435,409,1137]" captionTargetPageId="6" captionText="FIGURE 10. Generalized diagram of male and female genitalia. A. Male phallic complex, with membrane depicted as transparent dotted area. B. Female valvulae and surrounding components, with membrane depicted as opaque white area.Note that either the valvula I or the valvulaeII and III are artificially reversed.Dashed line, cutting line; circled line, folding line." httpUri="https://zenodo.org/record/1221518/files/figure.png" pageId="3" pageNumber="254">Fig. 10B</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3678E89C8690106419BFCD9FA4BFCB1" box="[199,1427,797,822]" pageId="3" pageNumber="254" type="distribution">
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<paragraph id="8BC2DD02C8690106419BFCD9FA4BFCB1" blockId="3.[151,1437,329,1506]" box="[199,1427,797,822]" pageId="3" pageNumber="254">
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<emphasis id="B9090110C8690106419BFCD9FEB8FCB1" bold="true" box="[199,352,797,822]" pageId="3" pageNumber="254">Distribution.</emphasis>
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Oriental, Australian and Ethiopian Regions.
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<figureCitation id="1346C187C86901064206FCD9FC70FCB1" box="[858,936,797,822]" captionStart="FIGURE 11" captionStartId="6.[151,250,1905,1927]" captionTargetBox="[151,1436,1291,1883]" captionTargetId="figure@6.[151,1436,1291,1883]" captionTargetPageId="6" captionText="FIGURE 11. Localities in southwestern China. For details see Appendix I. Map image: Google, Copernicus; data: SIO, NOAA, U.S. Navy,NGA, GEBCO." httpUri="https://zenodo.org/record/1221520/files/figure.png" pageId="3" pageNumber="254">Fig. 11</figureCitation>
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shows the localities in southwestern
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<collectingCountry id="F36A9D92C86901064415FCD9FA48FCB1" box="[1353,1424,797,822]" name="China" pageId="3" pageNumber="254">China</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3678E89C8690106419BFC85FCB7FA65" pageId="3" pageNumber="254" type="discussion">
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<paragraph id="8BC2DD02C8690106419BFC85FB7AFBD2" blockId="3.[151,1437,329,1506]" pageId="3" pageNumber="254">
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<emphasis id="B9090110C8690106419BFC85FEE3FCDD" bold="true" box="[199,315,833,858]" pageId="3" pageNumber="254">Remarks.</emphasis>
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<taxonomicName id="4C7DA681C8690106401BFC85FE04FCDD" authorityName="Audinet-Serville" authorityYear="1831" box="[327,476,833,858]" class="Insecta" family="Blaberidae" genus="Perisphaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
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<emphasis id="B9090110C8690106401BFC85FE04FCDD" box="[327,476,833,858]" italics="true" pageId="3" pageNumber="254">Perisphaerus</emphasis>
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</taxonomicName>
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is the
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<typeStatus id="54C663A0C86901064369FC86FDBDFCDD" box="[565,613,834,858]" pageId="3" pageNumber="254">type</typeStatus>
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genus of
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<taxonomicName id="4C7DA681C869010643BEFC85FC53FCDD" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[738,907,833,858]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
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. But unfortunately,
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<bibRefCitation id="EFECA0F3C8690106452BFC86FAF1FCDD" box="[1143,1321,833,858]" pageId="3" pageNumber="254" refId="ref11431" refString="Brunner von Wattenwyl, C. (1865) Nouveau Systeme des Blattaires. G. Braumuller, Vienna, 426 pp." type="book">Brunner (1865)</bibRefCitation>
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proposed
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<taxonomicName id="4C7DA681C869010641CBFCA1FEE4FCF9" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[151,316,869,894]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
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(originally as a tribe) based on the name of
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<taxonomicName id="4C7DA681C8690106422BFCA1FBD4FCF9" authorityName="Audinet-Serville" authorityYear="1831" box="[887,1036,869,894]" class="Insecta" family="Blaberidae" genus="Perisphaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
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<emphasis id="B9090110C8690106422BFCA1FBD4FCF9" box="[887,1036,869,894]" italics="true" pageId="3" pageNumber="254">Perisphaerus</emphasis>
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</taxonomicName>
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, whilst based on the concept of
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<taxonomicName id="4C7DA681C869010641CBFC4DFE5FFC26" authority="Lucas" authorityName="Lucas" authorityYear="1863" box="[151,391,905,930]" class="Insecta" family="Blaberidae" genus="Hyposphaeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
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<emphasis id="B9090110C869010641CBFC4DFEEEFC25" box="[151,310,905,930]" italics="true" pageId="3" pageNumber="254">Hyposphaeria</emphasis>
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Lucas
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</taxonomicName>
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, though he did not mention the latter (see
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<bibRefCitation id="EFECA0F3C869010642D9FC4DFB96FC25" box="[901,1102,905,930]" pageId="3" pageNumber="254" refId="ref11777" refString="Li, X. - R. & Wang, Z. - Q. (2016) Perisphaerinae Brunner von Wattenwyl and Hyposphaeria Lucas are valid names concealed by the unavailable names Perisphaeriinae and Perisphaeria Burmeister (Blattodea, Blaberidae). ZooKeys, 574, 75 - 80. https: // doi. org / 10.3897 / zookeys. 574.7063" type="journal article">Li & Wang 2016</bibRefCitation>
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||
for details).
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<bibRefCitation id="EFECA0F3C869010645B6FC4EFA44FC25" box="[1258,1436,905,930]" pageId="3" pageNumber="254" refId="ref11431" refString="Brunner von Wattenwyl, C. (1865) Nouveau Systeme des Blattaires. G. Braumuller, Vienna, 426 pp." type="book">Brunner (1865)</bibRefCitation>
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||
included the
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<typeStatus id="54C663A0C86901064065FC6AFEB1FC41" box="[313,361,942,966]" pageId="3" pageNumber="254">type</typeStatus>
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||
species of
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<taxonomicName id="4C7DA681C8690106435EFC69FD4FFC41" authorityName="Audinet-Serville" authorityYear="1831" box="[514,663,941,966]" class="Insecta" family="Blaberidae" genus="Perisphaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
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<emphasis id="B9090110C8690106435EFC69FD4FFC41" box="[514,663,941,966]" italics="true" pageId="3" pageNumber="254">Perisphaerus</emphasis>
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</taxonomicName>
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into
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<taxonomicName id="4C7DA681C869010643BEFC69FCA7FC41" box="[738,895,941,966]" class="Insecta" family="Blaberidae" genus="Derocalymma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
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<emphasis id="B9090110C869010643BEFC69FCA7FC41" box="[738,895,941,966]" italics="true" pageId="3" pageNumber="254">Derocalymma</emphasis>
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</taxonomicName>
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, and later he (
|
||
<bibRefCitation id="EFECA0F3C8690106451EFC6AFB31FC41" box="[1090,1257,942,966]" pageId="3" pageNumber="254" refId="ref11454" refString="Brunner von Wattenwyl, C. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 13, 5 - 230. https: // doi. org / 10.5962 / bhl. title. 5121" type="journal article">Brunner 1893</bibRefCitation>
|
||
) included two
|
||
<taxonomicName id="4C7DA681C869010641CBFC15FEF4FC6D" authorityName="Audinet-Serville" authorityYear="1831" box="[151,300,977,1002]" class="Insecta" family="Blaberidae" genus="Perisphaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C869010641CBFC15FEF4FC6D" box="[151,300,977,1002]" italics="true" pageId="3" pageNumber="254">Perisphaerus</emphasis>
|
||
</taxonomicName>
|
||
species into
|
||
<taxonomicName id="4C7DA681C86901064087FC15FD27FC6E" authority="Brunner" authorityName="Brunner" box="[475,767,977,1002]" class="Insecta" family="Blaberidae" genus="Pseudoglomeris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C86901064087FC15FD48FC6D" box="[475,656,977,1002]" italics="true" pageId="3" pageNumber="254">Pseudoglomeris</emphasis>
|
||
Brunner
|
||
</taxonomicName>
|
||
; the typification of
|
||
<taxonomicName id="4C7DA681C8690106455DFC15FB7EFC6D" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[1025,1190,977,1002]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
|
||
is therefore further complicated. To avoid confusion, we treat
|
||
<taxonomicName id="4C7DA681C86901064326FC31FCD7FB89" authorityName="Audinet-Serville" authorityYear="1831" box="[634,783,1013,1038]" class="Insecta" family="Blaberidae" genus="Perisphaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C86901064326FC31FCD7FB89" box="[634,783,1013,1038]" italics="true" pageId="3" pageNumber="254">Perisphaerus</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="54C663A0C8690106423DFC32FC49FB89" box="[865,913,1014,1038]" pageId="3" pageNumber="254">type</typeStatus>
|
||
genus of
|
||
<taxonomicName id="4C7DA681C8690106455EFC31FB72FB89" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[1026,1194,1013,1038]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
|
||
, nomenclaturally and actually. In addition, if the name
|
||
<taxonomicName id="4C7DA681C86901064346FBDDFCB4FBB5" authority="Brunner, 1865" authorityName="Brunner" authorityYear="1865" box="[538,876,1049,1074]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">
|
||
Perisphaerinae
|
||
<bibRefCitation id="EFECA0F3C86901064394FBDEFCB4FBB5" box="[712,876,1049,1074]" pageId="3" pageNumber="254" refId="ref11431" refString="Brunner von Wattenwyl, C. (1865) Nouveau Systeme des Blattaires. G. Braumuller, Vienna, 426 pp." type="book">Brunner, 1865</bibRefCitation>
|
||
</taxonomicName>
|
||
becomes misleading, then we recommend using
|
||
<taxonomicName id="4C7DA681C869010641CBFBF9FE03FBD1" authority="Walker, 1868" authorityName="Walker" authorityYear="1868" box="[151,475,1085,1110]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">
|
||
Perisphaerinae
|
||
<bibRefCitation id="EFECA0F3C8690106401FFBF9FE03FBD1" box="[323,475,1085,1110]" pageId="3" pageNumber="254" refId="ref12387" refString="Walker, F. (1868) Catalogue of the Specimens of Blattariae in the Collection of the British Museum. British Museum, London, 239 pp." type="book">Walker, 1868</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is based on both the name and concept of
|
||
<taxonomicName id="4C7DA681C8690106455AFBF9FB43FBD1" authorityName="Audinet-Serville" authorityYear="1831" box="[1030,1179,1085,1110]" class="Insecta" family="Blaberidae" genus="Perisphaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C8690106455AFBF9FB43FBD1" box="[1030,1179,1085,1110]" italics="true" pageId="3" pageNumber="254">Perisphaerus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC2DD02C8690106419BFBA5FCB7FA65" blockId="3.[151,1437,329,1506]" pageId="3" pageNumber="254">
|
||
<bibRefCitation id="EFECA0F3C8690106419BFBA5FE42FBFD" box="[199,410,1121,1146]" pageId="3" pageNumber="254" refId="ref11514" refString="Grandcolas, P. (1997) The monophyly of the subfamily Perisphaeriinae (Dictyoptera: Blattaria: Blaberidae). Systematic Entomology, 22 (2), 123 - 130. https: // doi. org / 10.1046 / j. 1365 - 3113.1997. d 01 - 28. x" type="journal article">Grandcolas (1997)</bibRefCitation>
|
||
suggested five synapomorphies for
|
||
<taxonomicName id="4C7DA681C86901064261FBA5FC3AFBFD" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[829,994,1121,1146]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
|
||
(see Introduction herein). The third is that the paratergite (tergal extension in
|
||
<figureCitation id="1346C187C8690106430CFB41FD6BFB19" box="[592,691,1157,1182]" captionStart="FIGURE 10" captionStartId="6.[151,250,1158,1180]" captionTargetBox="[153,1432,413,1131]" captionTargetId="figure@6.[151,1435,409,1137]" captionTargetPageId="6" captionText="FIGURE 10. Generalized diagram of male and female genitalia. A. Male phallic complex, with membrane depicted as transparent dotted area. B. Female valvulae and surrounding components, with membrane depicted as opaque white area.Note that either the valvula I or the valvulaeII and III are artificially reversed.Dashed line, cutting line; circled line, folding line." httpUri="https://zenodo.org/record/1221518/files/figure.png" pageId="3" pageNumber="254">Fig. 10B</figureCitation>
|
||
) of female genitalia has an unusual groove and thus a “lip” on the other side. This character, however, is found at least in
|
||
<taxonomicName id="4C7DA681C86901064213FB6DFB94FB45" authority="(Blaberinae)" authorityName="Stål" authorityYear="1874" baseAuthorityName="Blaberinae" box="[847,1100,1193,1218]" class="Insecta" family="Blaberidae" genus="Blaptica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C86901064213FB6DFC76FB45" box="[847,942,1193,1218]" italics="true" pageId="3" pageNumber="254">Blaptica</emphasis>
|
||
(Blaberinae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7DA681C86901064502FB6DFA40FB45" authority="(Diplopterinae)" baseAuthorityName="Diplopterinae" box="[1118,1432,1193,1218]" class="Insecta" family="Blaberidae" genus="Diploptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C86901064502FB6DFB01FB45" box="[1118,1241,1193,1218]" italics="true" pageId="3" pageNumber="254">Diploptera</emphasis>
|
||
(Diplopterinae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7DA681C869010641CBFB09FEFFFB61" box="[151,295,1229,1254]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Panesthiinae">Panesthiinae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7DA681C8690106401CFB09FE36FB61" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[320,494,1229,1254]" class="Insecta" family="Blaberidae" genus="Paranauphoeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C8690106401CFB09FE36FB61" box="[320,494,1229,1254]" italics="true" pageId="3" pageNumber="254">Paranauphoeta</emphasis>
|
||
</taxonomicName>
|
||
(subfamily uncertain) and
|
||
<taxonomicName id="4C7DA681C86901064208FB09FB0AFB61" authority="(Epilamprinae)" authorityName="Kirby" authorityYear="1903" baseAuthorityName="Epilamprinae" box="[852,1234,1229,1254]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090110C86901064208FB09FBCDFB61" box="[852,1045,1229,1254]" italics="true" pageId="3" pageNumber="254">Pseudophoraspis</emphasis>
|
||
(Epilamprinae)
|
||
</taxonomicName>
|
||
(XRL personal observation). This is common among
|
||
<taxonomicName id="4C7DA681C86901064316FB35FD1EFA8D" box="[586,710,1265,1290]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
|
||
, and likely extends beyond just
|
||
<taxonomicName id="4C7DA681C86901064565FB35FB6EFA8D" box="[1081,1206,1265,1290]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
|
||
. Besides, we do not consider that the tergal extension is useful in subfamilial taxonomy in
|
||
<taxonomicName id="4C7DA681C86901064295FAD1FB9EFAA9" box="[969,1094,1301,1326]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
|
||
. The fourth synapomorphy in
|
||
<bibRefCitation id="EFECA0F3C869010641CBFAFDFEB1FAD5" box="[151,361,1337,1362]" pageId="3" pageNumber="254" refId="ref11514" refString="Grandcolas, P. (1997) The monophyly of the subfamily Perisphaeriinae (Dictyoptera: Blattaria: Blaberidae). Systematic Entomology, 22 (2), 123 - 130. https: // doi. org / 10.1046 / j. 1365 - 3113.1997. d 01 - 28. x" type="journal article">Grandcolas (1997)</bibRefCitation>
|
||
is that the first valvifer arm (gonocoxa in
|
||
<figureCitation id="1346C187C86901064217FAFDFC77FAD5" box="[843,943,1337,1362]" captionStart="FIGURE 10" captionStartId="6.[151,250,1158,1180]" captionTargetBox="[153,1432,413,1131]" captionTargetId="figure@6.[151,1435,409,1137]" captionTargetPageId="6" captionText="FIGURE 10. Generalized diagram of male and female genitalia. A. Male phallic complex, with membrane depicted as transparent dotted area. B. Female valvulae and surrounding components, with membrane depicted as opaque white area.Note that either the valvula I or the valvulaeII and III are artificially reversed.Dashed line, cutting line; circled line, folding line." httpUri="https://zenodo.org/record/1221518/files/figure.png" pageId="3" pageNumber="254">Fig. 10B</figureCitation>
|
||
) of female genitalia has a small fold on the right side, which often separates the sclerite. This is, however, found in only two out of 15 species (and some undetermined specimens) examined, namely
|
||
<taxonomicName id="4C7DA681C869010643CBFA45FCCAFA1E" box="[663,786,1409,1434]" pageId="3" pageNumber="254" rank="subSpecies" subSpecies="aerea">
|
||
,
|
||
<emphasis id="B9090110C869010643F9FA46FCCAFA1E" box="[677,786,1410,1433]" italics="true" pageId="3" pageNumber="254">Ps. aerea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7DA681C8690106420CFA46FC16FA1D" authorityName="Li & Wang & Wang" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1958" box="[848,974,1409,1434]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B9090110C8690106420CFA46FC16FA1D" box="[848,974,1409,1434]" italics="true" pageId="3" pageNumber="254">Ps. sculpta</emphasis>
|
||
</taxonomicName>
|
||
. In addition, we agree with Anisyutkin (2003) about the first and second synapomorphies. Therefore, the first through fourth synapomorphies suggested by
|
||
<bibRefCitation id="EFECA0F3C869010641E6FA0DFE53FA65" box="[186,395,1481,1506]" pageId="3" pageNumber="254" refId="ref11514" refString="Grandcolas, P. (1997) The monophyly of the subfamily Perisphaeriinae (Dictyoptera: Blattaria: Blaberidae). Systematic Entomology, 22 (2), 123 - 130. https: // doi. org / 10.1046 / j. 1365 - 3113.1997. d 01 - 28. x" type="journal article">Grandcolas (1997)</bibRefCitation>
|
||
are not synapomorphies of
|
||
<taxonomicName id="4C7DA681C8690106439FFA0DFCB4FA65" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[707,876,1481,1506]" class="Insecta" family="Blaberidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subFamily" subFamily="Perisphaerinae">Perisphaerinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |