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<document id="915FEF4D57671B0360E6846D8D60A505" ID-DOI="10.5281/zenodo.198427" ID-GBIF-Dataset="1c32eb21-6978-41da-b589-c98453d4f572" ID-ISSN="1175-5326" ID-Zenodo-Dep="198427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460297110161" checkinUser="plazi" docAuthor="Short, Andrew E. Z. &amp; García, Mauricio" docDate="2010" docId="03CEA91F8C3CB900A2C066F7FE88F725" docLanguage="en" docName="zt02635p031.pdf" docOrigin="Zootaxa 2635" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oocyclus trujillo Short &amp; Garcia, sp. n." docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="FFF7D1678C27B91CA2576660FFD8F557" masterDocTitle="A review of the Oocyclus Sharp of Venezuela with description of twelve new species (Coleoptera: Hydrophilidae: Laccobiini)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="28" updateTime="1698252212253" updateUser="plazi">
<mods:mods id="7257A25B0BFBD6C5E24041217FBCC2F8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6CB269C16B221080AF6B8D93B4FC50A9">
<mods:title id="675C8585ABEEF8B5FF47341CE8A57AE1">A review of the Oocyclus Sharp of Venezuela with description of twelve new species (Coleoptera: Hydrophilidae: Laccobiini)</mods:title>
</mods:titleInfo>
<mods:name id="CD1F10B77C61B2A3F982ACBE60848BC6" type="personal">
<mods:role id="54686F1C76504065F9F7020042AAC199">
<mods:roleTerm id="53ED3A52245A2BF53C75D41232A4C239">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6791FB5199C62822AE16C244FA093AEE">Short, Andrew E. Z.</mods:namePart>
</mods:name>
<mods:name id="E17CE34D87EB4FB6C7DD4D04E7981703" type="personal">
<mods:role id="2CDE4743A70F538D539996CDEF85C5E7">
<mods:roleTerm id="D8E8F9DB5BF39A66614E06C49EB19FE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BD6BF5BBE580FA7808BB3EAA1DA9697">García, Mauricio</mods:namePart>
</mods:name>
<mods:typeOfResource id="FF2BFFC7884A2060867BAC1EF90F04D5">text</mods:typeOfResource>
<mods:relatedItem id="2F6E8B7B01043042C69A9E58EEF19F88" type="host">
<mods:titleInfo id="9B0DBEDBA9A0A25FA95C2BDA928B3FB7">
<mods:title id="E53C91DF4CF56BD8F927283FC1C67BDA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B2E62891175D08E313745CA0C9D3CD11">
<mods:date id="410D49D5863696C068F9F63E41FD4AF5">2010</mods:date>
<mods:detail id="8538BA015266C7805D28A5640CC1DF01" type="volume">
<mods:number id="71C0D728F4275ECEA27595765A302EEA">2635</mods:number>
</mods:detail>
<mods:extent id="F485275B771668557B7A6B98A9F05E78" unit="page">
<mods:start id="03860003E304C5A0FBD686460818ED88">1</mods:start>
<mods:end id="D36209A39B51ED897DC209FA3877FBAE">31</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BE54DC08507FA3BB9B12DCB1B5BCD68D">journal article</mods:classification>
<mods:identifier id="DC157AA3CEA6414AFAE1ACAE7D122036" type="DOI">10.5281/zenodo.198427</mods:identifier>
<mods:identifier id="2E3AA46C475DC4EFF9A248255F8E21D4" type="GBIF-Dataset">1c32eb21-6978-41da-b589-c98453d4f572</mods:identifier>
<mods:identifier id="8C449CF23D5ED65156D5EE9E923FD044" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="347F56C10D719FB7C50AB3494921FA29" type="Zenodo-Dep">198427</mods:identifier>
</mods:mods>
<treatment id="03CEA91F8C3CB900A2C066F7FE88F725" ID-DOI="http://doi.org/10.5281/zenodo.6205314" ID-GBIF-Taxon="119399264" ID-Zenodo-Dep="6205314" LSID="urn:lsid:plazi:treatment:03CEA91F8C3CB900A2C066F7FE88F725" httpUri="http://treatment.plazi.org/id/03CEA91F8C3CB900A2C066F7FE88F725" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="C37D4B828C3CB907A2C066F7FE82F583" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BD818098C3CB907A2C066F7FDABF5E6" blockId="27.[151,627,151,212]" box="[151,627,151,177]" pageId="27" pageNumber="28">
<heading id="D090AF658C3CB907A2C066F7FDABF5E6" bold="true" box="[151,627,151,177]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B913C41B8C3CB907A2C066F7FDABF5E6" bold="true" box="[151,627,151,177]" pageId="27" pageNumber="28">
<taxonomicName id="4C67638A8C3CB907A2C066F7FDF2F5E6" ID-CoL="49RX2" authority="Short &amp; Garcia" authorityName="Short &amp; Garcia" box="[151,554,151,177]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">
<emphasis id="B913C41B8C3CB907A2C066F7FEB8F5E6" bold="true" box="[151,352,151,177]" italics="true" pageId="27" pageNumber="28">Oocyclus trujillo</emphasis>
Short &amp; García
</taxonomicName>
<taxonomicNameLabel id="A22079608C3CB907A06666F7FDABF5E6" box="[561,627,151,177]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD818098C3CB907A2C066DAFE82F583" blockId="27.[151,627,151,212]" box="[151,346,186,212]" pageId="27" pageNumber="28">
(
<figureCitation id="135C048C8C3CB907A2C866DAFF29F583" box="[159,241,186,212]" captionStart="FIGURE 3" captionStartId="7.[151,255,1852,1876]" captionTargetBox="[158,1416,651,1818]" captionTargetId="figure@7.[151,1436,642,1830]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal and lateral habitus of Oocyclus species found in the Venezuelan Andes region. A) O. meridensis sp. n.; B) O. trujillo sp. n.; C) O. andinus sp. n.; D) O. pico sp. n. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/198430/files/figure.png" pageId="27" pageNumber="28">Figs. 3</figureCitation>
B, 5E, 7)
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C3CB907A2C06765FAA8F67D" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BD818098C3CB907A2C06765FAA8F67D" blockId="27.[151,1438,261,2004]" pageId="27" pageNumber="28">
<emphasis id="B913C41B8C3CB907A2C06765FC92F448" bold="true" box="[151,842,261,287]" pageId="27" pageNumber="28">
<typeStatus id="54DCA6AB8C3CB907A2C06765FF0CF448" box="[151,212,261,287]" pageId="27" pageNumber="28">Type</typeStatus>
material.
<typeStatus id="54DCA6AB8C3CB907A3046765FE11F448" box="[339,457,261,287]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F37058998C3CB907A3826765FD48F448" box="[469,656,261,287]" name="Venezuela" pageId="27" pageNumber="28">VENEZUELA</collectingCountry>
:
<taxonomicName id="4C67638A8C3CB907A0CB6765FD24F448" box="[668,764,261,287]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">Trujillo</taxonomicName>
State:
</emphasis>
male, “
<collectingCountry id="F37058998C3CB907A1F56765FB80F448" box="[930,1112,261,287]" name="Venezuela" pageId="27" pageNumber="28">VENEZUELA</collectingCountry>
:
<taxonomicName id="4C67638A8C3CB907A6356765FB63F448" box="[1122,1211,261,287]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">Trujillo</taxonomicName>
State/
<geoCoordinate id="EE537ECE8C3CB907A75C6765FA4FF448" box="[1291,1431,261,287]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="9.46325">9°27.795’N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C3CB907A2C0674AFEEEF413" box="[151,310,298,324]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-70.41893">70°25.136’W</geoCoordinate>
,
<quantity id="4C9FB5EC8C3CB907A316674AFE40F413" box="[321,408,298,324]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="28" unit="m" value="1100.0">1100 m</quantity>
/
<date id="FFD93EC98C3CB907A3FF674AFDC1F413" box="[424,537,298,324]" pageId="27" pageNumber="28" value="2009-01-21">21.i.2009</date>
; Seep, E. of Pampan/ leg. Short, García, &amp; Camacho/ VZ09-0121-
<date id="FFD93EC98C3CB907A77F674AFA84F413" box="[1320,1372,298,324]" pageId="27" pageNumber="28" value="2005-10">05X</date>
; rock seep”, “[barcode]/SM0844435/ KUNHM-ENT”, “
<typeStatus id="54DCA6AB8C3CB907A0AD6730FC44F43D" box="[762,924,336,362]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C67638A8C3CB907A1FA6730FB9FF43E" box="[941,1095,336,361]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B913C41B8C3CB907A1FA6730FB9FF43E" box="[941,1095,336,361]" italics="true" pageId="27" pageNumber="28">OOCYCLUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C67638A8C3CB907A60F6730FB75F43E" box="[1112,1197,336,361]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">
<emphasis id="B913C41B8C3CB907A60F6730FB75F43E" box="[1112,1197,336,361]" italics="true" pageId="27" pageNumber="28">trujillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C3CB907A6E16730FB36F43D" box="[1206,1262,336,362]" pageId="27" pageNumber="28" rank="species">sp.n.</taxonomicNameLabel>
/ des.
<bibRefCitation id="EFF665F88C3CB907A76D6730FEE0F4D8" author="Short" pageId="27" pageNumber="28" refString="Hall, W. E. &amp; Short, A. E. Z. (2010) A new species of Hydroscapha LeConte from northwestern Venezuela (Coleoptera: Myxophaga: Hydroscaphidae). Zootaxa, 2358, 49 - 56." type="journal article" year="2010">Short &amp; García 2010</bibRefCitation>
” (
<collectionCode id="ED7680CC8C3CB907A3026715FE75F4D8" LSID="urn:lsid:biocol.org:col:33834" box="[341,429,373,399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="27" pageNumber="28">MIZA</collectionCode>
).
<typeStatus id="54DCA6AB8C3CB907A3946715FD93F4D8" box="[451,587,373,399]" pageId="27" pageNumber="28" type="paratype">
<emphasis id="B913C41B8C3CB907A3946715FD93F4D8" bold="true" box="[451,587,373,399]" pageId="27" pageNumber="28">Paratypes</emphasis>
</typeStatus>
(398 exs.):
<emphasis id="B913C41B8C3CB907A0BE6715FB59F4D8" bold="true" box="[745,1153,373,399]" pageId="27" pageNumber="28">
<collectingCountry id="F37058998C3CB907A0BE6715FC68F4D8" box="[745,944,373,399]" name="Venezuela" pageId="27" pageNumber="28">VENEZUELA</collectingCountry>
: Barinas State:
</emphasis>
<quantity id="4C9FB5EC8C3CB907A6DD6715FB01F4D8" box="[1162,1241,373,399]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="27" pageNumber="28" unit="km" value="24.0">24 km</quantity>
NW Barinitas,
<date id="FFD93EC98C3CB907A2C067FAFECAF4E3" box="[151,274,410,436]" pageId="27" pageNumber="28" value="1969-02-23">23.ii.1969</date>
, leg. P. &amp; P. Spangler (143 exs.,
<collectionCode id="ED7680CC8C3CB907A0CC67FAFD2EF4E3" LSID="urn:lsid:biocol.org:col:34871" box="[667,758,410,436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="27" pageNumber="28">USNM</collectionCode>
);
<emphasis id="B913C41B8C3CB907A15D67FAFC61F4E3" bold="true" box="[778,953,410,436]" pageId="27" pageNumber="28">Mérida State:</emphasis>
Along road between Jaji &amp; Mérida, wet rock and cement at river crossing,
<geoCoordinate id="EE537ECE8C3CB907A07D67A0FD75F48D" box="[554,685,448,474]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="8.5857">8.58570°N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C3CB907A0EF67A0FC96F48D" box="[696,846,448,474]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-71.29927">71.29927°W</geoCoordinate>
,
<quantity id="4C9FB5EC8C3CB907A10E67A0FC6DF48D" box="[857,949,448,474]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="27" pageNumber="28" unit="m" value="1730.0">1730 m</quantity>
,
<date id="FFD93EC98C3CB907A19667A0FBE9F48D" box="[961,1073,448,474]" pageId="27" pageNumber="28" value="2006-01-16">16.i.2006</date>
, leg. A. Short &amp; P. Torres, AS- 06-033 (13 exs.); Along road between Jaji &amp; Mérida, small waterfall and stream pools,
<geoCoordinate id="EE537ECE8C3CB907A75E6785FA40F4A8" box="[1289,1432,485,511]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="8.58515">8.58515°N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C3CB907A2C0646AFEE9F773" box="[151,305,522,548]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-71.30103">71.30103°W</geoCoordinate>
,
<quantity id="4C9FB5EC8C3CB907A369646AFE45F773" box="[318,413,522,548]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="27" pageNumber="28" unit="m" value="1750.0">1750 m</quantity>
,
<date id="FFD93EC98C3CB907A3FC646AFDF8F773" box="[427,544,522,548]" pageId="27" pageNumber="28" value="2006-01-16">16.i.2006</date>
, leg. A. Short &amp; P. Torres, AS-06-034 (11 exs.).; ca.
<quantity id="4C9FB5EC8C3CB907A6FC646AFB31F773" box="[1195,1257,522,548]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="km" value="2.0">2 km</quantity>
S. La Azulita, Cascada la Palmita,
<date id="FFD93EC98C3CB907A3DF6450FDC9F71D" box="[392,529,560,586]" pageId="27" pageNumber="28" value="2009-07-21">21.vii.2009</date>
, leg. Short et al., VZ09-0721-01A (94 exs.);
<emphasis id="B913C41B8C3CB907A67E644FFB04F71E" bold="true" box="[1065,1244,559,585]" pageId="27" pageNumber="28">Táchira State:</emphasis>
Chorro el Indio National Park,
<geoCoordinate id="EE537ECE8C3CB907A31C6435FE2DF738" box="[331,501,597,623]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="7.7232804">7°43’23.81”N</geoCoordinate>
, 72°
<date id="FFD93EC98C3CB907A07D6435FD4CF738" box="[554,660,597,623]" pageId="27" pageNumber="28" value="13-11-31">11.31.13</date>
”W,
<quantity id="4C9FB5EC8C3CB907A09E6435FCFEF738" box="[713,806,597,623]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="27" pageNumber="28" unit="m" value="1225.0">1225 m</quantity>
,
<date id="FFD93EC98C3CB907A1646435FC66F738" box="[819,958,597,623]" pageId="27" pageNumber="28" value="2009-07-16">16.vii.2009</date>
, leg. Short &amp; García, wet rock and rock seeps, VZ09-0716-01A (26 exs.); Chorro el Indio National Park, stream road crossing,
<geoCoordinate id="EE537ECE8C3CB907A6A8641AFA40F7C3" box="[1279,1432,634,660]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="7.73145">7°43.887’N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C3CB907A2C064C0FEE0F7ED" box="[151,312,672,698]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-72.19521">72°11.713”W</geoCoordinate>
,
<quantity id="4C9FB5EC8C3CB907A31364C0FE79F7ED" box="[324,417,672,698]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="27" pageNumber="28" unit="m" value="1330.0">1330 m</quantity>
,
<date id="FFD93EC98C3CB907A3F964C0FDEEF7ED" box="[430,566,672,698]" pageId="27" pageNumber="28" value="2009-07-16">16.vii.2009</date>
, leg. Short &amp; Gustafson, wet cement next to stream, VZ09-0716-02A (50 exs.);
<emphasis id="B913C41B8C3CB907A28864A5FE57F788" bold="true" box="[223,399,709,735]" pageId="27" pageNumber="28">
<taxonomicName id="4C67638A8C3CB907A28864A5FEE7F788" box="[223,319,709,735]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">Trujillo</taxonomicName>
State:
</emphasis>
El Chorreron, road to Bocono,
<quantity id="4C9FB5EC8C3CB907A15F64A5FC8EF788" box="[776,854,709,735]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="28" unit="m" value="500.0">500 m</quantity>
,
<date id="FFD93EC98C3CB907A13564A5FC3BF788" box="[866,995,709,735]" pageId="27" pageNumber="28" value="2002-11-10">10.xi.2002</date>
, leg. M. García (2 exs.,
<collectionCode id="ED7680CC8C3CB907A75664A5FA92F788" box="[1281,1354,709,735]" httpUri="http://grbio.org/cool/hrb9-44fx" name="International Institute of Biological Control" pageId="27" pageNumber="28">CIBC</collectionCode>
), same data but
<date id="FFD93EC98C3CB907A2AC648AFEA0F653" box="[251,376,746,772]" pageId="27" pageNumber="28" value="2002-09-12">12.ix.2002</date>
(5 exs.,
<collectionCode id="ED7680CC8C3CB907A38C648AFDFCF653" box="[475,548,746,772]" httpUri="http://grbio.org/cool/hrb9-44fx" name="International Institute of Biological Control" pageId="27" pageNumber="28">CIBC</collectionCode>
); same data as
<typeStatus id="54DCA6AB8C3CB907A085648AFCE0F653" box="[722,824,746,772]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(29 exs.); E. of Pampan,
<geoCoordinate id="EE537ECE8C3CB907A637648AFB35F653" box="[1120,1261,746,772]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="9.456933">9°27.416’N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C3CB907A6AF648AFA4FF653" box="[1272,1431,746,772]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-70.4265">70°25.590’W</geoCoordinate>
,
<quantity id="4C9FB5EC8C3CB907A2C06570FF2AF67D" box="[151,242,784,810]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="28" unit="m" value="1100.0">1100 m</quantity>
, rock seep along road,
<date id="FFD93EC98C3CB907A3A86570FDA8F67D" box="[511,624,784,810]" pageId="27" pageNumber="28" value="2009-01-21">21.i.2009</date>
, leg. A. Short, M. García, J. Camacho, VZ09-0121-
<date id="FFD93EC98C3CB907A6996570FADAF67D" box="[1230,1282,784,810]" pageId="27" pageNumber="28" value="2006-10">06X</date>
(25 exs.)
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C3CB907A2926555FDEBF178" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BD818098C3CB907A2926555FDC1F6E8" blockId="27.[151,1438,261,2004]" pageId="27" pageNumber="28">
<emphasis id="B913C41B8C3CB907A2926555FE9CF618" bold="true" box="[197,324,821,847]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Smaller sized species (
<figureCitation id="135C048C8C3CB907A0026555FD42F618" box="[597,666,821,847]" captionStart="FIGURE 3" captionStartId="7.[151,255,1852,1876]" captionTargetBox="[158,1416,651,1818]" captionTargetId="figure@7.[151,1436,642,1830]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal and lateral habitus of Oocyclus species found in the Venezuelan Andes region. A) O. meridensis sp. n.; B) O. trujillo sp. n.; C) O. andinus sp. n.; D) O. pico sp. n. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/198430/files/figure.png" pageId="27" pageNumber="28">Fig. 3</figureCitation>
B). Posterolateral corners of pronotum spinose. Procoxae strongly spinose. Elytra with interrupted rows of larger systematic punctures, bearing dense short rows or clumps of erect setae. Elytral suture not raised in posterior third. Elytral margins not explanate, and with a marginal fringe of setae on posterior third.
</paragraph>
<paragraph id="8BD818098C3CB907A29265AAFDEBF178" blockId="27.[151,1438,261,2004]" pageId="27" pageNumber="28">
The smaller size (less than 4.0 mm) and spinose posterior pronotal corners will separate the species from all other Venezuelan taxa except
<taxonomicName id="4C67638A8C3CB907A04D6590FDA8F15E" box="[538,624,1008,1033]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pico">
<emphasis id="B913C41B8C3CB907A04D6590FDA8F15E" box="[538,624,1008,1033]" italics="true" pageId="27" pageNumber="28">
<collectingCountry id="F37058998C3CB907A04D6590FDEAF15E" box="[538,562,1008,1033]" name="Iceland" pageId="27" pageNumber="28">O</collectingCountry>
. pico
</emphasis>
</taxonomicName>
, which does not have clumps of erect setae on the elytra and is much more rarely collected in the Andes.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C3CB900A292625AFECBF4C5" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8BD818098C3CB907A292625AFA5DF103" blockId="27.[151,1438,261,2004]" box="[197,1413,1082,1108]" pageId="27" pageNumber="28">
<emphasis id="B913C41B8C3CB907A292625AFE85F103" bold="true" box="[197,349,1082,1108]" pageId="27" pageNumber="28">Description.</emphasis>
<emphasis id="B913C41B8C3CB907A333625AFDD5F104" box="[356,525,1082,1107]" italics="true" pageId="27" pageNumber="28">Size and form.</emphasis>
Body length = 3.0–
<quantity id="4C9FB5EC8C3CB907A0A0625AFC8CF103" box="[759,852,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="27" pageNumber="28" unit="mm" value="3.9">3.9 mm</quantity>
. TL/GW= 1.52. Body oval, moderately convex.
</paragraph>
<paragraph id="8BD818098C3CB907A2926200FB5FF058" blockId="27.[151,1438,261,2004]" pageId="27" pageNumber="28">
<emphasis id="B913C41B8C3CB907A2926200FECEF12E" box="[197,278,1120,1145]" italics="true" pageId="27" pageNumber="28">Color.</emphasis>
Dorsum of head, pronotum, and elytra black, usually with iridescent green reflections. Anterolateral margins of pronotum with white patch. Elytra sometimes with very faint iridescent green maculae. Maxillary and labial palps yellow, the former slightly darkened at apex of apical segment. Mentum and stipes light to reddish brown, distinctly paler than ventral face of head. Legs, epipleura, lateral margins of prosternum medium to yellowish brown. Abdominal ventrites medium to dark brown.
</paragraph>
<paragraph id="8BD818098C3CB907A292637BFE09F36D" blockId="27.[151,1438,261,2004]" pageId="27" pageNumber="28">
<emphasis id="B913C41B8C3CB907A292637BFED4F063" box="[197,268,1307,1332]" italics="true" pageId="27" pageNumber="28">Head.</emphasis>
Ground punctation on labrum, clypeus and frons moderately fine, distance between punctures 1.5– 3.0× the width of one puncture. Systematic punctures on labrum consisting of several indistinct punctures, sometimes bearing short setae. Frons with an irregular row of systematic punctures mesad of each eye, bearing fine setae. Clypeus with a few very indistinct systematic punctures along anterolateral margins, slightly larger than surrounding punctation. Maxillary palps short, slightly shorter than width of labrum; segment 2 slightly bulbous, apical segment slightly longer than penultimate. Labial palps three-fourths as long as width of mentum. Mentum quadrate, smooth, anterior margin very slightly convex, bearing very fine, sparse scattered punctures.
</paragraph>
<paragraph id="8BD818098C3CB900A2926025FCACF475" blockId="27.[151,1438,261,2004]" lastBlockId="28.[151,1437,152,626]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B913C41B8C3CB907A2926025FEF9F309" box="[197,289,1605,1630]" italics="true" pageId="27" pageNumber="28">Thorax.</emphasis>
Ground punctation on pronotum and elytra composed of very fine, evenly distributed punctures. Pronotal systematic punctures with short fine setae, distinctly larger than ground punctures, anterior and posterior series each forming irregular row. Lateral margins of pronotum densely set with setiferous punctures bearing moderately long yellow setae. Posterolateral corners of pronotum sharply spinose. Sutural punctation on elytra absent or unmodified from ground punctuation except for short row of dense coarse punctures bearing erect setae of same
<typeStatus id="54DCA6AB8C3CB907A3886160FDCAF24D" box="[479,530,1792,1818]" pageId="27" pageNumber="28">type</typeStatus>
as systematic punctures, appearing as if short section of row 1 of systematic punctures has migrated mesad between the rest of the row and the suture. Elytra with five very irregular, interrupted rows of systematic punctures; rows 1–3 appearing as either series of short rows of dense erect setae or coalesced into a series of clumps of punctures bearing erect setae. Rows 4–5 represented by broadly spaced punctures bearing fine decumbent setae. Prosternum with defined median carina; slightly elevated anteromedially, the elevation with two very long thickened spine-like setae. Elevated process of mesoventrite elongate, slightly longer than wide; surface nearly flat; set with 5 thickened spine-like setae. Metaventrite with small oval glabrous area posteromedially, slightly longer than wide, length of glabrous area slightly less than one-half length of metaventrite. Procoxae with dense, fine short pubescence and long coarse spine-like setae. Protibiae each with 8–12 spine-like setae on dorsal face.
</paragraph>
<paragraph id="8BD818098C3BB900A292674DFECBF4C5" blockId="28.[151,1437,152,626]" pageId="28" pageNumber="29">
<emphasis id="B913C41B8C3BB900A292674DFEE6F411" box="[197,318,301,326]" italics="true" pageId="28" pageNumber="29">Abdomen.</emphasis>
Ventrites with scattered setae of varying lengths; each ventrite with 2 irregular rows of very long erect setae which are more than twice as long as the longest setae on the metaventrite. Aedeagus as in
<figureCitation id="135C048C8C3BB900A2C06718FF21F4C5" box="[151,249,376,402]" captionStart="FIGURE 5" captionStartId="15.[151,255,1928,1952]" captionTargetBox="[174,1419,194,1908]" captionTargetId="figure@15.[174,1420,194,1909]" captionTargetPageId="15" captionText="FIGURE 5. Aedeagi of Oocyclus species. A) O. pittieri sp. n., B) O. maculatus Sharp; C) O. s ubstillus Short &amp; Perkins; D) O. floccus sp. n.; E) O. trujillo sp. n.; F) O. andinus sp. n.; G) O. meridensis sp. n.; H) O. sapphirus sp. n. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/198432/files/figure.png" pageId="28" pageNumber="29">Figure 5</figureCitation>
E.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C3BB900A29267FDFAA6F4E0" box="[197,1406,413,439]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BD818098C3BB900A29267FDFAA6F4E0" blockId="28.[151,1437,152,626]" box="[197,1406,413,439]" pageId="28" pageNumber="29">
<emphasis id="B913C41B8C3BB900A29267FDFE8AF4E0" bold="true" box="[197,338,413,439]" pageId="28" pageNumber="29">Etymology.</emphasis>
<taxonomicName id="4C67638A8C3BB900A30E67FDFE69F4E1" box="[345,433,413,438]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">
<emphasis id="B913C41B8C3BB900A30E67FDFE69F4E1" box="[345,433,413,438]" italics="true" pageId="28" pageNumber="29">Trujillo</emphasis>
</taxonomicName>
, after the Andean city and state of the same name. Treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C3BB900A29267A2FE32F755" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BD818098C3BB900A29267A2FE32F755" blockId="28.[151,1437,152,626]" pageId="28" pageNumber="29">
<emphasis id="B913C41B8C3BB900A29267A2FEB6F48B" bold="true" box="[197,366,450,476]" pageId="28" pageNumber="29">Distribution.</emphasis>
Widely distributed in the Venezuelan Andes, including the states of Táchira, Barinas, Mérida, and
<taxonomicName id="4C67638A8C3BB900A37B6788FE5DF755" box="[300,389,488,514]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">Trujillo</taxonomicName>
(
<figureCitation id="135C048C8C3BB900A3C36788FE04F755" box="[404,476,488,514]" captionStart="FIGURE 7" captionStartId="22.[151,255,1419,1443]" captionTargetBox="[151,1431,198,1376]" captionTargetId="figure@22.[151,1436,197,1398]" captionTargetPageId="22" captionText="FIGURE 7. Distribution of Oocyclus species in Venezuela." httpUri="https://zenodo.org/record/198434/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C3BB900A292646DFE88F725" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BD818098C3BB900A292646DFE88F725" blockId="28.[151,1437,152,626]" pageId="28" pageNumber="29">
<emphasis id="B913C41B8C3BB900A292646DFEE7F770" bold="true" box="[197,319,525,551]" pageId="28" pageNumber="29">Remarks.</emphasis>
This fairly common species has been collected at a variety of hygropetric habitats, including along road cuts, the sides of waterfalls, wet cement, and seepages. It frequently co-occurs with
<taxonomicName id="4C67638A8C3BB900A6AE6452FA44F71C" box="[1273,1436,562,587]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="meridensis">
<emphasis id="B913C41B8C3BB900A6AE6452FA44F71C" box="[1273,1436,562,587]" italics="true" pageId="28" pageNumber="29">
<collectingCountry id="F37058998C3BB900A6AE6452FAC9F71C" box="[1273,1297,562,587]" name="Iceland" pageId="28" pageNumber="29">O</collectingCountry>
. meridensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C67638A8C3BB900A29F6438FE91F726" box="[200,329,600,625]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B913C41B8C3BB900A29F6438FE91F726" box="[200,329,600,625]" italics="true" pageId="28" pageNumber="29">
<collectingCountry id="F37058998C3BB900A29F6438FF38F726" box="[200,224,600,625]" name="Iceland" pageId="28" pageNumber="29">O</collectingCountry>
. andinus
</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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