306 lines
37 KiB
XML
306 lines
37 KiB
XML
<document id="5AEC85D51E9FC1573C447596087A26EE" ID-CLB-Dataset="27750" ID-DOI="10.11646/zootaxa.4532.2.6" ID-GBIF-Dataset="e0e97feb-5a82-4ff0-9a41-db5e3efa8c4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615311" ID-ZooBank="B3EAD977-FD62-4E46-B576-DEBA250B0AB0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788062995" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2018" docId="03CFA37A2E58FF96FF36F888FAB74931" docLanguage="en" docName="zootaxa.4532.2.6.pdf" docOrigin="Zootaxa 4532 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraneseuthia ehimensis Jałoszyński 2018, sp. n." docType="treatment" docVersion="8" lastPageNumber="270" masterDocId="FFF6DB022E5AFF92FFA1FFAFFFBE4F39" masterDocTitle="Paraneseuthia Franz of Japan: two new species and rediscovery of the enigmatic P. holzneri Franz (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="272" masterPageNumber="266" pageNumber="268" updateTime="1698715034190" updateUser="ExternalLinkService">
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<mods:mods id="2E02CCAB77ADC40EB27BD9D7E45905F5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D5FFB35A6055F440BDEC7661DFF81CD1">
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<mods:title id="FEA6D9B4018A37ED0EA0253E8437896C">Paraneseuthia Franz of Japan: two new species and rediscovery of the enigmatic P. holzneri Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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</mods:titleInfo>
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<mods:name id="908885802B0B89E2433727B7702CD533" type="personal">
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<mods:roleTerm id="DDC03ADB5EC86BD8F9804BC0D3908856">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="036857CBA6ACAB306C2A334707C04459">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="9861095E8ECB7737DF06BF447DD304E8">text</mods:typeOfResource>
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<mods:title id="6C1AD8F4BA837FF99F606B2E837EB73D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CFAA5373CB616BE2BEA077897C49E13E">
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<mods:date id="786D4E8C598953A85DCC05E1ACA43E1B">2018</mods:date>
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<mods:detail id="DCC6D2D914BDEA967669EA49AF659F39" type="pubDate">
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<mods:number id="EF005A36AB81794A7A87D22EBCD72C74">2018-12-18</mods:number>
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<mods:detail id="B27A36DD7D2843F26CED835448458D55" type="volume">
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<mods:number id="A179F0819883254E56E2FFCCF94B5E84">4532</mods:number>
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</mods:detail>
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<mods:detail id="BF54DAD8C3D1D597C7D4665C8F596BE0" type="issue">
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<mods:number id="3DDD7330681248C58A6D1517EEEF1FC6">2</mods:number>
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<mods:classification id="9C1A01FE99D7E260A537D0533F7CDB34">journal article</mods:classification>
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<mods:identifier id="3CFFF369D1B918B860736EB070A7627F" type="CLB-Dataset">27750</mods:identifier>
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<mods:identifier id="3E008744C47B202A32E6C55E80B22628" type="DOI">10.11646/zootaxa.4532.2.6</mods:identifier>
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</mods:mods>
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<treatment id="03CFA37A2E58FF96FF36F888FAB74931" ID-DOI="http://doi.org/10.5281/zenodo.5950822" ID-GBIF-Taxon="154521294" ID-Zenodo-Dep="5950822" LSID="urn:lsid:plazi:treatment:03CFA37A2E58FF96FF36F888FAB74931" httpUri="http://treatment.plazi.org/id/03CFA37A2E58FF96FF36F888FAB74931" lastPageId="4" lastPageNumber="270" pageId="2" pageNumber="268">
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<subSubSection id="C37C41E72E58FF90FF36F888FDAE4878" box="[151,528,1831,1857]" pageId="2" pageNumber="268" type="nomenclature">
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<paragraph id="8BD9126C2E58FF90FF36F888FDAE4878" blockId="2.[151,528,1831,1891]" box="[151,528,1831,1857]" pageId="2" pageNumber="268">
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<heading id="D091A5002E58FF90FF36F888FDAE4878" bold="true" box="[151,528,1831,1857]" fontSize="11" level="1" pageId="2" pageNumber="268" reason="1">
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<emphasis id="B912CE7E2E58FF90FF36F888FDAE4878" bold="true" box="[151,528,1831,1857]" pageId="2" pageNumber="268">
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<taxonomicName id="4C6669EF2E58FF90FF36F888FE784878" ID-CoL="8Y7T4" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[151,454,1831,1857]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="species" species="ehimensis" status="sp. nov.">
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<emphasis id="B912CE7E2E58FF90FF36F888FE784878" bold="true" box="[151,454,1831,1857]" italics="true" pageId="2" pageNumber="268">Paraneseuthia ehimensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22173052E58FF90FE6FF888FDAE4878" box="[462,528,1831,1857]" pageId="2" pageNumber="268" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37C41E72E58FF90FF36F8E5FEC4485A" box="[151,378,1866,1891]" pageId="2" pageNumber="268" type="description">
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<paragraph id="8BD9126C2E58FF90FF36F8E5FEC4485A" blockId="2.[151,528,1831,1891]" box="[151,378,1866,1891]" pageId="2" pageNumber="268">
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(
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<figureCitation id="135D0EE92E58FF90FF3EF8E5FF59485A" box="[159,231,1866,1891]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[159,1424,864,1700]" captionTargetId="figure@2.[151,1436,844,1715]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of males. Paraneseuthia ishigakiana sp. n. (1); P. ehimensis sp. n. (2), P. holzneri (Franz) (3)." httpUri="https://zenodo.org/record/2615313/files/figure.png" pageId="2" pageNumber="268">Figs 2</figureCitation>
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,
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<figureCitation id="135D0EE92E58FF90FF52F8E5FE9E485A" box="[243,288,1866,1891]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1928,1951]" captionTargetBox="[161,1436,445,1888]" captionTargetId="figure@3.[151,1436,445,1907]" captionTargetPageId="3" captionText="FIGURES 4–9. Aedeagus in ventral view (4–7, 9) and endophallic sclerites in ventral view (8). Paraneseuthia ishigakiana sp. n. (4); P. ehimensis sp. n. (variability among studied specimens, 5–7), P. paradoxa (K. Sawada) (8), P. holzneri (Franz) (9)." httpUri="https://zenodo.org/record/2615315/files/figure.png" pageId="2" pageNumber="268">5–7</figureCitation>
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,
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<figureCitation id="135D0EE92E58FF90FE8AF8E5FECC485A" box="[299,370,1866,1891]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1743,1766]" captionTargetBox="[166,1418,426,1712]" captionTargetId="figure@5.[151,1436,411,1722]" captionTargetPageId="5" captionText="FIGURES 10–17. Aedeagus in ventral (10, 12, 14, 16) and lateral (11, 13, 15, 17) views. Paraneseuthia ishigakiana sp. n. (10–11); P. ehimensis sp. n. (extreme forms of variability among studied specimens, 12–15), P. holzneri (Franz) (16–17)." httpUri="https://zenodo.org/record/2615317/files/figure.png" pageId="2" pageNumber="268">12–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C41E72E58FF91FF36F83DFED54FED" lastPageId="3" lastPageNumber="269" pageId="2" pageNumber="268" type="materials_examined">
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<paragraph id="8BD9126C2E58FF90FF36F83DFA2248CA" blockId="2.[151,1436,1938,2035]" pageId="2" pageNumber="268">
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<emphasis id="B912CE7E2E58FF90FF36F83DFCBD4892" bold="true" box="[151,771,1938,1963]" pageId="2" pageNumber="268">
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<collectingCounty id="62B86AE02E58FF90FF36F83DFF6C4892" box="[151,210,1938,1963]" pageId="2" pageNumber="268">Type</collectingCounty>
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material.
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<materialsCitation id="3B0E18312E58FF90FEF6F83DFCBD4892" ID-GBIF-Occurrence="2118203563" box="[343,771,1938,1963]" collectingDate="1992-05-05" collectionCode="EUMJ" collectorName="M. Sakai" country="Japan" county="Pref." determinerName="P. Jaloszynski" location="Hiromi" municipality="Narukawa" pageId="2" pageNumber="268" specimenCount="1" specimenCount-male="1" stateProvince="Ehime" typeStatus="holotype">
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||
<typeStatus id="54DDACCE2E58FF90FEF6F83DFE764892" box="[343,456,1938,1963]" pageId="2" pageNumber="268" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F37152FC2E58FF90FE7BF83DFD8A4892" box="[474,564,1938,1963]" name="Japan" pageId="2" pageNumber="268">JAPAN</collectingCountry>
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||
(
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<collectingRegion id="49A2DC8E2E58FF90FDE9F83DFD144892" box="[584,682,1938,1963]" country="Japan" name="Ehime" pageId="2" pageNumber="268">EHIME</collectingRegion>
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<collectingCounty id="62B86AE02E58FF90FD16F83DFD484892" box="[695,758,1938,1963]" pageId="2" pageNumber="268">Pref.</collectingCounty>
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):
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</materialsCitation>
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||
</emphasis>
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<specimenCount id="9D60D9E52E58FF90FCB1F83CFC994893" box="[784,807,1939,1962]" pageId="2" pageNumber="268" type="male">♂</specimenCount>
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, two labels: "[
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<collectingRegion id="49A2DC8E2E58FF90FC74F83DFB9A4892" box="[981,1060,1938,1963]" country="Japan" name="Ehime" pageId="2" pageNumber="268">Ehime</collectingRegion>
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||
:
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<collectingCounty id="62B86AE02E58FF90FBE4F83CFB364893" box="[1093,1160,1939,1962]" pageId="2" pageNumber="268">Japan</collectingCounty>
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] /
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<collectingMunicipality id="6BBD88162E58FF90FB11F83DFA984892" box="[1200,1318,1938,1963]" pageId="2" pageNumber="268">Narukawa</collectingMunicipality>
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||
,
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||
<location id="8EB944B72E58FF90FA96F83DFA364892" LSID="urn:lsid:plazi:treatment:03CFA37A2E58FF96FF36F888FAB74931:8EB944B72E58FF90FA96F83DFA364892" box="[1335,1416,1938,1963]" country="Japan" county="Pref." municipality="Narukawa" name="Hiromi" pageId="2" pageNumber="268" stateProvince="Ehime">Hiromi</location>
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/
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<date id="FFD834AC2E58FF90FF36F819FF4548F6" box="[151,251,1974,1999]" pageId="2" pageNumber="268" value="1992-05-05">
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||
<collectingDate id="EF9CCD442E58FF90FF36F819FF4548F6" box="[151,251,1974,1999]" pageId="2" pageNumber="268" value="1992-05-05">5.V.1992</collectingDate>
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||
</date>
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||
<collectorName id="269377BA2E58FF90FEA2F819FEDF48F6" box="[259,353,1974,1999]" pageId="2" pageNumber="268">M.Sakai</collectorName>
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||
/
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||
<emphasis id="B912CE7E2E58FF90FED9F818FE4048F6" box="[376,510,1974,1999]" italics="true" pageId="2" pageNumber="268">in leaf litter</emphasis>
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" [white, printed]; "
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<taxonomicName id="4C6669EF2E58FF90FD43F818FC0648F7" authorityName="Franz" authorityYear="1986" box="[738,952,1975,1998]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="genus">
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<emphasis id="B912CE7E2E58FF90FD43F818FC0648F7" box="[738,952,1975,1998]" italics="true" pageId="2" pageNumber="268">PARANESEUTHIA</emphasis>
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</taxonomicName>
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||
/
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||
<taxonomicName id="4C6669EF2E58FF90FC6EF819FB8348F6" authorityName="Jałoszyński" authorityYear="2018" box="[975,1085,1974,1999]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="species" species="ehimensis">
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<emphasis id="B912CE7E2E58FF90FC6EF819FB8348F6" box="[975,1085,1974,1999]" italics="true" pageId="2" pageNumber="268">ehimensis</emphasis>
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</taxonomicName>
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m. / det.
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<determinerName id="7E05FEA82E58FF90FB0BF818FAF748F6" box="[1194,1353,1974,1999]" pageId="2" pageNumber="268">P. Jałoszyński</determinerName>
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, 2018 /
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<typeStatus id="54DDACCE2E58FF90FF36F874FEFF48CA" box="[151,321,2011,2035]" pageId="2" pageNumber="268" type="holotype">HOLOTYPUS</typeStatus>
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" [red, printed] (EUMJ).
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<materialsCitation id="3B0E18312E58FF90FDF2F875FA2248CA" ID-GBIF-Occurrence="2118197157" box="[595,1436,2010,2035]" country="Japan" county="Pref." location="Ehime" pageId="2" pageNumber="268" specimenCount="2" specimenCount-male="2" stateProvince="Ehime" typeStatus="paratype">
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<emphasis id="B912CE7E2E58FF90FDF2F875FD6D48CA" bold="true" box="[595,723,2010,2035]" pageId="2" pageNumber="268">
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<typeStatus id="54DDACCE2E58FF90FDF2F875FD7148CA" box="[595,719,2010,2035]" pageId="2" pageNumber="268" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D60D9E52E58FF90FD7BF874FCA548CB" box="[730,795,2011,2035]" pageId="2" pageNumber="268" type="male">2 ♂♂</specimenCount>
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, "(
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<collectingRegion id="49A2DC8E2E58FF90FC96F875FC3A48CA" box="[823,900,2010,2035]" country="Japan" name="Ehime" pageId="2" pageNumber="268">Ehime</collectingRegion>
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: Japan) / Namakusadani / ODAMIYAMA / 15.
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8BD9126C2E59FF91FF36FF37FED54FED" blockId="3.[151,1437,152,392]" pageId="3" pageNumber="269">
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X. 1994 /
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<materialsCitation id="3B0E18312E59FF91FEA7FF37FED94FED" ID-GBIF-Occurrence="2118191779" collectingDate="1972-09-23" collectionCode="EUMJ" collectorName="M. Sakai & S. Kinoshita" location="Ehime" pageId="3" pageNumber="269" specimenCount="1" specimenCount-male="1" stateProvince="Ehime" typeStatus="paratype">
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||
<collectorName id="269377BA2E59FF91FEA7FF37FED54F89" box="[262,363,152,176]" pageId="3" pageNumber="269">M. Sakai</collectorName>
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||
leg.";
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||
<specimenCount id="9D60D9E52E59FF91FE14FF37FE5E4F96" box="[437,480,152,176]" pageId="3" pageNumber="269" type="male">1 ♂</specimenCount>
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||
, [SHIKOKU] / Ooshima I. /
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||
<collectingRegion id="49A2DC8E2E59FF91FC82FF37FCD24F89" box="[803,876,152,176]" country="Japan" name="Ehime" pageId="3" pageNumber="269">Ehime</collectingRegion>
|
||
Pref. / 22~
|
||
<date id="FFD834AC2E59FF91FC4AFF37FBC84F89" box="[1003,1142,152,176]" pageId="3" pageNumber="269" value="1972-09-23">
|
||
<collectingDate id="EF9CCD442E59FF91FC4AFF37FBC84F89" box="[1003,1142,152,176]" pageId="3" pageNumber="269" value="1972-09-23">23. IX. 1972</collectingDate>
|
||
</date>
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||
/
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||
<collectorName id="269377BA2E59FF91FB2AFF37FAA74F89" box="[1163,1305,152,176]" pageId="3" pageNumber="269">S. Kinoshita</collectorName>
|
||
" (
|
||
<typeStatus id="54DDACCE2E59FF91FA8EFF37FA224F89" box="[1327,1436,152,176]" pageId="3" pageNumber="269" type="paratype">paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="ED778AA92E59FF91FF15FF12FF434FED" box="[180,253,189,212]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="3" pageNumber="269">EUMJ</collectionCode>
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||
and cPJ)
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||
</materialsCitation>
|
||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41E72E59FF91FF66FF70FC744E79" pageId="3" pageNumber="269" type="diagnosis">
|
||
<paragraph id="8BD9126C2E59FF91FF66FF70FC744E79" blockId="3.[151,1437,152,392]" pageId="3" pageNumber="269">
|
||
<emphasis id="B912CE7E2E59FF91FF66FF70FEFF4FC1" bold="true" box="[199,321,223,248]" pageId="3" pageNumber="269">Diagnosis.</emphasis>
|
||
Body small (BL ±
|
||
<quantity id="4C9EBF892E59FF91FDBCFF4FFDDD4FC1" box="[541,611,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="269" unit="mm" value="1.0">1 mm</quantity>
|
||
); protibia in male with a distinct mesal denticle at middle; aedeagus with broadly and shallowly emarginate apex of median lobe with a distinct small median subtriangular projection and a pair of short, curved sclerites protruding from ostium; parameres slender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41E72E59FF96FF66FEE3FE204B29" lastPageId="4" lastPageNumber="270" pageId="3" pageNumber="269" type="description">
|
||
<paragraph id="8BD9126C2E59FF91FF66FEE3FE1E4EB1" blockId="3.[151,1437,152,392]" pageId="3" pageNumber="269">
|
||
<emphasis id="B912CE7E2E59FF91FF66FEE3FEE74E5C" bold="true" box="[199,345,332,357]" pageId="3" pageNumber="269">Description.</emphasis>
|
||
BL
|
||
<quantity id="4C9EBF892E59FF91FE28FEE3FD954E5D" box="[393,555,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="10.3" metricValueMin="9.3" pageId="3" pageNumber="269" unit="mm" value="0.98" valueMax="1.03" valueMin="0.93">0.93–1.03 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E59FF91FDDFFEE3FD584E5D" box="[638,742,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="269" unit="mm" value="0.98">0.98 mm</quantity>
|
||
). Body of male (
|
||
<figureCitation id="135D0EE92E59FF91FC3EFEE3FC5C4E5D" box="[927,994,332,357]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[159,1424,864,1700]" captionTargetId="figure@2.[151,1436,844,1715]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of males. Paraneseuthia ishigakiana sp. n. (1); P. ehimensis sp. n. (2), P. holzneri (Franz) (3)." httpUri="https://zenodo.org/record/2615313/files/figure.png" pageId="3" pageNumber="269">Fig. 2</figureCitation>
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||
) elongate and moderately convex, light brown, setae yellowish.
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</paragraph>
|
||
<caption id="DF1942E42E59FF91FF36F827FAC64884" httpUri="https://zenodo.org/record/2615315/files/figure.png" pageId="3" pageNumber="269" startId="3.[151,264,1928,1951]" targetBox="[161,1436,445,1888]" targetPageId="3">
|
||
<paragraph id="8BD9126C2E59FF91FF36F827FAC64884" blockId="3.[151,1436,1928,1981]" pageId="3" pageNumber="269">
|
||
<emphasis id="B912CE7E2E59FF91FF36F827FE8448A4" bold="true" box="[151,314,1928,1951]" pageId="3" pageNumber="269">FIGURES 4–9.</emphasis>
|
||
Aedeagus in ventral view (4–7, 9) and endophallic sclerites in ventral view (8).
|
||
<taxonomicName id="4C6669EF2E59FF91FBC6F827FAC948A7" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[1127,1399,1928,1950]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="ishigakiana" status="sp. nov.">
|
||
<emphasis id="B912CE7E2E59FF91FBC6F827FAC948A7" box="[1127,1399,1928,1950]" italics="true" pageId="3" pageNumber="269">Paraneseuthia ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912CE7E2E59FF91FADCF826FF154885" bold="true" pageId="3" pageNumber="269">
|
||
<taxonomicNameLabel id="A22173052E59FF91FADCF826FF154885" pageId="3" pageNumber="269" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(4);
|
||
<taxonomicName id="4C6669EF2E59FF91FF7DF807FEE74884" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[220,345,1959,1981]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="ehimensis" status="sp. nov.">
|
||
<emphasis id="B912CE7E2E59FF91FF7DF807FEE74884" box="[220,345,1959,1981]" italics="true" pageId="3" pageNumber="269">P. ehimensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912CE7E2E59FF91FEFEF808FE264885" bold="true" box="[351,408,1959,1980]" pageId="3" pageNumber="269">
|
||
<taxonomicNameLabel id="A22173052E59FF91FEFEF808FE264885" box="[351,408,1959,1980]" pageId="3" pageNumber="269" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(variability among studied specimens, 5–7),
|
||
<taxonomicName id="4C6669EF2E59FF91FCC0F807FBDE4884" authority="(K. Sawada)" authorityName="K. Sawada" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[865,1120,1959,1981]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="paradoxa">
|
||
<emphasis id="B912CE7E2E59FF91FCC0F807FC654884" box="[865,987,1959,1981]" italics="true" pageId="3" pageNumber="269">P. paradoxa</emphasis>
|
||
(K. Sawada)
|
||
</taxonomicName>
|
||
(8),
|
||
<taxonomicName id="4C6669EF2E59FF91FB31F807FAF34884" authority="(Franz)" authorityName="Franz" baseAuthorityName="Franz" baseAuthorityYear="1976" box="[1168,1357,1959,1981]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="holzneri">
|
||
<emphasis id="B912CE7E2E59FF91FB31F807FB424884" box="[1168,1276,1959,1981]" italics="true" pageId="3" pageNumber="269">P. holzneri</emphasis>
|
||
(Franz)
|
||
</taxonomicName>
|
||
(9).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FF38FD854E5C" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="270">
|
||
Head broadest at large, strongly convex and coarsely faceted eyes, HL
|
||
<quantity id="4C9EBF892E5EFF96FC55FF37FB264F96" box="[1012,1176,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="4" pageNumber="270" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FB4EFF37FAE74F89" box="[1263,1369,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="270" unit="mm" value="0.13">0.13 mm</quantity>
|
||
), HW
|
||
<quantity id="4C9EBF892E5EFF96FF36FF13FE824FED" box="[151,316,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="4" pageNumber="270" unit="mm" value="0.235" valueMax="0.24" valueMin="0.23">0.23–0.24 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FE35FF13FE404FED" box="[404,510,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="270" unit="mm" value="0.23">0.23 mm</quantity>
|
||
); vertex and frons confluent and evenly, weakly convex; supraantennal tubercles small but distinct. Punctures on head dorsum inconspicuous; setae short, sparse and suberect. Antennae slender, with indistinctly delimited trimerous club, AnL
|
||
<quantity id="4C9EBF892E5EFF96FD16FEABFCE54E25" box="[695,859,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="4" pageNumber="270" unit="mm" value="0.45499999999999996" valueMax="0.48" valueMin="0.43">0.43–0.48 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FC10FEABFBA54E25" box="[945,1051,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="270" unit="mm" value="0.46">0.46 mm</quantity>
|
||
), antennomeres I–II elongate, III– VIII each indistinctly transverse, IX–X each distinctly transverse, XI broader than X, about as long as IX–X together, about 1.7× as long as broad.
|
||
</paragraph>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FEDFFB004D04" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="270">
|
||
Pronotum subquadrate in dorsal view, with anterior and lateral margins rounded and posterior margin nearly straight, broadest near anterior third (in
|
||
<figureCitation id="135D0EE92E5EFF96FDFFFE3BFD1E4E95" box="[606,672,404,429]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[159,1424,864,1700]" captionTargetId="figure@2.[151,1436,844,1715]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of males. Paraneseuthia ishigakiana sp. n. (1); P. ehimensis sp. n. (2), P. holzneri (Franz) (3)." httpUri="https://zenodo.org/record/2615313/files/figure.png" pageId="4" pageNumber="270">Fig. 2</figureCitation>
|
||
pronotum is slightly tilted anteroventrad and its shape differs from that seen in strictly dorsal view), sides weakly narrowing toward obtuse-angled and blunt posterior corners; PL
|
||
<quantity id="4C9EBF892E5EFF96FF36FE73FE844ECD" box="[151,314,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="4" pageNumber="270" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28–0.30 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FE2FFE73FE484ECD" box="[398,502,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="270" unit="mm" value="0.28">0.28 mm</quantity>
|
||
), PW
|
||
<quantity id="4C9EBF892E5EFF96FD9AFE73FD604ECD" box="[571,734,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="4" pageNumber="270" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.33–0.35 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FC93FE73FC244ECD" box="[818,922,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="270" unit="mm" value="0.33">0.33 mm</quantity>
|
||
); pronotal base with two lateral pairs of small pits and a short median variously distinct (from moderately distinct to barely discernible) transverse impression. Punctures on pronotal disc fine and inconspicuous; setae moderately long, sparse and suberect.
|
||
</paragraph>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FDE8FCC64DC9" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="270">
|
||
Elytra oval, broadest between middle and anterior third and moderately strongly narrowing posterad, evenly convex; EL
|
||
<quantity id="4C9EBF892E5EFF96FE87FDC3FE724DBD" box="[294,460,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="6.0" metricValueMin="5.3" pageId="4" pageNumber="270" unit="mm" value="0.565" valueMax="0.6" valueMin="0.53">0.53–0.60 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FD86FDC3FD2C4DBD" box="[551,658,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="4" pageNumber="270" unit="mm" value="0.57">0.57 mm</quantity>
|
||
), EW
|
||
<quantity id="4C9EBF892E5EFF96FD7FFDC3FC3A4DBD" box="[734,900,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="4" pageNumber="270" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
|
||
(mean
|
||
<quantity id="4C9EBF892E5EFF96FC7FFDC3FBF74DBD" box="[990,1097,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="270" unit="mm" value="0.43">0.43 mm</quantity>
|
||
), EI 1.29–1.41; humeral calli distinct but small, basal impressions indistinct; apices separately rounded. Punctures on elytra distinct but small and diffuse, dense, in anteromedian portion of elytra separated by spaces subequal to diameters of punctures; setae long, moderately dense and suberect. Hind wings well developed.
|
||
</paragraph>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FD53FAA84C2C" blockId="4.[151,1437,151,1545]" box="[199,1302,764,789]" pageId="4" pageNumber="270">Metaventrite with large, elongate but shallow and diffuse impression occupying most of its length.</paragraph>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FC8FFA964C01" blockId="4.[151,1437,151,1545]" box="[199,1320,799,824]" pageId="4" pageNumber="270">Legs moderately long and slender; protibiae distinctly bent at middle, with a distinct mesal denticle.</paragraph>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FCEBFC5E4CD4" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="270">
|
||
Aedeagus (
|
||
<figureCitation id="135D0EE92E5EFF96FEEBFCEBFE084C64" box="[330,438,836,861]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1928,1951]" captionTargetBox="[161,1436,445,1888]" captionTargetId="figure@3.[151,1436,445,1907]" captionTargetPageId="3" captionText="FIGURES 4–9. Aedeagus in ventral view (4–7, 9) and endophallic sclerites in ventral view (8). Paraneseuthia ishigakiana sp. n. (4); P. ehimensis sp. n. (variability among studied specimens, 5–7), P. paradoxa (K. Sawada) (8), P. holzneri (Franz) (9)." httpUri="https://zenodo.org/record/2615315/files/figure.png" pageId="4" pageNumber="270">Figs 5–7</figureCitation>
|
||
,
|
||
<figureCitation id="135D0EE92E5EFF96FE69FCEBFDB14C64" box="[456,527,836,861]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1743,1766]" captionTargetBox="[166,1418,426,1712]" captionTargetId="figure@5.[151,1436,411,1722]" captionTargetPageId="5" captionText="FIGURES 10–17. Aedeagus in ventral (10, 12, 14, 16) and lateral (11, 13, 15, 17) views. Paraneseuthia ishigakiana sp. n. (10–11); P. ehimensis sp. n. (extreme forms of variability among studied specimens, 12–15), P. holzneri (Franz) (16–17)." httpUri="https://zenodo.org/record/2615317/files/figure.png" pageId="4" pageNumber="270">12–15</figureCitation>
|
||
) moderately stout, AeL
|
||
<quantity id="4C9EBF892E5EFF96FC9AFCEBFC164C65" box="[827,936,836,860]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="270" unit="mm" value="0.23">0.23 mm</quantity>
|
||
; median lobe in ventral view broadest in subapical region of basal capsule, apical portion of median lobe with rounded sides, and a shallow and broad apical emargination bearing a small median subtriangular projection; endophallus with a pair of moderately long and weakly curved lateral projections protruding from ostium and not overlapping at middle; parameres slender, unmodified, each with one apical and one subapical seta of unequal length.
|
||
</paragraph>
|
||
<paragraph id="8BD9126C2E5EFF96FF66FC58FE204B29" blockId="4.[151,1437,151,1545]" box="[199,414,1015,1040]" pageId="4" pageNumber="270">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41E72E5EFF96FF66FBB3FE724B0C" box="[199,460,1052,1077]" pageId="4" pageNumber="270" type="distribution">
|
||
<paragraph id="8BD9126C2E5EFF96FF66FBB3FE724B0C" blockId="4.[151,1437,151,1545]" box="[199,460,1052,1077]" pageId="4" pageNumber="270">
|
||
<emphasis id="B912CE7E2E5EFF96FF66FBB3FEDE4B0C" bold="true" box="[199,352,1052,1077]" pageId="4" pageNumber="270">Distribution.</emphasis>
|
||
Shikoku.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41E72E5EFF96FF66FB90FD534B61" box="[199,749,1087,1112]" pageId="4" pageNumber="270" type="etymology">
|
||
<paragraph id="8BD9126C2E5EFF96FF66FB90FD534B61" blockId="4.[151,1437,151,1545]" box="[199,749,1087,1112]" pageId="4" pageNumber="270">
|
||
<emphasis id="B912CE7E2E5EFF96FF66FB90FEF04B61" bold="true" box="[199,334,1087,1112]" pageId="4" pageNumber="270">Etymology.</emphasis>
|
||
Locotypical, after
|
||
<collectingRegion id="49A2DC8E2E5EFF96FD83FB90FD544B61" box="[546,746,1087,1112]" country="Japan" name="Ehime" pageId="4" pageNumber="270">Ehime Prefecture</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41E72E5EFF96FF66FBCBFAB74931" pageId="4" pageNumber="270" type="discussion">
|
||
<paragraph id="8BD9126C2E5EFF96FF66FBCBFAB74931" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="270">
|
||
<emphasis id="B912CE7E2E5EFF96FF66FBCBFE854B44" bold="true" box="[199,315,1124,1149]" pageId="4" pageNumber="270">Remarks.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C6669EF2E5EFF96FE0AFBCBFD7C4B44" authorityName="Jałoszyński" authorityYear="2018" box="[427,706,1124,1149]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="ehimensis">
|
||
<emphasis id="B912CE7E2E5EFF96FE0AFBCBFD7C4B44" box="[427,706,1124,1149]" italics="true" pageId="4" pageNumber="270">Paraneseuthia ehimensis</emphasis>
|
||
</taxonomicName>
|
||
are externally indistinguishable from those of
|
||
<taxonomicName id="4C6669EF2E5EFF96FB6CFBCAFAEA4B44" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[1229,1364,1124,1149]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="paradoxa">
|
||
<emphasis id="B912CE7E2E5EFF96FB6CFBCAFAEA4B44" box="[1229,1364,1124,1149]" italics="true" pageId="4" pageNumber="270">P. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
; these are the only Japanese species with the aedeagal apical margin not bifurcate, but weakly and broadly emarginate and bearing a distinct median subtriangular projection. As all Japanese congeners,
|
||
<taxonomicName id="4C6669EF2E5EFF96FBB6FB02FB1C4BFC" authorityName="Jałoszyński" authorityYear="2018" box="[1047,1186,1196,1221]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="ehimensis">
|
||
<emphasis id="B912CE7E2E5EFF96FBB6FB02FB1C4BFC" box="[1047,1186,1196,1221]" italics="true" pageId="4" pageNumber="270">P. ehimensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6669EF2E5EFF96FB7CFB02FAD94BFC" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[1245,1383,1196,1221]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="paradoxa">
|
||
<emphasis id="B912CE7E2E5EFF96FB7CFB02FAD94BFC" box="[1245,1383,1196,1221]" italics="true" pageId="4" pageNumber="270">P. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
also have a pair of elongate endophallic sclerites, which are clearly different in each species. In the aedeagus of
|
||
<taxonomicName id="4C6669EF2E5EFF96FA26FB7EFEBC4A34" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="paradoxa">
|
||
<emphasis id="B912CE7E2E5EFF96FA26FB7EFEBC4A34" italics="true" pageId="4" pageNumber="270">P. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
, the endophallic sclerites are conspicuously long and strongly curved mesad, so that their distal halves overlap (
|
||
<figureCitation id="135D0EE92E5EFF96FF5AFAB8FEFE4A09" box="[251,320,1303,1328]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1928,1951]" captionTargetBox="[161,1436,445,1888]" captionTargetId="figure@3.[151,1436,445,1907]" captionTargetPageId="3" captionText="FIGURES 4–9. Aedeagus in ventral view (4–7, 9) and endophallic sclerites in ventral view (8). Paraneseuthia ishigakiana sp. n. (4); P. ehimensis sp. n. (variability among studied specimens, 5–7), P. paradoxa (K. Sawada) (8), P. holzneri (Franz) (9)." httpUri="https://zenodo.org/record/2615315/files/figure.png" pageId="4" pageNumber="270">Fig. 8</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C6669EF2E5EFF96FED7FAB6FDBF4A09" authorityName="Jałoszyński" authorityYear="2018" box="[374,513,1303,1328]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="ehimensis">
|
||
<emphasis id="B912CE7E2E5EFF96FED7FAB6FDBF4A09" box="[374,513,1303,1328]" italics="true" pageId="4" pageNumber="270">P. ehimensis</emphasis>
|
||
</taxonomicName>
|
||
, the endophallic sclerites are much shorter and only weakly curved. The sclerites show some variability in shape (
|
||
<figureCitation id="135D0EE92E5EFF96FDA0FA93FDD64A6C" box="[513,616,1340,1365]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1928,1951]" captionTargetBox="[161,1436,445,1888]" captionTargetId="figure@3.[151,1436,445,1907]" captionTargetPageId="3" captionText="FIGURES 4–9. Aedeagus in ventral view (4–7, 9) and endophallic sclerites in ventral view (8). Paraneseuthia ishigakiana sp. n. (4); P. ehimensis sp. n. (variability among studied specimens, 5–7), P. paradoxa (K. Sawada) (8), P. holzneri (Franz) (9)." httpUri="https://zenodo.org/record/2615315/files/figure.png" pageId="4" pageNumber="270">Figs 5–7</figureCitation>
|
||
). Endophallic sclerites are extricable structures; in a fully erect state they are entirely exposed and projecting from the aedeagal apex, the erection also requires the sclerites to rotate (e.g.,
|
||
<figureCitation id="135D0EE92E5EFF96FF36FA2BFF524AA5" box="[151,236,1412,1437]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[159,1424,864,1700]" captionTargetId="figure@2.[151,1436,844,1715]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of males. Paraneseuthia ishigakiana sp. n. (1); P. ehimensis sp. n. (2), P. holzneri (Franz) (3)." httpUri="https://zenodo.org/record/2615313/files/figure.png" pageId="4" pageNumber="270">Fig. 3G</figureCitation>
|
||
in
|
||
<bibRefCitation id="EFF76F9D2E5EFF96FEAEFA2BFDD34AA5" author="Jaloszynski, P. & Hoshina, H." box="[271,621,1412,1437]" pageId="4" pageNumber="270" pagination="133 - 143" refId="ref3305" refString="Jaloszynski, P. & Hoshina, H. (2004) Revision of Japanese species of Paraneseuthia Franz (Coleoptera, Scydmaenidae). Japanese Journal of Systematic Entomology, 10, 133 - 143." type="journal article" year="2004">Jałoszyński & Hoshina (2004))</bibRefCitation>
|
||
. Consequently, some of the variability observed in the studied specimens is caused not by true differences in shape, but by the rotation of the sclerites. However, in all studied specimens of
|
||
<taxonomicName id="4C6669EF2E5EFF96FF36FA62FE9A4ADC" authorityName="Jałoszyński" authorityYear="2018" box="[151,292,1484,1509]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="ehimensis">
|
||
<emphasis id="B912CE7E2E5EFF96FF36FA62FE9A4ADC" box="[151,292,1484,1509]" italics="true" pageId="4" pageNumber="270">P. ehimensis</emphasis>
|
||
</taxonomicName>
|
||
the sclerites are too short to overlap at middle, whereas those in
|
||
<taxonomicName id="4C6669EF2E5EFF96FB8CFA62FB064ADC" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[1069,1208,1484,1509]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="paradoxa">
|
||
<emphasis id="B912CE7E2E5EFF96FB8CFA62FB064ADC" box="[1069,1208,1484,1509]" italics="true" pageId="4" pageNumber="270">P. paradoxa</emphasis>
|
||
</taxonomicName>
|
||
are so long and so strongly curved that they cannot occupy the subapical region of the median lobe without overlapping.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |