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<mods:title id="F4CB8C12E29001DDBAF0BC2D6B5DF455">Paraneseuthia Franz of Japan: two new species and rediscovery of the enigmatic P. holzneri Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="03CFA37A2E5EFF97FF36F9F4FB904E5D" ID-DOI="http://doi.org/10.5281/zenodo.5950824" ID-GBIF-Taxon="154521292" ID-Zenodo-Dep="5950824" LSID="urn:lsid:plazi:treatment:03CFA37A2E5EFF97FF36F9F4FB904E5D" httpUri="http://treatment.plazi.org/id/03CFA37A2E5EFF97FF36F9F4FB904E5D" lastPageId="5" lastPageNumber="271" pageId="4" pageNumber="270">
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<paragraph id="8BD9126C2E5EFF96FF36F9F4FDA9494F" blockId="4.[151,535,1627,1687]" box="[151,535,1627,1654]" pageId="4" pageNumber="270">
<heading id="D091A5002E5EFF96FF36F9F4FDA9494F" bold="true" box="[151,535,1627,1654]" fontSize="11" level="1" pageId="4" pageNumber="270" reason="1">
<taxonomicName id="4C6669EF2E5EFF96FF36F9F4FDA9494F" ID-CoL="4D5YH" authority="(Franz)" authorityName="Franz" baseAuthorityName="Franz" baseAuthorityYear="1976" box="[151,535,1627,1654]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="holzneri">
<emphasis id="B912CE7E2E5EFF96FF36F9F4FDA9494F" bold="true" box="[151,535,1627,1654]" pageId="4" pageNumber="270">
<emphasis id="B912CE7E2E5EFF96FF36F9F4FE0F494C" bold="true" box="[151,433,1627,1653]" italics="true" pageId="4" pageNumber="270">Paraneseuthia holzneri</emphasis>
(Franz)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41E72E5EFF96FF36F9D1FEFC49AE" box="[151,322,1662,1687]" pageId="4" pageNumber="270" type="description">
<paragraph id="8BD9126C2E5EFF96FF36F9D1FEFC49AE" blockId="4.[151,535,1627,1687]" box="[151,322,1662,1687]" pageId="4" pageNumber="270">
(
<figureCitation id="135D0EE92E5EFF96FF3EF9D1FF5949AE" box="[159,231,1662,1687]" captionStart="FIGURES 13" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[159,1424,864,1700]" captionTargetId="figure@2.[151,1436,844,1715]" captionTargetPageId="2" captionText="FIGURES 13. Dorsal habitus of males. Paraneseuthia ishigakiana sp. n. (1); P. ehimensis sp. n. (2), P. holzneri (Franz) (3)." httpUri="https://zenodo.org/record/2615313/files/figure.png" pageId="4" pageNumber="270">Figs 3</figureCitation>
,
<figureCitation id="135D0EE92E5EFF96FF52F9D1FE8449AE" box="[243,314,1662,1687]" captionStart="FIGURES 1017" captionStartId="5.[151,264,1743,1766]" captionTargetBox="[166,1418,426,1712]" captionTargetId="figure@5.[151,1436,411,1722]" captionTargetPageId="5" captionText="FIGURES 1017. Aedeagus in ventral (10, 12, 14, 16) and lateral (11, 13, 15, 17) views. Paraneseuthia ishigakiana sp. n. (1011); P. ehimensis sp. n. (extreme forms of variability among studied specimens, 1215), P. holzneri (Franz) (1617)." httpUri="https://zenodo.org/record/2615317/files/figure.png" pageId="4" pageNumber="270">1617</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41E72E5EFF96FF36F969FD2E49C2" pageId="4" pageNumber="270" type="reference_group">
<paragraph id="8BD9126C2E5EFF96FF36F969FD9C49E5" blockId="4.[151,1037,1734,1818]" box="[151,546,1734,1756]" pageId="4" pageNumber="270">
<treatmentCitationGroup id="AB7635422E5EFF96FF36F969FD9C49E5" box="[151,546,1734,1756]" pageId="4" pageNumber="270">
<taxonomicName id="4C6669EF2E5EFF96FF36F969FE6549E5" ID-CoL="3DD67" authority="Franz, 1976: 51" authorityName="Franz" authorityPageNumber="51" authorityYear="1976" box="[151,475,1734,1756]" class="Insecta" family="Staphylinidae" genus="Euthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="holzneri">
<emphasis id="B912CE7E2E5EFF96FF36F969FE8A49E5" box="[151,308,1734,1756]" italics="true" pageId="4" pageNumber="270">Euthia holzneri</emphasis>
<treatmentCitation id="0AC7347D2E5EFF96FE9BF968FE6549E5" author="Franz, H." box="[314,475,1734,1756]" page="51" pageId="4" pageNumber="270" year="1976">
<bibRefCitation id="EFF76F9D2E5EFF96FE9BF968FE0849E5" author="Franz, H." box="[314,438,1734,1756]" pageId="4" pageNumber="270" pagination="51 - 60" refId="ref2840" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Blatter, 72, 51 - 60." type="journal article" year="1976">Franz, 1976</bibRefCitation>
: 51
</treatmentCitation>
</taxonomicName>
, fig. 1.
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</paragraph>
<paragraph id="8BD9126C2E5EFF96FF36F94AFD2E49C2" blockId="4.[151,1037,1734,1818]" box="[151,656,1765,1787]" pageId="4" pageNumber="270">
<treatmentCitationGroup id="AB7635422E5EFF96FF36F94AFD2E49C2" box="[151,656,1765,1787]" pageId="4" pageNumber="270">
<taxonomicName id="4C6669EF2E5EFF96FF36F94AFE3C49C2" authority="Franz" authorityName="Franz" box="[151,386,1765,1787]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="holzneri">
<emphasis id="B912CE7E2E5EFF96FF36F94AFE8149C2" box="[151,319,1765,1787]" italics="true" pageId="4" pageNumber="270">Eutheia holzneri</emphasis>
Franz
</taxonomicName>
;
<treatmentCitation id="0AC7347D2E5EFF96FE2DF94AFD3249C2" author="O'Keefe, S. T. &amp; Li, J. - K." box="[396,652,1765,1787]" page="157" pageId="4" pageNumber="270" year="1998">
<bibRefCitation id="EFF76F9D2E5EFF96FE2DF94AFDE749C2" author="O'Keefe, S. T. &amp; Li, J. - K." box="[396,601,1765,1787]" pageId="4" pageNumber="270" pagination="150 - 162" refId="ref3416" refString="O'Keefe, S. T. &amp; Li, J. - K. (1998) Review of the Scydmaenidae (Coleoptera: Scydmaenidae) of eastern Asia, with particular reference to Scydmaenus, and description of the first Scydmaenid from Hainan Island, China. Journal of the New York Entomological Society, 106 (4), 150 - 162." type="journal article" year="1998">O'Keefe &amp; Li, 1998</bibRefCitation>
: 157
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37C41E72E5EFF96FF36F8ABFBB34823" box="[151,1037,1796,1818]" pageId="4" pageNumber="270" type="description">
<paragraph id="8BD9126C2E5EFF96FF36F8ABFBB34823" blockId="4.[151,1037,1734,1818]" box="[151,1037,1796,1818]" pageId="4" pageNumber="270">
<treatmentCitationGroup id="AB7635422E5EFF96FF36F8ABFBB34823" box="[151,1037,1796,1818]" pageId="4" pageNumber="270">
<taxonomicName id="4C6669EF2E5EFF96FF36F8ABFE694823" ID-CoL="4D5YH" authority="(Franz)" authorityName="Franz" baseAuthorityName="Franz" baseAuthorityYear="1976" box="[151,471,1796,1818]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="holzneri">
<emphasis id="B912CE7E2E5EFF96FF36F8ABFE3A4823" box="[151,388,1796,1818]" italics="true" pageId="4" pageNumber="270">Paraneseuthia holzneri</emphasis>
(Franz)
</taxonomicName>
;
<treatmentCitation id="0AC7347D2E5EFF96FE43F8ABFCF44823" author="Jaloszynski, P. &amp; Hoshina, H." box="[482,842,1796,1818]" page="140" pageId="4" pageNumber="270" year="2004">
<bibRefCitation id="EFF76F9D2E5EFF96FE43F8ABFCA64823" author="Jaloszynski, P. &amp; Hoshina, H." box="[482,792,1796,1818]" pageId="4" pageNumber="270" pagination="133 - 143" refId="ref3305" refString="Jaloszynski, P. &amp; Hoshina, H. (2004) Revision of Japanese species of Paraneseuthia Franz (Coleoptera, Scydmaenidae). Japanese Journal of Systematic Entomology, 10, 133 - 143." type="journal article" year="2004">Jałoszyński &amp; Hoshina (2004)</bibRefCitation>
: 140
</treatmentCitation>
, figs 2D, G, 4AC)
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</paragraph>
</subSubSection>
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<emphasis id="B912CE7E2E5EFF96FF36F8E9FD384866" bold="true" box="[151,646,1862,1887]" pageId="4" pageNumber="270">
<collectingCounty id="62B86AE02E5EFF96FF36F8E9FF414866" box="[151,255,1862,1887]" pageId="4" pageNumber="270">Material</collectingCounty>
studied.
<materialsCitation id="3B0E18312E5EFF96FED6F8E9FD384866" ID-GBIF-Occurrence="2118194746" box="[375,646,1862,1887]" collectingDate="1980-08-01" collectionCode="MHNG" collectorName="I. Lobl" country="Japan" county="Pref." elevation="750" location="Pref." pageId="4" pageNumber="270" specimenCount="1" specimenCount-male="1" stateProvince="Gifu">
<collectingCountry id="F37152FC2E5EFF96FED6F8E9FE6F4866" box="[375,465,1862,1887]" name="Japan" pageId="4" pageNumber="270">JAPAN</collectingCountry>
(
<collectingRegion id="49A2DC8E2E5EFF96FE44F8E9FD934866" box="[485,557,1862,1887]" country="Japan" name="Gifu" pageId="4" pageNumber="270">GIFU</collectingRegion>
<collectingCounty id="62B86AE02E5EFF96FD9BF8E9FDC74866" box="[570,633,1862,1887]" pageId="4" pageNumber="270">Pref.</collectingCounty>
):
</materialsCitation>
</emphasis>
<specimenCount id="9D60D9E52E5EFF96FD35F8E8FD154867" box="[660,683,1863,1886]" pageId="4" pageNumber="270" type="male"></specimenCount>
,
<quantity id="4C9EBF892E5EFF96FD1CF8E8FD424866" box="[701,764,1862,1887]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="270" unit="km" value="8.0">8 km</quantity>
SE
<collectingRegion id="49A2DC8E2E5EFF96FC97F8E9FCC14866" box="[822,895,1862,1887]" country="Japan" name="Osaka" pageId="4" pageNumber="270">Ōsaka</collectingRegion>
,
<collectingRegion id="49A2DC8E2E5EFF96FC31F8E9FC7D4866" box="[912,963,1862,1887]" country="Japan" name="Gifu" pageId="4" pageNumber="270">Gifu</collectingRegion>
Pref.,
<quantity id="4C9EBF892E5EFF96FBBBF8E9FBD74867" box="[1050,1129,1862,1887]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="270" unit="m" value="750.0">
<elevation id="004BF55F2E5EFF96FBBBF8E9FBD74867" box="[1050,1129,1862,1887]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="270" unit="m" value="750.0">750 m</elevation>
</quantity>
,
<date id="FFD834AC2E5EFF96FBDAF8E8FABC4866" box="[1147,1282,1863,1887]" pageId="4" pageNumber="270" value="1980-08-01">
<collectingDate id="EF9CCD442E5EFF96FBDAF8E8FABC4866" box="[1147,1282,1863,1887]" pageId="4" pageNumber="270" value="1980-08-01">1.VIII.1980</collectingDate>
</date>
,
<collectorName id="269377BA2E5EFF96FAB5F8E8FAD84866" box="[1300,1382,1862,1887]" pageId="4" pageNumber="270">I. Löbl</collectorName>
leg. (MHNG).
</paragraph>
</subSubSection>
<subSubSection id="C37C41E72E5EFF97FF66F821FB904E5D" lastPageId="5" lastPageNumber="271" pageId="4" pageNumber="270" type="discussion">
<paragraph id="8BD9126C2E5EFF96FF66F821FA2248D6" blockId="4.[151,1436,1862,2031]" pageId="4" pageNumber="270">
<emphasis id="B912CE7E2E5EFF96FF66F821FE85489E" bold="true" box="[199,315,1934,1959]" pageId="4" pageNumber="270">Remarks.</emphasis>
Only the
<typeStatus id="54DDACCE2E5EFF96FE0BF821FDB2489E" box="[426,524,1934,1959]" pageId="4" pageNumber="270" type="holotype">holotype</typeStatus>
male of this species has been known so far, and this specimen has a fully erected and somewhat distorted aedeagus (illustrated in
<bibRefCitation id="EFF76F9D2E5EFF96FD7FF81DFBEB48F2" author="Jaloszynski, P. &amp; Hoshina, H." box="[734,1109,1970,1995]" pageId="4" pageNumber="270" pagination="133 - 143" refId="ref3305" refString="Jaloszynski, P. &amp; Hoshina, H. (2004) Revision of Japanese species of Paraneseuthia Franz (Coleoptera, Scydmaenidae). Japanese Journal of Systematic Entomology, 10, 133 - 143." type="journal article" year="2004">Jałoszyński &amp; Hoshina (2004))</bibRefCitation>
. The
<typeStatus id="54DDACCE2E5EFF96FB3EF81DFABF48F2" box="[1183,1281,1970,1995]" pageId="4" pageNumber="270" type="holotype">holotype</typeStatus>
comes from Okutama-machi env., probably from Mt. Mazukari (misspelled Badsukari on the label),
<collectingRegion id="49A2DC8E2E5EFF96FBDBF879FAFC48D6" box="[1146,1346,2006,2031]" country="Japan" name="Tokyo" pageId="4" pageNumber="270">Tōkyō Prefecture</collectingRegion>
. During
</paragraph>
<paragraph id="8BD9126C2E5FFF97FF36FF38FB904E5D" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="271">
several collecting trips to the region of Okutama, I was not able to find any additional specimens of this species. The male recorded here shares with the
<typeStatus id="54DDACCE2E5FFF97FDC2FF13FD7B4FEC" box="[611,709,188,213]" pageId="5" pageNumber="271" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6669EF2E5FFF97FD4FFF12FCDA4FEC" baseAuthorityName="Franz" baseAuthorityYear="1976" box="[750,868,188,213]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="holzneri">
<emphasis id="B912CE7E2E5FFF97FD4FFF12FCDA4FEC" box="[750,868,188,213]" italics="true" pageId="5" pageNumber="271">P. holzneri</emphasis>
</taxonomicName>
the protibiae, which have a very indistinct mesal expansion at middle, but lack the denticle which is characteristic of other Japanese
<taxonomicName id="4C6669EF2E5FFF97FBF3FF70FB4B4FC1" authorityName="Franz" authorityYear="1986" box="[1106,1269,223,248]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE7E2E5FFF97FBF3FF70FB4B4FC1" box="[1106,1269,223,248]" italics="true" pageId="5" pageNumber="271">Paraneseuthia</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EFF76F9D2E5FFF97FF36FEABFE524E25" author="Jaloszynski, P. &amp; Hoshina, H." box="[151,492,260,285]" pageId="5" pageNumber="271" pagination="133 - 143" refId="ref3305" refString="Jaloszynski, P. &amp; Hoshina, H. (2004) Revision of Japanese species of Paraneseuthia Franz (Coleoptera, Scydmaenidae). Japanese Journal of Systematic Entomology, 10, 133 - 143." type="journal article" year="2004">Jałoszyński &amp; Hoshina (2004)</bibRefCitation>
, fig. 2G), similar measurements, and the aedeagus with the lateral subapical lobes of the basal capsule, broadly and deeply emarginate apex of median lobe, a pair of long and divergent endophallic sclerites, and parameres narrowing distad, each bearing only one short apical seta.
</paragraph>
</subSubSection>
</treatment>
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