treatments-xml/data/03/D3/D4/03D3D43AE44BFFA82580F9B3FEE2F9AC.xml

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<mods:title id="08D6045F292E0915CDD45CF3ABF622F5">New enchytraeid species from Mount Hallasan (Jeju Island, Korea) (Enchytraeidae, Oligochaeta)</mods:title>
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<mods:namePart id="085EEC7A58A407D2242BD31775C07423">Dózsa-Farkas, Klára</mods:namePart>
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<mods:namePart id="3A439E9CD62968ECE43D3D91D817682A">Felföldi, Tamás</mods:namePart>
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<mods:namePart id="13652F3DD73920F48611A050CDCB0503">Nagy, Hajnalka</mods:namePart>
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<mods:namePart id="C0304D08CEA8E81079BF968D4EFCC0DA">Hong, Yong</mods:namePart>
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<heading id="D08DD240E44BFFAC2580F9B3FE2FF9AC" bold="true" box="[151,430,1589,1615]" fontSize="11" level="1" pageId="2" pageNumber="339" reason="1">
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<taxonomicName id="4C7A1EAFE44BFFAC2580F9B3FEE4F9AC" ID-CoL="85L4G" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[151,357,1589,1615]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="2" pageNumber="339" phylum="Annelida" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B90EB93EE44BFFAC2580F9B3FEE4F9AC" bold="true" box="[151,357,1589,1615]" italics="true" pageId="2" pageNumber="339">Achaeta koreana</emphasis>
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(
<figureCitation id="134179A9E44BFFAC2589F9D1FE88F993" box="[158,265,1623,1648]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="2" pageNumber="339">Figures 1</figureCitation>
, 
<figureCitation id="134179A9E44BFFAC2400F9DEFEA6F993" box="[279,295,1624,1648]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="2" pageNumber="339">2</figureCitation>
, 
<figureCitation id="134179A9E44BFFAC2424F9D1FEF6F993" box="[307,375,1623,1648]" captionStart="FIGURE 5" captionStartId="10.[151,249,1486,1508]" captionTargetBox="[219,1407,191,1452]" captionTargetId="figure@10.[151,1436,183,1472]" captionTargetPageId="10" captionText="FIGURE 5. Achaeta koreana sp. n. (A–C), Achaeta macroampullacea sp. n. (D–E), Bryodrilus hallasanensis sp. n. (F), Enchytronia seongpanakiensis sp. n. (G) and Chaemaedrilus baekrokdamensis sp. n. (H) A. Pharyngeal glands (dv=dorsal vessel, oesophageal appendages marked with arrows). B. Spermatheca. C. Sperm funnel. D. Male copulatory organ and granulocytes at ventral border of clitellum, E-H. Spermathecae. Scale bars 50 µm." httpUri="https://zenodo.org/record/1446861/files/figure.png" pageId="2" pageNumber="339">5A–C</figureCitation>
)
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material.
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:
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NIBRIV0000810584 slide No. 2210, adult, stained whole mounted specimen. 
<typeStatus id="54C1DB8EE44BFFAC2073F926FA1DF95B" box="[1380,1436,1696,1720]" pageId="2" pageNumber="339">Type</typeStatus>
locality: site 4, 
<location id="8EA533F7E44BFFAC2451F942FE54F93E" LSID="urn:lsid:plazi:treatment:03D3D43AE44BFFA82580F9B3FEE2F9AC:8EA533F7E44BFFAC2451F942FE54F93E" box="[326,469,1732,1757]" country="South Korea" county="North" municipality="Mt. Hallasan" name="Baekrokdam" pageId="2" pageNumber="339" stateProvince="Jeju">Baekrokdam</location>
crater on the summit of 
<collectingMunicipality id="6BA1FF56E44BFFAC27FBF943FC00F93E" box="[748,897,1732,1757]" pageId="2" pageNumber="339">Mt. Hallasan</collectingMunicipality>
, 
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Island, 
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, soil of grass on highland in 
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slope (N 33˚21'46.0&quot;, E 126˚31'58.0&quot;), 
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elevation, 0 9.06.2016, leg. Y. Hong. 
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<emphasis id="B90EB93EE44BFFAC21B8F96FFAA1F8E3" box="[1199,1312,1769,1792]" italics="true" pageId="2" pageNumber="339">Paratypes</emphasis>
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(in total 9 stained, adult specimens on slides and 
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in 70% ethanol): NIBRIV0000810585, slide No. 2198, site 1, NIBRIV0000811380, slide No. 2276, site 2, P.113.1. slide No. 2212, from 
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locality, P.113.2.1–113.2.3. slides No
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. 
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, site 6, P.113.3. slide No. 2196, site 5, P.113.4.1, slides No. 2197, site 1. 
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70% ethanol: P.113.5, from 
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locality 
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; P.113.6 site 2 seven specimens; P.113.7 site 5 eight specimens; P.113.8, site 6 twelve specimens.
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</paragraph>
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<emphasis id="B90EB93EE44AFFAD2584FDFAFF74FD73" bold="true" box="[147,245,636,656]" pageId="3" pageNumber="340">TABLE 1.</emphasis>
Detected enchytraeid species (and the two polychaetes) and their distribution at the study sites in Mt. Hallasan, Korea (site 1-4 Baekrokdam crater, 5-11 Gwaneumsa trail, 12 Gwaneumsa temple, 13-15 Seongpanak trail). New species described here are highlighted in bold, while species new for the Korean fauna are marked with an asterisk.
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<th id="769B0E12E44A0051211CFD49FB96FD00" box="[1035,1047,719,739]" gridcol="3" gridrow="0" pageId="3" pageNumber="340">3</th>
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<emphasis id="B90EB93EE44AFFAD2584FD72FEE6FCEB" bold="true" box="[147,359,756,776]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FD72FEB1FCEB" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,304,756,776]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FD72FEB1FCEB" bold="true" box="[147,304,756,776]" italics="true" pageId="3" pageNumber="340">Achaeta koreana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2421FD72FEE6FCEB" box="[310,359,756,776]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A0051266AFD72FC08FCEB" box="[893,905,756,776]" gridcol="1" gridrow="1" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005126D3FD72FC51FCEB" box="[964,976,756,776]" gridcol="2" gridrow="1" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051211CFD72FB96FCEB" box="[1035,1047,756,776]" gridcol="3" gridrow="1" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512145FD72FBDEFCEB" box="[1106,1119,756,776]" gridcol="4" gridrow="1" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051218EFD72FB24FCEB" box="[1177,1189,756,776]" gridcol="5" gridrow="1" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005121F7FD72FB6DFCEB" box="[1248,1260,756,776]" gridcol="6" gridrow="1" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512030FD72FAB2FCEB" box="[1319,1331,756,776]" gridcol="7" gridrow="1" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FC90F8EFFCC8" box="[147,1902,790,811]" gridrow="2" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FC90FD55FCC8" box="[147,724,790,811]" gridcol="0" gridrow="2" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FC90FE08FCC9" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,393,790,810]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="macroampullacea" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FC90FE08FCC9" bold="true" box="[147,393,790,810]" italics="true" pageId="3" pageNumber="340">Achaeta macroampullacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2498FC91FE40FCC8" box="[399,449,791,811]" pageId="3" pageNumber="340" rank="species">
<emphasis id="B90EB93EE44AFFAD2498FC91FE3AFCC8" bold="true" box="[399,443,791,811]" pageId="3" pageNumber="340">sp. n</emphasis>
.
</taxonomicNameLabel>
</th>
<td id="769B0E12E44A0051232AFC90F9D2FCC8" box="[1597,1619,790,811]" gridcol="11" gridrow="2" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FC90F918FCC8" box="[1667,1689,790,811]" gridcol="12" gridrow="2" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FCBCF8EFFCAD" box="[147,1902,826,846]" gridrow="3" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-3="1" rowspan-7="1" rowspan-9="1">
<th id="769B0E12E44A00512584FCBCFD55FCAD" box="[147,724,826,846]" gridcol="0" gridrow="3" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FCBCFE2EFCAD" bold="true" box="[147,431,826,846]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FCBCFEF6FCAD" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,375,826,846]" class="Clitellata" family="Enchytraeidae" genus="Bryodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FCBCFEF6FCAD" bold="true" box="[147,375,826,846]" italics="true" pageId="3" pageNumber="340">Bryodrilus hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2469FCBCFE2EFCAD" box="[382,431,826,846]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A005126D3FCBCFC51FCAD" box="[964,976,826,846]" gridcol="2" gridrow="3" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512145FCBCFBDEFCAD" box="[1106,1119,826,846]" gridcol="4" gridrow="3" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051218EFCBCFB24FCAD" box="[1177,1189,826,846]" gridcol="5" gridrow="3" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005121F7FCBCFB6DFCAD" box="[1248,1260,826,846]" gridcol="6" gridrow="3" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051207AFCBCFAF8FCAD" box="[1389,1401,826,846]" gridcol="8" gridrow="3" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFCBCF9D2FCAD" box="[1597,1619,826,846]" gridcol="11" gridrow="3" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FCBCF918FCAD" box="[1667,1689,826,846]" gridcol="12" gridrow="3" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FCD8F8EFFC91" box="[147,1902,862,882]" gridrow="4" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FCD8FD55FC91" box="[147,724,862,882]" gridcol="0" gridrow="4" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FCD8FE73FC91" bold="true" box="[147,498,862,882]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FCD8FE3AFC91" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,443,862,882]" class="Clitellata" family="Enchytraeidae" genus="Chamaedrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="baekrokdamensis" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FCD8FE3AFC91" bold="true" box="[147,443,862,882]" italics="true" pageId="3" pageNumber="340">Chamaedrilus baekrokdamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD24D7FCD8FE73FC91" box="[448,498,862,882]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A005126D3FCD8FC51FC91" box="[964,976,862,882]" gridcol="2" gridrow="4" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051211CFCD8FB96FC91" box="[1035,1047,862,882]" gridcol="3" gridrow="4" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFCD8F9D2FC91" box="[1597,1619,862,882]" gridcol="11" gridrow="4" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FC06F8EFFC77" box="[147,1902,896,916]" gridrow="5" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="769B0E12E44A00512584FC06FD55FC77" box="[147,724,896,916]" gridcol="0" gridrow="5" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FC07FE7CFC77" authority="Vejdovsky, 1879" authorityName="Vejdovsky" authorityYear="1879" box="[147,509,896,916]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" sensu="lato" species="buchholzi">
<emphasis id="B90EB93EE44AFFAD2584FC07FEDEFC77" box="[147,351,897,916]" italics="true" pageId="3" pageNumber="340">Enchytraeus buchholzi</emphasis>
Vejdovský, 1879
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2714FC07FDDAFC77" box="[515,603,896,916]" pageId="3" pageNumber="340" sensu="lato">sensu lato</taxonomicNameLabel>
</th>
<td id="769B0E12E44A0051207AFC06FAF8FC77" box="[1389,1401,896,916]" gridcol="8" gridrow="5" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFC06F9D2FC77" box="[1597,1619,896,916]" gridcol="11" gridrow="5" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FC06F918FC77" box="[1667,1689,896,916]" gridcol="12" gridrow="5" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FC22F8EFFC5B" box="[147,1902,932,952]" gridrow="6" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FC22FD55FC5B" box="[147,724,932,952]" gridcol="0" gridrow="6" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FC22FD96FC5B" authority="Rota &amp; Healy, 1994" authorityName="Rota &amp; Healy" authorityYear="1994" box="[147,535,932,952]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="dichaetus">
<emphasis id="B90EB93EE44AFFAD2584FC22FEDFFC54" box="[147,350,932,951]" italics="true" pageId="3" pageNumber="340">Enchytraeus dichaetus</emphasis>
Rota &amp; Healy, 1994
</taxonomicName>
</th>
<td id="769B0E12E44A00512394FC22F918FC5B" box="[1667,1689,932,952]" gridcol="12" gridrow="6" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005123DDFC22F961FC5B" box="[1738,1760,932,952]" gridcol="13" gridrow="6" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FC41F8EFFC3F" box="[147,1902,967,988]" gridrow="7" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FC41FD55FC3F" box="[147,724,967,988]" gridcol="0" gridrow="7" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FC41FE64FC3F" bold="true" box="[147,485,967,988]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FC41FE2FFC38" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,430,967,987]" class="Clitellata" family="Enchytraeidae" genus="Enchytronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="seongpanakiensis" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FC41FE2FFC38" bold="true" box="[147,430,967,987]" italics="true" pageId="3" pageNumber="340">Enchytronia seongpanakiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD24A3FC4EFE64FC3F" box="[436,485,968,988]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A005123DDFC41F961FC3F" box="[1738,1760,967,988]" gridcol="13" gridrow="7" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FC6CF8EFFC1D" box="[147,1902,1002,1022]" gridrow="8" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FC6CFD55FC1D" box="[147,724,1002,1022]" gridcol="0" gridrow="8" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FC6CFDD2FC1D" authority="Nielsen &amp; Christensen, 1959" authorityName="Nielsen &amp; Christensen" authorityYear="1959" box="[147,595,1002,1022]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="bulboides">
<emphasis id="B90EB93EE44AFFAD2584FC6CFECAFC1E" box="[147,331,1002,1021]" italics="true" pageId="3" pageNumber="340">Fridericia bulboides</emphasis>
Nielsen &amp; Christensen, 1959
</taxonomicName>
</th>
<td id="769B0E12E44A00512206FC6CF8A6FC1D" box="[1809,1831,1002,1022]" gridcol="14" gridrow="8" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FB8BF8EFFBC2" box="[147,1902,1037,1057]" gridrow="9" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FB8BFD55FBC2" box="[147,724,1037,1057]" gridcol="0" gridrow="9" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FB8BFD55FBC2" authority="Dozsa-Farkas, Felfoldi &amp; Hong, 1915" authorityName="Dozsa-Farkas, Felfoldi &amp; Hong" authorityYear="1915" box="[147,724,1037,1057]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="cusanicaformis">
<emphasis id="B90EB93EE44AFFAD2584FB8BFEFDFBC2" box="[147,380,1037,1057]" italics="true" pageId="3" pageNumber="340">Fridericia cusanicaformis</emphasis>
Dózsa-Farkas, Felföldi &amp; Hong, 1915
</taxonomicName>
</th>
<td id="769B0E12E44A00512145FB8BFBDEFBC2" box="[1106,1119,1037,1057]" gridcol="4" gridrow="9" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FB8BF918FBC2" box="[1667,1689,1037,1057]" gridcol="12" gridrow="9" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FBB7F8EFFBA6" box="[147,1902,1073,1093]" gridrow="10" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FBB7FD55FBA6" box="[147,724,1073,1093]" gridcol="0" gridrow="10" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FBB7FEEFFBA0" authority="Issel" authorityName="Issel" authorityYear="1904" box="[147,366,1073,1093]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="paroniana">
<emphasis id="B90EB93EE44AFFAD2584FBB7FF6EFBA7" box="[147,239,1073,1092]" italics="true" pageId="3" pageNumber="340">Fridericia</emphasis>
cf. 
<emphasis id="B90EB93EE44AFFAD2406FBB7FEEFFBA0" box="[273,366,1073,1091]" italics="true" pageId="3" pageNumber="340">paroniana</emphasis>
</taxonomicName>
<emphasis id="B90EB93EE44AFFAD2464FBB7FEFFFBA7" box="[371,382,1073,1092]" italics="true" pageId="3" pageNumber="340">*</emphasis>
Issel, 1904
</th>
<td id="769B0E12E44A00512206FBB7F8A6FBA6" box="[1809,1831,1073,1093]" gridcol="14" gridrow="10" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FBD2F8EFFB8B" box="[147,1902,1108,1128]" gridrow="11" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FBD2FD55FB8B" box="[147,724,1108,1128]" gridcol="0" gridrow="11" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FBD2FE76FB8B" authority="* (Vejdovsky, 1878)" authorityName="Vejdovsky" authorityYear="1878" box="[147,503,1108,1128]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="perrieri">
<emphasis id="B90EB93EE44AFFAD2584FBD2FECBFB84" box="[147,330,1108,1127]" italics="true" pageId="3" pageNumber="340">Fridericia perrieri *</emphasis>
(Vejdovský, 1878)
</taxonomicName>
</th>
<td id="769B0E12E44A00512206FBD2F8A6FB8B" box="[1809,1831,1108,1128]" gridcol="14" gridrow="11" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FBF1F8EFFB68" box="[147,1902,1143,1163]" gridrow="12" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FBF1FD55FB68" box="[147,724,1143,1163]" gridcol="0" gridrow="12" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FBF1FF6EFB69" box="[147,239,1143,1162]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="genus">
<emphasis id="B90EB93EE44AFFAD2584FBF1FF6EFB69" box="[147,239,1143,1162]" italics="true" pageId="3" pageNumber="340">Fridericia</emphasis>
</taxonomicName>
sp. 1
</th>
<td id="769B0E12E44A0051232AFBF1F9D2FB68" box="[1597,1619,1143,1163]" gridcol="11" gridrow="12" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FB1CF8EFFB4D" box="[147,1902,1178,1198]" gridrow="13" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FB1CFD55FB4D" box="[147,724,1178,1198]" gridcol="0" gridrow="13" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FB1DFF6EFB4D" box="[147,239,1179,1198]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="genus">
<emphasis id="B90EB93EE44AFFAD2584FB1DFF6EFB4D" box="[147,239,1179,1198]" italics="true" pageId="3" pageNumber="340">Fridericia</emphasis>
</taxonomicName>
sp. 2
</th>
<td id="769B0E12E44A0051232AFB1CF9D2FB4D" box="[1597,1619,1178,1198]" gridcol="11" gridrow="13" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FB1CF918FB4D" box="[1667,1689,1178,1198]" gridcol="12" gridrow="13" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FB3BF8EFFB32" box="[147,1902,1213,1233]" gridrow="14" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FB3BFD55FB32" box="[147,724,1213,1233]" gridcol="0" gridrow="14" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FB38FF6EFB32" box="[147,239,1214,1233]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="genus">
<emphasis id="B90EB93EE44AFFAD2584FB38FF6EFB32" box="[147,239,1214,1233]" italics="true" pageId="3" pageNumber="340">Fridericia</emphasis>
</taxonomicName>
sp. 3
</th>
<td id="769B0E12E44A00512145FB3BFBDEFB32" box="[1106,1119,1213,1233]" gridcol="4" gridrow="14" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FB3BF918FB32" box="[1667,1689,1213,1233]" gridcol="12" gridrow="14" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FB67F8EFFB16" box="[147,1902,1249,1269]" gridrow="15" pageId="3" pageNumber="340" rowspan-1="1" rowspan-15="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th id="769B0E12E44A00512584FB67FD55FB16" box="[147,724,1249,1269]" gridcol="0" gridrow="15" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FB67FD21FB16" authority="Dozsa-Farkas &amp; Hong, 2010" authorityName="Dozsa-Farkas &amp; Hong" authorityYear="2010" box="[147,672,1249,1269]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="jeonjuensis">
<emphasis id="B90EB93EE44AFFAD2584FB67FE18FB10" box="[147,409,1249,1268]" italics="true" pageId="3" pageNumber="340">Hemienchytraeus jeonjuensis</emphasis>
Dózsa-Farkas &amp; Hong, 2010
</taxonomicName>
</th>
<td id="769B0E12E44A0051211CFB67FB96FB16" box="[1035,1047,1249,1269]" gridcol="3" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051218EFB67FB24FB16" box="[1177,1189,1249,1269]" gridcol="5" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512030FB67FAB2FB16" box="[1319,1331,1249,1269]" gridcol="7" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051207AFB67FAF8FB16" box="[1389,1401,1249,1269]" gridcol="8" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005120E1FB67F98DFB16" box="[1526,1548,1249,1269]" gridcol="10" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFB67F9D2FB16" box="[1597,1619,1249,1269]" gridcol="11" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FB67F918FB16" box="[1667,1689,1249,1269]" gridcol="12" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005123DDFB67F961FB16" box="[1738,1760,1249,1269]" gridcol="13" gridrow="15" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512206FB67F8A6FB16" box="[1809,1831,1249,1269]" gridcol="14" gridrow="15" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FA82F8EFFAFB" box="[147,1902,1284,1304]" gridrow="16" pageId="3" pageNumber="340" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-7="1" rowspan-8="1">
<th id="769B0E12E44A00512584FA82FD55FAFB" box="[147,724,1284,1304]" gridcol="0" gridrow="16" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FA82FD17FAFB" authority="Dozsa-Farkas &amp; Hong, 2010" authorityName="Dozsa-Farkas &amp; Hong" authorityYear="2010" box="[147,662,1284,1304]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="quadratus">
<emphasis id="B90EB93EE44AFFAD2584FA82FE0EFAF4" box="[147,399,1284,1303]" italics="true" pageId="3" pageNumber="340">Hemienchytraeus quadratus</emphasis>
Dózsa-Farkas &amp; Hong, 2010
</taxonomicName>
</th>
<td id="769B0E12E44A0051266AFA82FC08FAFB" box="[893,905,1284,1304]" gridcol="1" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005126D3FA82FC51FAFB" box="[964,976,1284,1304]" gridcol="2" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051211CFA82FB96FAFB" box="[1035,1047,1284,1304]" gridcol="3" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512145FA82FBDEFAFB" box="[1106,1119,1284,1304]" gridcol="4" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051218EFA82FB24FAFB" box="[1177,1189,1284,1304]" gridcol="5" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005121F7FA82FB6DFAFB" box="[1248,1260,1284,1304]" gridcol="6" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005120A3FA82FA41FAFB" box="[1460,1472,1284,1304]" gridcol="9" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005120E1FA82F98DFAFB" box="[1526,1548,1284,1304]" gridcol="10" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFA82F9D2FAFB" box="[1597,1619,1284,1304]" gridcol="11" gridrow="16" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051224FFA82F8EFFAFB" box="[1880,1902,1284,1304]" gridcol="15" gridrow="16" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FAA1F8EFFAD8" box="[147,1902,1319,1339]" gridrow="17" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-13="1" rowspan-15="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="769B0E12E44A00512584FAA1FD55FAD8" box="[147,724,1319,1339]" gridcol="0" gridrow="17" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FAA1FD0CFAD8" authority="Dozsa-Farkas &amp; Hong, 2010" authorityName="Dozsa-Farkas &amp; Hong" authorityYear="2010" box="[147,653,1319,1339]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="koreanus">
<emphasis id="B90EB93EE44AFFAD2584FAA1FE06FAD9" box="[147,391,1319,1338]" italics="true" pageId="3" pageNumber="340">Hemienchytraeus koreanus</emphasis>
Dózsa-Farkas &amp; Hong, 2010
</taxonomicName>
</th>
<td id="769B0E12E44A005126D3FAA1FC51FAD8" box="[964,976,1319,1339]" gridcol="2" gridrow="17" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051211CFAA1FB96FAD8" box="[1035,1047,1319,1339]" gridcol="3" gridrow="17" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512145FAA1FBDEFAD8" box="[1106,1119,1319,1339]" gridcol="4" gridrow="17" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051207AFAA1FAF8FAD8" box="[1389,1401,1319,1339]" gridcol="8" gridrow="17" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFAA1F9D2FAD8" box="[1597,1619,1319,1339]" gridcol="11" gridrow="17" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FAA1F918FAD8" box="[1667,1689,1319,1339]" gridcol="12" gridrow="17" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512206FAA1F8A6FAD8" box="[1809,1831,1319,1339]" gridcol="14" gridrow="17" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FACCF8EFFABD" box="[147,1902,1354,1374]" gridrow="18" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FACCFD55FABD" box="[147,724,1354,1374]" gridcol="0" gridrow="18" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FACCFE2AFABD" authority="Friend, 1911" authorityName="Friend" authorityYear="1911" box="[147,427,1354,1374]" class="Clitellata" family="Enchytraeidae" genus="Henlea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="perpusilla">
<emphasis id="B90EB93EE44AFFAD2584FACCFEB5FABE" box="[147,308,1354,1373]" italics="true" pageId="3" pageNumber="340">Henlea perpusilla</emphasis>
Friend, 1911
</taxonomicName>
</th>
<td id="769B0E12E44A00512394FACCF918FABD" box="[1667,1689,1354,1374]" gridcol="12" gridrow="18" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FAE8F8EFFA61" box="[147,1902,1390,1410]" gridrow="19" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-3="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FAE8FD55FA61" box="[147,724,1390,1410]" gridcol="0" gridrow="19" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FAE8FE49FA61" bold="true" box="[147,456,1390,1410]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FAE8FE10FA61" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,401,1390,1410]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="jungsaihoi" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FAE8FE10FA61" bold="true" box="[147,401,1390,1410]" italics="true" pageId="3" pageNumber="340">Mesenchytraeus jungsaihoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2481FAE8FE49FA61" box="[406,456,1390,1410]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A005126D3FAE8FC51FA61" box="[964,976,1390,1410]" gridcol="2" gridrow="19" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512145FAE8FBDEFA61" box="[1106,1119,1390,1410]" gridcol="4" gridrow="19" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051218EFAE8FB24FA61" box="[1177,1189,1390,1410]" gridcol="5" gridrow="19" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFAE8F9D2FA61" box="[1597,1619,1390,1410]" gridcol="11" gridrow="19" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FAE8F918FA61" box="[1667,1689,1390,1410]" gridcol="12" gridrow="19" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FA17F8EFFA46" box="[147,1902,1425,1445]" gridrow="20" pageId="3" pageNumber="340" rowspan-1="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-9="1">
<th id="769B0E12E44A00512584FA17FD55FA46" box="[147,724,1425,1445]" gridcol="0" gridrow="20" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FA17FE01FA46" bold="true" box="[147,384,1425,1445]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FA17FEC9FA46" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,328,1425,1445]" class="Clitellata" family="Enchytraeidae" genus="Xetadrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="jejuensis" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FA17FEC9FA46" bold="true" box="[147,328,1425,1445]" italics="true" pageId="3" pageNumber="340">Xetadrilus jejuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2459FA17FE01FA46" box="[334,384,1425,1445]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A005121F7FA17FB6DFA46" box="[1248,1260,1425,1445]" gridcol="6" gridrow="20" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512030FA17FAB2FA46" box="[1319,1331,1425,1445]" gridcol="7" gridrow="20" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051207AFA17FAF8FA46" box="[1389,1401,1425,1445]" gridcol="8" gridrow="20" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005120E1FA17F98DFA46" box="[1526,1548,1425,1445]" gridcol="10" gridrow="20" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFA17F9D2FA46" box="[1597,1619,1425,1445]" gridcol="11" gridrow="20" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FA17F918FA46" box="[1667,1689,1425,1445]" gridcol="12" gridrow="20" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FA32F8EFFA2A" box="[147,1902,1460,1481]" gridrow="21" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-14="1" rowspan-15="1" rowspan-9="1">
<th id="769B0E12E44A00512584FA32FD55FA2A" box="[147,724,1460,1481]" gridcol="0" gridrow="21" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FA32FE18FA2A" bold="true" box="[147,409,1460,1481]" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584FA32FEE0FA2B" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[147,353,1460,1480]" class="Clitellata" family="Enchytraeidae" genus="Xetadrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="aphanoides" status="sp. nov.">
<emphasis id="B90EB93EE44AFFAD2584FA32FEE0FA2B" bold="true" box="[147,353,1460,1480]" italics="true" pageId="3" pageNumber="340">Xetadrilus aphanoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44AFFAD2470FA33FE18FA2A" box="[359,409,1461,1481]" pageId="3" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="769B0E12E44A005126D3FA32FC51FA2A" box="[964,976,1460,1481]" gridcol="2" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051211CFA32FB96FA2A" box="[1035,1047,1460,1481]" gridcol="3" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512145FA32FBDEFA2A" box="[1106,1119,1460,1481]" gridcol="4" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051218EFA32FB24FA2A" box="[1177,1189,1460,1481]" gridcol="5" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005121F7FA32FB6DFA2A" box="[1248,1260,1460,1481]" gridcol="6" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512030FA32FAB2FA2A" box="[1319,1331,1460,1481]" gridcol="7" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051207AFA32FAF8FA2A" box="[1389,1401,1460,1481]" gridcol="8" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A0051232AFA32F9D2FA2A" box="[1597,1619,1460,1481]" gridcol="11" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A00512394FA32F918FA2A" box="[1667,1689,1460,1481]" gridcol="12" gridrow="21" pageId="3" pageNumber="340">+</td>
<td id="769B0E12E44A005123DDFA32F961FA2A" box="[1738,1760,1460,1481]" gridcol="13" gridrow="21" pageId="3" pageNumber="340">+</td>
</tr>
<tr id="354A676EE44A00512584FA5EF8EFFA0F" box="[147,1902,1496,1516]" gridrow="22" pageId="3" pageNumber="340">
<th id="769B0E12E44A00512584FA5EFD55FA0F" box="[147,724,1496,1516]" gridcol="0" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FA5EFD84FA0F" bold="true" box="[147,517,1496,1516]" pageId="3" pageNumber="340">Enchytraeid species number (total: 21)</emphasis>
</th>
<td id="769B0E12E44A0051266AFA5EFC08FA0F" box="[893,905,1496,1516]" gridcol="1" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2669FA5EFC08FA0F" bold="true" box="[894,905,1496,1516]" pageId="3" pageNumber="340">2</emphasis>
</td>
<td id="769B0E12E44A005126D3FA5EFC51FA0F" box="[964,976,1496,1516]" gridcol="2" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD26D3FA5EFC4EFA0F" bold="true" box="[964,975,1496,1516]" pageId="3" pageNumber="340">7</emphasis>
</td>
<td id="769B0E12E44A0051211CFA5EFB96FA0F" box="[1035,1047,1496,1516]" gridcol="3" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD211BFA5EFB96FA0F" bold="true" box="[1036,1047,1496,1516]" pageId="3" pageNumber="340">6</emphasis>
</td>
<td id="769B0E12E44A00512145FA5EFBDEFA0F" box="[1106,1119,1496,1516]" gridcol="4" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2145FA5EFBDCFA0F" bold="true" box="[1106,1117,1496,1516]" pageId="3" pageNumber="340">8</emphasis>
</td>
<td id="769B0E12E44A0051218EFA5EFB24FA0F" box="[1177,1189,1496,1516]" gridcol="5" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD218DFA5EFB24FA0F" bold="true" box="[1178,1189,1496,1516]" pageId="3" pageNumber="340">6</emphasis>
</td>
<td id="769B0E12E44A005121F7FA5EFB6DFA0F" box="[1248,1260,1496,1516]" gridcol="6" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD21F7FA5EFB6AFA0F" bold="true" box="[1248,1259,1496,1516]" pageId="3" pageNumber="340">5</emphasis>
</td>
<td id="769B0E12E44A00512030FA5EFAB2FA0F" box="[1319,1331,1496,1516]" gridcol="7" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2030FA5EFAB3FA0F" bold="true" box="[1319,1330,1496,1516]" pageId="3" pageNumber="340">4</emphasis>
</td>
<td id="769B0E12E44A0051207AFA5EFAF8FA0F" box="[1389,1401,1496,1516]" gridcol="8" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2079FA5EFAF8FA0F" bold="true" box="[1390,1401,1496,1516]" pageId="3" pageNumber="340">6</emphasis>
</td>
<td id="769B0E12E44A005120A3FA5EFA41FA0F" box="[1460,1472,1496,1516]" gridcol="9" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD20A2FA5EFA41FA0F" bold="true" box="[1461,1472,1496,1516]" pageId="3" pageNumber="340">1</emphasis>
</td>
<td id="769B0E12E44A005120E1FA5EF98DFA0F" box="[1526,1548,1496,1516]" gridcol="10" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD20ECFA5EF987FA0F" bold="true" box="[1531,1542,1496,1516]" pageId="3" pageNumber="340">3</emphasis>
</td>
<td id="769B0E12E44A0051232AFA5EF9D2FA0F" box="[1597,1619,1496,1516]" gridcol="11" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD232AFA5EF9D2FA0F" bold="true" box="[1597,1619,1496,1516]" pageId="3" pageNumber="340">12</emphasis>
</td>
<td id="769B0E12E44A00512394FA5EF918FA0F" box="[1667,1689,1496,1516]" gridcol="12" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2394FA5EF918FA0F" bold="true" box="[1667,1689,1496,1516]" pageId="3" pageNumber="340">13</emphasis>
</td>
<td id="769B0E12E44A005123DDFA5EF961FA0F" box="[1738,1760,1496,1516]" gridcol="13" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD23C7FA5EF95AFA0F" bold="true" box="[1744,1755,1496,1516]" pageId="3" pageNumber="340">4</emphasis>
</td>
<td id="769B0E12E44A00512206FA5EF8A6FA0F" box="[1809,1831,1496,1516]" gridcol="14" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2200FA5EF8A3FA0F" bold="true" box="[1815,1826,1496,1516]" pageId="3" pageNumber="340">5</emphasis>
</td>
<td id="769B0E12E44A0051224FFA5EF8EFFA0F" box="[1880,1902,1496,1516]" gridcol="15" gridrow="22" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2249FA5EF8E8FA0F" bold="true" box="[1886,1897,1496,1516]" pageId="3" pageNumber="340">1</emphasis>
</td>
</tr>
<tr id="354A676EE44A00512584FA7CF8EFF9ED" box="[147,1902,1530,1550]" gridrow="23" pageId="3" pageNumber="340" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="769B0E12E44A00512584FA7CFD55F9ED" box="[147,724,1530,1550]" gridcol="0" gridrow="23" pageId="3" pageNumber="340">
<emphasis id="B90EB93EE44AFFAD2584FA7DFEFFF9ED" box="[147,382,1531,1550]" italics="true" pageId="3" pageNumber="340">Hrabeiella periglandulata</emphasis>
Pižl &amp; Chalupský, 1984
</th>
<td id="769B0E12E44A005121F7FA7CFB6DF9ED" box="[1248,1260,1530,1550]" gridcol="6" gridrow="23" pageId="3" pageNumber="340">1</td>
<td id="769B0E12E44A00512394FA7CF918F9ED" box="[1667,1689,1530,1550]" gridcol="12" gridrow="23" pageId="3" pageNumber="340">1</td>
<td id="769B0E12E44A005123DDFA7CF961F9ED" box="[1738,1760,1530,1550]" gridcol="13" gridrow="23" pageId="3" pageNumber="340">1</td>
</tr>
<tr id="354A676EE44A00512584F998F8EFF9D1" box="[147,1902,1566,1586]" gridrow="24" pageId="3" pageNumber="340" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="769B0E12E44A00512584F998FD55F9D1" box="[147,724,1566,1586]" gridcol="0" gridrow="24" pageId="3" pageNumber="340">
<taxonomicName id="4C7A1EAFE44AFFAD2584F998FE68F9D1" authority="Reisinger, 1925" authorityName="Resisinger" authorityYear="1925" box="[147,489,1566,1586]" class="Clitellata" family="Parergodrilidae" genus="Parergodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Apodadrilida" pageId="3" pageNumber="340" phylum="Annelida" rank="species" species="heideri">
<emphasis id="B90EB93EE44AFFAD2584F998FED7F9D2" box="[147,342,1566,1585]" italics="true" pageId="3" pageNumber="340">Parergodrilus heideri</emphasis>
Reisinger, 1925
</taxonomicName>
</th>
<td id="769B0E12E44A0051266AF998FC08F9D1" box="[893,905,1566,1586]" gridcol="1" gridrow="24" pageId="3" pageNumber="340">1</td>
<td id="769B0E12E44A0051207AF998FAF8F9D1" box="[1389,1401,1566,1586]" gridcol="8" gridrow="24" pageId="3" pageNumber="340">1</td>
<td id="769B0E12E44A00512394F998F918F9D1" box="[1667,1689,1566,1586]" gridcol="12" gridrow="24" pageId="3" pageNumber="340">1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC5652CE44DFFAA25D0FF11FAD4FF53" blockId="4.[151,1436,151,501]" box="[199,1365,151,176]" pageId="4" pageNumber="341">
<emphasis id="B90EB93EE44DFFAA25D0FF11FD92FF53" bold="true" box="[199,531,151,176]" pageId="4" pageNumber="341">Further material examined.</emphasis>
15 specimens investigated 
<emphasis id="B90EB93EE44DFFAA2650FF1EFC12FF4C" box="[839,915,152,175]" italics="true" pageId="4" pageNumber="341">in vivo</emphasis>
, 3 of them processed for DNA analysis.
</paragraph>
</subSubSection>
<subSubSection id="C36036A7E44DFFAA25D0FF3AFCC8FF36" box="[199,841,188,213]" pageId="4" pageNumber="341" type="etymology">
<paragraph id="8BC5652CE44DFFAA25D0FF3AFCC8FF36" blockId="4.[151,1436,151,501]" box="[199,841,188,213]" pageId="4" pageNumber="341">
<emphasis id="B90EB93EE44DFFAA25D0FF3AFECFFF36" bold="true" box="[199,334,188,213]" pageId="4" pageNumber="341">Etymology.</emphasis>
Named after the country where it was found.
</paragraph>
</subSubSection>
<subSubSection id="C36036A7E44DFFAA25D0FF59FB11FE17" pageId="4" pageNumber="341" type="diagnosis">
<paragraph id="8BC5652CE44DFFAA25D0FF59FB11FE17" blockId="4.[151,1436,151,501]" pageId="4" pageNumber="341">
<emphasis id="B90EB93EE44DFFAA25D0FF59FEC0FF1B" bold="true" box="[199,321,223,248]" pageId="4" pageNumber="341">Diagnosis.</emphasis>
The new species can be recognized by the following combination of characters: (1) small, stout worms (
<quantity id="4C82C8C9E44DFFAA25E2FE82FED7FEFF" box="[245,342,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="341" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long and 200–280 µm wide at clitellum 
<emphasis id="B90EB93EE44DFFAA2625FE83FC01FEFF" box="[818,896,261,284]" italics="true" pageId="4" pageNumber="341">in vivo</emphasis>
), segments 17–22; (2) no pyriform glands; (3) clitellum developed only laterally: dorso-lateral area with hyalocytes in disordered three or four longitudinal rows; (4) dorsal blood vessel from VI; (5) pharyngeal glands at 4/5–5/6 connected dorsally, at 6/7 separate dorsally, the first ventral lobes separate from the dorsal lobes in IV, secondary ventral glands absent; (6) three pairs of preclitellar nephridia; (7) pars tumida of midgut from XIII–XIV, extending over 2 segments; (8) sperm funnel small, barrel-shaped, collar narrower than funnel body; (9) male pores in XII, ventrolaterally, no glandular body, only the pore surrounded by small inconspicuous glands; (10) spermathecae confined to V.
</paragraph>
</subSubSection>
<caption id="DF0535A4E44DFFAA2580F8C9FEADF83C" httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="4" pageNumber="341" startId="4.[151,249,1871,1893]" targetBox="[188,1391,598,1801]" targetPageId="4">
<paragraph id="8BC5652CE44DFFAA2580F8C9FEADF83C" blockId="4.[151,1437,1871,2015]" pageId="4" pageNumber="341">
<emphasis id="B90EB93EE44DFFAA2580F8C9FE92F887" bold="true" box="[151,275,1871,1893]" pageId="4" pageNumber="341">FIGURE 1.</emphasis>
Micrographs of 
<taxonomicName id="4C7A1EAFE44DFFAA24D6F8C9FDECF886" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[449,621,1871,1893]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="4" pageNumber="341" phylum="Annelida" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B90EB93EE44DFFAA24D6F8C9FDECF886" box="[449,621,1871,1893]" italics="true" pageId="4" pageNumber="341">Achaeta koreana</emphasis>
</taxonomicName>
<emphasis id="B90EB93EE44DFFAA2762F8C9FD4EF887" bold="true" box="[629,719,1871,1892]" pageId="4" pageNumber="341">
<taxonomicNameLabel id="A23D0445E44DFFAA2762F8C9FD2EF887" box="[629,687,1871,1892]" pageId="4" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). 
<emphasis id="B90EB93EE44DFFAA2423F8E8FECBF860" bold="true" box="[308,330,1902,1923]" pageId="4" pageNumber="341">B.</emphasis>
Clitellar glands, dorsal view (interruption marked with arrow). 
<emphasis id="B90EB93EE44DFFAA26E6F8EBFB8BF860" bold="true" box="[1009,1034,1901,1923]" pageId="4" pageNumber="341">C.</emphasis>
Clitellar glands, lateral view (arrows: hyaline glands). 
<emphasis id="B90EB93EE44DFFAA2453F80AFEDCF842" bold="true" box="[324,349,1932,1953]" pageId="4" pageNumber="341">D.</emphasis>
Entire individual (head marked with arrow). 
<emphasis id="B90EB93EE44DFFAA262EF80AFCD1F842" bold="true" box="[825,848,1932,1953]" pageId="4" pageNumber="341">E.</emphasis>
Clitellar glands, lateral view. 
<emphasis id="B90EB93EE44DFFAA219BF80AFB3EF841" bold="true" box="[1164,1215,1932,1954]" pageId="4" pageNumber="341">F–G.</emphasis>
Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G 
<emphasis id="B90EB93EE44DFFAA26B1F82AFC6BF822" box="[934,1002,1964,1985]" italics="true" pageId="4" pageNumber="341">in vivo</emphasis>
, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm).
</paragraph>
</caption>
<caption id="DF0535A4E44CFFAB2580F950FC73F884" httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="5" pageNumber="342" startId="5.[151,249,1750,1772]" targetBox="[188,1388,195,1724]" targetPageId="5">
<paragraph id="8BC5652CE44CFFAB2580F950FC73F884" blockId="5.[151,1436,1750,1895]" pageId="5" pageNumber="342">
<emphasis id="B90EB93EE44CFFAB2580F950FE92F90F" bold="true" box="[151,275,1750,1772]" pageId="5" pageNumber="342">FIGURE 2.</emphasis>
Micrographs of 
<taxonomicName id="4C7A1EAFE44CFFAB24A8F950FDEAF90F" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[447,619,1750,1772]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="342" phylum="Annelida" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B90EB93EE44CFFAB24A8F950FDEAF90F" box="[447,619,1750,1772]" italics="true" pageId="5" pageNumber="342">Achaeta koreana</emphasis>
</taxonomicName>
<emphasis id="B90EB93EE44CFFAB2765F951FD4DF908" bold="true" box="[626,716,1750,1772]" pageId="5" pageNumber="342">
<taxonomicNameLabel id="A23D0445E44CFFAB2765F951FD2DF90F" box="[626,684,1751,1772]" pageId="5" pageNumber="342" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). 
<emphasis id="B90EB93EE44CFFAB27B6F970FD39F8E8" bold="true" box="[673,696,1782,1803]" pageId="5" pageNumber="342">B.</emphasis>
Midgut pars tumida XIII 
<emphasis id="B90EB93EE44CFFAB26AFF970FC44F8E8" bold="true" box="[952,965,1782,1803]" pageId="5" pageNumber="342">–</emphasis>
XV. 
<emphasis id="B90EB93EE44CFFAB26E4F973FB8AF8E8" bold="true" box="[1011,1035,1781,1803]" pageId="5" pageNumber="342">C.</emphasis>
Coelomocytes. 
<emphasis id="B90EB93EE44CFFAB21B8F970FB49F8E8" bold="true" box="[1199,1224,1782,1803]" pageId="5" pageNumber="342">D.</emphasis>
First nephridium. 
<emphasis id="B90EB93EE44CFFAB2093F970FA1AF8E8" bold="true" box="[1412,1435,1782,1803]" pageId="5" pageNumber="342">E.</emphasis>
Nephridium terminally. 
<emphasis id="B90EB93EE44CFFAB2487F892FE42F8C9" bold="true" box="[400,451,1812,1834]" pageId="5" pageNumber="342">F–G.</emphasis>
Sperm funnels. 
<emphasis id="B90EB93EE44CFFAB2765F892FD0CF8CA" bold="true" box="[626,653,1812,1833]" pageId="5" pageNumber="342">H.</emphasis>
Clitellar glands, lateral view (arrow: male opening). 
<emphasis id="B90EB93EE44CFFAB21AAF892FB4CF8CA" bold="true" box="[1213,1229,1812,1833]" pageId="5" pageNumber="342">I.</emphasis>
Male opening with small surrounding glands (arrow). 
<emphasis id="B90EB93EE44CFFAB24E8F8B5FDB1F8AB" bold="true" box="[511,560,1843,1864]" pageId="5" pageNumber="342">J–L.</emphasis>
Spermathecae (arrow in L: ectal gland). 
<emphasis id="B90EB93EE44CFFAB26C8F8B5FC7CF8AB" bold="true" box="[991,1021,1843,1864]" pageId="5" pageNumber="342">M.</emphasis>
Spermathecal ectal gland (marked with arrow). A–G, I–M 
<emphasis id="B90EB93EE44CFFAB2443F8D4FE19F884" box="[340,408,1874,1895]" italics="true" pageId="5" pageNumber="342">in vivo</emphasis>
, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm.
</paragraph>
</caption>
<subSubSection id="C36036A7E44CFFA825D0F808FD45FBF3" lastPageId="6" lastPageNumber="343" pageId="5" pageNumber="342" type="description">
<paragraph id="8BC5652CE44CFFA825D0F808FD43FE64" blockId="5.[151,1436,1934,2032]" lastBlockId="6.[151,1437,151,1616]" lastPageId="6" lastPageNumber="343" pageId="5" pageNumber="342">
<emphasis id="B90EB93EE44CFFAB25D0F808FED8F844" bold="true" box="[199,345,1934,1959]" pageId="5" pageNumber="342">Description.</emphasis>
Small worm (
<figureCitation id="134179A9E44CFFAB2717F808FDDDF844" box="[512,604,1934,1959]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="5" pageNumber="342">Fig. 1D</figureCitation>
). 
<typeStatus id="54C1DB8EE44CFFAB2764F808FD5AF844" box="[627,731,1934,1959]" pageId="5" pageNumber="342" type="holotype">Holotype</typeStatus>
<quantity id="4C82C8C9E44CFFAB27F2F809FCBFF844" box="[741,830,1935,1959]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="342" unit="mm" value="2.8">2.8 mm</quantity>
long, 275 µm wide at VIII and 315 µm at clitellum (fixed), 21 segments. Length of 
<typeStatus id="54C1DB8EE44CFFAB2716F835FDEDF828" box="[513,620,1971,1995]" pageId="5" pageNumber="342" type="paratype">paratypes</typeStatus>
<quantity id="4C82C8C9E44CFFAB2764F834FD7CF829" box="[627,765,1970,1995]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.8" metricValueMin="2.5" pageId="5" pageNumber="342" unit="mm" value="3.15" valueMax="3.8" valueMin="2.5">2.5–3.8 mm</quantity>
, width 200–250 µm at VIII and 220–280 µm at clitellum 
<emphasis id="B90EB93EE44CFFAB2091F835FF47F80C" italics="true" pageId="5" pageNumber="342">in vivo</emphasis>
, length of fixed specimens 
<quantity id="4C82C8C9E44CFFAB24EEF851FDD9F80C" box="[505,600,2007,2031]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="342" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, width 220–270 µm at VIII and 2 30–300 µm at clitellum, segments 17–22. No pyriform glands. Body wall thick, thickness dorsally 22–24 µm, cuticle 4–7 µm, ventrally 17–18 µm and 2–3 µm, respectively, 
<emphasis id="B90EB93EE44FFFA82473FF3BFE32FF37" box="[356,435,189,212]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
(
<figureCitation id="134179A9E44FFFA824D3FF3AFDA0FF37" box="[452,545,188,213]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Fig. 1A</figureCitation>
). Clitellum in 
<date id="FFC443ECE44FFFA827DAFF3AFC8EFF37" box="[717,783,188,212]" bridgedPair="-" pageId="6" pageNumber="343">XII–1</date>
/2 XIII developed only laterally, dorso-lateral areas with hyalocytes in two or three disordered longitudinal rows (
<figureCitation id="134179A9E44FFFA82630FF59FCFEFF1B" box="[807,895,223,248]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Fig. 1C</figureCitation>
), hyalocytes large and conspicuous, 28–32 µm high and 17–19 µm wide, granulocytes 20–24 by 16–14 µm (
<figureCitation id="134179A9E44FFFA82670FE82FC41FEFF" box="[871,960,260,285]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Fig. 1E</figureCitation>
). Ventro-laterally only granulocytes. The dorsal interruption 60–70 µm wide 
<emphasis id="B90EB93EE44FFFA8273CFEAEFDF6FEDC" box="[555,631,296,319]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
, dorsal borderline consisting of granular cells only in one longitudinal row (
<figureCitation id="134179A9E44FFFA825C5FECAFEA9FE87" box="[210,296,332,357]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Fig. 1B</figureCitation>
). Ventral borderlines inconspicuous. Head pore on the top of prostomium (
<figureCitation id="134179A9E44FFFA82179FECAFB71FE87" box="[1134,1264,332,357]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Figs. 1F–G</figureCitation>
). Spermathecal pores at 4/ 
<quantity id="4C82C8C9E44FFFA8241AFEE9FEB7FE6B" box="[269,310,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="343" unit="in" value="5.0">5 in</quantity>
lateral position. Male pores in XII.
</paragraph>
<paragraph id="8BC5652CE44FFFA825D0FE12FD45FBF3" blockId="6.[151,1437,151,1616]" pageId="6" pageNumber="343">
Brain egg-shaped 1.5 times longer than wide, 80–90 µm long, fixed (
<figureCitation id="134179A9E44FFFA826C3FE12FBA8FE4F" box="[980,1065,404,429]" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Fig. 1F</figureCitation>
). The prostomial ganglion (
<figureCitation id="134179A9E44FFFA82073FE12FF4FFE33" captionStart="FIGURE 1" captionStartId="4.[151,249,1871,1893]" captionTargetBox="[188,1391,598,1801]" captionTargetId="figure@4.[188,1398,598,1811]" captionTargetPageId="4" captionText="FIGURE 1. Micrographs of Achaeta koreana sp. n. A. Thick cuticle terminally dorsally thicker (white arrow) than ventrally (black arrow). B. Clitellar glands, dorsal view (interruption marked with arrow). C. Clitellar glands, lateral view (arrows: hyaline glands). D. Entire individual (head marked with arrow). E. Clitellar glands, lateral view. F–G. Head (white arrows: head pore; black arrows: prostomial ganglia; b=brain, ph=pharynx). A–E, G in vivo, F fixed, stained. Scale bars A–D, F, G 50 µm; E 20 µm)." httpUri="https://zenodo.org/record/1446853/files/figure.png" pageId="6" pageNumber="343">Figs. 1F,G</figureCitation>
) well visible, 37–40 µm long (fixed). Pharyngeal glands at 4/5–6/7 (
<figureCitation id="134179A9E44FFFA826D9FE31FBB3FE33" box="[974,1074,439,464]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2A</figureCitation>
, 
<figureCitation id="134179A9E44FFFA82129FE31FBE3FE33" box="[1086,1122,439,464]" captionStart="FIGURE 5" captionStartId="10.[151,249,1486,1508]" captionTargetBox="[219,1407,191,1452]" captionTargetId="figure@10.[151,1436,183,1472]" captionTargetPageId="10" captionText="FIGURE 5. Achaeta koreana sp. n. (A–C), Achaeta macroampullacea sp. n. (D–E), Bryodrilus hallasanensis sp. n. (F), Enchytronia seongpanakiensis sp. n. (G) and Chaemaedrilus baekrokdamensis sp. n. (H) A. Pharyngeal glands (dv=dorsal vessel, oesophageal appendages marked with arrows). B. Spermatheca. C. Sperm funnel. D. Male copulatory organ and granulocytes at ventral border of clitellum, E-H. Spermathecae. Scale bars 50 µm." httpUri="https://zenodo.org/record/1446861/files/figure.png" pageId="6" pageNumber="343">5A</figureCitation>
), all with ventral lobes. The first and secondary dorsal lobes united dorsally, the third pair separate dorsally. Three pairs of preclitellar nephridia at 6/7–8/9 constricted by septum. Length ratio anteseptale: postseptale 1: 2.5 preclitellarly, postseptale tapers gradually into efferent duct, with small terminal vesicle (
<figureCitation id="134179A9E44FFFA82603FDA2FC18FDDF" box="[788,921,548,573]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2D–E</figureCitation>
). Dorsal blood vessel from VI (
<figureCitation id="134179A9E44FFFA821EDFDA2FADCFDDF" box="[1274,1373,548,573]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2A</figureCitation>
, 
<figureCitation id="134179A9E44FFFA8207EFDA2FA0EFDDE" box="[1385,1423,548,573]" captionStart="FIGURE 5" captionStartId="10.[151,249,1486,1508]" captionTargetBox="[219,1407,191,1452]" captionTargetId="figure@10.[151,1436,183,1472]" captionTargetPageId="10" captionText="FIGURE 5. Achaeta koreana sp. n. (A–C), Achaeta macroampullacea sp. n. (D–E), Bryodrilus hallasanensis sp. n. (F), Enchytronia seongpanakiensis sp. n. (G) and Chaemaedrilus baekrokdamensis sp. n. (H) A. Pharyngeal glands (dv=dorsal vessel, oesophageal appendages marked with arrows). B. Spermatheca. C. Sperm funnel. D. Male copulatory organ and granulocytes at ventral border of clitellum, E-H. Spermathecae. Scale bars 50 µm." httpUri="https://zenodo.org/record/1446861/files/figure.png" pageId="6" pageNumber="343">5A</figureCitation>
). Coelomocytes about 40–53 µm long 
<emphasis id="B90EB93EE44FFFA8272BFDCEFD09FDBC" box="[572,648,584,607]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
, flat, cytoplasm pale, margin with indefinite contour in top view, but spindle-shaped in side view (
<figureCitation id="134179A9E44FFFA824CBFDEAFDB5FD67" box="[476,564,620,645]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Fig. 2C</figureCitation>
), the length measurable only in side view. Oesophageal appendages in V, well developed without canal in IV (
<figureCitation id="134179A9E44FFFA824EAFD09FDE3FD4B" box="[509,610,655,680]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2A</figureCitation>
, 
<figureCitation id="134179A9E44FFFA82779FD09FD15FD4B" box="[622,660,655,680]" captionStart="FIGURE 5" captionStartId="10.[151,249,1486,1508]" captionTargetBox="[219,1407,191,1452]" captionTargetId="figure@10.[151,1436,183,1472]" captionTargetPageId="10" captionText="FIGURE 5. Achaeta koreana sp. n. (A–C), Achaeta macroampullacea sp. n. (D–E), Bryodrilus hallasanensis sp. n. (F), Enchytronia seongpanakiensis sp. n. (G) and Chaemaedrilus baekrokdamensis sp. n. (H) A. Pharyngeal glands (dv=dorsal vessel, oesophageal appendages marked with arrows). B. Spermatheca. C. Sperm funnel. D. Male copulatory organ and granulocytes at ventral border of clitellum, E-H. Spermathecae. Scale bars 50 µm." httpUri="https://zenodo.org/record/1446861/files/figure.png" pageId="6" pageNumber="343">5A</figureCitation>
). Chloragocytes yellowish-brown, about 15 µm long 
<emphasis id="B90EB93EE44FFFA821F9FD16FAC3FD44" box="[1262,1346,656,679]" italics="true" pageId="6" pageNumber="343">in vivo.</emphasis>
Midgut pars tumida XIII–XIV (occupying 2 segments) (
<figureCitation id="134179A9E44FFFA827D5FD32FC9CFD2F" box="[706,797,692,717]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Fig. 2B</figureCitation>
). Sperm funnel small, barrel-shaped, 65–90 µm long 
<emphasis id="B90EB93EE44FFFA82091FD33FF47FD0C" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
(53–62 µm fixed), about 1.2–2 times longer than wide, collar narrower than funnel body (
<figureCitation id="134179A9E44FFFA821CBFD51FADFFD13" box="[1244,1374,727,752]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2F–G</figureCitation>
, 
<figureCitation id="134179A9E44FFFA8207CFD51FA11FD13" box="[1387,1424,727,752]" captionStart="FIGURE 5" captionStartId="10.[151,249,1486,1508]" captionTargetBox="[219,1407,191,1452]" captionTargetId="figure@10.[151,1436,183,1472]" captionTargetPageId="10" captionText="FIGURE 5. Achaeta koreana sp. n. (A–C), Achaeta macroampullacea sp. n. (D–E), Bryodrilus hallasanensis sp. n. (F), Enchytronia seongpanakiensis sp. n. (G) and Chaemaedrilus baekrokdamensis sp. n. (H) A. Pharyngeal glands (dv=dorsal vessel, oesophageal appendages marked with arrows). B. Spermatheca. C. Sperm funnel. D. Male copulatory organ and granulocytes at ventral border of clitellum, E-H. Spermathecae. Scale bars 50 µm." httpUri="https://zenodo.org/record/1446861/files/figure.png" pageId="6" pageNumber="343">5C</figureCitation>
). Sperm duct 6 µm thick 
<emphasis id="B90EB93EE44FFFA824B3FD7BFE71FCF7" box="[420,496,765,788]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
. Spermatozoa 75–90 µm, heads 30–38 µm long 
<emphasis id="B90EB93EE44FFFA8210FFD7BFBE4FCF7" box="[1048,1125,765,788]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
(52–60 µm and 17–23 µm, fixed). Seminal vesicle absent. Male copulatory organs small, widely separate ventro-laterally, no glandular body or bursa, only the pore surrounded by small inconspicuous glands (
<figureCitation id="134179A9E44FFFA8269FFCC2FC7CFCBF" box="[904,1021,836,861]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2H,I</figureCitation>
). Spermathecae small, free, confined to V. The short ectal duct (20–30 µm long 20–23 µm wide 
<emphasis id="B90EB93EE44FFFA8264CFCEEFC2AFC9C" box="[859,939,872,895]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
) has a small gland (
<figureCitation id="134179A9E44FFFA821B5FCE1FAA9FC63" box="[1186,1320,871,896]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2L,M</figureCitation>
). In welldeveloped specimens the duct widens slightly into a narrow dilation of ampulla (25–28 µm wide). After the dilation the connecting tube (about 60 µm long and 20 µm wide) ends in an elongated globular ental reservoir (35–50 µm long, 25–35 µm wide 
<emphasis id="B90EB93EE44FFFA82499FC53FE5BFC0F" box="[398,474,981,1004]" italics="true" pageId="6" pageNumber="343">in vivo</emphasis>
) in V. The surface of the reservoir is granulated, sperm is visible only in the ampullar dilation and the connecting tube (
<figureCitation id="134179A9E44FFFA82719FC71FD06FBF3" box="[526,647,1015,1040]" captionStart="FIGURE 2" captionStartId="5.[151,249,1750,1772]" captionTargetBox="[188,1388,195,1724]" captionTargetId="figure@5.[188,1398,195,1729]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Achaeta koreana sp. n. A. Pharyngeal glands, lateral view (marked with arrows, o= oesophageal appendage, dv= dorsal vessel, n=first nephridium). B. Midgut pars tumida XIII–XV. C. Coelomocytes. D. First nephridium. E. Nephridium terminally. F–G. Sperm funnels. H. Clitellar glands, lateral view (arrow: male opening). I. Male opening with small surrounding glands (arrow). J–L. Spermathecae (arrow in L: ectal gland). M. Spermathecal ectal gland (marked with arrow). A–G, I–M in vivo, H fixed, stained. Scale bars A–H, J–L 50 µm; I, M 20 µm." httpUri="https://zenodo.org/record/1446855/files/figure.png" pageId="6" pageNumber="343">Figs. 2J–L</figureCitation>
, 
<figureCitation id="134179A9E44FFFA82784FC71FD37FBF3" box="[659,694,1015,1040]" captionStart="FIGURE 5" captionStartId="10.[151,249,1486,1508]" captionTargetBox="[219,1407,191,1452]" captionTargetId="figure@10.[151,1436,183,1472]" captionTargetPageId="10" captionText="FIGURE 5. Achaeta koreana sp. n. (A–C), Achaeta macroampullacea sp. n. (D–E), Bryodrilus hallasanensis sp. n. (F), Enchytronia seongpanakiensis sp. n. (G) and Chaemaedrilus baekrokdamensis sp. n. (H) A. Pharyngeal glands (dv=dorsal vessel, oesophageal appendages marked with arrows). B. Spermatheca. C. Sperm funnel. D. Male copulatory organ and granulocytes at ventral border of clitellum, E-H. Spermathecae. Scale bars 50 µm." httpUri="https://zenodo.org/record/1446861/files/figure.png" pageId="6" pageNumber="343">5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36036A7E44FFFA825D0FB9AFB1DFBD7" box="[199,1180,1052,1077]" pageId="6" pageNumber="343" type="distribution">
<paragraph id="8BC5652CE44FFFA825D0FB9AFB1DFBD7" blockId="6.[151,1437,151,1616]" box="[199,1180,1052,1077]" pageId="6" pageNumber="343">
<emphasis id="B90EB93EE44FFFA825D0FB9AFE70FBD6" bold="true" box="[199,497,1052,1077]" pageId="6" pageNumber="343">Distribution and habitat.</emphasis>
In 
<collectingCountry id="F36D25BCE44FFFA82701FB9BFDDCFBD7" box="[534,605,1053,1076]" name="South Korea" pageId="6" pageNumber="343">Korea</collectingCountry>
, Mt. Hallasan, above 
<quantity id="4C82C8C9E44FFFA82646FB9AFC27FBD7" box="[849,934,1052,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.3059999999999998" pageId="6" pageNumber="343" unit="m" value="1306.0">1306 m</quantity>
elevation, at site 1–7.
</paragraph>
</subSubSection>
<subSubSection id="C36036A7E44FFFA825D0FBB9FEE2F9AC" pageId="6" pageNumber="343" type="diagnosis">
<paragraph id="8BC5652CE44FFFA825D0FBB9FEE2F9AC" blockId="6.[151,1437,151,1616]" pageId="6" pageNumber="343">
<emphasis id="B90EB93EE44FFFA825D0FBB9FE51FBBB" bold="true" box="[199,464,1087,1112]" pageId="6" pageNumber="343">Differential diagnosis.</emphasis>
Among the previously described 12 small 
<taxonomicName id="4C7A1EAFE44FFFA826D2FBC6FBA0FBBB" box="[965,1057,1088,1112]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="genus">
<emphasis id="B90EB93EE44FFFA826D2FBC6FBA0FBBB" box="[965,1057,1088,1112]" italics="true" pageId="6" pageNumber="343">Achaeta</emphasis>
</taxonomicName>
species without pyriform glands and the spermathecae confined to segments V–VI, none has the origin of the dorsal vessel from VI; moreover, secondary pharyngeal glands are, with one exception (
<taxonomicName id="4C7A1EAFE44FFFA827E3FB0EFBC1FB43" authority="Schmelz, 2008" authorityName="Schmelz" authorityYear="2008" box="[756,1088,1159,1184]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="hanagarthi">
<emphasis id="B90EB93EE44FFFA827E3FB0EFC11FB43" box="[756,912,1160,1184]" italics="true" pageId="6" pageNumber="343">A. hanagarthi</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA82681FB01FBC1FB43" box="[918,1088,1159,1184]" pageId="6" pageNumber="343" refId="ref20409" refString="Schmelz, R. M., Collado, R. &amp; Rombke, J. (2008) Mata Atlantica enchytraeids (Parana, Brazil): The genus Achaeta (Oligochaeta, Enchytraeidae), Zootaxa, 1809, 1 - 35." type="journal article">Schmelz, 2008</bibRefCitation>
</taxonomicName>
), present, and the spermathecal opening is without a small gland. In 
<taxonomicName id="4C7A1EAFE44FFFA82725FB2AFCFDFB27" authority="(Cognetti, 1899)" baseAuthorityName="Cognetti" baseAuthorityYear="1899" box="[562,892,1196,1221]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="camerani">
<emphasis id="B90EB93EE44FFFA82725FB2AFD3DFB27" box="[562,700,1196,1220]" italics="true" pageId="6" pageNumber="343">A. camerani</emphasis>
(Cognetti, 1899)
</taxonomicName>
the dorsal blood vessel originates in VIII and in the all other species in VII. 
<taxonomicName id="4C7A1EAFE44FFFA824F0FB56FC9AFB0B" authority="Graefe, 1980" authorityName="Graefe" authorityYear="1980" box="[487,795,1231,1256]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="brevivasa">
<emphasis id="B90EB93EE44FFFA824F0FB56FDF7FB0B" box="[487,630,1232,1256]" italics="true" pageId="6" pageNumber="343">A. brevivasa</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA82796FB49FC9AFB0B" box="[641,795,1231,1256]" pageId="6" pageNumber="343" refId="ref19177" refString="Graefe, U. (1980) Systematische Untersuchungen an der Gattung Achaeta (Enchytraeidae, Oligochaeta). I. Beschreibung von Achaeta brevivasa sp. n. und Achaeta camerani (Cognetti). Mitteilungen aus den hamburgischen zoologischen Muzeum und Institut, 77, 35 - 39." type="journal article">Graefe, 1980</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7A1EAFE44FFFA8263DFB56FBC2FB0B" authority="Graefe, 2007" authorityName="Graefe" authorityYear="2007" box="[810,1091,1231,1256]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="diddeni">
<emphasis id="B90EB93EE44FFFA8263DFB56FC20FB0B" box="[810,929,1232,1256]" italics="true" pageId="6" pageNumber="343">A. diddeni</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA826BBFB49FBC2FB0B" box="[940,1091,1231,1256]" pageId="6" pageNumber="343" refId="ref19270" refString="Graefe, U. (2007) Achaeta diddeni sp. nov. (Enchytraeidae, Clitellata) from the northern German lowlands. Folia Facultatis Scentiarum. Nauralium Universitatis Masarykianae Brunensis, Biologia, 110, 35 - 40." type="journal article">Graefe, 2007</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7A1EAFE44FFFA82142FB56FAF0FB0B" authority="Moller, 1976" authorityName="Moller" authorityYear="1976" box="[1109,1393,1231,1256]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="hallensis">
<emphasis id="B90EB93EE44FFFA82142FB56FB5CFB0B" box="[1109,1245,1232,1256]" italics="true" pageId="6" pageNumber="343">A. hallensis</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA821F0FB49FAF0FB0B" box="[1255,1393,1231,1256]" pageId="6" pageNumber="343" refId="ref19937" refString="Moller, F. (1976) Achaeta hallensis nom. nov. pro Achaeta segmentata Moller, 1974. Mitteilungen aus dem Zologischen Museum in Berlin, 52, 175. https: // doi. org / 10.1071 / IS 16042" type="journal article">Möller,1976</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7A1EAFE44FFFA82093FB56FD22FAEE" authority="Dozsa-Farkas &amp; Boros, 2005" authorityName="Dozsa-Farkas &amp; Boros" authorityYear="2005" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="antefolliculata">
<emphasis id="B90EB93EE44FFFA82093FB56FEBAFAEF" italics="true" pageId="6" pageNumber="343">A. antefolliculata</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA82450FB72FD22FAEE" box="[327,675,1268,1293]" pageId="6" pageNumber="343" refId="ref18787" refString="Dozsa-Farkas, K. &amp; Boros, G. (2005) Achaeta antefolliculata sp. n., a new enchytraeid species (Oligochaeta, Enchytraeidae) from the rock grassland of the Sas-Hegy in Hungary. Acta Zoologica Academiae scientiarum Hungaricae, 51, 279 - 285." type="journal article">Dózsa-Farkas &amp; Boros, 2005</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B90EB93EE44FFFA827A3FB72FCF4FAE8" box="[692,885,1268,1292]" italics="true" pageId="6" pageNumber="343">
<taxonomicName id="4C7A1EAFE44FFFA827A3FB72FCDBFAEF" box="[692,858,1268,1292]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="afolliculata">A. afolliculata</taxonomicName>
S
</emphasis>
esma &amp; 
<bibRefCitation id="EFEB18DDE44FFFA826C9FB72FB45FAEF" box="[990,1220,1268,1293]" pageId="6" pageNumber="343" refId="ref20518" refString="Sesma, V. &amp; Dozsa-Farkas, K. (1993) Description of seven new species of enchytraeidae (Oligochaeta) from Spain. Acta Zoologica Hungarica, 39, 249 - 265." type="journal article">Dózsa-Farkas, 1993</bibRefCitation>
and 
<taxonomicName id="4C7A1EAFE44FFFA82010FB72FEC1FAD3" authority="Schmelz, 2008" authorityName="Schmelz" authorityYear="2008" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="singularis">
<emphasis id="B90EB93EE44FFFA82010FB72FA1DFAEF" box="[1287,1436,1268,1292]" italics="true" pageId="6" pageNumber="343">A. singularis</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA82580FA91FEC1FAD3" box="[151,320,1303,1328]" pageId="6" pageNumber="343" refId="ref20409" refString="Schmelz, R. M., Collado, R. &amp; Rombke, J. (2008) Mata Atlantica enchytraeids (Parana, Brazil): The genus Achaeta (Oligochaeta, Enchytraeidae), Zootaxa, 1809, 1 - 35." type="journal article">Schmelz, 2008</bibRefCitation>
</taxonomicName>
have two pairs of preclitellar nephridia, further 
<taxonomicName id="4C7A1EAFE44FFFA82665FA9EFB91FAD3" authorityName="Schmelz" authorityYear="2008" box="[882,1040,1304,1328]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="hanagarthi">
<emphasis id="B90EB93EE44FFFA82665FA9EFB91FAD3" box="[882,1040,1304,1328]" italics="true" pageId="6" pageNumber="343">A. hanagarthi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A1EAFE44FFFA8215BFA9EFA1DFAD3" authority="Schmelz, 2008" authorityName="Schmelz" authorityYear="2008" box="[1100,1436,1303,1328]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="paranensis">
<emphasis id="B90EB93EE44FFFA8215BFA9EFB68FACC" box="[1100,1257,1304,1327]" italics="true" pageId="6" pageNumber="343">A. paranensis</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA821E4FA91FA1DFAD3" box="[1267,1436,1303,1328]" pageId="6" pageNumber="343" refId="ref20409" refString="Schmelz, R. M., Collado, R. &amp; Rombke, J. (2008) Mata Atlantica enchytraeids (Parana, Brazil): The genus Achaeta (Oligochaeta, Enchytraeidae), Zootaxa, 1809, 1 - 35." type="journal article">Schmelz, 2008</bibRefCitation>
</taxonomicName>
have only one pair. Only 
<taxonomicName id="4C7A1EAFE44FFFA824B8FABAFD5EFAB7" authority="Graefe, 1989" authorityName="Graefe" authorityYear="1989" box="[431,735,1340,1365]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="pannonica">
<emphasis id="B90EB93EE44FFFA824B8FABAFDC2FAB7" box="[431,579,1340,1364]" italics="true" pageId="6" pageNumber="343">A. pannonica</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA8275DFABAFD5EFAB7" box="[586,735,1340,1365]" pageId="6" pageNumber="343" refId="ref19228" refString="Graefe, U. (1989) Systematische Untersuchungen an der Gattung Achaeta (Enchytraeidae, Clitellata). 2. Beschreibung von vier neuen Arten. Mitteilungen aus dem hamburgischen zoologischen Museum und Institut, 86, 127 - 131." type="journal article">Graefe, 1989</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7A1EAFE44FFFA827FEFABAFC6EFAB7" authority="Graefe, 1989" authorityName="Graefe" authorityYear="1989" box="[745,1007,1340,1365]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="iberica">
<emphasis id="B90EB93EE44FFFA827FEFABAFCD9FAB7" box="[745,856,1340,1364]" italics="true" pageId="6" pageNumber="343">A. iberica</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA8264AFABAFC6EFAB7" box="[861,1007,1340,1365]" pageId="6" pageNumber="343" refId="ref19228" refString="Graefe, U. (1989) Systematische Untersuchungen an der Gattung Achaeta (Enchytraeidae, Clitellata). 2. Beschreibung von vier neuen Arten. Mitteilungen aus dem hamburgischen zoologischen Museum und Institut, 86, 127 - 131." type="journal article">Graefe, 1989</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7A1EAFE44FFFA82133FABAFA94FAB6" authority="Rota, 1995" authorityName="Rota" authorityYear="1995" box="[1060,1301,1340,1365]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="etrusca">
<emphasis id="B90EB93EE44FFFA82133FABAFB17FAB7" box="[1060,1174,1340,1364]" italics="true" pageId="6" pageNumber="343">A. etrusca</emphasis>
<bibRefCitation id="EFEB18DDE44FFFA8218CFABBFA94FAB6" box="[1179,1301,1340,1365]" pageId="6" pageNumber="343" refId="ref20271" refString="Rota, E. (1995) Italian Enchytraeidae (Oligochaeta). I. Bollettino di Zoologia, 62, 183 - 231. https: // doi. org / 10.1080 / 11250009509356067" type="journal article">Rota, 1995</bibRefCitation>
</taxonomicName>
similarly to 
<taxonomicName id="4C7A1EAFE44FFFA82580FAE6FE96FA9B" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" box="[151,279,1376,1400]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B90EB93EE44FFFA82580FAE6FE96FA9B" box="[151,279,1376,1400]" italics="true" pageId="6" pageNumber="343">A. koreana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44FFFA82433FAD9FEE8FA9B" box="[292,361,1375,1400]" pageId="6" pageNumber="343" rank="species">
<emphasis id="B90EB93EE44FFFA82433FAD9FEE8FA9B" bold="true" box="[292,361,1375,1400]" pageId="6" pageNumber="343">sp. n.</emphasis>
</taxonomicNameLabel>
have three pairs of preclitellar nephridia, as in the new species. But in 
<taxonomicName id="4C7A1EAFE44FFFA821C0FAE6FAF3FA94" authorityName="Graefe" authorityYear="1989" box="[1239,1394,1376,1399]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="pannonica">
<emphasis id="B90EB93EE44FFFA821C0FAE6FAF3FA94" box="[1239,1394,1376,1399]" italics="true" pageId="6" pageNumber="343">A. pannonica</emphasis>
</taxonomicName>
all reproductive organs except the spermathecae are shifted one segment forward, so the male pores are in XI, while in the new species, in 
<taxonomicName id="4C7A1EAFE44FFFA8246CFA2EFE6DFA23" authorityName="Graefe" authorityYear="1989" box="[379,492,1448,1472]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="iberica">
<emphasis id="B90EB93EE44FFFA8246CFA2EFE6DFA23" box="[379,492,1448,1472]" italics="true" pageId="6" pageNumber="343">A. iberica</emphasis>
</taxonomicName>
and in 
<taxonomicName id="4C7A1EAFE44FFFA82750FA2EFD3DFA23" authorityName="Rota" authorityYear="1995" box="[583,700,1448,1472]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="etrusca">
<emphasis id="B90EB93EE44FFFA82750FA2EFD3DFA23" box="[583,700,1448,1472]" italics="true" pageId="6" pageNumber="343">A. etrusca</emphasis>
</taxonomicName>
they are in the usual position in XII. 
<taxonomicName id="4C7A1EAFE44FFFA82163FA2EFB68FA23" authorityName="Rota" authorityYear="1995" box="[1140,1257,1448,1472]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="etrusca">
<emphasis id="B90EB93EE44FFFA82163FA2EFB68FA23" box="[1140,1257,1448,1472]" italics="true" pageId="6" pageNumber="343">A. etrusca</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7A1EAFE44FFFA82093FA2EFF73FA07" authority="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong, 2018" authorityName="Dózsa-Farkas &amp; Felföldi &amp; Nagy &amp; Hong" authorityYear="2018" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B90EB93EE44FFFA82093FA2EFF73FA07" italics="true" pageId="6" pageNumber="343">A. koreana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D0445E44FFFA825E8FA4AFEC7FA06" box="[255,326,1484,1509]" pageId="6" pageNumber="343" rank="species">
<emphasis id="B90EB93EE44FFFA825E8FA4AFEC7FA06" bold="true" box="[255,326,1484,1509]" pageId="6" pageNumber="343">sp. n.</emphasis>
</taxonomicNameLabel>
by his paired knob-like cutaneous gland structures dorsolateral in II–VI and by the shorter spermatozoa (40 µm long, sperm heads 15 µm (
<bibRefCitation id="EFEB18DDE44FFFA827A3FA76FCACF9EB" box="[692,813,1519,1544]" pageId="6" pageNumber="343" refId="ref20306" refString="Rota, E. (2015) Five new species of Enchytraeidae (Annelida: Clitellata) from Mediterranean woodlands of Italy and reaffirmed validity of Achaeta etrusca, Fridericia bulbosa and F. miraflores. Journal of Natural History, 49 (33), 1 - 34. https: // doi. org / 10.1080 / 00222933.2015.1009514" type="journal article">Rota 2015</bibRefCitation>
) vs. 75–90 µm and 30–38 µm, respectively), moreover the penial bulbs are compact. Finally, in 
<taxonomicName id="4C7A1EAFE44FFFA82793F992FD76F9CF" authorityName="Graefe" authorityYear="1989" box="[644,759,1556,1580]" class="Clitellata" family="Enchytraeidae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="343" phylum="Annelida" rank="species" species="iberica">
<emphasis id="B90EB93EE44FFFA82793F992FD76F9CF" box="[644,759,1556,1580]" italics="true" pageId="6" pageNumber="343">A. iberica</emphasis>
</taxonomicName>
, hyalocytes of the clitellar glands are lined in 4 distinct longitudinal rows.
</paragraph>
</subSubSection>
</treatment>
</document>