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<document id="A8B9753DE2AB4930F1170FDC54E1969B" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. &amp; Micó, E." docDate="2015" docId="03CFA3486B6DFFCFFD7AFB4A5E76FE6B" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala unilineata Filippini, Galante, and Mico, new species" docType="treatment" docVersion="3" lastPageNumber="475" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="474" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C345891D4B835018CD82E9C3E76AD74E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6DD58A06F8BBCC7FE481D374AA5D051">
<mods:title id="4E763043761F6E16EA92BB7E926FFCBF">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="BC470E11106DC1A739358B0BC5B3071E" type="personal">
<mods:role id="5AF1E53268E9819A53485BECAE015251">
<mods:roleTerm id="84F979BFEDDF93AF11F13AE339481579">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0D742E33446968857C178CB29E317F5">Filippini, V.</mods:namePart>
</mods:name>
<mods:name id="8628AAAF0007C4F44042B95F6571BA1D" type="personal">
<mods:role id="EEB6D4391768CA6B82A1E1870F1F3E7C">
<mods:roleTerm id="CC62C9DBE759AF279E82CAB2BE5D3691">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B2B96B126BFA57E6C8115EB8F358D61">Galante, E.</mods:namePart>
</mods:name>
<mods:name id="2CDE57AC970C4D73B5DA56B67DA7E96D" type="personal">
<mods:role id="9A24C1DEB8B3F69A894212D779CD06C3">
<mods:roleTerm id="7AF6F4C4CA5583C81CEDC193E6AB394C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6C4B0C4B1A4D72530460A15C1EA7E8F">Micó, E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="42B8331C7D6718A309DEFDFC86E06575">text</mods:typeOfResource>
<mods:relatedItem id="32F009FB0823BBEB6089F241FBA799E7" type="host">
<mods:titleInfo id="99A52C79E335E641B8BDE5FF7B2E1137">
<mods:title id="8D7C83B11A80B7F301A01BC1FB72ABB7">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="D693DB3190F5084E875AA76703559EF8">
<mods:date id="EFFBE077AE895640A29D37CFFFCE237E">2015</mods:date>
<mods:detail id="41AE7244D86F0B03E77750B7EC4CAFFC" type="pubDate">
<mods:number id="153801E6B4A0AF90F7D77EA30E9B834F">2015-09-30</mods:number>
</mods:detail>
<mods:detail id="0DA031D08A0B60B78866CBB19AA75DDC" type="volume">
<mods:number id="B94D2E1B2702A1059893E1A16EDB1200">69</mods:number>
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<mods:detail id="2AE274CF97A2A2EA58FFD27CBE0802B2" type="issue">
<mods:number id="6465E5019E6548C6BD9476A86805BE92">3</mods:number>
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<mods:url id="C339AD75851CC90FAF1640B3E07142DF">http://dx.doi.org/10.1649/0010-065x-69.3.463</mods:url>
</mods:location>
<mods:classification id="05CB3A9C2A03D6B112E2FEF30A0B6537">journal article</mods:classification>
<mods:identifier id="8D811562339ED4ABF91E9B8B0597CC18" type="DOI">10.1649/0010-065x-69.3.463</mods:identifier>
<mods:identifier id="AF9AC37838B21524B1DCFBB60AD44114" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="E1916F41B3609349051A487820548FA2" type="Zenodo-Dep">10106050</mods:identifier>
<mods:identifier id="DCF07D2F3F9E50D43134A780E8356193" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
</mods:mods>
<treatment id="03CFA3486B6DFFCFFD7AFB4A5E76FE6B" LSID="urn:lsid:plazi:treatment:03CFA3486B6DFFCFFD7AFB4A5E76FE6B" httpUri="http://treatment.plazi.org/id/03CFA3486B6DFFCFFD7AFB4A5E76FE6B" lastPageId="12" lastPageNumber="475" pageId="11" pageNumber="474">
<subSubSection id="C37C41D56B6DFFC8FD7AFB4A598EFB14" pageId="11" pageNumber="474" type="nomenclature">
<paragraph id="8BD9125E6B6DFFC8FD7AFB4A598EFB14" blockId="11.[708,1149,1187,1262]" pageId="11" pageNumber="474">
<heading id="D091A5326B6DFFC8FD7AFB4A598EFB14" centered="true" fontSize="10" level="2" pageId="11" pageNumber="474" reason="2">
<emphasis id="B912CE4C6B6DFFC8FD7AFB4A598EFB14" bold="true" pageId="11" pageNumber="474">
<taxonomicName id="4C6669DD6B6DFFC8FD7AFB4A5909FB14" authority="Filippini, Galante, and Mico" authorityName="Filippini, Galante, and Mico" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="474" phylum="Arthropoda" rank="species" species="unilineata" status="sp. nov.">
<emphasis id="B912CE4C6B6DFFC8FD7AFB4A59FAFB7E" bold="true" box="[708,905,1187,1209]" italics="true" pageId="11" pageNumber="474">Anomala unilineata</emphasis>
Filippini, Galante, and Micó
</taxonomicName>
,
<taxonomicNameLabel id="A22173376B6DFFC8FC39FB57598EFB14" box="[903,1021,1214,1235]" pageId="11" pageNumber="474" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6DFFC8FC95FB305E65FB29" box="[811,1046,1241,1262]" pageId="11" pageNumber="474" type="description">
<paragraph id="8BD9125E6B6DFFC8FC95FB305E65FB29" blockId="11.[708,1149,1187,1262]" box="[811,1046,1241,1262]" pageId="11" pageNumber="474">
(
<figureCitation id="135D0EDB6B6DFFC8FC8CFB30590BFB29" box="[818,888,1241,1262]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="11" pageNumber="474">Figs. 8</figureCitation>
,
<figureCitation id="135D0EDB6B6DFFC8FC3BFB3059ECFB29" box="[901,927,1241,1262]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="11" pageNumber="474">16</figureCitation>
,
<figureCitation id="135D0EDB6B6DFFC8FC15FB3059B6FB29" box="[939,965,1241,1262]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="11" pageNumber="474">24</figureCitation>
,
<figureCitation id="135D0EDB6B6DFFC8FC6EFB305999FB29" box="[976,1002,1241,1262]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="11" pageNumber="474">32</figureCitation>
,
<figureCitation id="135D0EDB6B6DFFC8FC48FB305E7CFB29" box="[1014,1039,1241,1262]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="11" pageNumber="474">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6DFFCFFD7BFAEA5BFBFBBE" lastPageId="12" lastPageNumber="475" pageId="11" pageNumber="474" type="materials_examined">
<paragraph id="8BD9125E6B6DFFC8FD7BFAEA5EE6FA7F" blockId="11.[685,1173,1283,1729]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFAEA592CFADF" bold="true" box="[709,863,1283,1304]" pageId="11" pageNumber="474">Type Material.</emphasis>
H o l o t y p e:
<materialsCitation id="3B0E18036B6DFFC8FC57FAEA58AAFA44" collectingDate="1980-05-17" collectingDateMax="1980-05-19" collectingDateMin="1980-05-17" collectionCode="MNCR" collectorName="W. Hallwachs" country="Costa Rica" location="National Park" pageId="11" pageNumber="474" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B6DFFC8FC57FAEA5988FADE" box="[1001,1019,1283,1305]" pageId="11" pageNumber="474" type="male">♂</specimenCount>
&quot;S a nt a R o s a
<location id="8EB944856B6DFFC8FD13FAF7594AFAF4" LSID="urn:lsid:plazi:treatment:03CFA3486B6DFFCFFD7AFB4A5E76FE6B:8EB944856B6DFFC8FD13FAF7594AFAF4" box="[685,825,1310,1331]" country="Costa Rica" name="National Park" pageId="11" pageNumber="474" stateProvince="Guanacaste">National Park</location>
<collectingRegion id="49A2DCBC6B6DFFC8FCFDFAF75980FAF4" box="[835,1011,1310,1331]" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="474">Guanacaste Prov.</collectingRegion>
<collectingCountry id="F37152CE6B6DFFC8FC45FAF75EE2FAF4" box="[1019,1169,1310,1331]" name="Costa Rica" pageId="11" pageNumber="474">COSTA RICA</collectingCountry>
.
<date id="FFD8349E6B6DFFC8FD13FAD15929FA8A" box="[685,858,1336,1358]" pageId="11" pageNumber="474" value="1980-05-17" valueMax="1980-05-19" valueMin="1980-05-17">
<collectingDate id="EF9CCD766B6DFFC8FD13FAD15929FA8A" box="[685,858,1336,1358]" pageId="11" pageNumber="474" value="1980-05-17" valueMax="1980-05-19" valueMin="1980-05-17">17–19 May 1980</collectingDate>
</date>
DH Janzen &amp;
<collectorName id="269377886B6DFFC8FC47FAD15EF1FA8A" box="[1017,1154,1336,1357]" pageId="11" pageNumber="474">W. Hallwachs</collectorName>
/ INBIOCRI001117054&quot; deposited at
<collectionCode id="ED778A9B6B6DFFC8FBB0FABA5E24FAAF" box="[1038,1111,1363,1384]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="474">MNCR</collectionCode>
.
<typeStatus id="54DDACFC6B6DFFC8FBDCFABA58AAFA44" pageId="11" pageNumber="474" type="paratype">Paratype</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B0E18036B6DFFC8FD56FA875EE2FA7F" collectingDate="1978-03-05" collectingDateMax="1978-03-12" collectingDateMin="1978-03-05" collectionCode="MNCR" collectorName="Costa Rica. D. H. Janzen" country="Costa Rica" location="Santa Rosa National Park" pageId="11" pageNumber="474" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B6DFFC8FD56FA875975FA44" box="[744,774,1389,1411]" pageId="11" pageNumber="474" type="male">1♂</specimenCount>
&quot;
<location id="8EB944856B6DFFC8FCA6FA875E6FFA44" LSID="urn:lsid:plazi:treatment:03CFA3486B6DFFCFFD7AFB4A5E76FE6B:8EB944856B6DFFC8FCA6FA875E6FFA44" box="[792,1052,1390,1411]" country="Costa Rica" name="Santa Rosa National Park" pageId="11" pageNumber="474" stateProvince="Guanacaste">Santa Rosa National Park</location>
<collectingRegion id="49A2DCBC6B6DFFC8FB98FA875971FA5A" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="474">Guanacaste Province</collectingRegion>
<collectorName id="269377886B6DFFC8FCB3FA615E63FA5A" box="[781,1040,1416,1437]" pageId="11" pageNumber="474">Costa Rica. D. H. Janzen</collectorName>
<date id="FFD8349E6B6DFFC8FBA4FA6158A8FA7F" pageId="11" pageNumber="474" value="1978-03-05" valueMax="1978-03-12" valueMin="1978-03-05">
<collectingDate id="EF9CCD766B6DFFC8FBA4FA6158A8FA7F" pageId="11" pageNumber="474" value="1978-03-05" valueMax="1978-03-12" valueMin="1978-03-05">5–12 March 1978</collectingDate>
</date>
/ INBIOCRI001118447&quot;, deposited at
<collectionCode id="ED778A9B6B6DFFC8FBF7FA4A5EE2FA7F" box="[1097,1169,1443,1464]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="474">MNCR</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9125E6B6DFFCFFD7BFA545B7CFB99" blockId="11.[685,1173,1283,1729]" lastBlockId="12.[173,660,194,1730]" lastPageId="12" lastPageNumber="475" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFA54595AFA15" bold="true" box="[709,809,1469,1490]" pageId="11" pageNumber="474">
<typeStatus id="54DDACFC6B6DFFC8FD7BFA545957FA15" box="[709,804,1469,1490]" pageId="11" pageNumber="474" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="135D0EDB6B6DFFC8FCC4FA5459C8FA15" box="[890,955,1469,1490]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="11" pageNumber="474">Fig. 8</figureCitation>
). Body oval. Length
<quantity id="4C9EBFBB6B6DFFC8FD13FA315978FA2A" box="[685,779,1496,1517]" metricMagnitude="-3" metricUnit="m" metricValue="8.52" pageId="11" pageNumber="474" unit="mm" value="8.52">8.52 mm</quantity>
, width
<quantity id="4C9EBFBB6B6DFFC8FCE2FA3159C9FA2A" box="[860,954,1496,1517]" metricMagnitude="-3" metricUnit="m" metricValue="4.27" pageId="11" pageNumber="474" unit="mm" value="4.27">4.27 mm</quantity>
. Head and pronotum dark bronze-brown with green luster. Scutellum dark bronze-brown. Elytra ochre with dark brown suture and median wavy band. Pygidium reddish brown. Legs from yellowish to dark brown. Venter reddish brown.
<emphasis id="B912CE4C6B6DFFC8FCF2F9B459F9F9B5" bold="true" box="[844,906,1629,1650]" pageId="11" pageNumber="474">Head:</emphasis>
Clypeus trapezoidal, with anterior angles widely curved, surface densely punctate-reticulate, convex. Anterior margin straight, thin in frontal view, width/length ratio 2.03. Frons punctate, less densely than clypeus, with a triangular concavity. Ocular canthus long, thin, with acute apex. Ratio of interocular width/width of eye 2.71. Ratio of antennal funiculus/club 0.7.
<emphasis id="B912CE4C6B6AFFCFFD9AFEFB58E0FEE0" bold="true" box="[548,659,274,295]" pageId="12" pageNumber="475">Pronotum:</emphasis>
Subtrapezoidal. Ratio width/length 1.57. Lateral margins regularly convex. Anterior angles acute, posterior angles obtuse. Basal margin complete. Surface with fine, dense punctures.
<emphasis id="B912CE4C6B6AFFCFFD9DFE9558E0FE56" bold="true" box="[547,659,380,401]" pageId="12" pageNumber="475">Scutellum:</emphasis>
Subtriangular, with rounded sides and blunt apex. Ratio width/length 1.16. Surface sparsely punctate.
<emphasis id="B912CE4C6B6AFFCFFF13FE255A85FE26" bold="true" box="[173,246,460,481]" pageId="12" pageNumber="475">Elytra:</emphasis>
With costae defined by regular rows of punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2
<superScript id="7C13BF166B6AFFCFFEDBFE155B04FDCD" attach="none" box="[357,375,508,522]" fontSize="6" pageId="12" pageNumber="475">nd</superScript>
–4
<superScript id="7C13BF166B6AFFCFFE30FE155BEFFDCD" attach="left" box="[398,412,508,522]" fontSize="6" pageId="12" pageNumber="475">th</superScript>
interstices with 1 irregular, secondary stria.
<emphasis id="B912CE4C6B6AFFCFFECDFDF25BA9FDF7" bold="true" box="[371,474,539,560]" pageId="12" pageNumber="475">Pygidium:</emphasis>
Strigate with long setae on posterior margin. Ratio width/length 1.64.
<emphasis id="B912CE4C6B6AFFCFFF59FDB85B43FDA1" bold="true" box="[231,304,593,614]" pageId="12" pageNumber="475">Venter:</emphasis>
Space between mesocoxae narrow, flat. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 2–4 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin.
<emphasis id="B912CE4C6B6AFFCFFE0EFD3C5B95FD2D" bold="true" box="[432,486,725,746]" pageId="12" pageNumber="475">Legs:</emphasis>
Protibia (
<figureCitation id="135D0EDB6B6AFFCFFDF8FD3C58FFFD2D" box="[582,652,725,746]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="12" pageNumber="475">Fig. 16</figureCitation>
) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, sharp, acute. Metatibia stout, fusiform. Ratio length/width 3.27. First external carina well-developed. Surface rugose. External protarsal claw strongly curved, internal claw bifurcate, with upper branch of same length and half width of the lower one, inferior margin sinuate.
<emphasis id="B912CE4C6B6AFFCFFE2DFC435859FC78" bold="true" box="[403,554,938,959]" pageId="12" pageNumber="475">Male genitalia:</emphasis>
Parameres with acute blunt apex in lateral view (
<figureCitation id="135D0EDB6B6AFFCFFD85FC2D58FBFC1E" box="[571,648,964,985]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="12" pageNumber="475">Fig. 24</figureCitation>
). Endophallus (
<figureCitation id="135D0EDB6B6AFFCFFE8BFC365B0CFC33" box="[309,383,991,1012]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="12" pageNumber="475">Fig. 32</figureCitation>
) elongate, dorsally covered with setae, and with 2 ventral patches of sclerotized setae. Sclerotized, flat plate present on ventral inflation, where ejaculatory duct opening is located.
</paragraph>
<paragraph id="8BD9125E6B6AFFCFFF7BFB8D5BFBFBBE" blockId="12.[173,660,194,1730]" box="[197,392,1124,1145]" pageId="12" pageNumber="475">
<emphasis id="B912CE4C6B6AFFCFFF7BFB8D5B66FBBE" bold="true" box="[197,277,1124,1145]" pageId="12" pageNumber="475">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6AFFCFFF7BFB965B3AFAAF" pageId="12" pageNumber="475" type="description">
<paragraph id="8BD9125E6B6AFFCFFF7BFB965B3AFAAF" blockId="12.[173,660,194,1730]" pageId="12" pageNumber="475">
<emphasis id="B912CE4C6B6AFFCFFF7BFB965B5AFB53" bold="true" box="[197,297,1151,1172]" pageId="12" pageNumber="475">Variation.</emphasis>
One
<typeStatus id="54DDACFC6B6AFFCFFED8FB965BC8FB53" box="[358,443,1151,1172]" pageId="12" pageNumber="475" type="paratype">paratype</typeStatus>
. Head, pronotum, and venter vary from reddish to dark brown. Body length
<quantity id="4C9EBFBB6B6AFFCFFF4EFB5D5B0CFB0E" box="[240,383,1203,1225]" metricMagnitude="-3" metricUnit="m" metricValue="8.129999999999999" metricValueMax="8.52" metricValueMin="7.74" pageId="12" pageNumber="475" unit="mm" value="8.129999999999999" valueMax="8.52" valueMin="7.74">7.74–8.52 mm</quantity>
, body width
<quantity id="4C9EBFBB6B6AFFCFFDBEFB5D58E3FB0E" box="[512,656,1203,1225]" metricMagnitude="-3" metricUnit="m" metricValue="4.275" metricValueMax="4.28" metricValueMin="4.27" pageId="12" pageNumber="475" unit="mm" value="4.275" valueMax="4.28" valueMin="4.27">4.27–4.28 mm</quantity>
. Clypeal width/length ratio 2.03–2.07. Ratio of interocular width/width of eye 2.71–2.83. Ratio of antennal funiculus/club 0.70–0.73. Pronotal width/length ratio 1.57–1.60. Scutellar width/ length ratio 1.16–1.22. Metatibial length/width ratio 2.96–3.27.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6AFFCFFF7BFA875BBBFA7F" pageId="12" pageNumber="475" type="etymology">
<paragraph id="8BD9125E6B6AFFCFFF7BFA875BBBFA7F" blockId="12.[173,660,194,1730]" pageId="12" pageNumber="475">
<emphasis id="B912CE4C6B6AFFCFFF7BFA875B44FA44" bold="true" box="[197,311,1390,1411]" pageId="12" pageNumber="475">Etymology.</emphasis>
From the Latin adjectives
<emphasis id="B912CE4C6B6AFFCFFDFEFA845AA0FA59" italics="true" pageId="12" pageNumber="475">unus, -a, -um</emphasis>
, one, and
<emphasis id="B912CE4C6B6AFFCFFEFDFA615B98FA59" box="[323,491,1416,1438]" italics="true" pageId="12" pageNumber="475">lineatus, -a, -um</emphasis>
, striped, for the single band across the elytra.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6AFFCFFF7BFA545BDEF9C0" pageId="12" pageNumber="475" type="distribution">
<paragraph id="8BD9125E6B6AFFCFFF7BFA545BDEF9C0" blockId="12.[173,660,194,1730]" pageId="12" pageNumber="475">
<emphasis id="B912CE4C6B6AFFCFFF7BFA545B34FA15" bold="true" box="[197,327,1469,1490]" pageId="12" pageNumber="475">Distribution.</emphasis>
This species occurs in the dry forest in northeastern
<collectingCountry id="F37152CE6B6AFFCFFE2BFA31587BFA2A" box="[405,520,1496,1517]" name="Costa Rica" pageId="12" pageNumber="475">Costa Rica</collectingCountry>
, from
<quantity id="4C9EBFBB6B6AFFCFFDEEFA315A99F9C0" metricMagnitude="2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="12" pageNumber="475" unit="m" value="260.0" valueMax="300.0" valueMin="220.0">220 to 300 m</quantity>
elevation (
<figureCitation id="135D0EDB6B6AFFCFFEE7FA1B5BD1F9C0" box="[345,418,1522,1543]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="12" pageNumber="475">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6AFFCFFF7BF9E45E76FE6B" pageId="12" pageNumber="475" type="diagnosis">
<paragraph id="8BD9125E6B6AFFCFFF7BF9E45B1FF94B" blockId="12.[173,660,194,1730]" pageId="12" pageNumber="475">
<emphasis id="B912CE4C6B6AFFCFFF7BF9E45B5EF9E5" bold="true" box="[197,301,1549,1570]" pageId="12" pageNumber="475">Diagnosis.</emphasis>
This species is recognized by its small size, pronotum dark bronze-brown, elytra ochre with dark brown, median, wavy band, and endophallus elongated with two ventral patches of sclerotized setae.
</paragraph>
<paragraph id="8BD9125E6B6AFFCFFF7BF97B5E76FE6B" blockId="12.[173,660,194,1730]" lastBlockId="12.[707,1194,194,428]" pageId="12" pageNumber="475">
Another species with one transverse band on the elytra is
<taxonomicName id="4C6669DD6B6AFFCFFE90F94558FCF906" authority="Stephens, 1830" authorityName="Stephens" authorityYear="1830" box="[302,655,1708,1730]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="donovani">
<emphasis id="B912CE4C6B6AFFCFFE90F9455B99F905" box="[302,490,1708,1730]" italics="true" pageId="12" pageNumber="475">Anomala donovani</emphasis>
Stephens, 1830
</taxonomicName>
, but it is larger, with a black pronotum, pigmented punctures on the elytra, and shaded apical portion of the elytra. Other small species, like
<taxonomicName id="4C6669DD6B6AFFCFFBEFFF1E59B3FEE0" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="B912CE4C6B6AFFCFFBEFFF1E5947FEE0" italics="true" pageId="12" pageNumber="475">Anomala discoidalis</emphasis>
Bates, 1888
</taxonomicName>
,
<taxonomicName id="4C6669DD6B6AFFCFFC71FEF8594BFE86" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="retusicollis">
<emphasis id="B912CE4C6B6AFFCFFC71FEF85EDAFEE0" box="[975,1193,273,295]" italics="true" pageId="12" pageNumber="475">Anomala retusicollis</emphasis>
Bates, 1888
</taxonomicName>
,
<taxonomicName id="4C6669DD6B6AFFCFFCF8FEC55886FE9B" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="tessellatipennis">
<emphasis id="B912CE4C6B6AFFCFFCF8FEC55E4BFE85" box="[838,1080,300,322]" italics="true" pageId="12" pageNumber="475">Anomala tessellatipennis</emphasis>
Blanchard, 1851
</taxonomicName>
, and
<taxonomicName id="4C6669DD6B6AFFCFFC93FEAF5EDAFE9B" authority="(Melsheimer, 1844)" baseAuthorityName="Melsheimer" baseAuthorityYear="1844" box="[813,1193,326,348]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B912CE4C6B6AFFCFFC93FEAF5992FE9B" box="[813,993,326,348]" italics="true" pageId="12" pageNumber="475">Anomala undulata</emphasis>
(Melsheimer, 1844)
</taxonomicName>
have the elytral pattern formed by isolated maculae, not forming a continuous band, and differently shaped parameres and endophalli.
</paragraph>
</subSubSection>
</treatment>
</document>
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