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<document id="ACD193F702F664D75767CCB9345D69A9" ID-DOI="10.26107/RBZ-2023-0046" ID-ISSN="2345-7600" ID-Zenodo-Dep="10271430" ID-ZooBank="8C4D372F-CB5D-4DF1-913C-F27B6D5EC32E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701819552854" checkinUser="felipe" docAuthor="Chen, Zhe-Yu &amp; Páll-Gergely, Barna" docDate="2023" docId="03D3E319B975E63DFC71C389FF6AF999" docLanguage="en" docName="RafflesBZool.71.597-605.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Anaglyphula cancellata Rensch 1932" docType="treatment" docVersion="2" lastPageNumber="602" masterDocId="FFEA9B61B977E638FFDFC554FFACFFE4" masterDocTitle="A new species of Anaglyphula Rensch, 1932 from an island off the coast of West Indonesia, with redescription of the type species (Gastropoda: Caenogastropoda: Assimineidae)" masterLastPageNumber="605" masterPageNumber="597" pageNumber="599" updateTime="1701820180598" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7165769B269E1BFD0B862F44A2A0B0D7">A new species of Anaglyphula Rensch, 1932 from an island off the coast of West Indonesia, with redescription of the type species (Gastropoda: Caenogastropoda: Assimineidae)</mods:title>
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<paragraph id="8BC5520FB975E63AFC71C389FAF9F911" blockId="2.[942,1365,1757,1813]" box="[942,1365,1757,1781]" pageId="2" pageNumber="599">
<heading id="D08DE563B975E63AFC71C389FAF9F911" bold="true" box="[942,1365,1757,1781]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="599" reason="2">
<taxonomicName id="4C7A298CB975E63AFC71C389FAF9F911" ID-CoL="66JXS" authority="Rensch, 1932" authorityName="Rensch" authorityYear="1932" box="[942,1365,1757,1781]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="599" phylum="Mollusca" rank="species" species="cancellata">
<emphasis id="B90E8E1DB975E63AFC71C389FAF9F911" bold="true" box="[942,1365,1757,1781]" pageId="2" pageNumber="599">
<emphasis id="B90E8E1DB975E63AFC71C389FB1AF911" bold="true" box="[942,1206,1757,1781]" italics="true" pageId="2" pageNumber="599">Anaglyphula cancellata</emphasis>
<bibRefCitation id="EFEB2FFEB975E63AFB61C389FAF9F911" author="Rensch B" box="[1214,1365,1757,1781]" pageId="2" pageNumber="599" pagination="1 - 130" refId="ref4322" refString="Rensch B (1932) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba II. Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 63: 1 - 130, 3 pls." type="journal article" year="1932">Rensch, 1932</bibRefCitation>
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<paragraph id="8BC5520FB975E63AFBFBC3A9FB73F8F1" blockId="2.[942,1365,1757,1813]" box="[1060,1247,1789,1813]" pageId="2" pageNumber="599">
(
<figureCitation id="13414E8AB975E63AFBF3C3A9FBD4F8F1" box="[1068,1144,1789,1813]" captionStart="Fig" captionStartId="1.[113,148,1947,1969]" captionTargetBox="[113,1454,306,1932]" captionTargetId="figure-9@1.[113,1454,306,1932]" captionTargetPageId="1" captionText="Fig. 1. Anaglyphula cancellata Rensch, 1932, syntype NHMUK 1897.8.3.150. A, Shell; B, Upper whorls; C, Structure on body whorl; D, Detailed structure on penultimate whorl." figureDoi="http://doi.org/10.5281/zenodo.10271433" httpUri="https://zenodo.org/record/10271433/files/figure.png" pageId="2" pageNumber="599">Figs. 1</figureCitation>
,
<figureCitation id="13414E8AB975E63AFB58C3A9FB06F8F1" box="[1159,1194,1789,1813]" captionStart="Fig" captionStartId="3.[113,148,2064,2086]" captionTargetBox="[125,1440,189,2049]" captionTargetId="figure-9@3.[125,1440,189,2049]" captionTargetPageId="3" captionText="Fig. 2. A, Anaglyphula cancellata Rensch, 1932, syntype NHMUK 1897.8.3.149; BC, Anaglyphula whitteni Vermeulen, 1996, Paratype. NHMUK 20000247; B, Shell; C, Ribbing structure details." figureDoi="http://doi.org/10.5281/zenodo.10271439" httpUri="https://zenodo.org/record/10271439/files/figure.png" pageId="2" pageNumber="599">2A</figureCitation>
,
<figureCitation id="13414E8AB975E63AFB68C3A9FB7BF8F1" box="[1207,1239,1789,1813]" captionStart="Fig" captionStartId="5.[113,148,606,628]" captionTargetBox="[113,1451,189,591]" captionTargetId="figure-9@5.[113,1451,189,591]" captionTargetPageId="5" captionText="Fig. 4. A, Putative constriction position of Anaglyphula yanseni, new species, holotype MZB.Gst. 23.829; B, Constriction of Anaglyphula cancellata Rensch, 1932, syntype NHMUK 1897.8.3.150; C, Constriction of Anaglyphula whitteni Vermeulen, 1996, paratype NHMUK 20000247. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.10271449" httpUri="https://zenodo.org/record/10271449/files/figure.png" pageId="2" pageNumber="599">4B</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB6A7521B975E63AFC9DC268FAD9F8B5" box="[834,1397,1852,1874]" pageId="2" pageNumber="599">
<treatmentCitation id="0ADB741EB975E63AFC9DC268FB46F8B5" author="Rensch B" box="[834,1258,1852,1874]" page="123" pageId="2" pageNumber="599" year="1932">
<taxonomicName id="4C7A298CB975E63AFC9DC268FB46F8B5" ID-CoL="66JXS" authority="Rensch, 1932: 123" authorityName="Rensch" authorityPageNumber="123" authorityYear="1932" box="[834,1258,1852,1874]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="599" phylum="Mollusca" rank="species" species="cancellata">
<emphasis id="B90E8E1DB975E63AFC9DC268FB85F8B5" box="[834,1065,1852,1873]" italics="true" pageId="2" pageNumber="599">Anaglyphula cancellata</emphasis>
<bibRefCitation id="EFEB2FFEB975E63AFBEEC268FB46F8B5" author="Rensch B" box="[1073,1258,1852,1874]" pageId="2" pageNumber="599" pagination="1 - 130" refId="ref4322" refString="Rensch B (1932) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba II. Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 63: 1 - 130, 3 pls." type="journal article" year="1932">Rensch, 1932: 123</bibRefCitation>
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, pl. 3, fig. 38.
</treatmentCitationGroup>
</paragraph>
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<paragraph id="8BC5520FB975E63AFC9DC22EFAE0F856" blockId="2.[834,1473,1914,1970]" pageId="2" pageNumber="599">
<emphasis id="B90E8E1DB975E63AFC9DC22EFC5FF876" bold="true" box="[834,1011,1914,1938]" pageId="2" pageNumber="599">Type material.</emphasis>
<materialsCitation id="3B125852B975E63AFC21C22EFB3AF856" collectionCode="NHMUK" country="Indonesia" elevation="1219" location="Flores" pageId="2" pageNumber="599" specimenCount="5" typeStatus="syntype">
<typeStatus id="54C1ECADB975E63AFC21C22EFBCBF876" box="[1022,1127,1914,1938]" pageId="2" pageNumber="599" type="syntype">Syntypes</typeStatus>
.
<collectionCode id="ED6BCACAB975E63AFBABC22FFB76F876" box="[1140,1242,1915,1938]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="599" type="Museum">NHMUK</collectionCode>
1897.8.3.14953 (
<specimenCount id="9D7C9986B975E63AFA6CC22EFC19F856" count="5" pageId="2" pageNumber="599" type="generic">5 specimens</specimenCount>
), S.
<collectingCountry id="F36D129FB975E63AFC37C2CEFB87F856" box="[1000,1067,1946,1970]" name="Indonesia" pageId="2" pageNumber="599">Flores</collectingCountry>
(
<quantity id="4C82FFEAB975E63AFBE5C2CEFB22F856" box="[1082,1166,1946,1970]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="2" pageNumber="599" unit="ft" value="4000.0">
<elevation id="0057B53CB975E63AFBE5C2CEFB22F856" box="[1082,1166,1946,1970]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="2" pageNumber="599" unit="ft" value="4000.0">4000 ft</elevation>
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)
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, leg. A. Everett.
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<subSubSection id="C3600184B975E63DFC9DC28EFED8FA39" lastPageId="5" lastPageNumber="602" pageId="2" pageNumber="599" type="description">
<paragraph id="8BC5520FB975E63DFC9DC28EFE3FFAB9" blockId="2.[833,1473,2010,2098]" lastBlockId="5.[113,755,741,1373]" lastPageId="5" lastPageNumber="602" pageId="2" pageNumber="599">
<emphasis id="B90E8E1DB975E63AFC9DC28EFC42F816" bold="true" box="[834,1006,2010,2034]" pageId="2" pageNumber="599">Redescription.</emphasis>
Shell small, light brown, conical-ovoid, rather thick, somewhat translucent, 4.375 moderately convex whorls. Protoconch consisting of 1.25 whorls, with fine growth-lines; boundary between protoconch and teleoconch clearly visible. Teleoconch surface with fine and regularly arranged ribs, approximately four ribs per 200 μm, extending from the suture to the umbilical region; ribs rather straight and parallel near the suture, and on the middle of the body whorl, but branched and fused to a pitted structure in the upper middle and lower regions of the whorl, width of about
<superScript id="7C0FFF47B972E63DFF66C690FF6DFC36" attach="right" box="[185,193,964,978]" fontSize="6" pageId="5" pageNumber="602">1</superScript>
/5 of the whorl height; margins of the rib-meshes with some gradually descending sharp ridges; depressions (pits) irregular, round to elongated ellipsoids, tend to be divided by elongated ridges; lower rib-mesh situated just below the suture, therefore not visible on the upper whorls but only on the body whorl. Aperture semilunar, strongly oblique to the shell axis. Peristome discontinuous, parietal callus indicated by a thin and transparent calcareous layer; peristome thickened and expanded, very slightly reflected. A constriction formed by a thin radial palatal fold internally present ½ whorl away from the aperture, sometimes visible from the aperture. Umbilicus open, narrow, approximately
<superScript id="7C0FFF47B972E63DFFAEC010FFD5FAB6" attach="none" box="[113,121,1348,1362]" fontSize="6" pageId="5" pageNumber="602">1</superScript>
/7 of shell major diameter.
</paragraph>
<caption id="DF050287B974E63BFFAECD44FD19F7A7" ID-DOI="http://doi.org/10.5281/zenodo.10271439" ID-Zenodo-Dep="10271439" httpUri="https://zenodo.org/record/10271439/files/figure.png" pageId="3" pageNumber="600" startId="3.[113,148,2064,2086]" targetBox="[125,1440,189,2049]" targetPageId="3" targetType="figure">
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Fig. 2. A,
<taxonomicName id="4C7A298CB974E63BFF0ACD44FDEAF7C1" authority="Rensch, 1932" authorityName="Rensch" authorityYear="1932" box="[213,582,2064,2086]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="600" phylum="Mollusca" rank="species" species="cancellata">
<emphasis id="B90E8E1DB974E63BFF0ACD44FE17F7C1" box="[213,443,2064,2085]" italics="true" pageId="3" pageNumber="600">Anaglyphula cancellata</emphasis>
<bibRefCitation id="EFEB2FFEB974E63BFE1ECD44FDEAF7C1" author="Rensch B" box="[449,582,2064,2086]" pageId="3" pageNumber="600" pagination="1 - 130" refId="ref4322" refString="Rensch B (1932) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba II. Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 63: 1 - 130, 3 pls." type="journal article" year="1932">Rensch, 1932</bibRefCitation>
</taxonomicName>
, syntype NHMUK 1897.8.3.149; BC,
<taxonomicName id="4C7A298CB974E63BFC16CD44FAE4F7C2" authority="Vermeulen, 1996" authorityName="Vermeulen" authorityYear="1996" box="[969,1352,2064,2086]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="600" phylum="Mollusca" rank="species" species="whitteni">
<emphasis id="B90E8E1DB974E63BFC16CD44FB3BF7C1" box="[969,1175,2064,2085]" italics="true" pageId="3" pageNumber="600">Anaglyphula whitteni</emphasis>
<bibRefCitation id="EFEB2FFEB974E63BFB41CD44FAE4F7C2" author="Vermeulen JJ" box="[1182,1352,2064,2086]" pageId="3" pageNumber="600" pagination="149 - 162" refId="ref4506" refString="Vermeulen JJ (1996). Notes on terrestrial molluscs of Java, Bali and Nusa Penida. Basteria. 59: 149 - 162." type="journal article" year="1996">Vermeulen, 1996</bibRefCitation>
</taxonomicName>
, Paratype. NHMUK 20000247; B, Shell; C, Ribbing structure details.
</paragraph>
</caption>
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<paragraph id="8BC5520FB973E63CFF58CD44FBF6F7A7" blockId="4.[135,1474,2064,2115]" pageId="4" pageNumber="601">
Fig. 3.
<taxonomicName id="4C7A298CB973E63CFF0FCD44FE37F7C1" authority="Chen &amp; Páll-Gergely, 2023" authorityName="Chen &amp; Páll-Gergely" authorityYear="2023" box="[208,411,2064,2085]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="601" phylum="Mollusca" rank="species" species="yanseni" status="sp. nov.">
<emphasis id="B90E8E1DB973E63CFF0FCD44FE37F7C1" box="[208,411,2064,2085]" italics="true" pageId="4" pageNumber="601">Anaglyphula yanseni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23D3366B973E63CFE75CD44FD8EF7C2" box="[426,546,2064,2086]" pageId="4" pageNumber="601" rank="species">new species</taxonomicNameLabel>
, holotype MZB.Gst. 23.829. AB, Shell; A, Visible light camera images; B, SEM images; C, Protoconch; D, Detailed structure on upper body whorl; E, Detailed structure on lower body whorl.
</paragraph>
</caption>
<caption id="DF050287B972E63DFFAEC70AFEF2FD4A" ID-DOI="http://doi.org/10.5281/zenodo.10271449" ID-Zenodo-Dep="10271449" httpUri="https://zenodo.org/record/10271449/files/figure.png" pageId="5" pageNumber="602" startId="5.[113,148,606,628]" targetBox="[113,1451,189,591]" targetPageId="5" targetType="figure">
<paragraph id="8BC5520FB972E63DFFAEC70AFEF2FD4A" blockId="5.[113,1453,606,686]" pageId="5" pageNumber="602">
Fig. 4. A, Putative constriction position of
<taxonomicName id="4C7A298CB972E63DFDCDC70AFD70FD90" authority="Chen &amp; Páll-Gergely, 2023" authorityName="Chen &amp; Páll-Gergely" authorityYear="2023" box="[530,732,606,628]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="602" phylum="Mollusca" rank="species" species="yanseni" status="sp. nov.">
<emphasis id="B90E8E1DB972E63DFDCDC70AFD70FD90" box="[530,732,606,628]" italics="true" pageId="5" pageNumber="602">Anaglyphula yanseni</emphasis>
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,
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, holotype MZB.Gst. 23.829; B, Constriction of
<taxonomicName id="4C7A298CB972E63DFAEDC70AFECFFD74" authority="Rensch, 1932" authorityName="Rensch" authorityYear="1932" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="602" phylum="Mollusca" rank="species" species="cancellata">
<emphasis id="B90E8E1DB972E63DFAEDC70AFF79FD74" italics="true" pageId="5" pageNumber="602">Anaglyphula cancellata</emphasis>
<bibRefCitation id="EFEB2FFEB972E63DFF02C72FFECFFD74" author="Rensch B" box="[221,355,635,657]" pageId="5" pageNumber="602" pagination="1 - 130" refId="ref4322" refString="Rensch B (1932) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba II. Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 63: 1 - 130, 3 pls." type="journal article" year="1932">Rensch, 1932</bibRefCitation>
</taxonomicName>
, syntype NHMUK 1897.8.3.150; C, Constriction of
<taxonomicName id="4C7A298CB972E63DFCB5C72FFB47FD75" authority="Vermeulen, 1996" authorityName="Vermeulen" authorityYear="1996" box="[874,1259,635,657]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="602" phylum="Mollusca" rank="species" species="whitteni">
<emphasis id="B90E8E1DB972E63DFCB5C72FFB96FD74" box="[874,1082,635,656]" italics="true" pageId="5" pageNumber="602">Anaglyphula whitteni</emphasis>
<bibRefCitation id="EFEB2FFEB972E63DFB9FC72FFB47FD75" author="Vermeulen JJ" box="[1088,1259,635,657]" pageId="5" pageNumber="602" pagination="149 - 162" refId="ref4506" refString="Vermeulen JJ (1996). Notes on terrestrial molluscs of Java, Bali and Nusa Penida. Basteria. 59: 149 - 162." type="journal article" year="1996">Vermeulen, 1996</bibRefCitation>
</taxonomicName>
, paratype NHMUK 20000247. Not to scale.
</paragraph>
</caption>
<paragraph id="8BC5520FB972E63DFFAEC0D1FED8FA39" blockId="5.[113,751,1413,1501]" pageId="5" pageNumber="602">
<emphasis id="B90E8E1DB972E63DFFAEC0D1FE3DFA79" bold="true" box="[113,401,1413,1437]" pageId="5" pageNumber="602">Measurements (in mm).</emphasis>
Shell height: 2.082.18, Shell width: 1.961.97, Aperture height: 0.960.99, Aperture width: 1.001.03 (n=2).
</paragraph>
</subSubSection>
<subSubSection id="C3600184B972E63DFFAEC351FD5DF9F9" box="[113,753,1541,1566]" pageId="5" pageNumber="602" type="distribution">
<paragraph id="8BC5520FB972E63DFFAEC351FD5DF9F9" blockId="5.[113,753,1541,1566]" box="[113,753,1541,1566]" pageId="5" pageNumber="602">
<emphasis id="B90E8E1DB972E63DFFAEC351FEA8F9F9" bold="true" box="[113,260,1541,1565]" pageId="5" pageNumber="602">Distribution.</emphasis>
The species is found in the
<typeStatus id="54C1ECADB972E63DFDF0C352FDF1F9FA" box="[559,605,1542,1566]" pageId="5" pageNumber="602">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
<subSubSection id="C3600184B972E63DFFAEC311FF6AF999" pageId="5" pageNumber="602" type="discussion">
<paragraph id="8BC5520FB972E63DFFAEC311FF6AF999" blockId="5.[113,752,1605,1661]" pageId="5" pageNumber="602">
<emphasis id="B90E8E1DB972E63DFFAEC311FF4CF9B9" bold="true" box="[113,224,1605,1629]" pageId="5" pageNumber="602">Remarks.</emphasis>
See the remarks for
<taxonomicName id="4C7A298CB972E63DFE10C311FD1FF9B9" authority="Chen &amp; Páll-Gergely, 2023" authorityName="Chen &amp; Páll-Gergely" authorityYear="2023" box="[463,691,1605,1629]" class="Gastropoda" family="Assimineidae" genus="Anaglyphula" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="602" phylum="Mollusca" rank="species" species="yanseni" status="sp. nov.">
<emphasis id="B90E8E1DB972E63DFE10C311FD1FF9B9" box="[463,691,1605,1629]" italics="true" pageId="5" pageNumber="602">Anaglyphula yanseni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23D3366B972E63DFD1BC312FF6DF999" pageId="5" pageNumber="602" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>