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<document id="E5468479342200ED2547A00F036AA98A" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. &amp; Micó, E." docDate="2015" docId="03CFA3486B6EFFCAFF08FD66580CFE9B" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala mersa Filippini, Galante, and Mico, new species" docType="treatment" docVersion="4" lastPageNumber="471" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="471" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="26D48441FC3881AE543260C1E2EABAC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07B61AA469E22F674669969B0E753B1D">
<mods:title id="704F2A149F9B38F7C90F907C11173B4B">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="3FED94E4025372FE1940FC65DF1F097C" type="personal">
<mods:role id="651C5B749A04DE840074EDF557F61AB7">
<mods:roleTerm id="15C66789A476FC693322F596D7BFC46D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AA95F31AE758061BB57AE5AE0365DDB">Filippini, V.</mods:namePart>
</mods:name>
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<mods:role id="B291F2D1691E63A72CA42DA107C7AA2D">
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</mods:role>
<mods:namePart id="FF6CB21D460B122AA8768E307ADD795E">Galante, E.</mods:namePart>
</mods:name>
<mods:name id="EB983C5689CB7019BE00ED19B7BC8380" type="personal">
<mods:role id="ABB6C005209A7BA1D3B6DB22DDFCA3A5">
<mods:roleTerm id="0C6A416FC2ED72CA228482D3562B480C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93E3ADD7A4AFEF4435177A15520D55BE">Micó, E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0853A2396C8CE16C0272DB48BAECEFA9">text</mods:typeOfResource>
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<mods:titleInfo id="E30AB049B6022F05B117A820DAE2D8A5">
<mods:title id="4DD6DDCE27413C3275DB891DF4F848E5">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="5031821B817C01686A35BF1073DBB958">
<mods:date id="5CD977E4D7CA52378C2DA799D84A77E6">2015</mods:date>
<mods:detail id="5A44A8CA28481BC97FC2FC97468AE07C" type="pubDate">
<mods:number id="EC50747E94BB9A38A23FD59743B3B88A">2015-09-30</mods:number>
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<mods:detail id="E1280EA819595042B58C88CB0B0D77B5" type="volume">
<mods:number id="9D594C6C1299D6DE7D8457FBC7753FCD">69</mods:number>
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<mods:detail id="2167DB9D6B131D4A0BC22B3AC8EEBD4F" type="issue">
<mods:number id="895B47099AC85D7AE07B5846E2E248F7">3</mods:number>
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<mods:url id="32798B7BA9114FA57E3BD738EC86C3A8">http://dx.doi.org/10.1649/0010-065x-69.3.463</mods:url>
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<mods:classification id="558F11A0BEDDE2744414478EE72C406C">journal article</mods:classification>
<mods:identifier id="666357ECCC242FFD55A19BC751842B6F" type="DOI">10.1649/0010-065x-69.3.463</mods:identifier>
<mods:identifier id="D7E35A11EB1432CE4B56E1E8F93B6A00" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="670906A8D1197D1B58B2F1BD69E34243" type="Zenodo-Dep">10106050</mods:identifier>
<mods:identifier id="D11F096A6BB0B38FB3B295FCED28203F" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
</mods:mods>
<treatment id="03CFA3486B6EFFCAFF08FD66580CFE9B" ID-DOI="http://doi.org/10.5281/zenodo.5413171" ID-Zenodo-Dep="5413171" LSID="urn:lsid:plazi:treatment:03CFA3486B6EFFCAFF08FD66580CFE9B" httpUri="http://treatment.plazi.org/id/03CFA3486B6EFFCAFF08FD66580CFE9B" lastPageId="9" lastPageNumber="471" pageId="8" pageNumber="471">
<subSubSection id="C37C41D56B6EFFCBFF08FD665BA9FD07" pageId="8" pageNumber="471" type="nomenclature">
<paragraph id="8BD9125E6B6EFFCBFF08FD665BA9FD07" blockId="8.[182,650,655,730]" pageId="8" pageNumber="471">
<heading id="D091A5326B6EFFCBFF08FD665BA9FD07" centered="true" fontSize="10" level="2" pageId="8" pageNumber="471" reason="2">
<emphasis id="B912CE4C6B6EFFCBFF08FD665BA9FD07" bold="true" pageId="8" pageNumber="471">
<taxonomicName id="4C6669DD6B6EFFCBFF08FD6658F6FD62" authority="Filippini, Galante, and Mico" authorityName="Filippini, Galante, and Mico" box="[182,645,655,677]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="mersa" status="sp. nov.">
<emphasis id="B912CE4C6B6EFFCBFF08FD665B21FD62" bold="true" box="[182,338,655,677]" italics="true" pageId="8" pageNumber="471">Anomala mersa</emphasis>
Filippini, Galante, and Micó
</taxonomicName>
,
<taxonomicNameLabel id="A22173376B6EFFCBFEDBFD425BA9FD07" box="[357,474,683,704]" pageId="8" pageNumber="471" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFE94FD2C5866FD1D" box="[298,533,709,730]" pageId="8" pageNumber="471" type="description">
<paragraph id="8BD9125E6B6EFFCBFE94FD2C5866FD1D" blockId="8.[182,650,655,730]" box="[298,533,709,730]" pageId="8" pageNumber="471">
(
<figureCitation id="135D0EDB6B6EFFCBFE8FFD2C5B0BFD1D" box="[305,376,709,730]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="8" pageNumber="471">Figs. 4</figureCitation>
,
<figureCitation id="135D0EDB6B6EFFCBFE3AFD2C5BEDFD1D" box="[388,414,709,730]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="8" pageNumber="471">12</figureCitation>
,
<figureCitation id="135D0EDB6B6EFFCBFE14FD2C5BB7FD1D" box="[426,452,709,730]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="8" pageNumber="471">20</figureCitation>
,
<figureCitation id="135D0EDB6B6EFFCBFE6EFD2C5B99FD1D" box="[464,490,709,730]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="8" pageNumber="471">28</figureCitation>
,
<figureCitation id="135D0EDB6B6EFFCBFE48FD2C587CFD1D" box="[502,527,709,730]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="8" pageNumber="471">34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFF7BFD1959EDFCA8" pageId="8" pageNumber="471" type="materials_examined">
<paragraph id="8BD9125E6B6EFFCBFF7BFD195872FBEE" blockId="8.[173,660,752,1729]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFF7BFD195B2FFCC2" bold="true" box="[197,348,752,773]" pageId="8" pageNumber="471">Type Material.</emphasis>
<materialsCitation id="3B0E18036B6EFFCBFED7FD19583CFCA8" collectingDate="1999-11-29" collectingDateMax="1999-12-15" collectingDateMin="1999-11-29" collectionCode="MNCR" collectorName="P. N. Palo Verde &amp; W. Porras. Manual" country="Costa Rica" elevation="25" location="Guanacaste" pageId="8" pageNumber="471" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54DDACFC6B6EFFCBFED7FD195BBBFCC2" box="[361,456,752,773]" pageId="8" pageNumber="471" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60D9D76B6EFFCBFE6BFD195B94FCC1" box="[469,487,752,774]" pageId="8" pageNumber="471" type="male">♂</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6EFFCBFE46FD1958FCFCC2" box="[504,655,752,773]" name="Costa Rica" pageId="8" pageNumber="471">COSTA RICA</collectingCountry>
, Prov.
<collectingRegion id="49A2DCBC6B6EFFCBFF5AFCE35B22FCD8" box="[228,337,778,799]" country="Costa Rica" name="Guanacaste" pageId="8" pageNumber="471">Guanacaste</collectingRegion>
, Bagaces,
<collectorName id="269377886B6EFFCBFE08FCE3583DFCD8" box="[438,590,778,799]" pageId="8" pageNumber="471">P.N. Palo Verde</collectorName>
, Sector Palo Verde.
<quantity id="4C9EBFBB6B6EFFCBFE93FCCC5B06FCFD" box="[301,373,805,827]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="471" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">
<elevation id="004BF56D6B6EFFCBFE93FCCC5B06FCFD" box="[301,373,805,827]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="471" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50m</elevation>
</quantity>
.
<date id="FFD8349E6B6EFFCBFE3CFCCC5818FCFD" box="[386,619,805,826]" pageId="8" pageNumber="471" value="1999-11-29" valueMax="1999-12-15" valueMin="1999-11-29">
<collectingDate id="EF9CCD766B6EFFCBFE3CFCCC5818FCFD" box="[386,619,805,826]" pageId="8" pageNumber="471" value="1999-11-29" valueMax="1999-12-15" valueMin="1999-11-29">29 NOV-15 DIC 1999</collectingDate>
</date>
.
<collectorName id="269377886B6EFFCBFDC6FCCC5B34FC92" pageId="8" pageNumber="471">W. Porras. Manual</collectorName>
L_N_260952_385020 #57243 / INB0003151362&quot; deposited at
<collectionCode id="ED778A9B6B6EFFCBFDBEFCB3583CFCA8" box="[512,591,858,879]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="8" pageNumber="471">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B0E18036B6EFFCBFDE2FCB35B8EFBEE" collectingDate="1999-11-29" collectingDateMax="1999-12-15" collectingDateMin="1999-11-29" collectionCode="MNCR" collectorName="P. N. Palo Verde &amp; W. Porras. Manual" country="Costa Rica" elevation="25" location="Universidad de Alicante" pageId="8" pageNumber="471" specimenCode="CEUA00106201" specimenCount="4" specimenCount-male="4" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54DDACFC6B6EFFCBFDE2FCB35A92FC4D" pageId="8" pageNumber="471" type="paratype">Paratypes</typeStatus>
(4):
<specimenCount id="9D60D9D76B6EFFCBFEAAFC9C5B41FC4D" box="[276,306,884,906]" pageId="8" pageNumber="471" type="male">4♂</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6EFFCBFEFCFC9C5BA5FC4D" box="[322,470,885,906]" name="Costa Rica" pageId="8" pageNumber="471">COSTA RICA</collectingCountry>
, Prov.
<collectingRegion id="49A2DCBC6B6EFFCBFDA3FC9C58E3FC4D" box="[541,656,885,906]" country="Costa Rica" name="Guanacaste" pageId="8" pageNumber="471">Guanacaste</collectingRegion>
, Bagaces,
<collectorName id="269377886B6EFFCBFEBAFC665BE5FC63" box="[260,406,911,932]" pageId="8" pageNumber="471">P.N. Palo Verde</collectorName>
, Sector Palo Verde.
<quantity id="4C9EBFBB6B6EFFCBFDF3FC6658E3FC63" box="[589,656,911,933]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="471" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">
<elevation id="004BF56D6B6EFFCBFDF3FC6658E3FC63" box="[589,656,911,933]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="471" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50m</elevation>
</quantity>
.
<date id="FFD8349E6B6EFFCBFF13FC435BFDFC78" box="[173,398,938,959]" pageId="8" pageNumber="471" value="1999-11-29" valueMax="1999-12-15" valueMin="1999-11-29">
<collectingDate id="EF9CCD766B6EFFCBFF13FC435BFDFC78" box="[173,398,938,959]" pageId="8" pageNumber="471" value="1999-11-29" valueMax="1999-12-15" valueMin="1999-11-29">29 NOV-15 DIC 1999</collectingDate>
</date>
.
<collectorName id="269377886B6EFFCBFE26FC435821FC78" box="[408,594,938,959]" pageId="8" pageNumber="471">W. Porras. Manual</collectorName>
L_N_ 260952_385020 #57243&quot; /
<specimenCode id="DBC0BA256B6EFFCBFE67FC2D58E3FC1E" box="[473,656,964,985]" collectionCode="CEUA" pageId="8" pageNumber="471">CEUA00106201</specimenCode>
, INB0003151360, INB0003151358 and INB000 3151412. Three
<typeStatus id="54DDACFC6B6EFFCBFEF2FC135BD5FBC8" box="[332,422,1018,1039]" pageId="8" pageNumber="471" type="paratype">paratypes</typeStatus>
are deposited at
<collectionCode id="ED778A9B6B6EFFCBFDF3FC1358E0FBC8" box="[589,659,1018,1039]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="8" pageNumber="471">MNCR</collectionCode>
and one at
<location id="8EB944856B6EFFCBFEA8FBFD5B8EFBEE" LSID="urn:lsid:plazi:treatment:03CFA3486B6EFFCAFF08FD66580CFE9B:8EB944856B6EFFCBFEA8FBFD5B8EFBEE" box="[278,509,1044,1065]" country="Costa Rica" name="Universidad de Alicante" pageId="8" pageNumber="471" stateProvince="Guanacaste">Universidad de Alicante</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9125E6B6EFFCBFF7BFBC6590BFC92" blockId="8.[173,660,752,1729]" lastBlockId="8.[707,1194,194,1729]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFF7BFBC65B5BFB83" bold="true" box="[197,296,1071,1092]" pageId="8" pageNumber="471">
<typeStatus id="54DDACFC6B6EFFCBFF7BFBC65B57FB83" box="[197,292,1071,1092]" pageId="8" pageNumber="471" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="135D0EDB6B6EFFCBFEC7FBC65BC8FB83" box="[377,443,1071,1092]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="8" pageNumber="471">Fig. 4</figureCitation>
). Body oval. Length
<quantity id="4C9EBFBB6B6EFFCBFF13FBA05B7AFB99" box="[173,265,1097,1118]" metricMagnitude="-3" metricUnit="m" metricValue="9.31" pageId="8" pageNumber="471" unit="mm" value="9.31">9.31 mm</quantity>
, width
<quantity id="4C9EBFBB6B6EFFCBFEE9FBA05BC7FB99" box="[343,436,1097,1118]" metricMagnitude="-3" metricUnit="m" metricValue="4.73" pageId="8" pageNumber="471" unit="mm" value="4.73">4.73 mm</quantity>
. Head reddish brown. Pronotum dark brown with wide, ochre sides, with green luster. Scutellum brown. Elytra dark brown with large ochre maculae on basal part and 2 small, ochre maculae at apex. Pygidium light brown with ochre apex and sides at base. Legs ochre with reddish brown tarsus, metatibia, and apical part of pro- and mesotibiae. Venter ochre on sternum, with darker median suture, and reddish brown abdominal sternites.
<emphasis id="B912CE4C6B6EFFCBFEC0FAD15BCEFA8A" bold="true" box="[382,445,1336,1357]" pageId="8" pageNumber="471">Head:</emphasis>
Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, clypeus thin in frontal view, width/ length ratio 1.98. Frons densely punctate, convex on suture with clypeus. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/ width of eye 3.46. Ratio antennal funiculus/club 0.71.
<emphasis id="B912CE4C6B6EFFCBFF57FA1B5B2BF9C0" bold="true" box="[233,344,1522,1543]" pageId="8" pageNumber="471">Pronotum:</emphasis>
Subtrapezoidal, basal portions of lateral margins nearly parallel. Ratio width/ length 1.66. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles quadrate. Basal margin sinuate, subapical bead complete. Surface with dense punctures.
<emphasis id="B912CE4C6B6EFFCBFE2CF97B5B8CF960" bold="true" box="[402,511,1682,1703]" pageId="8" pageNumber="471">Scutellum:</emphasis>
Subtriangular, with rounded sides and acute apex. Ratio width/ length 1.24. Surface densely punctate.
<emphasis id="B912CE4C6B6EFFCBFBE3FF2B5EDAFF10" bold="true" box="[1117,1193,194,215]" pageId="8" pageNumber="471">Elytra:</emphasis>
With striae defined by regular rows of punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2
<superScript id="7C13BF166B6EFFCBFCB6FEE55969FEDD" attach="none" box="[776,794,268,282]" fontSize="6" pageId="8" pageNumber="471">nd</superScript>
–4
<superScript id="7C13BF166B6EFFCBFC8FFEE5594CFEDD" attach="left" box="[817,831,268,282]" fontSize="6" pageId="8" pageNumber="471">th</superScript>
interstices with 1 irregular, secondary stria. Surface covered with secondary punctures.
<emphasis id="B912CE4C6B6EFFCBFD7DFEAE595EFE9B" bold="true" box="[707,813,327,348]" pageId="8" pageNumber="471">Pygidium:</emphasis>
Strigate with long setae on posterior margin. Ratio width/length 1.60.
<emphasis id="B912CE4C6B6EFFCBFBA4FE8B5E17FEB0" bold="true" box="[1050,1124,354,375]" pageId="8" pageNumber="471">Venter:</emphasis>
Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 5–6 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin.
<emphasis id="B912CE4C6B6EFFCBFC0CFDE85998FDD1" bold="true" box="[946,1003,513,534]" pageId="8" pageNumber="471">Legs:</emphasis>
Protibia (
<figureCitation id="135D0EDB6B6EFFCBFBE8FDE85ED2FDD1" box="[1110,1185,513,534]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="8" pageNumber="471">Fig. 12</figureCitation>
) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, sharp. Metatibia slightly narrower subapically. Ratio length/ width 2.92. First external carina with a row of sparse setae. Surface rugose. Internal protarsal claw bifurcate, with upper branch slightly shorter and 2/3 width of lower one, inferior margin curved.
<emphasis id="B912CE4C6B6EFFCBFBA9FD525EDAFD17" bold="true" box="[1047,1193,699,720]" pageId="8" pageNumber="471">Male genitalia:</emphasis>
Parameres with blunt apex in lateral view (
<figureCitation id="135D0EDB6B6EFFCBFBE6FD3C5EEDFD2D" box="[1112,1182,725,746]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="8" pageNumber="471">Fig. 20</figureCitation>
), ventral angle acute. Endophallus (
<figureCitation id="135D0EDB6B6EFFCBFBADFD195E28FCC2" box="[1043,1115,752,773]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="8" pageNumber="471">Fig. 28</figureCitation>
) with a long sacculus covered with short setae, a patch of sclerotized spines at base. Ejaculatory duct opening on lateral inflation.
</paragraph>
<paragraph id="8BD9125E6B6EFFCBFD65FCB359EDFCA8" blockId="8.[707,1194,194,1729]" box="[731,926,858,879]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFD65FCB35958FCA8" bold="true" box="[731,811,858,879]" pageId="8" pageNumber="471">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFD65FC9C59EBFB24" pageId="8" pageNumber="471" type="description">
<paragraph id="8BD9125E6B6EFFCBFD65FC9C59EBFB24" blockId="8.[707,1194,194,1729]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFD65FC9C594CFC4D" bold="true" box="[731,831,885,906]" pageId="8" pageNumber="471">Variation.</emphasis>
Four
<typeStatus id="54DDACFC6B6EFFCBFC3CFC9C5993FC4D" box="[898,992,885,906]" pageId="8" pageNumber="471" type="paratype">paratypes</typeStatus>
. Posterior margin of pronotum dark or with two ochre median maculae, size of ochre maculae on elytra variable, pygidium varies from ochre to ochre with light brown maculae on lateral depressions to light brown; venter varies from ochre with brown bands on abdominal sternites to mainly brown. Body length
<quantity id="4C9EBFBB6B6EFFCBFB9FFBFD5ED6FBEE" box="[1057,1189,1044,1066]" metricMagnitude="-3" metricUnit="m" metricValue="9.12" metricValueMax="9.31" metricValueMin="8.93" pageId="8" pageNumber="471" unit="mm" value="9.12" valueMax="9.31" valueMin="8.93">8.93–9.31 mm</quantity>
, body width 4.73–5.00 mm. Clypeal width/length ratio 1.80–2.02. Ratio of interocular width/width of eye 3.13–3.61. Ratio of antennal funiculus/club 0.57–0.71. Pronotal width/length ratio 1.66–1.70. Scutellar width/length ratio 1.21–1.37. Pygidial width/length ratio 1.54–1.62. Metatibial length/ width ratio 2.82–3.11.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFD65FB005956FA8A" pageId="8" pageNumber="471" type="etymology">
<paragraph id="8BD9125E6B6EFFCBFD65FB005956FA8A" blockId="8.[707,1194,194,1729]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFD65FB00593EFB39" bold="true" box="[731,845,1257,1278]" pageId="8" pageNumber="471">Etymology.</emphasis>
From the past participle of the Latin verb
<emphasis id="B912CE4C6B6EFFCBFD4AFAEA595DFADE" box="[756,814,1283,1305]" italics="true" pageId="8" pageNumber="471">mergo</emphasis>
, dipped, for the black apical portion of the elytra, which appears as if it was dipped into black dye.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFD65FABA59C1FA7F" pageId="8" pageNumber="471" type="distribution">
<paragraph id="8BD9125E6B6EFFCBFD65FABA59C1FA7F" blockId="8.[707,1194,194,1729]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFD65FABA592EFAAF" bold="true" box="[731,861,1363,1384]" pageId="8" pageNumber="471">Distribution.</emphasis>
<taxonomicName id="4C6669DD6B6EFFCBFCD5FABB5E76FAAF" box="[875,1029,1362,1384]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="mersa">
<emphasis id="B912CE4C6B6EFFCBFCD5FABB5E76FAAF" box="[875,1029,1362,1384]" italics="true" pageId="8" pageNumber="471">Anomala mersa</emphasis>
</taxonomicName>
inhabits the dry forest of Parque Nacional Palo Verde on the northern side of Golfo de Nicoya,
<collectingCountry id="F37152CE6B6EFFCBFC65FA615E30FA5A" box="[987,1091,1416,1437]" name="Costa Rica" pageId="8" pageNumber="471">Costa Rica</collectingCountry>
, from
<quantity id="4C9EBFBB6B6EFFCBFB3DFA615887FA7F" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="471" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0 to 50 m</quantity>
elevation (
<figureCitation id="135D0EDB6B6EFFCBFCE1FA4A59D4FA7F" box="[863,935,1443,1464]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="8" pageNumber="471">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCAFD65FA54580CFE9B" lastPageId="9" lastPageNumber="472" pageId="8" pageNumber="471" type="diagnosis">
<paragraph id="8BD9125E6B6EFFCAFD65FA54580CFE9B" blockId="8.[707,1194,194,1729]" lastBlockId="9.[151,639,194,348]" lastPageId="9" lastPageNumber="472" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFD65FA545937FA15" bold="true" box="[731,836,1469,1490]" pageId="8" pageNumber="471">Diagnosis.</emphasis>
This species is recognized by its small size, elytra dark brown with large, ochre maculae on the basal part, and parameres with blunt apex in lateral view and ventral angle acute. The particular elytral pattern easily sets this species apart from other
<taxonomicName id="4C6669DD6B6EFFCBFCDCF9AB59C6F99F" box="[866,949,1602,1624]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE4C6B6EFFCBFCDCF9AB59C6F99F" box="[866,949,1602,1624]" italics="true" pageId="8" pageNumber="471">Anomala</emphasis>
</taxonomicName>
. Species with similar patterns are dark specimens of
<taxonomicName id="4C6669DD6B6EFFCBFC78F9B55E75F94B" authority="Filippini, Mico, and Galante, 2013" authorityName="Filippini, Mico, and Galante" authorityYear="2013" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B912CE4C6B6EFFCBFC78F9B55EDAF9B5" box="[966,1193,1628,1650]" italics="true" pageId="8" pageNumber="471">Anomala pseudoeucoma</emphasis>
Filippini, Micó, and Galante, 2013
</taxonomicName>
(
<collectingCountry id="F37152CE6B6EFFCBFBACF99E5E09F94B" box="[1042,1146,1655,1676]" name="Costa Rica" pageId="8" pageNumber="471">Costa Rica</collectingCountry>
) and
<taxonomicName id="4C6669DD6B6EFFCBFD7DF9785ED5F960" authority="Ohaus, 1905" authorityName="Ohaus" authorityYear="1905" box="[707,1190,1681,1703]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="nigrosellata">
<emphasis id="B912CE4C6B6EFFCBFD7DF97859E9F960" box="[707,922,1681,1703]" italics="true" pageId="8" pageNumber="471">Anomala nigrosellata</emphasis>
Ohaus, 1905 (
<collectingCountry id="F37152CE6B6EFFCBFB8AF97B5EEDF960" box="[1076,1182,1682,1703]" name="Venezuela" pageId="8" pageNumber="471">Venezuela</collectingCountry>
)
</taxonomicName>
, which are much larger in size. The former species has the entire surface covered in dense setae and longer parameres with an acute apex and sinuate ventral margin. The latter species has a squared, dark macula on the elytra, hence the lateral elytral margins are ochre, coarser punctures on the elytra, and narrow parameres with a sinuate ventral margin.
</paragraph>
</subSubSection>
</treatment>
</document>
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