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<mods:title id="71BF95AEC8D8AF51B18426C30C139618">The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="2D1F5482A66833BC79CDE493CE6347DB">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="8713A535597D50323571C2FE4A8A8F10">Landau, Bernard M.</mods:namePart>
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<mods:affiliation id="D4BEA18844981A301680558DF5E085F8">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655</mods:affiliation>
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<taxonomicName id="4C675589FFC20C1FFF65FA6AED22FA25" authority="Harzhauser &amp; Landau, 2024" authorityName="Harzhauser &amp; Landau" authorityYear="2024" box="[151,343,1415,1442]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="zachosi" status="sp. nov.">
<emphasis id="B913F218FFC20C1FFF65FA6AED22FA25" bold="true" box="[151,343,1415,1442]" italics="true" pageId="88" pageNumber="89">Euthria zachosi</emphasis>
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,
<materialsCitation id="3B0F2457FFC20C1FFD92F9D1E82AF9D1" box="[608,1119,1596,1622]" collectionCode="MD" country="Austria" location="Niederleis" pageId="88" pageNumber="89" specimenCount="46" typeStatus="paratype">
<collectionCode id="ED76B6CFFFC20C1FFD92F9D1EEE4F9D1" box="[608,657,1596,1622]" country="Germany" name="Museum Donaueschingen" pageId="88" pageNumber="89">MD</collectionCode>
:
<quantity id="4C9F83EFFFC20C1FFD6EF9D1EF77F9D1" box="[668,770,1596,1622]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="88" pageNumber="89" unit="mm" value="19.6">19.6 mm</quantity>
,
<location id="8EB878D1FFC20C1FFCFFF9D1EFF5F9D1" LSID="urn:lsid:plazi:treatment:03CE9F1CFFC20C1EFF65FA6AED71FD02:8EB878D1FFC20C1FFCFFF9D1EFF5F9D1" box="[781,896,1596,1622]" country="Austria" name="Niederleis" pageId="88" pageNumber="89">Niederleis</location>
(
<collectingCountry id="F3706E9AFFC20C1FFC7DF9D1EF92F9D1" box="[911,999,1596,1622]" name="Austria" pageId="88" pageNumber="89">Austria</collectingCountry>
),
<figureCitation id="135C328FFFC20C1FFC09F9D1E82EF9D1" box="[1019,1115,1596,1622]" captionStart="FIGURE 46" captionStartId="88.[151,250,1201,1225]" captionTargetBox="[151,1435,181,1177]" captionTargetId="figure-30@88.[151,1435,181,1177]" captionTargetPageId="88" captionText="FIGURE 46. Euthria viciani Kovács, 2018. A1A2. PAL 2018.1.1. HNHM, holotype, Bánd, (Hungary). B1B. PAL 2018.2.1. HNHM, paratype, Bánd (Hungary). C. Coll.V.2017.0203, Collection Vicián, paratype, Bánd (Hungary). Euthria zachosi nov. sp. D1D2. NHMW 2023/0350/0003, holotype, Bad Vöslau (Austria). E1E2. NHMW 2023/0350/0003, paratype, Bad Vöslau (Austria). F. NHMW 2023/0347/0002, paratype, Niederleis (Austria)." figureDoi="http://doi.org/10.5281/zenodo.10841165" httpUri="https://zenodo.org/record/10841165/files/figure.png" pageId="88" pageNumber="89">
Fig.
<specimenCount id="9D61E583FFC20C1FFBDCF9D1E82EF9D1" box="[1070,1115,1596,1622]" count="46" pageId="88" pageNumber="89" type="generic">46F</specimenCount>
</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD82E0AFFC20C1FFF35F98DEF73F9FD" blockId="88.[151,1436,1523,2018]" box="[199,774,1631,1658]" pageId="88" pageNumber="89">
<materialsCitation id="3B0F2457FFC20C1FFF35F98DEF77F9FD" box="[199,770,1631,1658]" country="Austria" location="Basin" municipality="Bad Voslau" pageId="88" pageNumber="89" specimenCount="1" stateProvince="Wien" typeStatus="holotype">
<emphasis id="B913F218FFC20C1FFF35F98DED13F9FE" bold="true" box="[199,358,1631,1658]" pageId="88" pageNumber="89">
<typeStatus id="54DC90A8FFC20C1FFF35F98DED77F9FD" box="[199,258,1632,1658]" pageId="88" pageNumber="89">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="6BBCB470FFC20C1FFE9EF98DED9AF9FD" box="[364,495,1632,1658]" pageId="88" pageNumber="89">Bad Vöslau</collectingMunicipality>
(
<collectingCountry id="F3706E9AFFC20C1FFE0CF98DEE23F9FD" box="[510,598,1632,1658]" name="Austria" pageId="88" pageNumber="89">Austria</collectingCountry>
),
<collectingRegion id="49A3E0E8FFC20C1FFD9BF98DEECDF9FD" box="[617,696,1632,1658]" country="Austria" name="Wien" pageId="88" pageNumber="89">Vienna</collectingRegion>
<location id="8EB878D1FFC20C1FFD4DF98DEF77F9FD" LSID="urn:lsid:plazi:treatment:03CE9F1CFFC20C1EFF65FA6AED71FD02:8EB878D1FFC20C1FFD4DF98DEF77F9FD" box="[703,770,1632,1658]" country="Austria" municipality="Bad Voslau" name="Basin" pageId="88" pageNumber="89" stateProvince="Wien">Basin</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD82E0AFFC20C1FFF35F969EE34F919" blockId="88.[151,1436,1523,2018]" box="[199,577,1668,1694]" pageId="88" pageNumber="89">
<emphasis id="B913F218FFC20C1FFF35F969ED1AF919" bold="true" box="[199,367,1668,1694]" pageId="88" pageNumber="89">
<typeStatus id="54DC90A8FFC20C1FFF35F969ED77F919" box="[199,258,1668,1694]" pageId="88" pageNumber="89">Type</typeStatus>
stratum.
</emphasis>
Baden Formation.
</paragraph>
<paragraph id="8BD82E0AFFC20C1FFF35F94AEF60F945" blockId="88.[151,1436,1523,2018]" box="[199,789,1703,1730]" pageId="88" pageNumber="89">
<emphasis id="B913F218FFC20C1FFF35F94AEC88F946" bold="true" box="[199,253,1703,1729]" pageId="88" pageNumber="89">Age.</emphasis>
Middle Miocene, middle Badenian (Langhian).
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFC20C1FFF35F921EFE7F961" box="[199,914,1740,1766]" pageId="88" pageNumber="89" type="etymology">
<paragraph id="8BD82E0AFFC20C1FFF35F921EFE7F961" blockId="88.[151,1436,1523,2018]" box="[199,914,1740,1766]" pageId="88" pageNumber="89">
<emphasis id="B913F218FFC20C1FFF35F921ED3BF961" bold="true" box="[199,334,1740,1766]" pageId="88" pageNumber="89">Etymology.</emphasis>
In honor of Franz Zachos, zoologist at the NHMW.
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFC20C1FFF35F902EFD0F8D5" pageId="88" pageNumber="89" type="diagnosis">
<paragraph id="8BD82E0AFFC20C1FFF35F902EFD0F8D5" blockId="88.[151,1436,1523,2018]" pageId="88" pageNumber="89">
<emphasis id="B913F218FFC20C1FFF35F902ED37F88E" bold="true" box="[199,322,1775,1801]" pageId="88" pageNumber="89">Diagnosis.</emphasis>
Medium-sized, moderately broad, ovate shell with slightly coeloconoid early spire passing to cyrtoconoid on later whorls. Anal canal incised with broad, not very parietal denticle; indistinct swelling in area of anal denticle. Outer lip strongly thickened with about eight long lirae.
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFC20C1EFF35F8B6EF32FEE2" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="description">
<paragraph id="8BD82E0AFFC20C1EFF35F8B6EF32FEE2" blockId="88.[151,1436,1523,2018]" lastBlockId="89.[151,1437,151,645]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<emphasis id="B913F218FFC20C1FFF35F8B6ED2CF8F2" bold="true" box="[199,345,1883,1909]" pageId="88" pageNumber="89">Description.</emphasis>
Medium-sized, moderately broad, ovate shell of up to seven teleoconch whorls with slightly coeloconoid early spire passing to cyrtoconoid on later whorls; apical angle ~60°. Protoconch unknown. First two teleoconch whorls with weak axial ribs and delicate spiral cords. Later teleoconch whorls flat-sided, smooth, separated by indistinct suture. Last whorl smooth, ovate, broad, very high, attaining ~80% of total height, subsutural ramp slightly concave, hardly delimited, broadly convex below; base moderately constricted bearing distinct spiral cords; fasciole slightly swollen.Aperture moderately small, ovate. Columella broadly and weakly excavated, weakly angled at transition to siphonal canal. Columellar callus weakly delimited. Anal canal incised, accentuated by broad, not very parietal denticle. Indistinct swelling in area of anal denticle. Outer lip strongly thickened adapically, thinning below, with about eight long lirae starting some distance behind peristome. Siphonal canal moderately long, relatively wide, deflected to the left, shallowly notched.
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFC30C1EFF35FE83EFDBFDBA" pageId="89" pageNumber="90" type="discussion">
<paragraph id="8BD82E0AFFC30C1EFF35FE83E8EFFE72" blockId="89.[151,1437,151,645]" pageId="89" pageNumber="90">
<emphasis id="B913F218FFC30C1EFF35FE83ED39FE0F" bold="true" box="[199,332,366,392]" pageId="89" pageNumber="90">Discussion.</emphasis>
The specimens are smaller than
<taxonomicName id="4C675589FFC30C1EFD4BFE82E877FE0E" authority="Kovacs, 2018" authorityName="Kovacs" authorityYear="2018" box="[697,1026,367,393]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="viciani">
<emphasis id="B913F218FFC30C1EFD4BFE82EF14FE0F" box="[697,865,367,393]" italics="true" pageId="89" pageNumber="90">Euthria viciani</emphasis>
<bibRefCitation id="EFF653FBFFC30C1EFC9AFE82E877FE0E" author="Kovacs, Z." box="[872,1026,367,393]" pageId="89" pageNumber="90" pagination="179 - 182" refId="ref86477" refString="Kovacs, Z. (2018) New records of the genus Euthria (Mollusca, Buccinidae) in the Miocene Paratethys. Foldtani Kozlony, 148 (2), 179 - 182. https: // doi. org / 10.23928 / foldt. kozl. 2018.148.2.179" type="journal article" year="2018">Kovács, 2018</bibRefCitation>
</taxonomicName>
and lack the typical color pattern. In addition, the adapical part of the siphonal canal is wider in
<taxonomicName id="4C675589FFC30C1EFCC7FE7EEF93FE2A" authorityName="Harzhauser &amp; Landau" authorityYear="2024" box="[821,998,403,429]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="zachosi">
<emphasis id="B913F218FFC30C1EFCC7FE7EEF93FE2A" box="[821,998,403,429]" italics="true" pageId="89" pageNumber="90">Euthria zachosi</emphasis>
</taxonomicName>
but subparallel in
<taxonomicName id="4C675589FFC30C1EFB4BFE7EE917FE2B" authorityName="Kovacs" authorityYear="2018" box="[1209,1378,403,429]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="viciani">
<emphasis id="B913F218FFC30C1EFB4BFE7EE917FE2B" box="[1209,1378,403,429]" italics="true" pageId="89" pageNumber="90">Euthria viciani</emphasis>
</taxonomicName>
. The base and siphonal canal are sculptured with broad, but low, spiral cords whereas
<taxonomicName id="4C675589FFC30C1EFBDAFE5AE8A4FE57" authorityName="Kovacs" authorityYear="2018" box="[1064,1233,439,465]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="viciani">
<emphasis id="B913F218FFC30C1EFBDAFE5AE8A4FE57" box="[1064,1233,439,465]" italics="true" pageId="89" pageNumber="90">Euthria viciani</emphasis>
</taxonomicName>
has a smooth and glossy base. Lirae in
<taxonomicName id="4C675589FFC30C1EFE76FE36EE58FE73" authorityName="Kovacs" authorityYear="2018" box="[388,557,475,501]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="viciani">
<emphasis id="B913F218FFC30C1EFE76FE36EE58FE73" box="[388,557,475,501]" italics="true" pageId="89" pageNumber="90">Euthria viciani</emphasis>
</taxonomicName>
are short but reach far into aperture in
<taxonomicName id="4C675589FFC30C1EFC11FE36E8E6FE72" authorityName="Harzhauser &amp; Landau" authorityYear="2024" box="[995,1171,475,501]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="zachosi">
<emphasis id="B913F218FFC30C1EFC11FE36E8E6FE72" box="[995,1171,475,501]" italics="true" pageId="89" pageNumber="90">Euthria zachosi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD82E0AFFC30C1EFF35FE13EFDBFDBA" blockId="89.[151,1437,151,645]" pageId="89" pageNumber="90">
<emphasis id="B913F218FFC30C1EFF35FE13EDDDFD9F" bold="true" box="[199,424,510,536]" pageId="89" pageNumber="90">Paleoenvironment.</emphasis>
The occurrences in the Baden Formation of the
<collectingRegion id="49A3E0E8FFC30C1EFC14FE12E840FD9E" box="[998,1077,511,537]" country="Austria" name="Wien" pageId="89" pageNumber="90">Vienna</collectingRegion>
Basin suggest middle to outer neritic environments in up to
<quantity id="4C9F83EFFFC30C1EFE10FDCFEE5CFDBA" box="[482,553,546,573]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="89" pageNumber="90" unit="m" value="250.0">250 m</quantity>
water depth (
<bibRefCitation id="EFF653FBFFC30C1EFD33FDCEEFD4FDBA" author="Kranner, M. &amp; Harzhauser, M. &amp; Mandic, O. &amp; Strauss, P. &amp; Siedl, W. &amp; Piller, W. E." box="[705,929,547,573]" pageId="89" pageNumber="90" pagination="105187" refId="ref86893" refString="Kranner, M., Harzhauser, M., Mandic, O., Strauss, P., Siedl, W. &amp; Piller, W. E. (2021) Early and Middle Miocene paleobathymetry of the Vienna Basin (Austria). Marine and Petroleum Geology, 132, 105187. https: // doi. org / 10.1016 / j. marpetgeo. 2021.105187" type="journal article" year="2021">
Kranner
<emphasis id="B913F218FFC30C1EFCD4FDCEEF2BFDBA" box="[806,862,547,573]" italics="true" pageId="89" pageNumber="90">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFC30C1EFF35FDABED71FD02" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8BD82E0AFFC30C1EFF35FDABED71FD02" blockId="89.[151,1437,151,645]" pageId="89" pageNumber="90">
<emphasis id="B913F218FFC30C1EFF35FDABEE0CFDE6" bold="true" box="[199,633,582,609]" pageId="89" pageNumber="90">Distribution in Central Paratethys.</emphasis>
Badenian (Middle Miocene):
<emphasis id="B913F218FFC30C1EFC12FDAAE8F7FDE7" box="[992,1154,583,608]" italics="true" pageId="89" pageNumber="90">
<collectingRegion id="49A3E0E8FFC30C1EFC12FDAAE858FDE7" box="[992,1069,583,608]" country="Austria" name="Wien" pageId="89" pageNumber="90">Vienna</collectingRegion>
Basin:
</emphasis>
Niederleis, Bad Vöslau (
<collectingCountry id="F3706E9AFFC30C1EFF6DFD86EC8DFD02" box="[159,248,619,645]" name="Austria" pageId="89" pageNumber="90">Austria</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>