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<document id="020BBBAF75C1124527154E45A6AC29AA" ID-DOI="10.11646/zootaxa.3635.1.9" ID-GBIF-Dataset="c2e1b77b-2a89-46f8-8b24-64ae71254792" ID-ISSN="1175-5326" ID-Zenodo-Dep="283539" ID-ZooBank="3B9C28A8-B64D-46AB-BE68-BF6CA85DBCAF" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460585776088" checkinUser="plazi" docAuthor="Shear, William A." docDate="2013" docId="03CE87AC9906FFD2F0C31157FE3964FD" docLanguage="en" docName="zt03635p093.pdf" docOrigin="Zootaxa 3635 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cleidogona austrina Loomis" docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="FFF7FFD49904FFD1F054164EFFBE657A" masterDocTitle="A new miniature species of the milliped genus Cleidogona from Costa Rica (Diplopoda, Chordeumatida, Cleidogonidae), with a review of Central American Cleidogonidae" masterLastPageNumber="93" masterPageNumber="87" pageNumber="89" updateTime="1698325116444" updateUser="plazi">
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<mods:title id="9BE8E2F27E8B2537DEA42E74AD82534C">A new miniature species of the milliped genus Cleidogona from Costa Rica (Diplopoda, Chordeumatida, Cleidogonidae), with a review of Central American Cleidogonidae</mods:title>
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<mods:name id="5BAD6BA4192ADABB4C46B38FF15463B4" type="personal">
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<mods:namePart id="FCA46E95F9738E49B6E1108C7F9D529C">Shear, William A.</mods:namePart>
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<mods:title id="5DE518A849C73E0EE31C2A1C35D16C76">Zootaxa</mods:title>
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<mods:date id="F2EC31147164052BA96A09509F575DC3">2013</mods:date>
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<mods:identifier id="D09B575741552F45237F0E2BDC5AD56A" type="DOI">10.11646/zootaxa.3635.1.9</mods:identifier>
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<treatment id="03CE87AC9906FFD2F0C31157FE3964FD" ID-DOI="http://doi.org/10.5281/zenodo.6164048" ID-GBIF-Taxon="127663651" ID-Zenodo-Dep="6164048" LSID="urn:lsid:plazi:treatment:03CE87AC9906FFD2F0C31157FE3964FD" httpUri="http://treatment.plazi.org/id/03CE87AC9906FFD2F0C31157FE3964FD" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
<subSubSection id="C37D65319906FFD3F0C31157FDFF62EE" pageId="2" pageNumber="89" type="nomenclature">
<paragraph id="8BD836BA9906FFD3F0C31157FDBB6248" blockId="2.[151,517,1816,1843]" box="[151,517,1816,1843]" pageId="2" pageNumber="89">
<heading id="D09081D69906FFD3F0C31157FDBB6248" bold="true" box="[151,517,1816,1843]" fontSize="11" level="1" pageId="2" pageNumber="89" reason="1">
<taxonomicName id="4C674D399906FFD3F0C31157FDBB6248" ID-CoL="5Z8XM" authority="Loomis" authorityName="Loomis" box="[151,517,1816,1843]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="austrina">
<emphasis id="B913EAA89906FFD3F0C31157FDBB6248" bold="true" box="[151,517,1816,1843]" pageId="2" pageNumber="89">
<emphasis id="B913EAA89906FFD3F0C31157FE306249" bold="true" box="[151,398,1817,1843]" italics="true" pageId="2" pageNumber="89">Cleidogona austrina</emphasis>
(Loomis)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD836BA9906FFD3F0C3112EFDFF62EE" blockId="2.[151,577,1888,1941]" pageId="2" pageNumber="89">
<taxonomicName id="4C674D399906FFD3F0C3112EFE48620C" authority="Loomis, 1964" authorityName="Loomis" authorityYear="1964" box="[151,502,1888,1910]" class="Diplopoda" family="Cleidogonidae" genus="Akacandra" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="austrina">
<emphasis id="B913EAA89906FFD3F0C3112EFEDF620C" box="[151,353,1888,1910]" italics="true" pageId="2" pageNumber="89">Akacandra austrina</emphasis>
Loomis, 1964
</taxonomicName>
, p. 99.
<taxonomicName id="4C674D399906FFD3F0C31131FE4F62EE" authority="Shear, 1972" authorityName="Shear" authorityYear="1972" box="[151,497,1918,1941]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="austrina">
<emphasis id="B913EAA89906FFD3F0C31131FECE62EF" box="[151,368,1919,1941]" italics="true" pageId="2" pageNumber="89">Cleidogona austrina:</emphasis>
Shear, 1972
</taxonomicName>
, p. 158.
</paragraph>
</subSubSection>
<subSubSection id="C37D65319906FFD2F0C3118CFE3964FD" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89" type="materials_examined">
<paragraph id="8BD836BA9906FFD3F0C3118CFCF162A3" blockId="2.[151,847,1985,2010]" box="[151,847,1985,2010]" pageId="2" pageNumber="89">
<emphasis id="B913EAA89906FFD3F0C3118CFE1962A0" bold="true" box="[151,423,1985,2010]" pageId="2" pageNumber="89">
<typeStatus id="54DC88189906FFD3F0C3118CFF6C62A0" box="[151,210,1986,2010]" pageId="2" pageNumber="89">Type</typeStatus>
and only locality:
</emphasis>
Cerro Punta, Chiriqui Prov.,
<collectingCountry id="F370762A9906FFD3F2BB118CFCF562A3" box="[751,843,1986,2009]" name="Panama" pageId="2" pageNumber="89">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8BD836BA9907FFD2F09316D6FE3964FD" blockId="3.[151,1436,151,391]" pageId="3" pageNumber="90">
At the
<typeStatus id="54DC88189907FFD2F14A16D6FEF065CA" box="[286,334,152,176]" pageId="3" pageNumber="90">type</typeStatus>
locality, this species would be sympatric with two species of
<emphasis id="B913EAA89907FFD2F46B16D6FA8165CA" box="[1087,1343,152,176]" italics="true" pageId="3" pageNumber="90">
<taxonomicName id="4C674D399907FFD2F46B16D6FB2165CA" box="[1087,1183,152,176]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Dybasia</taxonomicName>
,
<taxonomicName id="4C674D399907FFD2F4E416D7FA8165CA" box="[1200,1343,152,176]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="chiriquia">D. chiriquia</taxonomicName>
</emphasis>
and
<emphasis id="B913EAA89907FFD2F5D516D7FEB765AE" italics="true" pageId="3" pageNumber="90">
<taxonomicName id="4C674D399907FFD2F5D516D7FEB865AE" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="divergens">D. divergens</taxonomicName>
,
</emphasis>
but Loomis records of the latter two from Cerro Punta are based on females and juveniles and all three species are close in general appearance and size. Unfortunately the
<typeStatus id="54DC88189907FFD2F3FC1691FBB4658D" box="[936,1034,223,247]" pageId="3" pageNumber="90" type="holotype">holotype</typeStatus>
is the only available male and the gonopods are not to be found in the vial (Shear 1972). The illustrations in the original description, especially of the ninth legpair, suggest that
<taxonomicName id="4C674D399907FFD2F1911766FDF8643A" box="[453,582,296,320]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="austrina">
<emphasis id="B913EAA89907FFD2F1911766FDF8643A" box="[453,582,296,320]" italics="true" pageId="3" pageNumber="90">C. austrina</emphasis>
</taxonomicName>
might eventually find a better home in
<emphasis id="B913EAA89907FFD2F4441766FBCB643A" box="[1040,1141,296,320]" italics="true" pageId="3" pageNumber="90">
<taxonomicName id="4C674D399907FFD2F4441766FBCF643A" box="[1040,1137,296,320]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Dybasia</taxonomicName>
.
</emphasis>
I did not place it there in 1972 because I was not able to study the gonopods, which Loomis depicted as very small, very simple, and lacking colpocoxites entirely.
</paragraph>
</subSubSection>
</treatment>
</document>