258 lines
32 KiB
XML
258 lines
32 KiB
XML
<document id="3D327B104D68595EC81AC3F74303CD38" ID-CLB-Dataset="20176" ID-DOI="10.11646/zootaxa.5116.2.5" ID-GBIF-Dataset="dc6e6f10-3b06-448e-bf5f-8bc181884366" ID-ISSN="1175-5326" ID-Zenodo-Dep="6367323" ID-ZooBank="2558C8B0-FABB-4104-B103-56C0830AF277" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647595653746" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Chigray, Ivan A., Ntatsopoulos, Konstantinos & Papadopoulou, Anna" docDate="2022" docId="03D0A51AFFD9FFBE9C97AE77CD2E91D0" docLanguage="en" docName="zootaxa.5116.2.5.pdf" docOrigin="Zootaxa 5116 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blaps lethifera Marsham 1802" docType="treatment" docVersion="7" lastPageNumber="288" masterDocId="FFE9DD62FFCAFFAB9C00A841CF16931E" masterDocTitle="A key to Russian and Eastern European species of Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae: Blaptinae) with the description of a new species from the North Caucasus supported by morphological and molecular data" masterLastPageNumber="291" masterPageNumber="267" pageNumber="286" updateTime="1699318281145" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9B840CD0A13889E9FB1350A9ADEBE4CF">A key to Russian and Eastern European species of Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae: Blaptinae) with the description of a new species from the North Caucasus supported by morphological and molecular data</mods:title>
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<mods:namePart id="7A71DFC0980C8FD0EB1C08A7058E67BE">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="FCA603B63E66CEE8858E697C627C8B53">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia & A. N. Severtsov Institute of ecology and evolution of the Russian Academy of Sciences, Leninsky Prospect 33, 119071 Moscow, Russia Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb., 1, Saint Petersburg 199034 Russia. chigray 93 @ bk. ru; https: // orcid. org / 0000 - 0002 - 3830 - 6860</mods:affiliation>
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<mods:namePart id="D7CBCEFFE6B56AF8F56FF3B59C380F09">Chigray, Ivan A.</mods:namePart>
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<mods:nameIdentifier id="F9A3032558739EDF7F31540DFB44FBA3" type="email">chigray93@bk.ru</mods:nameIdentifier>
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<mods:namePart id="2D1F0100849812C77C7BCF44D881A55B">Ntatsopoulos, Konstantinos</mods:namePart>
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<mods:affiliation id="C7EE71171AB2E1640C629C719AD92DD3">Department of Biological Sciences, University of Cyprus, 1678 Nicosia P. O. Box 20537, Cyprus. & ntatsopoulos. konstantinos @ ucy. ac. cy; https: // orcid. org / 0000 - 0003 - 0673 - 2416</mods:affiliation>
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<mods:nameIdentifier id="D4ACEE1D7021F04D9D76314DEE5D143E" type="email">ntatsopoulos.konstantinos@ucy.ac.cy</mods:nameIdentifier>
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<mods:namePart id="4C05B918C2F0A844D235750FF0870668">Papadopoulou, Anna</mods:namePart>
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<mods:affiliation id="271A3A475DE7CBEAACFF960B63AA6358">Department of Biological Sciences, University of Cyprus, 1678 Nicosia P. O. Box 20537, Cyprus. & papadopoulou. g. anna @ ucy. ac. cy; https: // orcid. org / 0000 - 0002 - 4656 - 4894</mods:affiliation>
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<mods:nameIdentifier id="0196D93B1C9464C87E9F8620D620FD06" type="email">papadopoulou.g.anna@ucy.ac.cy</mods:nameIdentifier>
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<mods:title id="0B3C95EF88738DD75BBD98686891856C">Zootaxa</mods:title>
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<mods:date id="796698A006808C8B588D4FF0AAD1A20E">2022</mods:date>
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<treatment id="03D0A51AFFD9FFBE9C97AE77CD2E91D0" ID-DOI="http://doi.org/10.5281/zenodo.6369181" ID-GBIF-Taxon="194098880" ID-Zenodo-Dep="6369181" LSID="urn:lsid:plazi:treatment:03D0A51AFFD9FFBE9C97AE77CD2E91D0" httpUri="http://treatment.plazi.org/id/03D0A51AFFD9FFBE9C97AE77CD2E91D0" lastPageId="21" lastPageNumber="288" pageId="19" pageNumber="286">
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<subSubSection id="C3634787FFD9FFB89C97AE77CD03954E" box="[151,533,1590,1617]" pageId="19" pageNumber="286" type="nomenclature">
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<paragraph id="8BC6140CFFD9FFB89C97AE77CD03954E" blockId="19.[151,533,1590,1653]" box="[151,533,1590,1617]" pageId="19" pageNumber="286">
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<heading id="D08EA360FFD9FFB89C97AE77CD03954E" bold="true" box="[151,533,1590,1617]" fontSize="11" level="1" pageId="19" pageNumber="286" reason="1">
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<taxonomicName id="4C796F8FFFD9FFB89C97AE77CD03954E" authority="Marsham, 1802" authorityName="Marsham" authorityYear="1802" box="[151,533,1590,1617]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="lethifera">
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<emphasis id="B90DC81EFFD9FFB89C97AE77CD03954E" bold="true" box="[151,533,1590,1617]" pageId="19" pageNumber="286">
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<emphasis id="B90DC81EFFD9FFB89C97AE77CE5C954F" bold="true" box="[151,330,1590,1617]" italics="true" pageId="19" pageNumber="286">Blaps lethifera</emphasis>
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Marsham, 1802
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3634787FFD9FFB89C97AE1ACE3E956B" box="[151,296,1627,1653]" pageId="19" pageNumber="286" type="description">
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<paragraph id="8BC6140CFFD9FFB89C97AE1ACE3E956B" blockId="19.[151,533,1590,1653]" box="[151,296,1627,1653]" pageId="19" pageNumber="286">
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(
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<figureCitation id="13420889FFD9FFB89CA0AE1ACE36956B" box="[160,288,1627,1653]" captionStart="FIGURE 23" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[162,1430,186,1912]" captionTargetId="figure-23@20.[154,1433,181,1920]" captionTargetPageId="20" captionText="FIGURE 23. Blaps spp., habitus. A = non-rugose male of B. verrucosa from Baku, Azerbaijan, (the type locality of B. scabriuscula) ZM MSU; B = rugose male of B. verrucosa from Georgia (Kazbegi), ZIN; C = lectotype labels of B. subalpina, D = lectotype of B. subalpina, male (ZIN); E = lectotype labels of B. pterotapha; F = lectotype of B. pterotapha, female (ZIN)." figureDoi="http://doi.org/10.5281/zenodo.6367379" httpUri="https://zenodo.org/record/6367379/files/figure.png" pageId="19" pageNumber="286">Fig. 23E, F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3634787FFD9FFBE9C97AEDECD0E927B" lastPageId="21" lastPageNumber="288" pageId="19" pageNumber="286" type="reference_group">
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<paragraph id="8BC6140CFFD9FFB89C97AEDECD4E95A9" blockId="19.[151,600,1695,1719]" box="[151,600,1695,1719]" pageId="19" pageNumber="286">
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<emphasis id="B90DC81EFFD9FFB89C97AEDECFB395A9" bold="true" box="[151,165,1695,1719]" pageId="19" pageNumber="286">=</emphasis>
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<taxonomicName id="4C796F8FFFD9FFB89CACAEDECD1E95A9" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[172,520,1695,1719]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="pterotapha">
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<emphasis id="B90DC81EFFD9FFB89CACAEDECE4C95A9" box="[172,346,1695,1719]" italics="true" pageId="19" pageNumber="286">Blaps pterotapha</emphasis>
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<bibRefCitation id="EFE869FDFFD9FFB89D61AEDECD1E95A9" author="Menetries, E." box="[353,520,1695,1719]" pageId="19" pageNumber="286" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
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||
</taxonomicName>
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||
,
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||
<emphasis id="B90DC81EFFD9FFB89E12AEDECD4E95A9" bold="true" box="[530,600,1695,1719]" pageId="19" pageNumber="286">
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<taxonomicNameLabel id="A23E7565FFD9FFB89E12AEDECD4E95A9" box="[530,600,1695,1719]" pageId="19" pageNumber="286">syn. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
<paragraph id="8BC6140CFFD9FFBE9C97AE9ECD0E927B" blockId="19.[151,1437,1759,2037]" lastBlockId="21.[151,1437,151,718]" lastPageId="21" lastPageNumber="288" pageId="19" pageNumber="286">
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The species
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<taxonomicName id="4C796F8FFFD9FFB89D1DAE9ECD9995E7" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[285,655,1759,1785]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="pterotapha">
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<emphasis id="B90DC81EFFD9FFB89D1DAE9ECECE95E7" box="[285,472,1759,1785]" italics="true" pageId="19" pageNumber="286">Blaps pterotapha</emphasis>
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<bibRefCitation id="EFE869FDFFD9FFB89DDEAE9ECD9995E7" author="Menetries, E." box="[478,655,1759,1785]" pageId="19" pageNumber="286" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
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||
</taxonomicName>
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was described from Baku and long time was interpreted as a subspecies of
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<taxonomicName id="4C796F8FFFD9FFB89CBAAF42CE769403" authorityName="Marsham" authorityYear="1802" box="[186,352,1795,1821]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="lethifera">
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<emphasis id="B90DC81EFFD9FFB89CBAAF42CE769403" box="[186,352,1795,1821]" italics="true" pageId="19" pageNumber="286">Blaps lethifera</emphasis>
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</taxonomicName>
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(see full bibliography in Abdurakhamnov & Nabozhenko (2011)). Transcaucasian populations named “
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<emphasis id="B90DC81EFFD9FFB89CF4AF66CEFA945F" box="[244,492,1831,1857]" italics="true" pageId="19" pageNumber="286">
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<taxonomicName id="4C796F8FFFD9FFB89CF4AF66CE78945F" authorityName="Marsham" authorityYear="1802" box="[244,366,1831,1857]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="lethifera">B. lethifera</taxonomicName>
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<taxonomicName id="4C796F8FFFD9FFB89D75AF66CEFA945F" authorityName="Menetries" authorityYear="1832" box="[373,492,1831,1857]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="pterotapha">pterotapha</taxonomicName>
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</emphasis>
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” differ from the European populations only by weakly emarginated lateral margins of the pronotum at the base. This character is very variable, and specimens with emarginated or weakly emarginated or non-emarginated lateral pronotum margins can be found even in one Transcaucasian micropopulation. This unreliable character’s entire spectrum of variability can be observed in specimens from the east Caspian coast to western
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<collectingCountry id="F36E549CFFD9FFB89949AFD2CA8F94B3" box="[1353,1433,1939,1965]" name="Turkey" pageId="19" pageNumber="286">Turkey</collectingCountry>
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. The extensive variability of this species has been beautifully illustrated in the work of
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<bibRefCitation id="EFE869FDFFD9FFB89827AFF9CBD794CF" author="Sustek, Z." box="[1063,1217,1975,2003]" pageId="19" pageNumber="286" pagination="143 - 153" refId="ref15463" refString="Sustek, Z. (1982) Contribution to the synonymy of Blaps lethifera Marsham, 1802 (Coleoptera Tenebrionidae). Acta entomologica bohemoslovaca, 79, 143 - 153." type="journal article" year="1982">Šustek (1982)</bibRefCitation>
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. Another illustrative example is two very unusual specimens from the Don River delta area. These specimens were collected after deep excavation by an excavator in sandy alluvial soils near the banks of Don River in the burrows of
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<taxonomicName id="4C796F8FFFDFFFBE989BA8D6CE7493CB" ID-CoL="4YDZ5" authority="Guldenstadt, 1770" authorityName="Guldenstadt" authorityYear="1770" class="Mammalia" family="Spalacidae" genus="Spalax" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="288" phylum="Chordata" rank="species" species="microphthalmus">
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<emphasis id="B90DC81EFFDFFFBE989BA8D6CA8A93AF" box="[1179,1436,151,177]" italics="true" pageId="21" pageNumber="288">Spalax microphthalmus</emphasis>
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Güldenstädt, 1770
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</taxonomicName>
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. The first two authors interpreted them as an isolated, distant micropopulation of
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<taxonomicName id="4C796F8FFFDFFFBE98CCA8FACE9C93E7" authority="(Chigray et al. 2015)" baseAuthorityName="Chigray" baseAuthorityYear="2015" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="scabriuscula">
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<emphasis id="B90DC81EFFDFFFBE98CCA8FACA8A93CB" box="[1228,1436,187,213]" italics="true" pageId="21" pageNumber="288">Blaps scabriuscula</emphasis>
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||
(
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<bibRefCitation id="EFE869FDFFDFFFBE9C9FA89ECE9793E7" author="Chigray, I. A. & Abdurakhmanov, G. M. & Nabozhenko, M. V. & Shmatko, V. Yu." box="[159,385,222,249]" pageId="21" pageNumber="288" pagination="59 - 68" refId="ref13195" refString="Chigray, I. A., Abdurakhmanov, G. M., Nabozhenko, M. V. & Shmatko, V. Yu. (2015) Morphological diversity and distribution of Blaps scabriuscula Menetries, 1832 (Coleoptera: Tenebrionidae). Yug Rossii: Ekologiya, Razvitiye, 10 (4), 59 - 68. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2015 - 4 - 59 - 68" type="journal article" year="2015">
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Chigray
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<emphasis id="B90DC81EFFDFFFBE9D03A89ECE2B93E7" box="[259,317,223,249]" italics="true" pageId="21" pageNumber="288">et al.</emphasis>
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2015
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</bibRefCitation>
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)
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</taxonomicName>
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. However, COI analysis showed that this Don micropopulation is genetically highly similar with specimens assigned to
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<taxonomicName id="4C796F8FFFDFFFBE9DCFA942CD659203" authorityName="Marsham" authorityYear="1802" box="[463,627,259,285]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="lethifera">
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<emphasis id="B90DC81EFFDFFFBE9DCFA942CD659203" box="[463,627,259,285]" italics="true" pageId="21" pageNumber="288">Blaps lethifera</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13420889FFDFFFBE9E83A942CDC39203" box="[643,725,259,285]" captionStart="FIGURE 22" captionStartId="15.[151,250,1093,1117]" captionTargetBox="[151,1429,433,1070]" captionTargetId="figure-325@15.[151,1436,433,1070]" captionTargetPageId="15" captionText="FIGURE 22. ML phylogenetic tree reconstruction of Blaps species from the Northern Caucasus in IQ-TREE. Numbers above branches represent bootstrap support values (values <50% and the ones corresponding to short intraspecific branches are omitted)." figureDoi="http://doi.org/10.5281/zenodo.6367377" httpUri="https://zenodo.org/record/6367377/files/figure.png" pageId="21" pageNumber="288">Fig. 22</figureCitation>
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||
), despite significant morphological differences. Considering the information mentioned above, we propose the following new synonymy:
|
||
<taxonomicName id="4C796F8FFFDFFFBE9FDBA966CA2C925F" authority="Marsham, 1802" authorityName="Marsham" authorityYear="1802" box="[987,1338,295,321]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="lethifera">
|
||
<emphasis id="B90DC81EFFDFFFBE9FDBA966CB96925F" box="[987,1152,295,321]" italics="true" pageId="21" pageNumber="288">Blaps lethifera</emphasis>
|
||
Marsham, 1802
|
||
</taxonomicName>
|
||
<emphasis id="B90DC81EFFDFFFBE9944A966CA42925F" bold="true" box="[1348,1364,295,321]" pageId="21" pageNumber="288">=</emphasis>
|
||
<taxonomicName id="4C796F8FFFDFFFBE995EA966CED1927B" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="pterotapha">
|
||
<emphasis id="B90DC81EFFDFFFBE995EA966CE18927B" italics="true" pageId="21" pageNumber="288">Blaps pterotapha</emphasis>
|
||
<bibRefCitation id="EFE869FDFFDFFFBE9D15A90ACED1927B" author="Menetries, E." box="[277,455,331,357]" pageId="21" pageNumber="288" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B90DC81EFFDFFFBE9DCDA90ACD0E927B" bold="true" box="[461,536,331,357]" pageId="21" pageNumber="288">
|
||
<taxonomicNameLabel id="A23E7565FFDFFFBE9DCDA90ACD0E927B" box="[461,536,331,357]" pageId="21" pageNumber="288">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF064484FFDEFFBF9C97AFD6CA9094E6" ID-DOI="http://doi.org/10.5281/zenodo.6367379" ID-Zenodo-Dep="6367379" httpUri="https://zenodo.org/record/6367379/files/figure.png" pageId="20" pageNumber="287" startId="20.[151,250,1943,1967]" targetBox="[162,1430,186,1912]" targetPageId="20">
|
||
<paragraph id="8BC6140CFFDEFFBF9C97AFD6CA9094E6" blockId="20.[151,1437,1943,2040]" pageId="20" pageNumber="287">
|
||
<emphasis id="B90DC81EFFDEFFBF9C97AFD6CE3294B1" bold="true" box="[151,292,1943,1967]" pageId="20" pageNumber="287">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C796F8FFFDEFFBF9D2FAFD9CEB694B1" box="[303,416,1944,1968]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90DC81EFFDEFFBF9D2FAFD9CE7F94AE" box="[303,361,1944,1968]" italics="true" pageId="20" pageNumber="287">Blaps</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, habitus. A = non-rugose male of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9F15AFD9CC8C94B1" authorityName="Adams" authorityYear="1817" box="[789,922,1944,1967]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="verrucosa">
|
||
<emphasis id="B90DC81EFFDEFFBF9F15AFD9CC8C94B1" box="[789,922,1944,1967]" italics="true" pageId="20" pageNumber="287">B. verrucosa</emphasis>
|
||
</taxonomicName>
|
||
from Baku, Azerbaijan, (the type locality of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9986AFD9CE0E94CA" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="scabriuscula">
|
||
<emphasis id="B90DC81EFFDEFFBF9986AFD9CE0E94CA" italics="true" pageId="20" pageNumber="287">B. scabriuscula</emphasis>
|
||
</taxonomicName>
|
||
) ZM MSU; B = rugose male of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9E5CAFFDCDCA94CD" authorityName="Adams" authorityYear="1817" box="[604,732,1980,2003]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="verrucosa">
|
||
<emphasis id="B90DC81EFFDEFFBF9E5CAFFDCDCA94CD" box="[604,732,1980,2003]" italics="true" pageId="20" pageNumber="287">B. verrucosa</emphasis>
|
||
</taxonomicName>
|
||
from Georgia (Kazbegi), ZIN; C = lectotype labels of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9901AFFDCA9794CA" authorityName="Menetries" authorityYear="1832" box="[1281,1409,1980,2004]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="subalpina">
|
||
<emphasis id="B90DC81EFFDEFFBF9901AFFDCA0E94CD" box="[1281,1304,1980,2003]" italics="true" pageId="20" pageNumber="287">B.</emphasis>
|
||
subalpina
|
||
</taxonomicName>
|
||
, D = lectotype of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9D2AAFA1CEBD94E6" authorityName="Menetries" authorityYear="1832" box="[298,427,2016,2040]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="subalpina">
|
||
<emphasis id="B90DC81EFFDEFFBF9D2AAFA1CEBD94E6" box="[298,427,2016,2040]" italics="true" pageId="20" pageNumber="287">B. subalpina</emphasis>
|
||
</taxonomicName>
|
||
, male (ZIN); E = lectotype labels of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9F22AFA1CCBA94E6" authorityName="Menetries" authorityYear="1832" box="[802,940,2016,2040]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="pterotapha">
|
||
<emphasis id="B90DC81EFFDEFFBF9F22AFA1CCBA94E6" box="[802,940,2016,2040]" italics="true" pageId="20" pageNumber="287">B. pterotapha</emphasis>
|
||
</taxonomicName>
|
||
; F = lectotype of
|
||
<taxonomicName id="4C796F8FFFDEFFBF9861AFA1CBFD94E6" authorityName="Menetries" authorityYear="1832" box="[1121,1259,2016,2040]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="287" phylum="Arthropoda" rank="species" species="pterotapha">
|
||
<emphasis id="B90DC81EFFDEFFBF9861AFA1CBFD94E6" box="[1121,1259,2016,2040]" italics="true" pageId="20" pageNumber="287">B. pterotapha</emphasis>
|
||
</taxonomicName>
|
||
, female (ZIN).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3634787FFDFFFBE9CC7A92ECC6B92CF" pageId="21" pageNumber="288" type="description">
|
||
<paragraph id="8BC6140CFFDFFFBE9CC7A92ECC6B92CF" blockId="21.[151,1437,151,718]" pageId="21" pageNumber="288">
|
||
In addition, we examined the specimen, which was listed as
|
||
<taxonomicName id="4C796F8FFFDFFFBE9F83A92ECA8A9297" authority="Fischer von Waldheim, 1844" authorityName="Fischer von Waldheim" authorityYear="1844" box="[899,1436,367,393]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="pterosticha">
|
||
<emphasis id="B90DC81EFFDFFFBE9F83A92ECB5F9297" box="[899,1097,367,393]" italics="true" pageId="21" pageNumber="288">Blaps pterosticha</emphasis>
|
||
Fischer von Waldheim, 1844
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49BDDAEEFFDFFFBE9CD4A9D2CE8092B3" box="[212,406,403,429]" country="Russia" name="Orenburg" pageId="21" pageNumber="288">Orenburg Region</collectingRegion>
|
||
of
|
||
<collectingCountry id="F36E549CFFDFFFBE9DB9A9D2CD1292B3" box="[441,516,403,429]" name="Russia" pageId="21" pageNumber="288">Russia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE869FDFFDFFFBE9E12A9D2CC0392B3" author="Shapovalov, A. M. & Grigoriev, V. E. & Nemkov, V. A. & Rusakov, A. V. & Kazakov, E. P." box="[530,789,403,429]" pageId="21" pageNumber="288" pagination="39 - 48" refId="ref15084" refString="Shapovalov, A. M., Grigoriev, V. E., Nemkov, V. A., Rusakov, A. V. & Kazakov, E. P. (2011) Interesting finds of beetles (Insecta, Coleoptera) in the Orenburg Region. Trudy Orenburgskogo otdeleniya Russkogo entomologicheskogo obshchestva, 1, 39 - 48. [in Russian]" type="journal article" year="2011">
|
||
Shapovalov
|
||
<emphasis id="B90DC81EFFDFFFBE9E9EA9D2CDC392B3" box="[670,725,403,429]" italics="true" pageId="21" pageNumber="288">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE869FDFFDFFFBE9F20A9D2CCFE92B3" author="Kozminykh, V. O." box="[800,1000,402,429]" pageId="21" pageNumber="288" pagination="16 - 42" refId="ref14016" refString="Kozminykh, V. O. (2015) Faunistic data on tenebrionid beetles (Coleoptera, Tenebrionidae) of the Orenburg Region. Vestnik of Orenburg State Pedagogical University. Electronic Scientific Journal, 1 (13), 16 - 42. [in Russian]" type="journal article" year="2015">Kozminykh 2015</bibRefCitation>
|
||
). The specimen is a typical
|
||
<taxonomicName id="4C796F8FFFDFFFBE9918A9D2CA8392B3" authorityName="Marsham" authorityYear="1802" box="[1304,1429,403,429]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="lethifera">
|
||
<emphasis id="B90DC81EFFDFFFBE9918A9D2CA8392B3" box="[1304,1429,403,429]" italics="true" pageId="21" pageNumber="288">B. lethifera</emphasis>
|
||
</taxonomicName>
|
||
, so
|
||
<taxonomicName id="4C796F8FFFDFFFBE9CB7A9F6CE4592CF" authorityName="Fischer von Waldheim" authorityYear="1844" box="[183,339,439,465]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="pterosticha">
|
||
<emphasis id="B90DC81EFFDFFFBE9CB7A9F6CE4592CF" box="[183,339,439,465]" italics="true" pageId="21" pageNumber="288">B. pterosticha</emphasis>
|
||
</taxonomicName>
|
||
must be excluded from the Russian faunistic list.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3634787FFDFFFBE9CC7A99ACD2E91D0" pageId="21" pageNumber="288" type="reference_group">
|
||
<paragraph id="8BC6140CFFDFFFBE9CC7A99ACD2E91D0" blockId="21.[151,1437,151,718]" pageId="21" pageNumber="288">
|
||
<taxonomicName id="4C796F8FFFDFFFBE9CC7A99ACD2292EB" authority="Solier, 1848" authorityName="Solier" authorityYear="1848" box="[199,564,475,501]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="sinuatocollis">
|
||
<emphasis id="B90DC81EFFDFFFBE9CC7A99ACEB692EB" box="[199,416,475,501]" italics="true" pageId="21" pageNumber="288">Blaps sinuatocollis</emphasis>
|
||
<bibRefCitation id="EFE869FDFFDFFFBE9DAAA99ACD2292EB" author="Solier, A. J. J." box="[426,564,475,501]" pageId="21" pageNumber="288" pagination="149 - 370" refId="ref15399" refString="Solier, A. J. J. (1848) Essai sur les collapterides. 14 e Tribu. Blapsites. In: Baudi di Selve: Studi Entomologici. Pubblicati per cura di Flaminio Baudi e di Eugenio Truqui. Tomo 1. Torino: Stamperia Degli Artisti Tipografi, pp. 149 - 370, pls. iv - xv." type="book chapter" year="1848">Solier, 1848</bibRefCitation>
|
||
</taxonomicName>
|
||
(recorded as nominotypical subspecies from Southern
|
||
<collectingCountry id="F36E549CFFDFFFBE98B1A99ACBEA92EB" box="[1201,1276,475,501]" name="Russia" pageId="21" pageNumber="288">Russia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE869FDFFDFFFBE990DA99ACE479107" author="Nabozhenko, M. V. & Chigray, I. A." pageId="21" pageNumber="288" pagination="1815" refId="ref14579" refString="Nabozhenko, M. V. & Chigray, I. A. (2020) Tribe Blaptini Leach, 1815. In: Iwan, D. & Lobl, I. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 5. Tenebrionoidea. Leiden, Brill, pp. 268 - 296. https: // doi. org / 10.1163 / 9789004434998 _ 004" type="book chapter" year="2020">Nabozhenko & Chigray 2020</bibRefCitation>
|
||
)) was described as the variation B of
|
||
<taxonomicName id="4C796F8FFFDFFFBE9F06A9BECCC79107" box="[774,977,511,537]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="reflexicollis">
|
||
<emphasis id="B90DC81EFFDFFFBE9F06A9BECCC79107" box="[774,977,511,537]" italics="true" pageId="21" pageNumber="288">Blaps reflexicollis</emphasis>
|
||
</taxonomicName>
|
||
(now a junior synonym of
|
||
<taxonomicName id="4C796F8FFFDFFFBE990CA9BECA9A9107" authorityName="Marsham" authorityYear="1802" box="[1292,1420,511,537]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="lethifera">
|
||
<emphasis id="B90DC81EFFDFFFBE990CA9BECA9A9107" box="[1292,1420,511,537]" italics="true" pageId="21" pageNumber="288">B. lethifera</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<typeStatus id="54C2AAAEFFDFFFBE9C97AA62CFCF9123" box="[151,217,547,573]" pageId="21" pageNumber="288">Types</typeStatus>
|
||
of both taxa are lost (personal communication with Antoine Mantilleri (MNHN)). In our opinion, this name is infrasubspecific and must be excluded from zoological nomenclature because Solier’s description unambiguously reveals that the name was proposed for an infrasubspecific entity (article 45.6.4 of ICZN). Solier indicated that var. B (
|
||
<taxonomicName id="4C796F8FFFDFFFBE9CBAAACECE5F91B7" authorityName="Solier" authorityYear="1848" box="[186,329,655,681]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="288" phylum="Arthropoda" rank="species" species="sinuatocollis">
|
||
<emphasis id="B90DC81EFFDFFFBE9CBAAACECE5F91B7" box="[186,329,655,681]" italics="true" pageId="21" pageNumber="288">sinuatocollis</emphasis>
|
||
</taxonomicName>
|
||
) is just
|
||
<specimenCount id="9D7FDF85FFDFFFBE9DA3AACECD1F91B7" box="[419,521,655,681]" pageId="21" pageNumber="288" type="male">one male</specimenCount>
|
||
deviating from the typical form, and if its characters were stable, then it could be a species (
|
||
<bibRefCitation id="EFE869FDFFDFFFBE9D0AAAF2CE8791D3" author="Solier, A. J. J." box="[266,401,691,717]" pageId="21" pageNumber="288" pagination="149 - 370" refId="ref15399" refString="Solier, A. J. J. (1848) Essai sur les collapterides. 14 e Tribu. Blapsites. In: Baudi di Selve: Studi Entomologici. Pubblicati per cura di Flaminio Baudi e di Eugenio Truqui. Tomo 1. Torino: Stamperia Degli Artisti Tipografi, pp. 149 - 370, pls. iv - xv." type="book chapter" year="1848">Solier 1848</bibRefCitation>
|
||
, pp. 315, 316).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |