244 lines
27 KiB
XML
244 lines
27 KiB
XML
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<mods:title id="873E5F8528216D09A1F6BE902078C617">Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth-death process</mods:title>
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<mods:namePart id="BE189315747FB119151DC626FBABC6E5">Jouault, Corentin</mods:namePart>
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<mods:namePart id="47805A088D30AAC9002B382E6BF218F6">Maréchal, Arthur</mods:namePart>
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<mods:namePart id="A2E51ECB027BAEC8F6F37419735A3399">Condamine, Fabien L.</mods:namePart>
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<mods:namePart id="8D87075C63063AAC2D3A2301A8E1D236">Wang, Bo</mods:namePart>
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<mods:namePart id="C6793626B4AC125A38DA6C1F2C5F3E61">Nel, André</mods:namePart>
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<mods:namePart id="FB9F7FDBBE4C5A748CE5EE4B7B8366DF">Legendre, Frédéric</mods:namePart>
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<mods:namePart id="5FB8164DA29B2FC02DC3FE71A7BA43B0">Perrichot, Vincent</mods:namePart>
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<mods:typeOfResource id="F598B1CA2AF487005E5CE15ECA2C70A9">text</mods:typeOfResource>
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<mods:title id="6959A84A4BF93B921E5AAB0C6D2B0719">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="0B17CEBDA62D1FD74FC460D759C33736">
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<mods:date id="1529DBB35B21352F336EC226A049EECD">2022</mods:date>
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<mods:number id="3AF17671293FC537062904A4AAA09BB3">2021-07-13</mods:number>
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<mods:number id="1342D91E3CC595BE4A6FB63E1EADD05A">194</mods:number>
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<mods:url id="275D54DF4C04C52984439AABC89BFFAD">https://doi.org/10.1093/zoolinnean/zlab034</mods:url>
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<mods:identifier id="D3E56DF703297E23754FDE44758D119D" type="DOI">10.1093/zoolinnean/zlab034</mods:identifier>
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<treatment id="03CE87D7FFFF4721DCE4C077FC467975" ID-DOI="http://doi.org/10.5281/zenodo.6458545" ID-GBIF-Taxon="194594350" ID-Zenodo-Dep="6458545" LSID="urn:lsid:plazi:treatment:03CE87D7FFFF4721DCE4C077FC467975" httpUri="http://treatment.plazi.org/id/03CE87D7FFFF4721DCE4C077FC467975" lastPageId="9" lastPageNumber="1405" pageId="7" pageNumber="1403">
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<subSubSection id="C37D654AFFFF472FDCE4C077FDC57910" pageId="7" pageNumber="1403" type="nomenclature">
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<paragraph id="8BD836C1FFFF472FDCE4C077FDC57910" blockId="7.[157,749,561,616]" pageId="7" pageNumber="1403">
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<heading id="D09081ADFFFF472FDCE4C077FDC57910" centered="true" fontSize="9" level="2" pageId="7" pageNumber="1403" reason="3">
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<taxonomicName id="4C674D42FFFF472FDCE4C077FDB3791F" authority="Jouault & Maréchal & Wang & Perrichot, 2021" authorityName="Jouault & Maréchal & Wang & Perrichot" authorityYear="2021" class="Insecta" family="Othniodellithidae" genus="Keratodellitha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="1403" phylum="Arthropoda" rank="species" species="basilisci" status="sp. nov.">
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<emphasis id="B913EAD3FFFF472FDCE4C077FE58793F" bold="true" box="[157,493,561,584]" italics="true" pageId="7" pageNumber="1403">KERATODELLITHA BASILISCI</emphasis>
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JOUAULT, MARÉCHAL, WANG & PERRICHOT
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</taxonomicName>
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<emphasis id="B913EAD3FFFF472FDE77C015FDDC791E" bold="true" box="[526,617,591,615]" pageId="7" pageNumber="1403">SP. NOV</emphasis>
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.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D654AFFFF472FDDF0C030FD5F79AB" pageId="7" pageNumber="1403" type="description">
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<paragraph id="8BD836C1FFFF472FDDF0C030FDB579F7" blockId="7.[393,512,630,655]" box="[393,512,630,655]" pageId="7" pageNumber="1403">
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(
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<figureCitation id="135C2A44FFFF472FDDEBC030FE6979F7" box="[402,476,630,655]" captionStart="Figure S" captionStartId="27.[145,231,290,312]" captionText="Figure S1. Strict cladogram generated from parsimony analysis. Tree length, 829 steps; consistency index (CI), 0.1496; homoplasy index (HI), 0.8504; retention index (RI), 0.6597). Values above branches represent bootstrap percentages> 50% (majority-rule consensus). Appendix S1. Nexus files + trees + summary post analyses. Appendix S2. Calibration and ages used in tip-dating analyses." pageId="7" pageNumber="1403">FIGS 1</figureCitation>
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,
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<figureCitation id="135C2A44FFFF472FDD93C030FE4D79F7" box="[490,504,630,655]" captionStart="Figure 2" captionStartId="8.[163,241,1805,1827]" captionTargetBox="[211,1396,197,1764]" captionTargetId="figure-26@8.[209,1396,195,1765]" captionTargetPageId="8" captionText="Figure 2. Keratodellitha basilisci. Holotype NIGP174738 (female).A, head in full-face view. B, head in right profile view. C, head in dorsal view. D, wings. E, line drawing of wing venation with nomenclature. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6458543" httpUri="https://zenodo.org/record/6458543/files/figure.png" pageId="7" pageNumber="1403">2</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD836C1FFFF472FDCE8C0D9FD5F79AB" blockId="7.[145,762,670,723]" pageId="7" pageNumber="1403">
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<emphasis id="B913EAD3FFFF472FDCE8C0D9FE6A79CC" box="[145,479,670,692]" italics="true" pageId="7" pageNumber="1403">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n: l s i d: z o o b a n k. org:act:
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<uuid id="FFC10C14FFFF472FDC99C0FBFD5F79AB" box="[224,746,701,723]" pageId="7" pageNumber="1403">C67E5240-F1BC-4836-86F7-3B9D1D375AA8</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D654AFFFF472FDCE8C0B0FB357A60" pageId="7" pageNumber="1403" type="materials_examined">
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<paragraph id="8BD836C1FFFF472FDCE8C0B0FD607853" blockId="7.[145,760,758,811]" pageId="7" pageNumber="1403">
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<materialsCitation id="3B0F3C9CFFFF472FDCE8C0B0FD607853" ID-GBIF-Occurrence="3751309301" collectionCode="NIGP" pageId="7" pageNumber="1403" specimenCode="NIGP174738" specimenCount="1" typeStatus="holotype">
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<emphasis id="B913EAD3FFFF472FDCE8C0B0FE127874" box="[145,423,758,780]" italics="true" pageId="7" pageNumber="1403">
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<typeStatus id="54DC8863FFFF472FDCE8C0B0FF4F7873" box="[145,250,758,779]" pageId="7" pageNumber="1403" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DBC19EBAFFFF472FDD77C0B1FE127874" box="[270,423,759,780]" pageId="7" pageNumber="1403">NIGP174738</specimenCode>
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</emphasis>
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, preserved in a rectangular piece of amber measuring
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<quantity id="4C9F9B24FFFF472FDDC5C153FDBF7853" box="[444,522,789,811]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="1403" unit="mm" value="17.0">17 mm</quantity>
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||
×
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<quantity id="4C9F9B24FFFF472FDE5CC153FDC67853" box="[549,627,789,811]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="1403" unit="mm" value="10.0">10 mm</quantity>
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×
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<quantity id="4C9F9B24FFFF472FDEF7C153FD647853" box="[654,721,789,811]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="1403" unit="mm" value="6.0">6 mm</quantity>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD836C1FFFF472FDF50C280FB357A60" blockId="7.[809,1425,197,280]" pageId="7" pageNumber="1403">
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<emphasis id="B913EAD3FFFF472FDF50C280FBE47BA2" box="[809,1105,197,219]" italics="true" pageId="7" pageNumber="1403">
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<typeStatus id="54DC8863FFFF472FDF50C280FCEB7BA3" box="[809,862,198,219]" pageId="7" pageNumber="1403">Type</typeStatus>
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locality and horizon:
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</emphasis>
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Noije Bum Hill, Hukawng Valley,
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<collectingRegion id="49A3F823FFFF472FDF00C2A2FBA57B81" box="[889,1040,228,250]" country="Myanmar" name="Kachin" pageId="7" pageNumber="1403">Kachin State</collectingRegion>
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,
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<collectingCountry id="F3707651FFFF472FD865C2A2FB3B7B81" box="[1052,1166,228,249]" name="Myanmar" pageId="7" pageNumber="1403">Myanmar</collectingCountry>
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; upper Albian to lower Cenomanian, mid-Cretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D654AFFFF472FDF50C307FAA37AA9" pageId="7" pageNumber="1403" type="etymology">
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<paragraph id="8BD836C1FFFF472FDF50C307FAA37AA9" blockId="7.[809,1425,321,466]" pageId="7" pageNumber="1403">
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<emphasis id="B913EAD3FFFF472FDF50C307FC1D7A2E" box="[809,936,321,342]" italics="true" pageId="7" pageNumber="1403">Etymology:</emphasis>
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The specific epithet refers to the basilisk (Latin
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<emphasis id="B913EAD3FFFF472FDF0CC326FC517A0D" box="[885,996,352,373]" italics="true" pageId="7" pageNumber="1403">basiliscus</emphasis>
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), a mythical animal that intermingled the traits of several animals, and is a general allusion to the fierce and enigmatic habitus of the species. The specific epithet is a noun in a genitive case.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D654AFFFF472FDF50C045FC407997" pageId="7" pageNumber="1403" type="diagnosis">
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<paragraph id="8BD836C1FFFF472FDF50C045FC407997" blockId="7.[809,1425,514,751]" pageId="7" pageNumber="1403">
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<emphasis id="B913EAD3FFFF472FDF50C045FC157960" box="[809,928,515,536]" italics="true" pageId="7" pageNumber="1403">Diagnosis:</emphasis>
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Scape long, ~3.5 times longer than pedicel; outer (external) pretarsal claw of foreleg with two preapical teeth; fore wing vein Rs+M as long as 2Rs; third submarginal cell as long as second one; 1rs-m meeting Rs slightly before r-rs; 2rs-m tubular; second discal cell short (twice as long as wide); hind wing with six hamuli; gaster laterally compressed (as in
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<taxonomicName id="4C674D42FFFF472FD976C0FCFA3079A8" box="[1295,1413,698,720]" class="Insecta" family="Evaniidae" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="1403" phylum="Arthropoda" rank="family">Evaniidae</taxonomicName>
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), rounded in shape.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D654AFFFF4721DF50C166FC467975" lastPageId="9" lastPageNumber="1405" pageId="7" pageNumber="1403" type="description">
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<paragraph id="8BD836C1FFFF472FDF50C166FBB97F53" blockId="7.[809,1425,800,1864]" pageId="7" pageNumber="1403">
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<emphasis id="B913EAD3FFFF472FDF50C166FC74784D" box="[809,961,800,821]" italics="true" pageId="7" pageNumber="1403">Description:</emphasis>
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Female. Total length as preserved
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<quantity id="4C9F9B24FFFF472FDF50C179FCC5782C" box="[809,880,831,852]" metricMagnitude="0" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="1403" unit="m" value="4.4">4.4 m</quantity>
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m (e x c l u d i n g a n t e n n a e a n d o v i p o s i t o r); fore wing
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<quantity id="4C9F9B24FFFF472FDFE0C11BFC5B780B" box="[921,1006,861,883]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="1403" unit="mm" value="2.7">2.7 mm</quantity>
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||
long; integument dark brown except metasoma lighter; body largely glabrous except legs bearing short, thick setae; integument without pronounced sculpturing or punctation, largely finely imbricate; propodeum coarsely and strongly areolate; wings clear and hyaline, veins brown to light brown in preserved colour.
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</paragraph>
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<paragraph id="8BD836C1FFFF472FDF38C672FB157C30" blockId="7.[809,1425,800,1864]" pageId="7" pageNumber="1403">
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Head slightly shorter than wide in frontal view, width
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and length
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; genal width
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||
<quantity id="4C9F9B24FFFF472FDF50C637FC307FFE" box="[809,901,1137,1158]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="1403" unit="mm" value="0.2">0.2 mm</quantity>
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; compound eyes ovoid, without prominent circumocular carina,
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||
<quantity id="4C9F9B24FFFF472FD84DC6D6FB287FDD" box="[1076,1181,1168,1190]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="1403" unit="mm" value="0.51">0.51 mm</quantity>
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long,
|
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<quantity id="4C9F9B24FFFF472FD895C6D6FAF37FDD" box="[1260,1350,1168,1189]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="1403" unit="mm" value="0.3">0.3 mm</quantity>
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wide; ocelli present and situated in a small triangle on top of vertex above compound eyes and immediately behind facial prominence, ocelli separated by approximately their diameter and from posterior border of head by about twice their diameter. Prominent facial horn projecting to a length of
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||
<quantity id="4C9F9B24FFFF472FD819C70EFB7D7E25" box="[1120,1224,1352,1373]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="1403" unit="mm" value="0.37">0.37 mm</quantity>
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in profile (from anterior toruli margin to apex of the horn). Antenna with scape ~3.5 times pedicel length,
|
||
<quantity id="4C9F9B24FFFF472FD893C7C3FAE57EE2" box="[1258,1360,1413,1434]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="7" pageNumber="1403" unit="mm" value="0.43">0.43 mm</quantity>
|
||
long,
|
||
<quantity id="4C9F9B24FFFF472FDF50C7E2FC247EC1" box="[809,913,1444,1465]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="7" pageNumber="1403" unit="mm" value="0.07">0.07 mm</quantity>
|
||
wide; pedicel more than twice as long as wide; flagellomeres each longer than wide except two apical most; flagellomere I the longest (
|
||
<quantity id="4C9F9B24FFFF472FD952C7A7FA257E8E" box="[1323,1424,1505,1526]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="1403" unit="mm" value="0.31">0.31 mm</quantity>
|
||
long). Clypeal base indistinct from frons (epistomal sulcus lacking), with strong medial longitudinal ridge, ridge extends posteriorly with a small sharp tooth and anteriorly projects as small triangular prominence between mandibles. Palp formula (maxillary–labial) at least 6-3. Mandible massive, square shaped in frontal view, with straight outer and apical margins meeting in a rounded orthogonal angle; inner margin with ridges demarcating blunt ‘teeth’, apical tooth well projected without overlapping opposing mandible.
|
||
</paragraph>
|
||
<caption id="DF186649FFFF472FDCE8C4B1FDC87C30" ID-DOI="http://doi.org/10.5281/zenodo.6458541" ID-Zenodo-Dep="6458541" httpUri="https://zenodo.org/record/6458541/files/figure.png" pageId="7" pageNumber="1403" startId="7.[145,161,1783,1804]" targetBox="[145,766,848,1744]" targetPageId="7">
|
||
<paragraph id="8BD836C1FFFF472FDCE8C4B1FDC87C30" blockId="7.[145,760,1783,1864]" pageId="7" pageNumber="1403">
|
||
<emphasis id="B913EAD3FFFF472FDCE8C4B1FDC67C74" bold="true" box="[145,627,1783,1805]" pageId="7" pageNumber="1403">
|
||
F i g u r e 1.
|
||
<emphasis id="B913EAD3FFFF472FDD58C4BEFDC67C74" bold="true" box="[289,627,1783,1805]" italics="true" pageId="7" pageNumber="1403">K e r a t o d e l l i t h a b a s i l i s c i</emphasis>
|
||
</emphasis>
|
||
. H o l o t y p e
|
||
<emphasis id="B913EAD3FFFF472FDCE8C553FEA97C53" box="[145,284,1813,1835]" italics="true" pageId="7" pageNumber="1403">NIGP174738</emphasis>
|
||
(female). A, habitus in right lateral view. B, habitus in left lateral view. Scale bars:
|
||
<quantity id="4C9F9B24FFFF472FDE5EC574FDCD7C30" box="[551,632,1842,1864]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="1403" unit="mm" value="0.5">0.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF186649FFF04720DCDAC54BFB457C38" ID-DOI="http://doi.org/10.5281/zenodo.6458543" ID-Zenodo-Dep="6458543" httpUri="https://zenodo.org/record/6458543/files/figure.png" pageId="8" pageNumber="1404" startId="8.[163,241,1805,1827]" targetBox="[211,1396,197,1764]" targetPageId="8">
|
||
<paragraph id="8BD836C1FFF04720DCDAC54BFB457C38" blockId="8.[163,1444,1805,1856]" pageId="8" pageNumber="1404">
|
||
<emphasis id="B913EAD3FFF04720DCDAC54BFD9D7C5B" bold="true" box="[163,552,1805,1827]" pageId="8" pageNumber="1404">
|
||
Figure 2.
|
||
<taxonomicName id="4C674D42FFF04720DD6CC54BFD9D7C5B" authorityName="Jouault, Marechal, Wang & Perrichot" authorityYear="2022" box="[277,552,1805,1827]" class="Insecta" family="Othniodellithidae" genus="Keratodellitha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1404" phylum="Arthropoda" rank="species" species="basilisci">
|
||
<emphasis id="B913EAD3FFF04720DD6CC54BFD9D7C5B" bold="true" box="[277,552,1805,1827]" italics="true" pageId="8" pageNumber="1404">Keratodellitha basilisci</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
. Holotype
|
||
<emphasis id="B913EAD3FFF04720DEEFC54BFCAB7C5B" box="[662,798,1805,1827]" italics="true" pageId="8" pageNumber="1404">NIGP174738</emphasis>
|
||
(female). A, head in full-face view. B, head in right profile view. C, head in dorsal view. D, wings. E, line drawing of wing venation with nomenclature. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836C1FFF14721DCD0C280FD4478C3" blockId="9.[145,761,197,1905]" pageId="9" pageNumber="1405">
|
||
Mesosoma laterally compressed, longer than high, length
|
||
<quantity id="4C9F9B24FFF14721DC98C2A2FE8D7B81" box="[225,312,228,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="1405" unit="mm" value="2.2">2.2 mm</quantity>
|
||
, height
|
||
<quantity id="4C9F9B24FFF14721DDEDC2A2FE4C7B81" box="[404,505,228,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="9" pageNumber="1405" unit="mm" value="1.41">1.41 mm</quantity>
|
||
; pronotum prominent, with well-developed concave anterior surface, with poorly defined raised posterior surface, medially longer than wide, lateral surfaces large and slightly depressed; propleura long but not projecting anterior to pronotum; mesoscutum
|
||
<quantity id="4C9F9B24FFF14721DD9DC33BFDFB7AEB" box="[484,590,381,403]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="1405" unit="mm" value="0.53">0.53 mm</quantity>
|
||
long, notauli distinct; mesoscutellum convex in profile,
|
||
<quantity id="4C9F9B24FFF14721DEEBC3DAFD4D7AC9" box="[658,760,412,433]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="1405" unit="mm" value="0.27">0.27 mm</quantity>
|
||
in its medial length; mesopleuron with small, oblique row of faint areolae demarcating border with pronotal lateral surface and more distinct, single row of areolae along border with metepisternum; propodeum weakly areolate, with prominent dorsal surface, dorsal length between metanotum and articulation with petiole equal to
|
||
<quantity id="4C9F9B24FFF14721DD30C035FE1479F0" box="[329,417,627,648]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="1405" unit="mm" value="0.3">0.3 mm</quantity>
|
||
. Legs long, with slender tarsi; meso- and metacoxae closer to each other than to procoxa; metafemur
|
||
<quantity id="4C9F9B24FFF14721DD02C0F6FE6879BD" box="[379,477,688,709]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="9" pageNumber="1405" unit="mm" value="1.81">1.81 mm</quantity>
|
||
long, metatibia
|
||
<quantity id="4C9F9B24FFF14721DEEEC0F6FD4C79BD" box="[663,761,688,709]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="9" pageNumber="1405" unit="mm" value="1.71">1.71 mm</quantity>
|
||
long; metabasitarsus elongate but slightly shorter than combined lengths of remaining tarsomeres, metabasitarsus length
|
||
<quantity id="4C9F9B24FFF14721DDD9C14AFE4D7859" box="[416,504,780,801]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="1405" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, lengths of remaining tarsomeres (from base to apex), 0.34, 0.3, 0.16 and
|
||
<quantity id="4C9F9B24FFF14721DCE8C10FFF4E7827" box="[145,251,841,863]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="1405" unit="mm" value="0.23">0.23 mm</quantity>
|
||
. Formula of tibial spurs 1-2-2. Apex of all tarsi with a plantula. Outer pretarsal claw of foreleg tridentate (bearing two small pre-apical teeth), all other claws bidentate (with a single pre-apical tooth).
|
||
</paragraph>
|
||
<paragraph id="8BD836C1FFF14721DCD0C182FDCB7C09" blockId="9.[145,761,197,1905]" pageId="9" pageNumber="1405">Fore wing with costal space apically slightly broader than pterostigma; pterostigma longer than wide, tapering gradually in width to acute apex, margin inside marginal cell faintly convex; vein M+Cu forking after Sc+R midlength; 1M straight, nearly aligned with and longer than 1Rs; 1Rs originating prior to pterostigma a distance inferior to its length; 1rs-m meeting Rs slightly before r-rs; 2Rs longer than 1Rs; 2M small, meeting 1m-cu, then M abscissa extending toward wing apex; 2Rs slightly curved; r-rs originating from basal third of pterostigmal length, elongate, softly sinuous, as long as 2Rs; marginal cell broad, three time as long as wide, greatest width near tangent of pterostigmal apex; 3Rs mostly straight for majority of marginal cell length; two rs-m cross-veins present; second submarginal cell about as long as third and slightly widening towards the wing margin; third submarginal cell slightly wider posteriorly than anteriorly; first discal cell almost forming strongly slanted rhomboid (not distinctly pentagonal owing to exceptionally short 2M); 1m-cu straight and subparallel to 1M; cu-a originating at M+Cu fork, nearly aligned with 1M; 2m-cu present but weaker than surrounding veins; A2 present and meeting Cu apically in a small 2cu-a vein (demarcating second subbasal cell); vein 2A present. Hind wing with venation nearly complete, lacking only C; R with six distal hamuli and not meeting 2Rs apically; rs-m oblique, about twice as long as 1Rs, shorter than first free abscissa M; Cu+cu-a present, nearly orthogonal to M+Cu; apical abscissa Cu not visible if present; jugal lobe lacking.</paragraph>
|
||
<paragraph id="8BD836C1FFF14721DF38C283FB9C7A97" blockId="9.[809,1425,197,525]" pageId="9" pageNumber="1405">
|
||
Metasoma with first segment forming tubular petiole
|
||
<quantity id="4C9F9B24FFF14721DFFEC2A5FC457B81" box="[903,1008,227,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="9" pageNumber="1405" unit="mm" value="0.57">0.57 mm</quantity>
|
||
long, tergum and sternum fused without apparent indication of individual sclerites; gaster
|
||
<quantity id="4C9F9B24FFF14721DF05C367FC577A4E" box="[892,994,289,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="9" pageNumber="1405" unit="mm" value="1.72">1.72 mm</quantity>
|
||
long, compressed laterally, ovoid in profile, obviously longer than high, maximum height
|
||
<quantity id="4C9F9B24FFF14721DF50C318FC3A7A0C" box="[809,911,350,372]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="9" pageNumber="1405" unit="mm" value="0.84">0.84 mm</quantity>
|
||
, widening to the third gastral segment and then narrowing toward apex; ovipositor exerted, ≥
|
||
<quantity id="4C9F9B24FFF14721DF45C3DAFCC97AC9" box="[828,892,412,433]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="1405" unit="mm" value="1.0">1 mm</quantity>
|
||
long (not totally visible), with five minute teeth present on apical declivitous face; sheaths slightly longer than ovipositor.
|
||
</paragraph>
|
||
<paragraph id="8BD836C1FFF14721DF38C3B1FC467975" blockId="9.[809,1425,197,525]" box="[833,1011,503,525]" pageId="9" pageNumber="1405">Male unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |