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<document id="869C481F6504F1B5CFE3BE379AF831AE" ID-DOI="10.5281/zenodo.5318944" ID-ISSN="0374-1036" ID-Zenodo-Dep="5318944" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132583789" checkinUser="felipe" docAuthor="Švihla, Vladimír" docDate="2009" docId="03D387E4EE31FFBEFED9478FFEE5FAFE" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.191-216.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Malthodes bucakensis Wittmer 1970, stat. nov." docType="treatment" docVersion="6" lastPageNumber="214" masterDocId="FFEAFF9CEE26FFA9FF8B4575FF8EFFF0" masterDocTitle="New species of the subfamily Malthininae (Coleoptera: Cantharidae) from the western Palaearctic Region" masterLastPageNumber="216" masterPageNumber="191" pageNumber="214" updateTime="1699162932936" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BB489FD9D29AF30DC48DF5481994DB69">New species of the subfamily Malthininae (Coleoptera: Cantharidae) from the western Palaearctic Region</mods:title>
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<mods:namePart id="BA8BE638236C75D59D0B567EE9A50C30">Švihla, Vladimír</mods:namePart>
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<mods:title id="EC2D9F5205DBC1ECF8533BC783A37783">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="24E61E844BC4BA775DED57DE0AF3ECD8">2009</mods:date>
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<subSubSection id="C3606579EE31FFBEFED9478FFCEEFCE4" box="[338,864,762,788]" pageId="23" pageNumber="214" type="nomenclature">
<paragraph id="8BC536F2EE31FFBEFED9478FFCEEFCE4" blockId="23.[338,864,762,788]" box="[338,864,762,788]" pageId="23" pageNumber="214">
<heading id="D08D819EEE31FFBEFED9478FFCEEFCE4" bold="true" box="[338,864,762,788]" centered="true" fontSize="8" level="2" pageId="23" pageNumber="214" reason="3">
<emphasis id="B90EEAE0EE31FFBEFED9478FFCEEFCE4" bold="true" box="[338,864,762,788]" pageId="23" pageNumber="214">
<taxonomicName id="4C7A4D71EE31FFBEFED9478FFD61FCE3" authority="Wittmer, 1970" authorityName="Wittmer" authorityYear="1970" box="[338,751,762,787]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="species" species="bucakensis" status="stat. nov.">
<emphasis id="B90EEAE0EE31FFBEFED9478FFDCBFCE3" bold="true" box="[338,581,762,787]" italics="true" pageId="23" pageNumber="214">Malthodes bucakensis</emphasis>
<bibRefCitation id="EFEB4B03EE31FFBEFDC7478EFD61FCE3" author="WITTMER W." box="[588,751,763,787]" pageId="23" pageNumber="214" pagination="13 - 107" refId="ref10580" refString="WITTMER W. 1970: Zur Kenntnis der Gattung Malthodes (Coleoptera, Cantharidae) (49. Beitrag zur Kenntnis der palaearktischen Cantharidae). Entomologischen Arbeiten aus dem Museum G. Frey 21: 13 - 107." type="journal article" year="1970">Wittmer, 1970</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A23D579BEE31FFBEFD704789FCEEFCE4" box="[763,864,764,788]" pageId="23" pageNumber="214">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3606579EE31FFBEFDBF466FFDF0FCDD" box="[564,638,794,813]" pageId="23" pageNumber="214" type="description">
<paragraph id="8BC536F2EE31FFBEFDBF466FFDF0FCDD" blockId="23.[113,1090,794,1023]" box="[564,638,794,813]" pageId="23" pageNumber="214">
(
<figureCitation id="13412A77EE31FFBEFDB1466FFDF6FCDD" box="[570,632,794,813]" captionStart="Figs" captionStartId="8.[151,190,1542,1561]" captionTargetBox="[174,1105,219,1355]" captionTargetId="figure-10@8.[173,1105,219,1356]" captionTargetPageId="8" captionText="Figs. 1720.Aedeagus, dorsal view.17 Malthodes paveli sp. nov. (holotype); 18 M. lycicus sp. nov. (paratype); 19 M. besucheti Wittmer, 1970 (paratype); 20 M. bucakensis Wittmer, 1970 (Gülükdağı). Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5363039" httpUri="https://zenodo.org/record/5363039/files/figure.png" pageId="23" pageNumber="214">Fig. 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3606579EE31FFBEFFFA4648FDA5FCBF" box="[113,555,828,847]" pageId="23" pageNumber="214" type="reference_group">
<paragraph id="8BC536F2EE31FFBEFFFA4648FDA5FCBF" blockId="23.[113,1090,794,1023]" box="[113,555,828,847]" pageId="23" pageNumber="214">
<treatmentCitationGroup id="AB6A11DCEE31FFBEFFFA4648FDA5FCBF" box="[113,555,828,847]" pageId="23" pageNumber="214">
<treatmentCitation id="0ADB10E3EE31FFBEFFFA4648FDA9FCBF" author="WITTMER W." box="[113,551,828,847]" page="39" pageId="23" pageNumber="214" year="1970">
<taxonomicName id="4C7A4D71EE31FFBEFFFA4648FDA9FCBF" authority="Wittmer, 1970: 39" authorityName="Wittmer" authorityPageNumber="39" authorityYear="1970" box="[113,551,828,847]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="subSpecies" species="besucheti" subSpecies="bucakensis">
<emphasis id="B90EEAE0EE31FFBEFFFA4648FE0BFCBF" box="[113,389,829,847]" italics="true" pageId="23" pageNumber="214">Malthodes besucheti bucakensis</emphasis>
<bibRefCitation id="EFEB4B03EE31FFBEFE024649FDA9FCBF" author="WITTMER W." box="[393,551,828,847]" pageId="23" pageNumber="214" pagination="13 - 107" refId="ref10580" refString="WITTMER W. 1970: Zur Kenntnis der Gattung Malthodes (Coleoptera, Cantharidae) (49. Beitrag zur Kenntnis der palaearktischen Cantharidae). Entomologischen Arbeiten aus dem Museum G. Frey 21: 13 - 107." type="journal article" year="1970">Wittmer, 1970: 39</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3606579EE31FFBEFFFA4613FE1CFC0E" pageId="23" pageNumber="214" type="materials_examined">
<paragraph id="8BC536F2EE31FFBEFFFA4613FE64FC5E" blockId="23.[113,1090,794,1023]" pageId="23" pageNumber="214">
<emphasis id="B90EEAE0EE31FFBEFFFA4613FED4FC89" bold="true" box="[113,346,869,889]" pageId="23" pageNumber="214">Type material examined.</emphasis>
<taxonomicName id="4C7A4D71EE31FFBEFEE94612FDFEFC89" authority="besucheti" box="[354,624,871,889]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="subSpecies" species="besucheti" subSpecies="besucheti">
<emphasis id="B90EEAE0EE31FFBEFEE94612FDFEFC89" box="[354,624,871,889]" italics="true" pageId="23" pageNumber="214">Malthodes besucheti besucheti</emphasis>
</taxonomicName>
:
<typeStatus id="54C18850EE31FFBEFDF44613FD40FC88" box="[639,718,870,889]" pageId="23" pageNumber="214" type="paratype">PARATYPE</typeStatus>
: J,
<collectingCountry id="F36D7662EE31FFBEFD764613FCB4FC89" box="[765,826,870,889]" name="Turkey" pageId="23" pageNumber="214">Türkei</collectingCountry>
[=
<collectingCountry id="F36D7662EE31FFBEFCD64613FC11FC89" box="[861,927,870,889]" name="Turkey" pageId="23" pageNumber="214">Turkey</collectingCountry>
], W.Wittmer [lgt.] [white label, printed] // Nur Dag [= prov.
<collectingRegion id="49BEF810EE31FFBEFE5546F5FDB9FC63" box="[478,567,896,915]" country="Turkey" name="Gaziantep" pageId="23" pageNumber="214">Gaziantep</collectingRegion>
, Nurdağı],
<date id="FFC41032EE31FFBEFD1246F5FD63FC63" box="[665,749,896,915]" pageId="23" pageNumber="214" value="1969-05-14">14.v.1969</date>
[white label, printed and handwriten //
<typeStatus id="54C18850EE31FFBEFFFA46EEFF67FC5E" box="[113,233,923,942]" pageId="23" pageNumber="214" type="paratype">PARATYPUS</typeStatus>
[red label, printed] (
<collectionCode id="ED6BAE37EE31FFBEFE1446EEFE6EFC5E" box="[415,480,923,942]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="214">NMPC</collectionCode>
).
</paragraph>
<paragraph id="8BC536F2EE31FFBEFFFA46C0FE1CFC0E" blockId="23.[113,1090,794,1023]" pageId="23" pageNumber="214">
<emphasis id="B90EEAE0EE31FFBEFFFA46C0FE1EFC39" bold="true" box="[113,400,949,969]" pageId="23" pageNumber="214">Additional material examined.</emphasis>
<materialsCitation id="3B123CAFEE31FFBEFE1346C3FF4EFC14" collectingDate="1999-04-07" collectionCode="AKKG" collectorName="A. Weigel" country="Turkey" location="Gulukdagi" pageId="23" pageNumber="214" specimenCount="1" stateProvince="Antalya">
<emphasis id="B90EEAE0EE31FFBEFE1346C3FDDAFC39" bold="true" box="[408,596,949,969]" pageId="23" pageNumber="214">
<collectingCountry id="F36D7662EE31FFBEFE1346C3FE7BFC39" box="[408,501,950,969]" name="Turkey" pageId="23" pageNumber="214">TURKEY</collectingCountry>
:
<collectingRegion id="49BEF810EE31FFBEFD8A46C0FDC3FC38" box="[513,589,949,969]" country="Turkey" name="Antalya" pageId="23" pageNumber="214">ANTALYA</collectingRegion>
:
</emphasis>
<location id="8EA56029EE31FFBEFDD746C3FD39FC39" LSID="urn:lsid:plazi:treatment:03D387E4EE31FFBEFED9478FFEE5FAFE:8EA56029EE31FFBEFDD746C3FD39FC39" box="[604,695,950,969]" country="Turkey" name="Gulukdagi" pageId="23" pageNumber="214" stateProvince="Antalya">Gülükdağı</location>
(Termessos),
<date id="FFC41032EE31FFBEFCBD46C3FC06FC39" box="[822,904,950,969]" pageId="23" pageNumber="214" value="1999-04-07">
<collectingDate id="EF80E9DAEE31FFBEFCBD46C3FC06FC39" box="[822,904,950,969]" pageId="23" pageNumber="214" value="1999-04-07">7.iv.1999</collectingDate>
</date>
,
<collectorName id="268F5324EE31FFBEFC1846C3FC65FC39" box="[915,1003,950,969]" pageId="23" pageNumber="214">A. Weigel</collectorName>
lgt., 1 J (
<collectionCode id="ED6BAE37EE31FFBEFFFC46A4FF37FC14" box="[119,185,977,996]" pageId="23" pageNumber="214">AKKG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B123CAFEE31FFBEFF4146A5FDF4FC14" box="[202,634,972,997]" collectingDate="1990-04-30" collectingDateMax="1990-05-06" collectingDateMin="1990-04-30" collectionCode="NMPC" collectorName="I. Rydh" country="Turkey" location="Yazir" pageId="23" pageNumber="214" specimenCount="1" stateProvince="Antalya">
<location id="8EA56029EE31FFBEFF4146A5FF77FC13" LSID="urn:lsid:plazi:treatment:03D387E4EE31FFBEFED9478FFEE5FAFE:8EA56029EE31FFBEFF4146A5FF77FC13" box="[202,249,976,995]" country="Turkey" name="Yazir" pageId="23" pageNumber="214" stateProvince="Antalya">Yazir</location>
,
<date id="FFC41032EE31FFBEFE8846A5FE09FC13" box="[259,391,976,995]" pageId="23" pageNumber="214" value="1990-04-30" valueMax="1990-05-06" valueMin="1990-04-30">
<collectingDate id="EF80E9DAEE31FFBEFE8846A5FE09FC13" box="[259,391,976,995]" pageId="23" pageNumber="214" value="1990-04-30" valueMax="1990-05-06" valueMin="1990-04-30">30.iv.6.v.1990</collectingDate>
</date>
,
<collectorName id="268F5324EE31FFBEFE1946A4FE5DFC13" box="[402,467,976,996]" pageId="23" pageNumber="214">I. Rydh</collectorName>
lgt., 1 J (
<collectionCode id="ED6BAE37EE31FFBEFDB946A4FDFDFC14" box="[562,627,977,996]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="214">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B123CAFEE31FFBEFD0846A4FE00FC0E" collectingDate="2005-05-17" collectionCode="NMPC" collectorName="P. Kment" country="Turkey" latitude="40.233334" location="Pazar" longLatPrecision="1164" longitude="36.3" pageId="23" pageNumber="214" specimenCount="5" specimenCount-female="5" stateProvince="Tokat">
<emphasis id="B90EEAE0EE31FFBEFD0846A4FD4AFC14" bold="true" box="[643,708,977,996]" pageId="23" pageNumber="214">
<collectingRegion id="49BEF810EE31FFBEFD0846A4FD33FC12" box="[643,701,977,996]" country="Turkey" name="Tokat" pageId="23" pageNumber="214">TOKAT</collectingRegion>
:
</emphasis>
<location id="8EA56029EE31FFBEFD4046A4FD74FC14" LSID="urn:lsid:plazi:treatment:03D387E4EE31FFBEFED9478FFEE5FAFE:8EA56029EE31FFBEFD4046A4FD74FC14" box="[715,762,977,996]" country="Turkey" latitude="40.233334" longLatPrecision="1164" longitude="36.3" name="Pazar" pageId="23" pageNumber="214" stateProvince="Tokat">Pazar</location>
env.,
<geoCoordinate id="EE4E5035EE31FFBEFCBA46A4FCF6FC14" box="[817,888,976,996]" degrees="40" direction="north" minutes="14" orientation="latitude" pageId="23" pageNumber="214" precision="925" value="40.233334">40°14N</geoCoordinate>
<geoCoordinate id="EE4E5035EE31FFBEFC0B46A5FC48FC14" box="[896,966,976,996]" degrees="36" direction="east" minutes="18" orientation="longitude" pageId="23" pageNumber="214" precision="925" value="36.3">36°18E</geoCoordinate>
,
<date id="FFC41032EE31FFBEFC5A46A5FBA9FC13" box="[977,1063,976,995]" pageId="23" pageNumber="214" value="2005-05-17">
<collectingDate id="EF80E9DAEE31FFBEFC5A46A5FBA9FC13" box="[977,1063,976,995]" pageId="23" pageNumber="214" value="2005-05-17">17.v.2005</collectingDate>
</date>
,
<collectorName id="268F5324EE31FFBEFBB946A4FF25FC0F" pageId="23" pageNumber="214">P. Kment</collectorName>
lgt., 3 JJ
<specimenCount id="9D7CFD7BEE31FFBEFE87469EFEB5FC0E" box="[268,315,997,1022]" pageId="23" pageNumber="214" type="female">5 ♀♀</specimenCount>
(
<collectionCode id="ED6BAE37EE31FFBEFECD469EFE09FC0E" box="[326,391,1003,1022]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="214">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3606579EE31FFBEFFFA4163FEE5FAFE" pageId="23" pageNumber="214" type="discussion">
<paragraph id="8BC536F2EE31FFBEFFFA4163FEE5FAFE" blockId="23.[113,1089,1046,1295]" pageId="23" pageNumber="214">
<emphasis id="B90EEAE0EE31FFBEFFFA4163FF7DFBDE" bold="true" box="[113,243,1046,1070]" pageId="23" pageNumber="214">Comments.</emphasis>
Differences in the shape of the last abdominal segments in males of
<taxonomicName id="4C7A4D71EE31FFBEFC5B416DFECDFBBF" authority="besucheti" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="subSpecies" species="besucheti" subSpecies="besucheti">
<emphasis id="B90EEAE0EE31FFBEFC5B416DFECDFBBF" italics="true" pageId="23" pageNumber="214">Malthodes besucheti besucheti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A4D71EE31FFBEFEF3414DFDF0FBBF" authorityName="Wittmer" authorityYear="1970" box="[376,638,1080,1103]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="subSpecies" species="besucheti" subSpecies="bucakensis">
<emphasis id="B90EEAE0EE31FFBEFEF3414DFDF0FBBF" box="[376,638,1080,1103]" italics="true" pageId="23" pageNumber="214">M. besucheti bucakensis</emphasis>
</taxonomicName>
were described and illustrated already by
<bibRefCitation id="EFEB4B03EE31FFBEFFFA4122FE92FB9F" author="WITTMER W." box="[113,284,1111,1135]" pageId="23" pageNumber="214" pagination="13 - 107" refId="ref10580" refString="WITTMER W. 1970: Zur Kenntnis der Gattung Malthodes (Coleoptera, Cantharidae) (49. Beitrag zur Kenntnis der palaearktischen Cantharidae). Entomologischen Arbeiten aus dem Museum G. Frey 21: 13 - 107." type="journal article" year="1970">WITTMER (1970)</bibRefCitation>
, who wrote [...] indem der Kopulationsaparat ein wenig in der Form abweicht und ca. 20/22 % grösser ist als bei dieser [
<taxonomicName id="4C7A4D71EE31FFBEFDC9410DFD42FB7F" authorityName="Wittmer" authorityYear="1970" box="[578,716,1144,1167]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B90EEAE0EE31FFBEFDC9410DFD42FB7F" box="[578,716,1144,1167]" italics="true" pageId="23" pageNumber="214">M. besucheti</emphasis>
</taxonomicName>
]. The aedeagus of
<taxonomicName id="4C7A4D71EE31FFBEFC2D410DFBCFFB7F" authorityName="Wittmer" authorityYear="1970" baseAuthorityName="Wittmer" baseAuthorityYear="1970" box="[934,1089,1144,1167]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="species" species="bucakensis">
<emphasis id="B90EEAE0EE31FFBEFC2D410DFBCFFB7F" box="[934,1089,1144,1167]" italics="true" pageId="23" pageNumber="214">M. bucakensis</emphasis>
</taxonomicName>
differs significantly from that of
<taxonomicName id="4C7A4D71EE31FFBEFE5341EDFDEDFB5F" authorityName="Wittmer" authorityYear="1970" box="[472,611,1176,1199]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B90EEAE0EE31FFBEFE5341EDFDEDFB5F" box="[472,611,1176,1199]" italics="true" pageId="23" pageNumber="214">M. besucheti</emphasis>
</taxonomicName>
by the narrower and apically turned laterophyses and different apex of the ventral part of the aedeagus (
<figureCitation id="13412A77EE31FFBEFC8F41C2FC09FB3F" box="[772,903,1207,1231]" captionStart="Figs" captionStartId="8.[151,190,1542,1561]" captionTargetBox="[174,1105,219,1355]" captionTargetId="figure-10@8.[173,1105,219,1356]" captionTargetPageId="8" captionText="Figs. 1720.Aedeagus, dorsal view.17 Malthodes paveli sp. nov. (holotype); 18 M. lycicus sp. nov. (paratype); 19 M. besucheti Wittmer, 1970 (paratype); 20 M. bucakensis Wittmer, 1970 (Gülükdağı). Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5363039" httpUri="https://zenodo.org/record/5363039/files/figure.png" pageId="23" pageNumber="214">Figs. 1920</figureCitation>
). I consider these differences as being of specific value and raise
<taxonomicName id="4C7A4D71EE31FFBEFDEA41ADFC39FB1F" authorityName="Wittmer" authorityYear="1970" box="[609,951,1240,1263]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="subSpecies" species="besucheti" subSpecies="bucakensis">
<emphasis id="B90EEAE0EE31FFBEFDEA41ADFC39FB1F" box="[609,951,1240,1263]" italics="true" pageId="23" pageNumber="214">Malthodes besucheti bucakensis</emphasis>
</taxonomicName>
to a separate species,
<taxonomicName id="4C7A4D71EE31FFBEFF47418DFEEAFAFF" authorityName="Wittmer" authorityYear="1970" baseAuthorityName="Wittmer" baseAuthorityYear="1970" box="[204,356,1272,1295]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="214" phylum="Arthropoda" rank="species" species="bucakensis">
<emphasis id="B90EEAE0EE31FFBEFF47418DFEEAFAFF" box="[204,356,1272,1295]" italics="true" pageId="23" pageNumber="214">M. bucakensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>