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<mods:title id="5020F42448C0F92D395EF52C76606CCC">Electrogena brulini sp. nov. and E. vipavensis Zurwerra &amp; Tomka, 1986 syn. nov. revealed by integrative taxonomy of E. gridellii (Grandi, 1953) (Ephemeroptera: Heptageniidae)</mods:title>
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<treatment id="03CDF150FFC2961175A5466CFBDAF9BE" ID-DOI="http://doi.org/10.5281/zenodo.6051313" ID-GBIF-Taxon="137627684" ID-Zenodo-Dep="6051313" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE" httpUri="http://treatment.plazi.org/id/03CDF150FFC2961175A5466CFBDAF9BE" lastPageId="20" lastPageNumber="379" pageId="16" pageNumber="375">
<subSubSection id="C37E13CDFFC2961575A5466CFD9CF8C9" box="[151,590,1892,1919]" pageId="16" pageNumber="375" type="nomenclature">
<paragraph id="8BDB4046FFC2961575A5466CFD9CF8C9" blockId="16.[151,590,1892,1919]" box="[151,590,1892,1919]" pageId="16" pageNumber="375">
<heading id="D093F72AFFC2961575A5466CFD9CF8C9" bold="true" box="[151,590,1892,1919]" fontSize="11" level="1" pageId="16" pageNumber="375" reason="1">
<taxonomicName id="4C643BC5FFC2961575A5466CFD9CF8C9" ID-CoL="396SS" authority="(Grandi, 1953)" baseAuthorityName="Grandi" baseAuthorityYear="1953" box="[151,590,1892,1919]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="375" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC2961575A5466CFD9CF8C9" bold="true" box="[151,590,1892,1919]" pageId="16" pageNumber="375">
<emphasis id="B9109C54FFC2961575A5466CFE5DF8C8" bold="true" box="[151,399,1892,1918]" italics="true" pageId="16" pageNumber="375">Electrogena gridellii</emphasis>
(
<bibRefCitation id="EFF53DB7FFC2961574AC466DFD94F8C9" author="Grandi" box="[414,582,1893,1919]" pageId="16" pageNumber="375" refString="Grandi, M. (1953) Contributi allo studio degli Efemeroidei italiani XVII. Ecdyonuridae. Bolletino dell'Istituto di Entomologia dell'Universita di Bologna, 19, 307 - 386." type="journal article" year="1953">Grandi, 1953</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37E13CDFFC2961475A546A5FD77FE79" lastPageId="17" lastPageNumber="376" pageId="16" pageNumber="375" type="reference_group">
<paragraph id="8BDB4046FFC2961475A546A5FD17FF18" blockId="16.[151,768,1964,2016]" lastBlockId="17.[151,1367,152,358]" lastPageId="17" lastPageNumber="376" pageId="16" pageNumber="375">
<taxonomicName id="4C643BC5FFC2961575A546A5FE20F874" authority=": Grandi 1953" authorityName="Grandi" authorityYear="1953" box="[151,498,1964,1986]" class="Insecta" family="Heptageniidae" genus="Heptagenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="375" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC2961575A546A5FEB7F874" box="[151,357,1964,1986]" italics="true" pageId="16" pageNumber="375">Heptagenia gridellii</emphasis>
:
<bibRefCitation id="EFF53DB7FFC29615744046A4FE20F874" author="Grandi" box="[370,498,1964,1986]" pageId="16" pageNumber="375" refString="Grandi, M. (1953) Contributi allo studio degli Efemeroidei italiani XVII. Ecdyonuridae. Bolletino dell'Istituto di Entomologia dell'Universita di Bologna, 19, 307 - 386." type="journal article" year="1953">Grandi 1953</bibRefCitation>
</taxonomicName>
male and female imagines
<taxonomicName id="4C643BC5FFC2961575A546C2FDD3F856" authority=": Belfiore 1981" authorityName="Belfiore" authorityYear="1981" box="[151,513,1994,2016]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="375" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC2961575A546C2FEBAF856" box="[151,360,1994,2016]" italics="true" pageId="16" pageNumber="375">Ecdyonurus gridellii</emphasis>
:
<bibRefCitation id="EFF53DB7FFC29615744746C2FDD3F856" author="Belfiore" box="[373,513,1994,2016]" pageId="16" pageNumber="375" refString="Belfiore, C. (1981) On the Italian species of the Ecdyonurus lateralis group (Ephemeroptera, Heptageniidae). Aquatic Insects, 3, 171 - 178. https: // doi. org / 10.1080 / 01650428109361059" type="journal article" year="1981">Belfiore 1981</bibRefCitation>
</taxonomicName>
<typeStatus id="54DFFEE4FFC29615773546C2FDB6F856" box="[519,612,1994,2016]" pageId="16" pageNumber="375" type="lectotype">lectotype</typeStatus>
designation
<taxonomicName id="4C643BC5FFC3961475A54190FDD3FF18" authority=": Belfiore 1982" authorityName="Belfiore" authorityYear="1982" box="[151,513,152,174]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC3961475A54190FEBAFF18" box="[151,360,152,174]" italics="true" pageId="17" pageNumber="376">Ecdyonurus gridellii</emphasis>
:
<bibRefCitation id="EFF53DB7FFC3961474474190FDD3FF18" author="Belfiore" box="[373,513,152,174]" pageId="17" pageNumber="376" refString="Belfiore, C. (1982) The nymphs of Italian species of the Ecdyonurus lateralis group, with a description of the nymph of E. gridellii (Grandi, 1953) (Ephemeroptera, Heptageniidae). Fragmenta Entomologica, 16, 125 - 131." type="journal article" year="1982">Belfiore 1982</bibRefCitation>
</taxonomicName>
nymph description
</paragraph>
<paragraph id="8BDB4046FFC3961475A541BEFD07FF7A" blockId="17.[151,1367,152,358]" box="[151,725,182,204]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475A541BEFF69FF7A" bold="true" box="[151,187,182,204]" pageId="17" pageNumber="376">nec</emphasis>
<taxonomicName id="4C643BC5FFC3961475F041BEFD07FF7A" authority="sensu Zurwerra &amp; Tomka 1984" authorityName="Zurwerra &amp; Tomka" authorityYear="1984" box="[194,725,182,204]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC3961475F041BEFE41FF7A" box="[194,403,182,204]" italics="true" pageId="17" pageNumber="376">Ecdyonurus gridellii</emphasis>
sensu
<bibRefCitation id="EFF53DB7FFC3961474E541BFFD07FF7A" author="Zurwerra" box="[471,725,182,204]" pageId="17" pageNumber="376" refString="Zurwerra, A. &amp; Tomka, I. (1984) Beitrag zur Kenntnis der Eintagsfliegenfauna der Schweiz (Insecta, Ephemeroptera). Bulletin de la Societe fribourgeoise des Sciences naturelles, 73, 132 - 146." type="journal article" year="1984">Zurwerra &amp; Tomka 1984</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BDB4046FFC3961475A541DDFD10FF5D" blockId="17.[151,1367,152,358]" box="[151,706,213,235]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475A541DDFF69FF5D" bold="true" box="[151,187,213,235]" pageId="17" pageNumber="376">nec</emphasis>
<taxonomicName id="4C643BC5FFC3961475F041DDFD10FF5D" authority="sensu Sartori &amp; Dethier 1985" authorityName="Sartori &amp; Dethier" authorityYear="1985" box="[194,706,213,235]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC3961475F041DDFE41FF5D" box="[194,403,213,235]" italics="true" pageId="17" pageNumber="376">Ecdyonurus gridellii</emphasis>
sensu
<bibRefCitation id="EFF53DB7FFC3961474E541DDFD10FF5D" author="Sartori" box="[471,706,213,235]" pageId="17" pageNumber="376" refString="Sartori, M. &amp; Dethier, M. (1985) Faune aquatique du Canton de Geneve: II. Ephemeropteres (Insecta, Ephemeroptera). Mitteilungen der Schweizerischen entomologischen Gesellschaft, 58, 493 - 510." type="journal article" year="1985">Sartori &amp; Dethier 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BDB4046FFC3961475A541FCFD85FE9E" blockId="17.[151,1367,152,358]" pageId="17" pageNumber="376">
<taxonomicName id="4C643BC5FFC3961475A541FCFDA6FEBC" ID-CoL="396SS" authority=": Zurwerra &amp; Tomka 1985" authorityName="Zurwerra &amp; Tomka" authorityYear="1985" box="[151,628,244,266]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="gridellii" status="comb. nov.">
<emphasis id="B9109C54FFC3961475A541FCFEBBFEBC" box="[151,361,244,266]" italics="true" pageId="17" pageNumber="376">Electrogena gridellii</emphasis>
:
<bibRefCitation id="EFF53DB7FFC39614744441FDFDA6FEBC" author="Zurwerra" box="[374,628,244,266]" pageId="17" pageNumber="376" refString="Zurwerra, A. &amp; Tomka, I. (1985) Electrogena gen. nov. eine Gattung der Heptageniidae (Ephemeroptera). Entomologische Berichte, Luzern, 13, 99 - 104." type="journal article" year="1985">Zurwerra &amp; Tomka 1985</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A223212FFFC39614774841FDFCFBFEBC" box="[634,809,244,266]" pageId="17" pageNumber="376">new combination</taxonomicNameLabel>
<emphasis id="B9109C54FFC3961475A5401AFF69FE9E" bold="true" box="[151,187,274,296]" pageId="17" pageNumber="376">nec</emphasis>
<taxonomicName id="4C643BC5FFC3961475F0401AFD85FE9E" ID-CoL="396SS" authority="sensu Sartori 1987" authorityName="Sartori" authorityYear="1987" box="[194,599,274,296]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC3961475F0401AFE47FE9E" box="[194,405,274,296]" italics="true" pageId="17" pageNumber="376">Electrogena gridellii</emphasis>
sensu
<bibRefCitation id="EFF53DB7FFC3961474EB401AFD85FE9E" author="Sartori" box="[473,599,274,296]" pageId="17" pageNumber="376" refString="Sartori, M. (1987) Contribution a l'etude taxonomique et eco-faunistique des Ephemeropteres de Suisse (Insecta; Ephemeroptera). University of Lausanne, Lausanne, 561 pp." type="book" year="1987">Sartori 1987</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BDB4046FFC3961475A54039FD1AFEF1" blockId="17.[151,1367,152,358]" box="[151,712,305,327]" pageId="17" pageNumber="376">
<taxonomicName id="4C643BC5FFC3961475A54039FDF5FEF1" ID-CoL="396SS" authority=": Gaino et al. 1987" authorityName="Gaino et al." authorityYear="1987" box="[151,551,305,327]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC3961475A54039FEBBFEF1" box="[151,361,305,327]" italics="true" pageId="17" pageNumber="376">Electrogena gridellii</emphasis>
:
<bibRefCitation id="EFF53DB7FFC3961474444039FDF5FEF1" author="Gaino" box="[374,551,305,327]" pageId="17" pageNumber="376" refString="Gaino, E., Belfiore, C. &amp; Mazzini, M. (1987) Ootaxonomic investigation of the Italian species of the genus Electrogena (Ephemeroptera, Heptageniidae). Bolletino di Zoologia, 54, 169 - 175. https: // doi. org / 10.1080 / 11250008709355578" type="journal article" year="1987">
Gaino
<emphasis id="B9109C54FFC396147489403AFE3CFEF1" box="[443,494,305,327]" italics="true" pageId="17" pageNumber="376">et al.</emphasis>
1987
</bibRefCitation>
</taxonomicName>
egg description
</paragraph>
<paragraph id="8BDB4046FFC3961475A54058FA85FED0" blockId="17.[151,1367,152,358]" box="[151,1367,336,358]" pageId="17" pageNumber="376">
<taxonomicName id="4C643BC5FFC3961475A54058FD5EFED0" ID-CoL="396TJ" authority=": Zurwerra &amp; Tomka 1986" authorityName=": Zurwerra &amp; Tomka" authorityYear="1986" box="[151,652,336,358]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="vipavensis">
<emphasis id="B9109C54FFC3961475A54058FE53FED0" box="[151,385,336,358]" italics="true" pageId="17" pageNumber="376">Electrogena vipavensis</emphasis>
:
<bibRefCitation id="EFF53DB7FFC3961474BD4059FD5EFED0" author="Zurwerra" box="[399,652,336,358]" pageId="17" pageNumber="376" refString="Zurwerra, A. &amp; Tomka, I. (1986) Drei neue Arten der Gattung Electrogena Zurwerra &amp; Tomka, 1985, aus Sudeuropa (Ephemeroptera, Heptageniidae). Bulletin de la Societe fribourgeoise des Sciences naturelles, 75, 216 - 230." type="journal article" year="1986">Zurwerra &amp; Tomka 1986</bibRefCitation>
</taxonomicName>
female imago, male subimago, nymph exuvia, egg NEW SYNONYM
</paragraph>
<paragraph id="8BDB4046FFC3961475A5409AFD77FE79" blockId="17.[151,1437,402,2011]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475A5409AFE3EFE1D" bold="true" box="[151,492,402,427]" pageId="17" pageNumber="376">Nymph description. Length:</emphasis>
Male: body
<quantity id="4C9CEDA3FFC3961477B2409AFCD6FE1C" box="[640,772,402,427]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="17" pageNumber="376" unit="mm" value="10.25" valueMax="12.0" valueMin="8.5">
8.5
<quantity id="4C9CEDA3FFC396147781409AFCD6FE1C" box="[691,772,402,427]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="376" unit="mm" value="12.0">12 mm</quantity>
</quantity>
, cerci:
<quantity id="4C9CEDA3FFC39614766A409AFC07FE1C" box="[856,981,402,427]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="17" pageNumber="376" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">1116 mm</quantity>
. Female: body
<quantity id="4C9CEDA3FFC3961471B7409BFAD0FE1C" box="[1157,1282,403,427]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="17" pageNumber="376" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
, cerci:
<quantity id="4C9CEDA3FFC396147064409BFF14FE78" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="17" pageNumber="376" unit="mm" value="15.0" valueMax="17.0" valueMin="13.0">1317 mm</quantity>
. General colouration dull, little contrasted.
</paragraph>
</subSubSection>
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<paragraph id="8BDB4046FFC3961475F540D2FC04FC15" blockId="17.[151,1437,402,2011]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475F540D2FEC2FE45" bold="true" box="[199,272,474,499]" pageId="17" pageNumber="376">Head:</emphasis>
surface little contrasted (
<figureCitation id="135F5CC3FFC39614777840D2FCF2FE45" box="[586,800,474,499]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Figs 7, 9, 11, 13</figureCitation>
); noteworthy the presence of a double sagittal faint (sometimes lacking) triangular marking on the anterior margin of head
<emphasis id="B9109C54FFC3961476D140F7FBD6FDA0" box="[995,1028,511,534]" italics="true" pageId="17" pageNumber="376">(a)</emphasis>
and a lateral light zone ahead the compound eyes
<emphasis id="B9109C54FFC396147467432BFEA7FD8C" box="[341,373,547,570]" italics="true" pageId="17" pageNumber="376">(c)</emphasis>
(
<figureCitation id="135F5CC3FFC3961474BA432AFE03FD8D" box="[392,465,546,571]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 7</figureCitation>
); no contrasted pattern on the posterior part (
<figureCitation id="135F5CC3FFC3961476DC432AFB91FD8D" box="[1006,1091,546,571]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 11</figureCitation>
), only a greyish band on the posterior margin; a dark marking touching the compound eyes, no colouration behind the ocelli (
<figureCitation id="135F5CC3FFC396147020434EFA5DFDE9" box="[1298,1423,582,607]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Figs 9, 13</figureCitation>
). Labrum rather slender R_LBR = 4.24.6. Mandible with a prostheca bearing 920 bristles (N_PRO), the first bristles thick, then the size of them tapering, often ending with very tiny hairy bristles (not counted) (
<figureCitation id="135F5CC3FFC39614700A4386FA42FD11" box="[1336,1424,654,679]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="17" pageNumber="376">Fig. 43</figureCitation>
). Addition of right and left N_PRO: 2239 bristles. Maxillae with a row of 1926 comb-shaped setae on anterior margin (N_CBS), the 5th (starting from inner side) with 1015 teeth (N_TCB), outer margin of galea-lacinia bearing 1143 setae (N_OUT), inner margin and ventral face of the first segment of palpus bearing 2446 thin setae (N_PLP). Hypopharynx with laterally expanded superlinguae, densely covered with long thin setae, apex of lobes with long setae (
<figureCitation id="135F5CC3FFC3961474A7424AFE3BFCED" box="[405,489,834,859]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="17" pageNumber="376">Fig. 49</figureCitation>
), sometimes missing when worn or very difficult to see because obscured by dirt as
<taxonomicName id="4C643BC5FFC3961475A5426FFED6FCC8" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[151,260,870,894]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC3961475A5426FFED6FCC8" box="[151,260,870,894]" italics="true" pageId="17" pageNumber="376">E. brulini</emphasis>
</taxonomicName>
in
<figureCitation id="135F5CC3FFC39614741E426EFE52FCC9" box="[300,384,870,895]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="17" pageNumber="376">Fig. 48</figureCitation>
. Labium, ventral face, bristles reaching about half of the width of the paraglossae (
<figureCitation id="135F5CC3FFC396147008426EFA5DFCC9" box="[1338,1423,870,895]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="17" pageNumber="376">Fig. 47</figureCitation>
), inner distal margin can be concave, R_GLA = 3.15.2, R_GLB = 2.53.0.
</paragraph>
<paragraph id="8BDB4046FFC3961475F542A6FE24FA2D" blockId="17.[151,1437,402,2011]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475F542A6FEFBFC71" bold="true" box="[199,297,942,967]" pageId="17" pageNumber="376">Thorax:</emphasis>
Pronotum without a pattern, faint, but presence of a reddish-brown spot
<emphasis id="B9109C54FFC39614714642A6FB46FC70" box="[1140,1172,942,966]" italics="true" pageId="17" pageNumber="376">(h)</emphasis>
, usually hidden by the head, the lateral margins regularly rounded with a faint greyish line
<emphasis id="B9109C54FFC39614769A42DBFC1BFC5C" box="[936,969,979,1002]" italics="true" pageId="17" pageNumber="376">(g)</emphasis>
(
<figureCitation id="135F5CC3FFC3961476E842DAFBE2FC5D" box="[986,1072,978,1003]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 17</figureCitation>
). Mesonotum with several light brown markings and an oval-shaped spot in young instars (arrow in
<figureCitation id="135F5CC3FFC3961476F242FEFBC4FBB9" box="[960,1046,1014,1039]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 19</figureCitation>
), becoming an oblique semilunar marking (arrow in
<figureCitation id="135F5CC3FFC3961474474512FE1EFB85" box="[373,460,1050,1075]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 21</figureCitation>
). Coxa little contrasted: proximal half white, distal part greyish (
<figureCitation id="135F5CC3FFC3961471FB4512FAF2FB85" box="[1225,1312,1050,1075]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Fig. 31</figureCitation>
). Thoracic sterna: sclerites light, imaginal colouration becoming brown during nymph development. Legs: faint ornamentation rather similar between fore-, mid and hind legs (
<figureCitation id="135F5CC3FFC396147658456AFC11FBCD" box="[874,963,1122,1147]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Figs 29</figureCitation>
,
<figureCitation id="135F5CC3FFC3961476FD456BFC3DFBCD" box="[975,1007,1123,1147]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="17" pageNumber="376">34</figureCitation>
). Femora dorsal face with four (some can be joined) less contrasted brownish spots delimiting an often incomplete light cross, the weak contrast disappearing in the last instars (
<figureCitation id="135F5CC3FFC3961474C545A2FD9EFB75" box="[503,588,1194,1219]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Fig. 29</figureCitation>
); ventral face of femora very rarely with a greyish cuticular suffusion in last instars; all femora with long, narrow and pointed bristles on the dorsal face (
<figureCitation id="135F5CC3FFC39614712545C6FB4DFB51" box="[1047,1183,1230,1255]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="17" pageNumber="376">Figs 38, 39</figureCitation>
); only one (can miss) strong spine in the middle of the ventral face of all femora, near the outer margins (N_BVF). Tibia faint, when contrasted, only the apex lighter; pectinate setae scarce on foretibia, numerous on mid and hind tibia (as
<taxonomicName id="4C643BC5FFC396147002441FFA4EFA98" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1328,1436,1302,1326]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="376" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC396147002441FFA4EFA98" box="[1328,1436,1302,1326]" italics="true" pageId="17" pageNumber="376">E. brulini</emphasis>
</taxonomicName>
<figureCitation id="135F5CC3FFC3961475A54432FF3BFAE5" box="[151,233,1338,1363]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="17" pageNumber="376">Fig. 35</figureCitation>
, inset). Tarsi distal quarter slightly darkened (mainly on dorsal face), exceptionally forming a less or more visible ring on distal part, colouration not strengthening toward the proximal part; claws with 2 teeth, rarely 1 or 3, exceptionally 0 or 4 (N_CLW).
</paragraph>
<paragraph id="8BDB4046FFC3961475F544AEFF24F825" blockId="17.[151,1437,402,2011]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475F544AEFE97FA09" bold="true" box="[199,325,1446,1471]" pageId="17" pageNumber="376">Abdomen:</emphasis>
Terga with little contrasted markings: terga (III) IVVI (VIIVIII) anterior margin: grey slender triangle in the middle (
<figureCitation id="135F5CC3FFC3961474A944C2FE0DFA55" box="[411,479,1482,1507]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 3</figureCitation>
); terga (IVVI and VIIIX) posterior margin: grey line, not visible when fused with entirely dark terga; terga IIVIII anterolateral epidermal brown spot larger than the posterolateral one (
<figureCitation id="135F5CC3FFC39614700B44E6FA5DF9B1" box="[1337,1423,1518,1543]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Fig. 23</figureCitation>
), anterolateral spot hidden under the preceding terga in small nymphs, growing up to last instar (
<figureCitation id="135F5CC3FFC3961471FF471AFAF4F99D" box="[1229,1318,1554,1579]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Figs 23</figureCitation>
); terga II VII posterolaterally with a reddish-brown spot (
<figureCitation id="135F5CC3FFC396147788473EFCC2F9F9" box="[698,784,1590,1615]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Fig. 23</figureCitation>
); in the last instars, tergum VII (darker) contrasting with terga VIIIX (
<figureCitation id="135F5CC3FFC3961474094752FE53F9C5" box="[315,385,1626,1651]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 3</figureCitation>
), tergum X anterior half brownish with a light area in the middle, posterior half white with only a slight line on the posterior margin (
<figureCitation id="135F5CC3FFC39614775D4776FD10F921" box="[623,706,1662,1687]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="17" pageNumber="376">Fig. 15</figureCitation>
); posterior margin of each tergum with thin and elongated spines with microdenticulations at their bases. Sterna: no cuticular colouration except a light greyish stain, epidermis light in young nymphs, becoming brown in last instars (
<figureCitation id="135F5CC3FFC3961477FF47CEFCF2F969" box="[717,800,1734,1759]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Fig. 25</figureCitation>
); ganglionic chain brownish pigmented, thoracic ganglia a bit darker, in last instars, ganglia hardly visible due to the dark epidermal colouration (
<figureCitation id="135F5CC3FFC3961471B147E2FB0AF8B5" box="[1155,1240,1770,1795]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="17" pageNumber="376">Fig. 25</figureCitation>
). In the male last instar nymphs, styliger processes and titillator already visible through sternum 9, but titillators not enough formed to be diagnostic. Gill I tongue-shaped, gills IIVI broad and asymmetrical, gill VII elongated, narrow at apex and slightly pointed (
<figureCitation id="135F5CC3FFC39614746B465EFE7FF8D9" box="[345,429,1878,1903]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="17" pageNumber="376">Fig. 51</figureCitation>
). Paracercus and cerci light, becoming darker only in the last instars, colouration fading apically.
</paragraph>
<paragraph id="8BDB4046FFC3961475F54696FBECF800" blockId="17.[151,1437,402,2011]" box="[199,1086,1950,1975]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475F54696FE10F801" bold="true" box="[199,450,1950,1975]" pageId="17" pageNumber="376">Male imago: Length:</emphasis>
body
<quantity id="4C9CEDA3FFC39614773A4697FD51F800" box="[520,643,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="17" pageNumber="376" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
, forewing
<quantity id="4C9CEDA3FFC3961477C84697FCA7F800" box="[762,885,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="17" pageNumber="376" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
, cerci
<quantity id="4C9CEDA3FFC39614768C4696FBE8F800" box="[958,1082,1950,1975]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="17" pageNumber="376" unit="mm" value="33.5" valueMax="36.0" valueMin="31.0">3136 mm</quantity>
.
</paragraph>
<paragraph id="8BDB4046FFC3961475F546CAFA45F86D" blockId="17.[151,1437,402,2011]" box="[199,1431,1986,2011]" pageId="17" pageNumber="376">
<emphasis id="B9109C54FFC3961475F546CAFEC2F86D" bold="true" box="[199,272,1986,2011]" pageId="17" pageNumber="376">Head:</emphasis>
Eyes clearly separated, sclerite joining the compound eyes W-shaped, generally well-marked (
<figureCitation id="135F5CC3FFC39614700446CAFA58F86D" box="[1334,1418,1986,2011]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="17" pageNumber="376">Fig. 85</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775F5419FFC63FD8B" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
<emphasis id="B9109C54FFC0961775F5419FFEFBFF06" bold="true" box="[199,297,151,176]" pageId="18" pageNumber="377">Thorax:</emphasis>
dark brown (
<figureCitation id="135F5CC3FFC0961774F5419FFD9CFF06" box="[455,590,151,176]" captionStart="FIGURES5255" captionStartId="12.[151,264,1973,1995]" captionTargetBox="[151,1436,193,1952]" captionTargetId="figure@12.[151,1436,193,1952]" captionTargetPageId="12" captionText="FIGURES5255.Male imagines' habitus of Electrogena brulini (Figs 52, 54)and Electrogena gridellii(Figs 53, 55).Figs 52 53:dorsal view; Figs 5455: lateral view. Arrows: markings on the tergite VI." httpUri="https://zenodo.org/record/1094924/files/figure.png" pageId="18" pageNumber="377">Figs 53, 55</figureCitation>
). Wings veins all dark brown (
<figureCitation id="135F5CC3FFC096177685419FFBDCFF06" box="[951,1038,151,176]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 65</figureCitation>
). Central part or entire forefemora darkened (
<figureCitation id="135F5CC3FFC09617743C41B4FE41FF63" box="[270,403,188,213]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Figs 6263</figureCitation>
); mid and hind femora dorsally with a very light transversal brown band (
<figureCitation id="135F5CC3FFC0961771D041B4FAEAFF63" box="[1250,1336,188,213]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 62</figureCitation>
); ventral face at most with a barely visible band (
<figureCitation id="135F5CC3FFC09617776441D7FD78FF4E" box="[598,682,223,248]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 63</figureCitation>
).
<emphasis id="B9109C54FFC09617778F41D7FCE9FF4E" bold="true" box="[701,827,223,248]" pageId="18" pageNumber="377">Abdomen:</emphasis>
Terga IIVII (VIII) anterior margin brown with a pair of large subtriangular markings pointed backwards reaching or not the posterior margin and a dark marking between them, posterior margin darkened (as female in
<figureCitation id="135F5CC3FFC09617763C402FFCB3FEF6" box="[782,865,295,320]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">Fig. 87</figureCitation>
), these markings can be fused, covering the dorsal surface (
<figureCitation id="135F5CC3FFC0961775CA4044FE99FED3" box="[248,331,332,357]" captionStart="FIGURES5255" captionStartId="12.[151,264,1973,1995]" captionTargetBox="[151,1436,193,1952]" captionTargetId="figure@12.[151,1436,193,1952]" captionTargetPageId="12" captionText="FIGURES5255.Male imagines' habitus of Electrogena brulini (Figs 52, 54)and Electrogena gridellii(Figs 53, 55).Figs 52 53:dorsal view; Figs 5455: lateral view. Arrows: markings on the tergite VI." httpUri="https://zenodo.org/record/1094924/files/figure.png" pageId="18" pageNumber="377">Fig. 53</figureCitation>
); in lateral view, the subtriangular markings are fused with the anterolateral and the posterolateral markings of the preceding tergum building a large oblique band (
<figureCitation id="135F5CC3FFC0961776B64067FC08FE3E" box="[900,986,367,392]" captionStart="FIGURES5255" captionStartId="12.[151,264,1973,1995]" captionTargetBox="[151,1436,193,1952]" captionTargetId="figure@12.[151,1436,193,1952]" captionTargetPageId="12" captionText="FIGURES5255.Male imagines' habitus of Electrogena brulini (Figs 52, 54)and Electrogena gridellii(Figs 53, 55).Figs 52 53:dorsal view; Figs 5455: lateral view. Arrows: markings on the tergite VI." httpUri="https://zenodo.org/record/1094924/files/figure.png" pageId="18" pageNumber="377">Fig. 55</figureCitation>
). Sterna pale with a large subtriangular light orange-brown marking covering more or less the entire face, ganglia violet (
<figureCitation id="135F5CC3FFC0961771B8409CFB34FE1B" box="[1162,1254,404,429]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 59</figureCitation>
). Cerci brown, colouration fading to the apex. Penis lobes subrhomboidal, narrowed towards the apex, slightly bent inwards (
<figureCitation id="135F5CC3FFC09617705D40BFFF64FE43" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="18" pageNumber="377">Fig. 69</figureCitation>
), only a few small dorsal spines basolaterally on penis lobes (
<figureCitation id="135F5CC3FFC09617765A40D4FC6EFE42" box="[872,956,476,501]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="18" pageNumber="377">Fig. 73</figureCitation>
). Titillator convex before the unique tooth, than concave until a more or less visible basal tip (
<figureCitation id="135F5CC3FFC0961777EB40F7FC8EFDAE" box="[729,860,511,536]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="18" pageNumber="377">Figs 7981</figureCitation>
). Styliger almost straight with lateral humps (as
<taxonomicName id="4C643BC5FFC0961770B64309FF31FD8A" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961770B64309FF31FD8A" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
on
<figureCitation id="135F5CC3FFC09617743F432CFE8DFD8B" box="[269,351,548,573]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 66</figureCitation>
), but humps can be very inconspicuous as in
<figureCitation id="135F5CC3FFC096177669432CFC7FFD8B" box="[859,941,548,573]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 67</figureCitation>
.
</paragraph>
<paragraph id="8BDB4046FFC0961775F5434FFBA2FDE9" blockId="18.[151,1437,151,1472]" box="[199,1136,583,608]" pageId="18" pageNumber="377">
<emphasis id="B9109C54FFC0961775F5434FFE0FFDD6" bold="true" box="[199,477,583,608]" pageId="18" pageNumber="377">Female imago: Length:</emphasis>
body:
<quantity id="4C9CEDA3FFC0961777194340FD77FDE9" box="[555,677,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="18" pageNumber="377" unit="mm" value="12.5" valueMax="14.0" valueMin="11.0">1114 mm</quantity>
, forewing:
<quantity id="4C9CEDA3FFC0961776164340FC4DFDE9" box="[804,927,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="18" pageNumber="377" unit="mm" value="15.5" valueMax="17.0" valueMin="14.0">1417 mm</quantity>
, cerci:
<quantity id="4C9CEDA3FFC0961776C2434FFBBEFDE9" box="[1008,1132,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="18" pageNumber="377" unit="mm" value="28.5" valueMax="32.0" valueMin="25.0">2532 mm</quantity>
.
</paragraph>
<paragraph id="8BDB4046FFC0961775A54364FBA1FD46" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
Same general colouration as in male; only thorax somewhat lighter (
<figureCitation id="135F5CC3FFC0961776924364FBF6FD32" box="[928,1060,620,645]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">Figs 87, 89</figureCitation>
). Wings veins all medium brown (
<figureCitation id="135F5CC3FFC0961775AD4387FF21FD1E" box="[159,243,655,680]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">Fig. 91</figureCitation>
). Light area behind the pair of ocelli
<emphasis id="B9109C54FFC0961777AF4398FD6CFD11" box="[669,702,656,679]" italics="true" pageId="18" pageNumber="377">(n)</emphasis>
, area between the pair of ocelli and the compound eyes dark
<emphasis id="B9109C54FFC0961770484398FA47FD1E" box="[1402,1429,656,680]" italics="true" pageId="18" pageNumber="377">(l)</emphasis>
, sclerite joining the compound eyes with an extended dark brown area (
<figureCitation id="135F5CC3FFC09617768743BCFBDAFD7A" box="[949,1032,692,717]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">Fig. 99</figureCitation>
, m). Subgenital plate covering ¾ of the sternum VIII surface, subanal plate in ventral and apical view rounded (
<figureCitation id="135F5CC3FFC0961776D643DFFBB5FD46" box="[996,1127,727,752]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">Figs 96, 97</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775F543F4FAB3FCA3" blockId="18.[151,1437,151,1472]" box="[199,1377,764,789]" pageId="18" pageNumber="377">
<emphasis id="B9109C54FFC0961775F543F4FDC3FCA3" bold="true" box="[199,529,764,789]" pageId="18" pageNumber="377">Male and female subimago:</emphasis>
Same colouration as imago, but paler. Wings without pattern, entirely grey.
</paragraph>
<paragraph id="8BDB4046FFC0961775F54217FCF9FC8E" blockId="18.[151,1437,151,1472]" box="[199,811,799,824]" pageId="18" pageNumber="377">
<emphasis id="B9109C54FFC0961775F54217FF24FC8E" bold="true" box="[199,246,799,824]" pageId="18" pageNumber="377">Egg</emphasis>
: Dimensions: 155/120 µm; micropyle: 11/6.5 µm.
</paragraph>
<paragraph id="8BDB4046FFC0961775F5424CFDF9FC12" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
Chorion entirely and regularly covered with relatively dense microgranules, KCTs evenly distributed over the surface, more concentrated only on a very restricted area of one pole (
<figureCitation id="135F5CC3FFC096177697426FFBD5FC36" box="[933,1031,871,896]" captionStart="FIGURES 100103" captionStartId="16.[151,264,1770,1792]" captionTargetBox="[154,1436,196,1749]" captionTargetId="figure@16.[151,1436,193,1749]" captionTargetPageId="16" captionText="FIGURES 100103. Eggs of Electrogena brulini (Fig. 100101) and Electrogena gridellii (Figs 102103). Figs 100, 102: general view; Figs 101, 103: emphasis on the micropyle." httpUri="https://zenodo.org/record/1094932/files/figure.png" pageId="18" pageNumber="377">Fig. 102</figureCitation>
). Micropyle bordered with the same microgranules as surface (
<figureCitation id="135F5CC3FFC09617748F4284FDCDFC12" box="[445,543,908,933]" captionStart="FIGURES 100103" captionStartId="16.[151,264,1770,1792]" captionTargetBox="[154,1436,196,1749]" captionTargetId="figure@16.[151,1436,193,1749]" captionTargetPageId="16" captionText="FIGURES 100103. Eggs of Electrogena brulini (Fig. 100101) and Electrogena gridellii (Figs 102103). Figs 100, 102: general view; Figs 101, 103: emphasis on the micropyle." httpUri="https://zenodo.org/record/1094932/files/figure.png" pageId="18" pageNumber="377">Fig. 103</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E13CDFFC0961675F542A7FDA9F9E6" lastPageId="19" lastPageNumber="378" pageId="18" pageNumber="377" type="diagnosis">
<paragraph id="8BDB4046FFC0961775F542A7FA4EFBA6" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
<emphasis id="B9109C54FFC0961775F542A7FE93FC7E" bold="true" box="[199,321,943,968]" pageId="18" pageNumber="377">Diagnosis.</emphasis>
Nymph: general colouration faded, femora without spot on the ventral face, two teeth on tarsal claws; (0) 1 bristle on ventral face of femora, near the outer margin; bristles of femora long, thin and pointed; prostheca consisting in 920 bristles (
<figureCitation id="135F5CC3FFC09617770F42FFFD42FBA6" box="[573,656,1015,1040]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="18" pageNumber="377">Fig. 43</figureCitation>
), terga anterolateral spot smaller than the posterolateral one (
<figureCitation id="135F5CC3FFC09617700942FFFA5DFBA6" box="[1339,1423,1015,1040]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="18" pageNumber="377">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775F54515FBFBFB16" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
Imago: dorsal face of mid and hind femora with an inconspicuous band (
<figureCitation id="135F5CC3FFC0961776C44514FB99FB83" box="[1014,1099,1052,1077]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 62</figureCitation>
); ventral face of mid and hind femora without colouration (
<figureCitation id="135F5CC3FFC0961774ED4537FDE7FBEE" box="[479,565,1087,1112]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 63</figureCitation>
); wings veins dark brown in male (
<figureCitation id="135F5CC3FFC0961776E24537FBF4FBEE" box="[976,1062,1087,1112]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">Fig. 65</figureCitation>
), medium brown in female (
<figureCitation id="135F5CC3FFC09617705D4537FF64FBCA" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">Fig. 91</figureCitation>
); terga well contrasted, white and dark brown, posterolateral and anterolateral spots well visible forming an oblique streak (
<figureCitation id="135F5CC3FFC096177471458FFE48FB16" box="[323,410,1159,1184]" captionStart="FIGURES5255" captionStartId="12.[151,264,1973,1995]" captionTargetBox="[151,1436,193,1952]" captionTargetId="figure@12.[151,1436,193,1952]" captionTargetPageId="12" captionText="FIGURES5255.Male imagines' habitus of Electrogena brulini (Figs 52, 54)and Electrogena gridellii(Figs 53, 55).Figs 52 53:dorsal view; Figs 5455: lateral view. Arrows: markings on the tergite VI." httpUri="https://zenodo.org/record/1094924/files/figure.png" pageId="18" pageNumber="377">Figs 55</figureCitation>
,
<figureCitation id="135F5CC3FFC096177494458FFE17FB16" box="[422,453,1159,1184]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="18" pageNumber="377">57</figureCitation>
,
<figureCitation id="135F5CC3FFC0961774E24580FE3DFB16" box="[464,495,1160,1184]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="18" pageNumber="377">89</figureCitation>
); titillators with 1 apico-lateral tooth (
<figureCitation id="135F5CC3FFC0961776A8458FFBCFFB16" box="[922,1053,1159,1184]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="18" pageNumber="377">Figs 7981</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775F545A4FCE1FACE" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
Although many differences in colorations make it possible to easily separate
<taxonomicName id="4C643BC5FFC09617710B45A5FB74FB72" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1081,1190,1196,1220]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC09617710B45A5FB74FB72" box="[1081,1190,1196,1220]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
from
<taxonomicName id="4C643BC5FFC0961771DC45A5FABBFB72" box="[1262,1385,1196,1220]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC0961771DC45A5FABBFB72" box="[1262,1385,1196,1220]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
, the two species have always been confused because the value of colouration characters is underrated (or because of the use of old faded specimens); therefore those characters are generally omitted. Among colouration characters are true patterns, but also simple colourations which seem to have even less taxonomic importance. However, some of these are conservative characters, visible both in nymphs and adults, and have been positively tested as to their discriminating value to separate
<taxonomicName id="4C643BC5FFC0961777314469FDBCFACE" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[515,622,1376,1400]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961777314469FDBCFACE" box="[515,622,1376,1400]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
from
<emphasis id="B9109C54FFC0961777804469FCE1FACE" box="[690,819,1376,1400]" italics="true" pageId="18" pageNumber="377">
<taxonomicName id="4C643BC5FFC0961777804469FCFDFACE" box="[690,815,1376,1400]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">E. gridellii</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDB4046FFC0961775F5448CFDE2FA76" blockId="18.[151,1437,151,1472]" pageId="18" pageNumber="377">
Only the discriminating characters between
<taxonomicName id="4C643BC5FFC0961777F8448DFC9AFA2A" box="[714,840,1412,1436]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC0961777F8448DFC9AFA2A" box="[714,840,1412,1436]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C643BC5FFC0961776B4448DFC27FA2A" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[902,1013,1412,1436]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961776B4448DFC27FA2A" box="[902,1013,1412,1436]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
tested and easily usable in the vast majority of cases are presented here.
</paragraph>
<paragraph id="8BDB4046FFC0961775A544F8FE5CF9BF" blockId="18.[151,1436,1519,2013]" box="[151,398,1519,1545]" pageId="18" pageNumber="377">At the nymphal stage:</paragraph>
<paragraph id="8BDB4046FFC0961775A5471DFC0BF99B" blockId="18.[151,1436,1519,2013]" box="[151,985,1556,1581]" pageId="18" pageNumber="377">
- all colourations of
<taxonomicName id="4C643BC5FFC0961774A8471DFDD7F99A" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[410,517,1556,1580]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961774A8471DFDD7F99A" box="[410,517,1556,1580]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
contrasted, those of
<taxonomicName id="4C643BC5FFC0961777DF471DFCB4F99A" box="[749,870,1556,1580]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC0961777DF471DFCB4F99A" box="[749,870,1556,1580]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
very dull.
</paragraph>
<paragraph id="8BDB4046FFC0961775A54731FBD9F9C2" blockId="18.[151,1436,1519,2013]" pageId="18" pageNumber="377">
- pattern and different markings contrasted on the head in
<taxonomicName id="4C643BC5FFC09617760D4731FC78F9E6" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[831,938,1592,1616]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC09617760D4731FC78F9E6" box="[831,938,1592,1616]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC09617768B473FFBE8F9E6" box="[953,1082,1591,1616]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="18" pageNumber="377">Figs 6,8,10</figureCitation>
, af), no pattern and only some pale markings (notice the absence of spot
<emphasis id="B9109C54FFC0961777AB4754FD68F9C2" box="[665,698,1628,1652]" italics="true" pageId="18" pageNumber="377">(b)</emphasis>
) in
<taxonomicName id="4C643BC5FFC0961777D54755FCB2F9C2" box="[743,864,1628,1652]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC0961777D54755FCB2F9C2" box="[743,864,1628,1652]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC09617765D4754FC2CF9C2" box="[879,1022,1628,1653]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="18" pageNumber="377">Figs 7, 9, 11</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775A54789FD4FF956" blockId="18.[151,1436,1519,2013]" pageId="18" pageNumber="377">
- each prostheca consisting in 5-8 bristles (all bristles evenly thick, at the most the basal one a little thinner) in
<taxonomicName id="4C643BC5FFC0961770B64789FEDDF90A" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961770B64789FEDDF90A" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC09617741247ACFEA4F90A" box="[288,374,1700,1725]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="18" pageNumber="377">Fig. 42</figureCitation>
), 920 bristles (the first bristles thick, then the size of them tapering, often ending with very tiny hairy bristles) in
<taxonomicName id="4C643BC5FFC09617748647C1FDFFF956" box="[436,557,1736,1760]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC09617748647C1FDFFF956" box="[436,557,1736,1760]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC09617770F47CFFD5CF956" box="[573,654,1735,1760]" captionStart="FIGURES4051" captionStartId="11.[151,264,1873,1895]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@11.[151,1436,193,1851]" captionTargetPageId="11" captionText="FIGURES4051.Mouth parts and gills of Electrogena brulini (Figs 40, 42, 44, 46, 48, 50) and Electrogena gridellii (Figs 41, 43, 47, 49, 51). Figs 4041: dorsal face of labrum; Figs 4243: mandible with prostheca; Figs 4445: anterior margin of dorsal face of labrum, central part; Figs 4647 ventral face of glossae; Figs 4849: hypopharynx in dorsal view; Figs 5051: gills I VII, without the bunch of filaments." httpUri="https://zenodo.org/record/1094922/files/figure.png" pageId="18" pageNumber="377">Fig. 43</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775A547E5FCD0F8FA" blockId="18.[151,1436,1519,2013]" pageId="18" pageNumber="377">
- pronotum with a contrasted pattern but no reddish-brown spot
<emphasis id="B9109C54FFC0961776B647E4FC76F8B2" box="[900,932,1772,1796]" italics="true" pageId="18" pageNumber="377">(h)</emphasis>
, the lateral margins with a dark line
<emphasis id="B9109C54FFC09617700D47E4FAB2F8B5" box="[1343,1376,1772,1795]" italics="true" pageId="18" pageNumber="377">(g)</emphasis>
in
<taxonomicName id="4C643BC5FFC0961770B647E5FEDDF89E" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961770B647E5FEDDF89E" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC09617742C4607FEA3F89E" box="[286,369,1807,1832]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="18" pageNumber="377">Fig. 16</figureCitation>
), faint, but presence of a reddish-brown spot
<emphasis id="B9109C54FFC09617765F4618FC5CF89E" box="[877,910,1808,1832]" italics="true" pageId="18" pageNumber="377">(h)</emphasis>
, usually hidden by the head, the lateral margin with a faint greyish line
<emphasis id="B9109C54FFC0961774E6463CFE27F8FD" box="[468,501,1844,1867]" italics="true" pageId="18" pageNumber="377">(g)</emphasis>
in
<taxonomicName id="4C643BC5FFC09617772A463DFD43F8FA" box="[536,657,1844,1868]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC09617772A463DFD43F8FA" box="[536,657,1844,1868]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC096177792463CFD20F8FA" box="[672,754,1844,1869]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="18" pageNumber="377">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775A54650FB9FF822" blockId="18.[151,1436,1519,2013]" pageId="18" pageNumber="377">
- mesonotum with brown semilunar vertical markings in all instars in
<taxonomicName id="4C643BC5FFC0961776EE4651FB98F8C6" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[988,1098,1880,1904]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC0961776EE4651FB98F8C6" box="[988,1098,1880,1904]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
(arrow in
<figureCitation id="135F5CC3FFC0961771F4465FFA99F8C6" box="[1222,1355,1879,1904]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="18" pageNumber="377">Figs 18, 20</figureCitation>
), ovalshaped spot becoming an oblique semilunar marking in
<taxonomicName id="4C643BC5FFC0961776074675FC7CF822" box="[821,942,1916,1940]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC0961776074675FC7CF822" box="[821,942,1916,1940]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC09617768C4674FBEDF822" box="[958,1087,1916,1941]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="18" pageNumber="377">Figs 19, 21</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC0961775A546A9FC87F86A" blockId="18.[151,1436,1519,2013]" pageId="18" pageNumber="377">
- femora ventral face always with a visible dark (generally double) spot in
<taxonomicName id="4C643BC5FFC09617713D46A9FBAEF80E" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1039,1148,1952,1976]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC09617713D46A9FBAEF80E" box="[1039,1148,1952,1976]" italics="true" pageId="18" pageNumber="377">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC0961771BF46A8FB33F80E" box="[1165,1249,1952,1977]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="18" pageNumber="377">Fig. 27</figureCitation>
), at most with a greyish cuticular suffusion in the last instars in
<taxonomicName id="4C643BC5FFC0961777E646CDFC9CF86A" box="[724,846,1988,2012]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="377" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC0961777E646CDFC9CF86A" box="[724,846,1988,2012]" italics="true" pageId="18" pageNumber="377">E. gridellii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB4046FFC1961675A54190FB13FF62" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="378">
- bristles of dorsal face medium sized blunt pointed on forefemora, more peaked in mid and hind femora in
<taxonomicName id="4C643BC5FFC1961670B64191FEDDFF62" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961670B64191FEDDFF62" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616742C41B4FE72FF62" box="[286,416,188,213]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="19" pageNumber="378">Figs 36, 37</figureCitation>
), thin, long and pointed on all femora long in
<taxonomicName id="4C643BC5FFC19616769141B5FBCEFF62" box="[931,1052,188,212]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC19616769141B5FBCEFF62" box="[931,1052,188,212]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616711941B4FB67FF62" box="[1067,1205,188,213]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="19" pageNumber="378">Figs. 38, 39</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A541E8FE5FFEAA" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="378">
- coxa bicoloured, proximal half white, distal half dark grey in
<taxonomicName id="4C643BC5FFC19616764A41E9FC31FF4E" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[888,995,224,248]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC19616764A41E9FC31FF4E" box="[888,995,224,248]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961676C041D7FBA1FF4E" box="[1010,1139,223,248]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Figs 28, 30</figureCitation>
,
<figureCitation id="135F5CC3FFC19616714D41E8FB4CFF4E" box="[1151,1182,224,248]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="19" pageNumber="378">32</figureCitation>
), little contrasted in
<taxonomicName id="4C643BC5FFC1961670B641E9FECFFEAA" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961670B641E9FECFFEAA" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616741E400CFEADFEAA" box="[300,383,260,285]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A54020FC86FED2" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="378">
- tibia contrasted, distal quarter and proximal third light, middle darker in
<taxonomicName id="4C643BC5FFC1961671344021FBA0FEF6" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1030,1138,296,320]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961671344021FBA0FEF6" box="[1030,1138,296,320]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961671B6402FFB0BFEF6" box="[1156,1241,295,320]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Fig. 28</figureCitation>
,
<figureCitation id="135F5CC3FFC1961671D44020FAE2FEF6" box="[1254,1328,296,320]" captionStart="FIGURES 3239" captionStartId="10.[151,264,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 3239. Nymph of Electrogena brulini (Fig. 32); legs and parts of legs of Electrogena brulini (Figs 33, 3537) and Electrogenagridellii (Figs 34, 38,39).Fig.32:habitus of ayoung nymph in latero-dorsal view; Figs3334:dorsal face of fore-, mid and hind legs; Fig 35: ventral face of fore-, mid and hind legs, emphasis on a pectinate setae; Figs 36, 38: dorsal face of forefemora, emphasis on bristles; Figs 37, 39: dorsal face of hind femora, emphasis on bristles. Arrows in Fig. 32: example of different markings already visible in young nymphs." httpUri="https://zenodo.org/record/1094920/files/figure.png" pageId="19" pageNumber="378">32, 33</figureCitation>
), entirely faint, when contrasted, only the apex lighter in
<taxonomicName id="4C643BC5FFC1961677E64045FC9FFED2" box="[724,845,332,356]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961677E64045FC9FFED2" box="[724,845,332,356]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB4046FFC1961675A54078FA87FE1B" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="378">
- tergum X dark with a light marking on each side in
<taxonomicName id="4C643BC5FFC1961676354079FCA0FE3E" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[775,882,368,392]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961676354079FCA0FE3E" box="[775,882,368,392]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961676B34067FC01FE3E" box="[897,979,367,392]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="19" pageNumber="378">Fig. 14</figureCitation>
), anterior half brownish with a light area in the middle, posterior half white with only a slight line on the posterior margin in
<taxonomicName id="4C643BC5FFC196167159409DFB37FE1A" box="[1131,1253,404,428]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC196167159409DFB37FE1A" box="[1131,1253,404,428]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961671C6409CFA95FE1B" box="[1268,1351,404,429]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="19" pageNumber="378">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A540B0FABFFE43" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="378">
- sterna with an arch-shaped cuticular pattern, epidermis always entirely light in
<taxonomicName id="4C643BC5FFC19616716E40B1FB1BFE66" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1116,1225,440,464]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC19616716E40B1FB1BFE66" box="[1116,1225,440,464]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961671E940BFFAE2FE66" box="[1243,1328,439,464]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Fig. 24</figureCitation>
), without cuticular pattern, epidermis light in young nymphs, becoming brown in last instars in
<taxonomicName id="4C643BC5FFC1961671B140D5FB2EFE42" box="[1155,1276,476,500]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961671B140D5FB2EFE42" box="[1155,1276,476,500]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616703E40D4FA8DFE43" box="[1292,1375,476,501]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A5432CFEBDFD8B" blockId="19.[151,1436,548,1149]" box="[151,367,548,573]" pageId="19" pageNumber="378">At imaginal stages:</paragraph>
<paragraph id="8BDB4046FFC1961675A54340FABDFDD6" blockId="19.[151,1436,548,1149]" box="[151,1391,583,608]" pageId="19" pageNumber="378">
- general colouration of abdomen uniform in
<taxonomicName id="4C643BC5FFC1961677834341FCCEFDD6" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[689,796,584,608]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961677834341FCCEFDD6" box="[689,796,584,608]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC196167619434FFC50FDD6" box="[811,898,583,608]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Figs 56</figureCitation>
,
<figureCitation id="135F5CC3FFC1961676BF434FFC7EFDD6" box="[909,940,583,608]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">86</figureCitation>
), contrasted in
<taxonomicName id="4C643BC5FFC19616716A4341FB03FDD6" box="[1112,1233,584,608]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC19616716A4341FB03FDD6" box="[1112,1233,584,608]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961671D2434FFAE5FDD6" box="[1248,1335,583,608]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Figs 57</figureCitation>
,
<figureCitation id="135F5CC3FFC1961670714340FAB1FDD6" box="[1347,1379,584,608]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">87</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A54365FBBDFD1E" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- wings veins yellowish-brown; costa, subcosta and radius anterior brighter yellow in
<taxonomicName id="4C643BC5FFC1961671B44365FB26FD32" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1158,1268,620,644]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961671B44365FB26FD32" box="[1158,1268,620,644]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961670364364FAB0FD33" box="[1284,1378,620,645]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Figs. 64</figureCitation>
,
<figureCitation id="135F5CC3FFC1961670424364FA5DFD32" box="[1392,1423,620,644]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">90</figureCitation>
), regularly dark brown in male, medium brown in female in
<taxonomicName id="4C643BC5FFC1961676654399FC02FD1E" box="[855,976,656,680]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961676654399FC02FD1E" box="[855,976,656,680]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961676D24387FBE4FD1E" box="[992,1078,655,680]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Figs 65</figureCitation>
,
<figureCitation id="135F5CC3FFC1961671704398FBB3FD1E" box="[1090,1121,656,680]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">91</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A543BDFAEFFD46" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- mid and hind femora dorsally with a brown spot at the distal third and a second spot, variably visible, in the proximal third in
<taxonomicName id="4C643BC5FFC1961674B843D1FE27FD46" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[394,501,728,752]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961674B843D1FE27FD46" box="[394,501,728,752]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616773643DFFD8AFD46" box="[516,600,727,752]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Fig. 60</figureCitation>
), with a very light transversal brown band in
<taxonomicName id="4C643BC5FFC19616716643D1FB1FFD46" box="[1108,1229,728,752]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC19616716643D1FB1FFD46" box="[1108,1229,728,752]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961671EF43DFFAFCFD46" box="[1245,1326,727,752]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A543F5FD88FC8E" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- mid and hind femora ventrally with a brown spot at the distal third in
<taxonomicName id="4C643BC5FFC1961676E243F5FBE9FCA2" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[976,1083,764,788]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961676E243F5FBE9FCA2" box="[976,1083,764,788]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616717943F4FB4FFCA3" box="[1099,1181,764,789]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Fig. 61</figureCitation>
), at most with a barely visible band in
<taxonomicName id="4C643BC5FFC1961674434229FE38FC8E" box="[369,490,800,824]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961674434229FE38FC8E" box="[369,490,800,824]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961674CB4217FD9FFC8E" box="[505,589,799,824]" captionStart="FIGURES 5667" captionStartId="13.[151,264,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@13.[151,1436,193,1884]" captionTargetPageId="13" captionText="FIGURES 5667. Details of male imagines of Electrogena brulini (Figs 56, 58, 60, 61, 64, 66) and Electrogena gridellii (Figs 57, 59, 62, 63, 65, 67).Figs 5657: terga; Figs 5859:terga; Figs 60, 62:dorsal face of fore- mid and hind femora; Figs 61, 63: ventral face of fore- mid and hind femora; Figs6465:forewing; Figs 6667: sterna VIIIIXwith styliger." httpUri="https://zenodo.org/record/1094926/files/figure.png" pageId="19" pageNumber="378">Fig. 63</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A5424DFE19FC36" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- male: sclerite joining the compound eyes weakly rounded forming a wide V in
<taxonomicName id="4C643BC5FFC196167175424DFB61FCEA" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1095,1203,836,860]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC196167175424DFB61FCEA" box="[1095,1203,836,860]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961671F1424CFAC5FCEA" box="[1219,1303,836,861]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="19" pageNumber="378">Fig. 84</figureCitation>
), W-shaped in
<taxonomicName id="4C643BC5FFC1961675D04261FE89FC36" box="[226,347,872,896]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961675D04261FE89FC36" box="[226,347,872,896]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC196167458426FFE6CFC36" box="[362,446,871,896]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="19" pageNumber="378">Fig. 85</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A54285FECEFC5A" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- male: titillator massive, straight or concave before the two teeth (rarely one or three), but no basal tip in
<taxonomicName id="4C643BC5FFC1961670B64285FEC3FC7E" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961670B64285FEC3FC7E" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616741342A7FE76FC7E" box="[289,420,943,968]" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="19" pageNumber="378">Figs 7578</figureCitation>
), convex before the unique tooth, than concave until a more or less visible basal tip (
<figureCitation id="135F5CC3FFC19616705842A7FEDDFC5A" captionStart="FIGURES6885" captionStartId="14.[151,264,1884,1906]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@14.[151,1436,193,1863]" captionTargetPageId="14" captionText="FIGURES6885.Electrogena brulini (Figs 68, 72, 7578, 84), Electrogena gridellii (Figs 69, 73, 7981, 85)and Electrogena grandiae(Figs 70,71, 74, 82, 83). Figs6874:penis inventral view;Figs 7583:titillators;Figs 8485:head withemphasis on the posterior sclerit between the eyes." httpUri="https://zenodo.org/record/1094928/files/figure.png" pageId="19" pageNumber="378">Figs 7981</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A542F0FDFEFB82" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- female: subgenital plate covering about half of sternum VIII surface in
<taxonomicName id="4C643BC5FFC1961676DC42F1FB88FBA6" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[1006,1114,1016,1040]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961676DC42F1FB88FBA6" box="[1006,1114,1016,1040]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616715942FFFB6DFBA6" box="[1131,1215,1015,1040]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Fig. 95</figureCitation>
), covering ¾ of the sternum in
<taxonomicName id="4C643BC5FFC1961674714515FE6EFB82" box="[323,444,1052,1076]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961674714515FE6EFB82" box="[323,444,1052,1076]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961674FE4514FDF2FB82" box="[460,544,1052,1077]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Fig. 97</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A54548FA99FBCA" blockId="19.[151,1436,548,1149]" pageId="19" pageNumber="378">
- female: subanal plate tapering to apex, in apical view narrow and lateral margins rolled up towards the inside in
<taxonomicName id="4C643BC5FFC1961675D0456DFE9FFBCA" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[226,333,1124,1148]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961675D0456DFE9FFBCA" box="[226,333,1124,1148]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616746E456CFE0FFBCB" box="[348,477,1124,1149]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Figs 94, 95</figureCitation>
), subanal plate in ventral and apical view rounded in
<taxonomicName id="4C643BC5FFC196167106456DFB7FFBCA" box="[1076,1197,1124,1148]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC196167106456DFB7FFBCA" box="[1076,1197,1124,1148]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616718E456CFAECFBCA" box="[1212,1342,1124,1149]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Figs 96, 97</figureCitation>
).
</paragraph>
<paragraph id="8BDB4046FFC1961675A545A4FEEDFB73" blockId="19.[151,1436,1196,1616]" box="[151,319,1196,1221]" pageId="19" pageNumber="378">At both stages:</paragraph>
<paragraph id="8BDB4046FFC1961675A545D8FD8BFAE2" blockId="19.[151,1436,1196,1616]" pageId="19" pageNumber="378">
- nymph pattern (
<figureCitation id="135F5CC3FFC19616744A45C7FE19FB5E" box="[376,459,1231,1256]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="19" pageNumber="378">Fig. 10</figureCitation>
, d - f) disappearing, only narrow coloured lines remaining laterally on posterior margin in female imagines in
<taxonomicName id="4C643BC5FFC1961674F145FDFDE2FABA" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[451,560,1268,1292]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961674F145FDFDE2FABA" box="[451,560,1268,1292]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616777045FCFD44FABA" box="[578,662,1268,1293]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Fig. 98</figureCitation>
, m), nymph posterior part of the head with only a greyish band (so faint that it is not visible on photos) on the posterior margin, becoming a large centered brown band in female imagines in
<taxonomicName id="4C643BC5FFC19616747E4435FE17FAE2" box="[332,453,1340,1364]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC19616747E4435FE17FAE2" box="[332,453,1340,1364]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961674E64434FDF5FAE2" box="[468,551,1340,1365]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Fig. 99</figureCitation>
, m).
</paragraph>
<paragraph id="8BDB4046FFC1961675A54469FEB6FA52" blockId="19.[151,1436,1196,1616]" pageId="19" pageNumber="378">
- a light line along the inner margins of compound eyes, a dark colouration behind the ocelli (
<figureCitation id="135F5CC3FFC1961670204457FA5CFACE" box="[1298,1422,1375,1400]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="19" pageNumber="378">Figs 8, 12</figureCitation>
); characters still present in female imagines in
<taxonomicName id="4C643BC5FFC1961677F0448DFCFFFA2A" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[706,813,1412,1436]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC1961677F0448DFCFFFA2A" box="[706,813,1412,1436]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616760F448CFC43FA2A" box="[829,913,1412,1437]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Fig. 98</figureCitation>
,
<figureCitation id="135F5CC3FFC1961676AF448CFC79FA2A" box="[925,939,1412,1436]" captionStart="FIGURE 1" captionStartId="4.[151,250,1924,1946]" captionTargetBox="[151,1436,852,1899]" captionTargetId="figure@4.[151,1436,852,1903]" captionTargetPageId="4" captionText="FIGURE 1. Bayesian majority-rule consensus tree of Electrogena reconstructed from the CO1 data set, and designed in iTOL v. 3.5.3 (Letunic &amp; Bork 2016). Electrogena brulini is highlighted in green. Circles on branches indicate Bayesian posterior probabilities&gt; 0.9." httpUri="https://zenodo.org/record/1094914/files/figure.png" pageId="19" pageNumber="378">1</figureCitation>
, n), a dark marking touching the compound eyes, no colouration behind the ocelli (
<figureCitation id="135F5CC3FFC1961677B644AFFD2EFA76" box="[644,764,1447,1472]" captionStart="FIGURES 221" captionStartId="8.[151,264,1830,1852]" captionTargetBox="[151,1436,193,1808]" captionTargetId="figure@8.[151,1436,193,1808]" captionTargetPageId="8" captionText="FIGURES 221. General overview of the nymphs with emphasis on different markings and colourations of Electrogena brulini(Figs 2,4, 6,8, 10, 12, 14, 16, 18,20)and Electrogena gridellii(Figs 3, 5, 7,9,11, 13,15, 17, 19, 21).Figs 23:habitus, dorsal view; Figs 45: habitus, ventral view; Figs 67:head infrontal view; Figs 89: head in dorsal view; Figs 1011: head in postero-dorsal view; Figs 1213: ocelli; Figs 1415: tergites IXX; Figs 1617: pronotum; Figs 1821: mesonotum. Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094916/files/figure.png" pageId="19" pageNumber="378">Figs 9, 13</figureCitation>
); characters still present in female imagines in
<taxonomicName id="4C643BC5FFC19616701244A1FA4EFA76" box="[1312,1436,1448,1472]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC19616701244A1FA4EFA76" box="[1312,1436,1448,1472]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC1961675F944C4FECCFA52" box="[203,286,1484,1509]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">Fig. 99</figureCitation>
,
<figureCitation id="135F5CC3FFC19616741844C4FEEAFA52" box="[298,312,1484,1508]" captionStart="FIGURE 1" captionStartId="4.[151,250,1924,1946]" captionTargetBox="[151,1436,852,1899]" captionTargetId="figure@4.[151,1436,852,1903]" captionTargetPageId="4" captionText="FIGURE 1. Bayesian majority-rule consensus tree of Electrogena reconstructed from the CO1 data set, and designed in iTOL v. 3.5.3 (Letunic &amp; Bork 2016). Electrogena brulini is highlighted in green. Circles on branches indicate Bayesian posterior probabilities&gt; 0.9." httpUri="https://zenodo.org/record/1094914/files/figure.png" pageId="19" pageNumber="378">1</figureCitation>
, n).
</paragraph>
<paragraph id="8BDB4046FFC1961675A544F9FDA9F9E6" blockId="19.[151,1436,1196,1616]" pageId="19" pageNumber="378">
<emphasis id="B9109C54FFC1961675A544F9FF72F9BE" box="[151,160,1521,1544]" italics="true" pageId="19" pageNumber="378">-</emphasis>
terga anterolateral spot small and hidden, posterolateral bigger in
<taxonomicName id="4C643BC5FFC19616769B44F9FBC6F9BE" authorityName="Wagner &amp; Vuataz &amp; Sartori" authorityYear="2017" box="[937,1044,1520,1544]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="brulini">
<emphasis id="B9109C54FFC19616769B44F9FBC6F9BE" box="[937,1044,1520,1544]" italics="true" pageId="19" pageNumber="378">E. brulini</emphasis>
</taxonomicName>
(
<figureCitation id="135F5CC3FFC19616711644E7FBAEF9BE" box="[1060,1148,1519,1544]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Figs 22</figureCitation>
,
<figureCitation id="135F5CC3FFC1961671BA44E7FB74F9BE" box="[1160,1190,1519,1544]" captionStart="FIGURES5255" captionStartId="12.[151,264,1973,1995]" captionTargetBox="[151,1436,193,1952]" captionTargetId="figure@12.[151,1436,193,1952]" captionTargetPageId="12" captionText="FIGURES5255.Male imagines' habitus of Electrogena brulini (Figs 52, 54)and Electrogena gridellii(Figs 53, 55).Figs 52 53:dorsal view; Figs 5455: lateral view. Arrows: markings on the tergite VI." httpUri="https://zenodo.org/record/1094924/files/figure.png" pageId="19" pageNumber="378">54</figureCitation>
,
<figureCitation id="135F5CC3FFC19616718044F8FB29F9BE" box="[1202,1275,1520,1544]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">88, 92</figureCitation>
), anterolateral spot large, posterolateral spot little developed in nymphs, grows and merges with dorsal coloration in
<taxonomicName id="4C643BC5FFC1961670B6471DFECFF9E6" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961670B6471DFECFF9E6" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
imagines (
<figureCitation id="135F5CC3FFC1961674AA473FFE3DF9E6" box="[408,495,1591,1616]" captionStart="FIGURES 2231" captionStartId="9.[151,264,1704,1726]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@9.[151,1436,193,1683]" captionTargetPageId="9" captionText="FIGURES 2231. Nymph details of Electrogena brulini (Figs 22, 24, 27, 28, 30), Electrogena gridellii (Figs 23, 25, 29, 31) and Electrogena ujhelyii (Fig. 26). Figs 2223: abdominal segments VIIX in lateral view; Figs 2426: abdomen in ventral view; Fig. 27: hind femora in postero-ventral view; Fig. 28: latero-ventral general overview; Fig. 29: dorsal face of femora; Figs 3031:thorax in lateral view, emphasis on coxa. Arrows:specific colouration details." httpUri="https://zenodo.org/record/1094918/files/figure.png" pageId="19" pageNumber="378">Figs 23</figureCitation>
,
<figureCitation id="135F5CC3FFC1961674C9473FFDC8F9E6" box="[507,538,1591,1616]" captionStart="FIGURES5255" captionStartId="12.[151,264,1973,1995]" captionTargetBox="[151,1436,193,1952]" captionTargetId="figure@12.[151,1436,193,1952]" captionTargetPageId="12" captionText="FIGURES5255.Male imagines' habitus of Electrogena brulini (Figs 52, 54)and Electrogena gridellii(Figs 53, 55).Figs 52 53:dorsal view; Figs 5455: lateral view. Arrows: markings on the tergite VI." httpUri="https://zenodo.org/record/1094924/files/figure.png" pageId="19" pageNumber="378">55</figureCitation>
,
<figureCitation id="135F5CC3FFC1961677174730FDBCF9E6" box="[549,622,1592,1616]" captionStart="FIGURES 8699" captionStartId="15.[151,264,1800,1822]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@15.[151,1436,193,1778]" captionTargetPageId="15" captionText="FIGURES 8699. Details of female imagines of Electrogenabrulini (Figs 86, 88, 90, 92, 94, 95, 98) and Electrogena gridellii (Figs 87, 89, 91, 93, 96, 97, 99). Figs 8687: terga; Figs 8889: habitus, lateral view; Figs 8889: forewing; Figs 9293: subanal and subgenital plate in lateral view; Figs 94, 96: subanal plate in apical view; Figs95, 97:subanal andsubgenital plate; Figs 9899:head and pronotum.Arrows: specific colouration details." httpUri="https://zenodo.org/record/1094930/files/figure.png" pageId="19" pageNumber="378">89, 93</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E13CDFFC1961175F54777FB85FBE1" lastPageId="20" lastPageNumber="379" pageId="19" pageNumber="378" type="description">
<paragraph id="8BDB4046FFC1961675F54777FDC2F90B" blockId="19.[151,1436,1663,1725]" pageId="19" pageNumber="378">
<emphasis id="B9109C54FFC1961675F54777FD7BF92E" bold="true" box="[199,681,1663,1688]" pageId="19" pageNumber="378">Rationales for the proposed synonymy.</emphasis>
The type material of
<taxonomicName id="4C643BC5FFC1961676804789FB98F921" authorityName=": Zurwerra &amp; Tomka" authorityYear="1986" box="[946,1098,1664,1688]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="vipavensis">
<emphasis id="B9109C54FFC1961676804789FB98F921" box="[946,1098,1664,1688]" italics="true" pageId="19" pageNumber="378">E. vipavensis</emphasis>
</taxonomicName>
currently housed in MZL is composed of the following items:
</paragraph>
<paragraph id="8BDB4046FFC1961175A547E5FAA1FF19" blockId="19.[151,1437,1772,2013]" lastBlockId="20.[151,1437,151,861]" lastPageId="20" lastPageNumber="379" pageId="19" pageNumber="378">
- the
<typeStatus id="54DFFEE4FFC1961675CB47E4FE8CF8B3" box="[249,350,1772,1797]" pageId="19" pageNumber="378" type="holotype">holotype</typeStatus>
, a male subimago (N° 4155) with corresponding exuvia on slide (N° 4155_D_a_male). Subimago: sclerite joining the compound eyes W-shaped, styliger straight with lateral humps, all the colouration faded, but the drawings in the original description fits with our contrasted
<taxonomicName id="4C643BC5FFC1961671BA463DFAD3F8FA" box="[1160,1281,1844,1868]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="378" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC1961671BA463DFAD3F8FA" box="[1160,1281,1844,1868]" italics="true" pageId="19" pageNumber="378">E. gridellii</emphasis>
</taxonomicName>
subimagines: anterior margin of terga IIIV with a pair of subtriangular markings pointed backwards and a spot between them, posterior margin darkened, posterolateral part of terga IIIV with a dark marking near the border fused with the anterior margin marking of next tergum, an oval white area oblique from the anterolateral corner. Exuvia: all femora with long, narrow and pointed bristles on the dorsal face, N_BVF: 0, N_CLW: 2, N_PRO: 11 / 11. According to all observed characters we found no difference between the
<typeStatus id="54DFFEE4FFC696117168419FFB6EFF06" box="[1114,1212,151,176]" pageId="20" pageNumber="379" type="holotype">holotype</typeStatus>
and
<taxonomicName id="4C643BC5FFC6961171C14191FABEFF06" box="[1267,1388,152,176]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC6961171C14191FABEFF06" box="[1267,1388,152,176]" italics="true" pageId="20" pageNumber="379">E. gridellii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB4046FFC6961175A541B5FDF0FE42" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="379">
-
<typeStatus id="54DFFEE4FFC6961175F541B5FEF5FF63" box="[199,295,189,213]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
female abdominal segments VIIX (N° 4175) with corresponding exuvia on slide (N° 4175_a_fem) (assumed to be the
<typeStatus id="54DFFEE4FFC69611749741E8FDD7FF4E" box="[421,517,224,248]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
drawn in the original description because of the precision « fem » on the exuvia slide) and the forewings on slide (N° 4175_b). According to the original drawing, the pattern of the terga IIIV in lateral view is composed by the following elements: a large oval white area oblique from anterolateral corner, an anterolateral spot, posterolateral spots, a lateral dark streak at base of terga, central part of the subanal plate tapering towards apex. Exuvia: all femora with long, narrow parallel and truncate bristles on the dorsal face, N_BVF: numerous, N_CLW: 2, N_PRO: 9 / 9. According to all observed characters we found no difference between this
<typeStatus id="54DFFEE4FFC6961174E040B0FDE1FE66" box="[466,563,440,464]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
and
<taxonomicName id="4C643BC5FFC69611775A40B1FD0EFE66" baseAuthorityName="Sowa" baseAuthorityYear="1981" box="[616,732,440,464]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="ujhelyii">
<emphasis id="B9109C54FFC69611775A40B1FD0EFE66" box="[616,732,440,464]" italics="true" pageId="20" pageNumber="379">E. ujhelyii</emphasis>
</taxonomicName>
except that the 23 posterolateral spots of the original drawing are usually fused in
<taxonomicName id="4C643BC5FFC69611749A40D5FDC9FE42" baseAuthorityName="Sowa" baseAuthorityYear="1981" box="[424,539,476,500]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="ujhelyii">
<emphasis id="B9109C54FFC69611749A40D5FDC9FE42" box="[424,539,476,500]" italics="true" pageId="20" pageNumber="379">E. ujhelyii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB4046FFC6961175A54308FDF2FDE9" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="379">
-
<typeStatus id="54DFFEE4FFC6961175F54308FEF5FDAE" box="[199,295,512,536]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
female abdominal segments VIIX (N°4291) with corresponding exuvia on slide (slide 4291_a). Exuvia: all femora with long, narrow and pointed bristles on the dorsal face, N_BVF: 0, N_CLW:2, N_PRO: 9. New identification:
<taxonomicName id="4C643BC5FFC6961174954340FDCBFDD6" box="[423,537,584,608]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC6961174954340FDCBFDD6" box="[423,537,584,608]" italics="true" pageId="20" pageNumber="379">E.gridellii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB4046FFC6961175A54365FC40FD11" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="379">
-
<typeStatus id="54DFFEE4FFC6961175F54365FEF5FD33" box="[199,295,621,645]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
female abdominal segments VIIX (N° 4223). Central part of the subanal plate tapering towards apex, but not sufficient for a secure identification:
<emphasis id="B9109C54FFC6961177CF4399FC59FD1E" box="[765,907,656,680]" italics="true" pageId="20" pageNumber="379">
E. cf
<taxonomicName id="4C643BC5FFC6961176054398FC59FD1E" baseAuthorityName="Sowa" baseAuthorityYear="1981" box="[823,907,656,680]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="ujhelyii">ujhelyii</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="8BDB4046FFC6961175A543BDFC67FD59" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="379">
-
<typeStatus id="54DFFEE4FFC6961175F543BDFEF5FD7B" box="[199,295,693,717]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
female abdominal segments VIIX (N° 4198) plus a second sample with 5 legs and 1 forewing (same number N° 4198). Not sufficient for identification:
<emphasis id="B9109C54FFC69611763443D0FC7CFD46" box="[774,942,728,752]" italics="true" pageId="20" pageNumber="379">
<taxonomicName id="4C643BC5FFC69611763443D0FC5CFD46" authorityName="Zurwerra &amp; Tomka" authorityYear="1985" box="[774,910,728,752]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="genus">Electrogena</taxonomicName>
sp
</emphasis>
.
</paragraph>
<paragraph id="8BDB4046FFC6961175A543F4FD83FCEA" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="379">
Together with the
<typeStatus id="54DFFEE4FFC69611745C43F5FE4CFCA3" box="[366,414,765,789]" pageId="20" pageNumber="379">type</typeStatus>
material and from the same sampling data: one exuvia named
<taxonomicName id="4C643BC5FFC6961171B743F5FAD0FCA2" baseAuthorityName="Tanasijevic" baseAuthorityYear="1975" box="[1157,1282,765,788]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="ozrensis">
<emphasis id="B9109C54FFC6961171B743F5FAD0FCA2" box="[1157,1282,765,788]" italics="true" pageId="20" pageNumber="379">E. ozrensis</emphasis>
</taxonomicName>
on slide (N° 4167_a). All femora with long, narrow and pointed bristles on the dorsal face, N_BVF: 0, N_CLW:2, N_PRO: 14. New identification:
<taxonomicName id="4C643BC5FFC6961174E3424DFD98FCEA" box="[465,586,836,860]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC6961174E3424DFD98FCEA" box="[465,586,836,860]" italics="true" pageId="20" pageNumber="379">E. gridellii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB4046FFC6961175F54284FB4EFBA6" blockId="20.[151,1437,908,1544]" pageId="20" pageNumber="379">
Our examination of the
<typeStatus id="54DFFEE4FFC6961174E84285FDD8FC13" box="[474,522,909,933]" pageId="20" pageNumber="379">type</typeStatus>
material of
<taxonomicName id="4C643BC5FFC6961177AA4285FCFCFC12" authorityName=": Zurwerra &amp; Tomka" authorityYear="1986" box="[664,814,909,932]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="vipavensis">
<emphasis id="B9109C54FFC6961177AA4285FCFCFC12" box="[664,814,909,932]" italics="true" pageId="20" pageNumber="379">E. vipavensis</emphasis>
</taxonomicName>
indicates that the
<typeStatus id="54DFFEE4FFC6961171304284FBB5FC13" box="[1026,1127,908,933]" pageId="20" pageNumber="379" type="holotype">holotype</typeStatus>
, male subimago (N° 4155) and its exuvia on slide (N° 4155_D_a_male) and the female
<typeStatus id="54DFFEE4FFC69611767642B8FC77FC7E" box="[836,933,944,968]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
exuvia on slide (N° 4291_a) are conspecific with
<taxonomicName id="4C643BC5FFC6961175EA42DDFE8BFC5A" box="[216,345,980,1004]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC6961175EA42DDFE8BFC5A" box="[216,345,980,1004]" italics="true" pageId="20" pageNumber="379">E. gridellii</emphasis>
</taxonomicName>
. Therefore, we propose that
<taxonomicName id="4C643BC5FFC6961177F742DCFACAFC5B" authority="Zurwerra &amp; Tomka, 1986" authorityName=": Zurwerra &amp; Tomka" authorityYear="1986" box="[709,1304,980,1005]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="vipavensis">
<emphasis id="B9109C54FFC6961177F742DCFC03FC5A" box="[709,977,980,1004]" italics="true" pageId="20" pageNumber="379">Electrogena vipavensis</emphasis>
<bibRefCitation id="EFF53DB7FFC6961176D242DDFACAFC5B" author="Zurwerra" box="[992,1304,980,1005]" pageId="20" pageNumber="379" refString="Zurwerra, A. &amp; Tomka, I. (1986) Drei neue Arten der Gattung Electrogena Zurwerra &amp; Tomka, 1985, aus Sudeuropa (Ephemeroptera, Heptageniidae). Bulletin de la Societe fribourgeoise des Sciences naturelles, 75, 216 - 230." type="journal article" year="1986">Zurwerra &amp; Tomka, 1986</bibRefCitation>
</taxonomicName>
should be considered as a subjective junior synonym of
<taxonomicName id="4C643BC5FFC6961177A542F0FBFCFBA6" authority="Grandi, 1953" authorityName="Grandi" authorityYear="1953" box="[663,1070,1015,1040]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="gridellii">
<emphasis id="B9109C54FFC6961177A542F0FC52FBA6" box="[663,896,1016,1040]" italics="true" pageId="20" pageNumber="379">Electrogena gridellii</emphasis>
(
<bibRefCitation id="EFF53DB7FFC6961176BD42FFFBF7FBA6" author="Grandi" box="[911,1061,1015,1040]" pageId="20" pageNumber="379" refString="Grandi, M. (1953) Contributi allo studio degli Efemeroidei italiani XVII. Ecdyonuridae. Bolletino dell'Istituto di Entomologia dell'Universita di Bologna, 19, 307 - 386." type="journal article" year="1953">Grandi, 1953</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A223212FFFC69611710642FFFB4EFBA6" box="[1076,1180,1015,1040]" pageId="20" pageNumber="379">
<emphasis id="B9109C54FFC69611710642FFFB4EFBA6" bold="true" box="[1076,1180,1015,1040]" pageId="20" pageNumber="379">syn. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDB4046FFC6961175F54514FB85FBE1" blockId="20.[151,1437,908,1544]" pageId="20" pageNumber="379">
After comparison of the original drawing with our material and re-investigation of the female
<typeStatus id="54DFFEE4FFC6961171DB4515FA98FB83" box="[1257,1354,1053,1077]" pageId="20" pageNumber="379" type="paratype">paratype</typeStatus>
exuvia slide (N° 4175_a_fem), we consider that this specimen is conspecific with
<taxonomicName id="4C643BC5FFC6961176EF4549FB82FBEE" baseAuthorityName="Sowa" baseAuthorityYear="1981" box="[989,1104,1088,1112]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="ujhelyii">
<emphasis id="B9109C54FFC6961176EF4549FB82FBEE" box="[989,1104,1088,1112]" italics="true" pageId="20" pageNumber="379">E. ujhelyii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E13CDFFC6961175F5456CFBDAF9BE" pageId="20" pageNumber="379" type="materials_examined">
<paragraph id="8BDB4046FFC6961175F5456CFCF1FA76" blockId="20.[151,1437,908,1544]" pageId="20" pageNumber="379">
<materialsCitation id="3B0C4A1BFFC6961175F5456CFE3AFA76" ID-GBIF-Occurrence="1703280313" collectingDate="2015-04-06" collectingDateMax="2015-05-23" collectingDateMin="2015-04-06" collectionCode="MZL" collectorName="A. Wagner &amp; Manzano &amp; Fiume Natisone &amp; Oleis &amp; F. Desio" county="Material" elevation="91" latitude="46.02" location="Rosandra" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" pageId="20" pageNumber="379" specimenCount="212" specimenCount-female="84" specimenCount-male="128" stateProvince="Koper">
<emphasis id="B9109C54FFC6961175F5456CFE68FBCB" bold="true" box="[199,442,1124,1149]" pageId="20" pageNumber="379">
<collectingCounty id="62BA38CAFFC6961175F5456CFEFDFBCB" box="[199,303,1124,1149]" pageId="20" pageNumber="379">Material</collectingCounty>
examined:
</emphasis>
<specimenCount id="9D628BCFFFC6961174FA456CFDFAFBCB" box="[456,552,1124,1149]" pageId="20" pageNumber="379" type="male">97 male</specimenCount>
imagines,
<specimenCount id="9D628BCFFFC696117783456CFCFAFBCB" box="[689,808,1124,1149]" pageId="20" pageNumber="379" type="female">45 female</specimenCount>
imagines, 8 nymphs, 140 nymph exuviae,
<specimenCount id="9D628BCFFFC69611700E456CFA4EFBCB" box="[1340,1436,1124,1149]" pageId="20" pageNumber="379" type="male">31 male</specimenCount>
subimagines,
<specimenCount id="9D628BCFFFC6961174014580FE76FB16" box="[307,420,1159,1184]" pageId="20" pageNumber="379" type="female">39 female</specimenCount>
subimagines: IT / FV,
<collectingMunicipality id="6BBFDA3CFFC69611779C4580FC6FFB16" box="[686,957,1159,1184]" pageId="20" pageNumber="379">San Dorligo della Valle</collectingMunicipality>
,
<location id="8EBB169DFFC6961176FB458FFBE4FB16" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961176FB458FFBE4FB16" box="[969,1078,1159,1184]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Rosandra" pageId="20" pageNumber="379" stateProvince="Koper">Rosandra</location>
,
<location id="8EBB169DFFC696117171458FFAC7FB16" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC696117171458FFAC7FB16" box="[1091,1301,1159,1184]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Bagnoli Superiore" pageId="20" pageNumber="379" stateProvince="Koper">Bagnoli Superiore</location>
,
<geoCoordinate id="EE502681FFC696117010458FFA4EFB16" box="[1314,1436,1159,1184]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="45.6195">45.6195°N</geoCoordinate>
<geoCoordinate id="EE502681FFC6961175A545A4FEDDFB73" box="[151,271,1196,1221]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.8654">13.8654°E</geoCoordinate>
<elevation id="0049A775FFC69611742745A4FE83FB72" box="[277,337,1196,1220]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="20" pageNumber="379" unit="m" value="91.0">
<quantity id="4C9CEDA3FFC69611742745A4FE83FB72" box="[277,337,1196,1220]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="20" pageNumber="379" unit="m" value="91.0">91 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC69611746F45A4FE19FB73" box="[349,459,1196,1221]" pageId="20" pageNumber="379" value="2015-04-09">
<date id="FFDA6686FFC69611746F45A4FE19FB73" box="[349,459,1196,1221]" pageId="20" pageNumber="379" value="2015-04-09">9.IV.2015</date>
</collectingDate>
(emerged
<collectingDate id="EF9E9F6EFFC69611777145A4FD29FB73" box="[579,763,1196,1221]" pageId="20" pageNumber="379" value="2015-04-09" valueMax="2015-05-23" valueMin="2015-04-09">
<date id="FFDA6686FFC69611777145A4FD29FB73" box="[579,763,1196,1221]" pageId="20" pageNumber="379" value="2015-04-09" valueMax="2015-05-23" valueMin="2015-04-09">9.IV23.V.2015</date>
</collectingDate>
); 75 nymphs: same locality,
<collectingDate id="EF9E9F6EFFC69611710E45A4FB7CFB73" box="[1084,1198,1196,1221]" pageId="20" pageNumber="379" value="2015-04-07">
<date id="FFDA6686FFC69611710E45A4FB7CFB73" box="[1084,1198,1196,1221]" pageId="20" pageNumber="379" value="2015-04-07">7.IV.2015</date>
</collectingDate>
; 11 nymphs: IT / FV,
<location id="8EBB169DFFC6961175A545C7FED7FB5E" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961175A545C7FED7FB5E" box="[151,261,1231,1256]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Povoletto" pageId="20" pageNumber="379" stateProvince="Koper">Povoletto</location>
,
<location id="8EBB169DFFC69611742245C7FE66FB5E" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC69611742245C7FE66FB5E" box="[272,436,1231,1256]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Via Reclusane" pageId="20" pageNumber="379" stateProvince="Koper">Via Reclusane</location>
,
<geoCoordinate id="EE502681FFC6961174F245C7FDE8FB5E" box="[448,570,1231,1256]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="46.2658">46.2658°N</geoCoordinate>
<geoCoordinate id="EE502681FFC69611777345C7FD6BFB5E" box="[577,697,1231,1256]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.3126">13.3126°E</geoCoordinate>
<elevation id="0049A775FFC6961177F245C7FCDBFB51" box="[704,777,1231,1256]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="20" pageNumber="379" unit="m" value="158.0">
<quantity id="4C9CEDA3FFC6961177F245C7FCDBFB51" box="[704,777,1231,1256]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="20" pageNumber="379" unit="m" value="158.0">158 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC69611762745C7FC55FB5E" box="[789,903,1231,1256]" pageId="20" pageNumber="379" value="2015-04-06">
<date id="FFDA6686FFC69611762745C7FC55FB5E" box="[789,903,1231,1256]" pageId="20" pageNumber="379" value="2015-04-06">6.IV.2015</date>
</collectingDate>
; 1 nymph: IT / FV,
<location id="8EBB169DFFC69611715645C7FB69FB5E" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC69611715645C7FB69FB5E" box="[1124,1211,1231,1256]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Attimis" pageId="20" pageNumber="379" stateProvince="Koper">Attimis</location>
,
<location id="8EBB169DFFC6961171F445C7FAE2FB5E" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961171F445C7FAE2FB5E" box="[1222,1328,1231,1256]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Rio Valle" pageId="20" pageNumber="379" stateProvince="Koper">Rio Valle</location>
tributary,
<geoCoordinate id="EE502681FFC6961175A545FCFEC3FABB" box="[151,273,1268,1293]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="46.1748">46.1748°N</geoCoordinate>
<geoCoordinate id="EE502681FFC69611742A45FCFE5CFABA" box="[280,398,1268,1292]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.3211">13.3211°E</geoCoordinate>
<elevation id="0049A775FFC6961174A745FCFE0CFABA" box="[405,478,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="1.77" pageId="20" pageNumber="379" unit="m" value="177.0">
<quantity id="4C9CEDA3FFC6961174A745FCFE0CFABA" box="[405,478,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="1.77" pageId="20" pageNumber="379" unit="m" value="177.0">177 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC6961174DB45FCFD89FABB" box="[489,603,1268,1293]" pageId="20" pageNumber="379" value="2015-04-06">
<date id="FFDA6686FFC6961174DB45FCFD89FABB" box="[489,603,1268,1293]" pageId="20" pageNumber="379" value="2015-04-06">6.IV.2015</date>
</collectingDate>
; 6 nymphs: IT / FV,
<location id="8EBB169DFFC69611767245FCFC45FABB" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC69611767245FCFC45FABB" box="[832,919,1268,1293]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Attimis" pageId="20" pageNumber="379" stateProvince="Koper">Attimis</location>
,
<location id="8EBB169DFFC69611769345FDFB4BFABB" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC69611769345FDFB4BFABB" box="[929,1177,1268,1293]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Torrente Racchiusano" pageId="20" pageNumber="379" stateProvince="Koper">Torrente Racchiusano</location>
,
<geoCoordinate id="EE502681FFC69611719645FCFACCFABB" box="[1188,1310,1268,1293]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="46.1733">46.1733°N</geoCoordinate>
<geoCoordinate id="EE502681FFC69611701745FCFA4EFABA" box="[1317,1436,1268,1292]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.333">13.3330°E</geoCoordinate>
<elevation id="0049A775FFC6961175A54410FF0DFA86" box="[151,223,1304,1328]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="379" unit="m" value="230.0">
<quantity id="4C9CEDA3FFC6961175A54410FF0DFA86" box="[151,223,1304,1328]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="379" unit="m" value="230.0">230 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC6961175D8441FFE8EFA86" box="[234,348,1303,1328]" pageId="20" pageNumber="379" value="2015-04-06">
<date id="FFDA6686FFC6961175D8441FFE8EFA86" box="[234,348,1303,1328]" pageId="20" pageNumber="379" value="2015-04-06">6.IV.2015</date>
</collectingDate>
; 1 nymph: IT / FV,
<location id="8EBB169DFFC6961177064410FD5EFA86" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961177064410FD5EFA86" box="[564,652,1304,1328]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Stregna" pageId="20" pageNumber="379" stateProvince="Koper">Stregna</location>
,
<location id="8EBB169DFFC6961177A44411FC88FA86" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961177A44411FC88FA86" box="[662,858,1303,1328]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Torrente Erbezzo" pageId="20" pageNumber="379" stateProvince="Koper">Torrente Erbezzo</location>
,
<geoCoordinate id="EE502681FFC696117656441FFC0CFA86" box="[868,990,1303,1328]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="46.1287">46.1287°N</geoCoordinate>
<geoCoordinate id="EE502681FFC6961176D7441FFB8EFA86" box="[997,1116,1303,1328]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.5699">13.5699°E</geoCoordinate>
<elevation id="0049A775FFC6961171504410FB79FA86" box="[1122,1195,1304,1328]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="20" pageNumber="379" unit="m" value="232.0">
<quantity id="4C9CEDA3FFC6961171504410FB79FA86" box="[1122,1195,1304,1328]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="20" pageNumber="379" unit="m" value="232.0">232 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC696117184441FFAFAFA86" box="[1206,1320,1303,1328]" pageId="20" pageNumber="379" value="2015-04-06">
<date id="FFDA6686FFC696117184441FFAFAFA86" box="[1206,1320,1303,1328]" pageId="20" pageNumber="379" value="2015-04-06">6.IV.2015</date>
</collectingDate>
; 1 nymph: IT / FV,
<location id="8EBB169DFFC6961175CE4435FE8CFAE2" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961175CE4435FE8CFAE2" box="[252,350,1341,1364]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Prepotto" pageId="20" pageNumber="379" stateProvince="Koper">Prepotto</location>
,
<location id="8EBB169DFFC69611745E4434FDD4FAE3" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC69611745E4434FDD4FAE3" box="[364,518,1340,1365]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Fiume Judrio" pageId="20" pageNumber="379" stateProvince="Koper">Fiume Judrio</location>
,
<geoCoordinate id="EE502681FFC6961177214434FD5FFAE3" box="[531,653,1340,1365]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="46.0826">46.0826°N</geoCoordinate>
<geoCoordinate id="EE502681FFC6961177A54434FCDDFAE3" box="[663,783,1340,1365]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.5434">13.5434°E</geoCoordinate>
<elevation id="0049A775FFC6961176254434FCB1FAE2" box="[791,867,1340,1365]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="20" pageNumber="379" unit="m" value="145.0">
<quantity id="4C9CEDA3FFC6961176254434FCB1FAE2" box="[791,867,1340,1365]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="20" pageNumber="379" unit="m" value="145.0">145 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC6961176434434FC31FAE3" box="[881,995,1340,1365]" pageId="20" pageNumber="379" value="2015-04-06">
<date id="FFDA6686FFC6961176434434FC31FAE3" box="[881,995,1340,1365]" pageId="20" pageNumber="379" value="2015-04-06">6.IV.2015</date>
</collectingDate>
; 15 nymphs: IT / FV,
<location id="8EBB169DFFC6961171D74435FA95FAE2" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961171D74435FA95FAE2" box="[1253,1351,1341,1364]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Prepotto" pageId="20" pageNumber="379" stateProvince="Koper">Prepotto</location>
,
<location id="8EBB169DFFC6961170664434FF0FFACE" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961170664434FF0FFACE" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Fiume Judrio" pageId="20" pageNumber="379" stateProvince="Koper">Fiume Judrio</location>
tributary,
<geoCoordinate id="EE502681FFC6961174634457FE19FACE" box="[337,459,1375,1400]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="46.0827">46.0827°N</geoCoordinate>
<geoCoordinate id="EE502681FFC6961174E14457FD98FACE" box="[467,586,1375,1400]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.5427">13.5427°E</geoCoordinate>
<elevation id="0049A775FFC6961177634468FD49FACE" box="[593,667,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="20" pageNumber="379" unit="m" value="147.0">
<quantity id="4C9CEDA3FFC6961177634468FD49FACE" box="[593,667,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="20" pageNumber="379" unit="m" value="147.0">147 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC6961177944457FCCAFACE" box="[678,792,1375,1400]" pageId="20" pageNumber="379" value="2015-04-06">
<date id="FFDA6686FFC6961177944457FCCAFACE" box="[678,792,1375,1400]" pageId="20" pageNumber="379" value="2015-04-06">6.IV.2015</date>
</collectingDate>
; 1 nymph: SL / IS,
<collectingRegion id="49A08EA4FFC6961176C64469FBEEFACE" box="[1012,1084,1377,1400]" country="Slovenia" name="Koper-Capodistria" pageId="20" pageNumber="379">Koper</collectingRegion>
,
<location id="8EBB169DFFC6961171754457FB45FACE" LSID="urn:lsid:plazi:treatment:03CDF150FFC2961175A5466CFBDAF9BE:8EBB169DFFC6961171754457FB45FACE" box="[1095,1175,1375,1400]" county="Material" latitude="46.02" longLatPrecision="675" longitude="13.38" municipality="San Dorligo della Valle" name="Rizana" pageId="20" pageNumber="379" stateProvince="Koper">Rizana</location>
,
<geoCoordinate id="EE502681FFC6961171914457FACFFACE" box="[1187,1309,1375,1400]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="5" value="45.5328">45.5328°N</geoCoordinate>
<geoCoordinate id="EE502681FFC6961170174468FA4EFACE" box="[1317,1436,1376,1400]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="5" value="13.8729">13.8729°E</geoCoordinate>
<elevation id="0049A775FFC6961175A5448CFF04FA2A" box="[151,214,1412,1437]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="20" pageNumber="379" unit="m" value="66.0">
<quantity id="4C9CEDA3FFC6961175A5448CFF04FA2A" box="[151,214,1412,1437]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="20" pageNumber="379" unit="m" value="66.0">66 m</quantity>
</elevation>
,
<collectingDate id="EF9E9F6EFFC6961175D4448CFE84FA2B" box="[230,342,1412,1437]" pageId="20" pageNumber="379" value="2015-04-07">
<date id="FFDA6686FFC6961175D4448CFE84FA2B" box="[230,342,1412,1437]" pageId="20" pageNumber="379" value="2015-04-07">7.IV.2015</date>
</collectingDate>
,
<collectorName id="26912590FFC696117454448CFE30FA2B" box="[358,482,1412,1437]" pageId="20" pageNumber="379">A. Wagner</collectorName>
leg. [
<collectionCode id="ED75D883FFC69611771B448DFDBFFA2A" box="[553,621,1413,1436]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="20" pageNumber="379">MZL</collectionCode>
]; 1 nymph: IT / FV,
<collectorName id="26912590FFC696117655448DFC00FA2A" box="[871,978,1413,1436]" pageId="20" pageNumber="379">Manzano</collectorName>
,
<collectorName id="26912590FFC6961176D0448CFB4BFA2B" box="[994,1177,1412,1437]" pageId="20" pageNumber="379">Fiume Natisone</collectorName>
basin,
<collectorName id="26912590FFC6961171C3448CFAFDFA2B" box="[1265,1327,1412,1437]" pageId="20" pageNumber="379">Oleis</collectorName>
,
<geoCoordinate id="EE502681FFC69611700C448CFA4FFA2B" box="[1342,1437,1412,1437]" direction="north" orientation="latitude" pageId="20" pageNumber="379" precision="555" value="46.02">46.02°N</geoCoordinate>
<geoCoordinate id="EE502681FFC6961175A544A0FF20FA76" box="[151,242,1448,1472]" direction="east" orientation="longitude" pageId="20" pageNumber="379" precision="555" value="13.38">13.38°E</geoCoordinate>
<elevation id="0049A775FFC6961175CB44AFFEE6FA76" box="[249,308,1447,1472]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="379" unit="m" value="75.0">
<quantity id="4C9CEDA3FFC6961175CB44AFFEE6FA76" box="[249,308,1447,1472]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="379" unit="m" value="75.0">75 m</quantity>
</elevation>
,
<collectorName id="26912590FFC69611747244A1FE4EFA76" box="[320,412,1447,1472]" pageId="20" pageNumber="379">F. Desio</collectorName>
leg, E
</materialsCitation>
. Bauernfeind det. [NHMW]
</paragraph>
<paragraph id="8BDB4046FFC6961175F544C5FBDAF9BE" blockId="20.[151,1437,908,1544]" pageId="20" pageNumber="379">
<taxonomicName id="4C643BC5FFC6961175F544C5FE8FFA52" authorityName=": Zurwerra &amp; Tomka" authorityYear="1986" box="[199,349,1485,1508]" class="Insecta" family="Heptageniidae" genus="Electrogena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="379" phylum="Arthropoda" rank="species" species="vipavensis">
<emphasis id="B9109C54FFC6961175F544C5FE8FFA52" box="[199,349,1485,1508]" italics="true" pageId="20" pageNumber="379">E. vipavensis</emphasis>
</taxonomicName>
: one male subimago
<typeStatus id="54DFFEE4FFC69611776944C4FD6FFA53" box="[603,701,1484,1509]" pageId="20" pageNumber="379" type="holotype">holotype</typeStatus>
(N° 4155 with exuvia on slide),
<specimenCount id="9D628BCFFFC69611717644C4FB7AFA53" box="[1092,1192,1484,1509]" pageId="20" pageNumber="379" type="female">4 female</specimenCount>
<typeStatus id="54DFFEE4FFC69611718144C5FAF3FA53" box="[1203,1313,1485,1509]" pageId="20" pageNumber="379" type="paratype">paratypes</typeStatus>
, SL / GO,
<collectingRegion id="49A08EA4FFC6961175A544E7FEC8F9BE" box="[151,282,1519,1544]" country="Slovenia" name="Ajdovscina" pageId="20" pageNumber="379">Ajdovscina</collectingRegion>
, Stream Vipava,
<date id="FFDA6686FFC6961174E544E7FD84F9BE" box="[471,598,1519,1544]" pageId="20" pageNumber="379" value="1983-04-16">16.IV.1983</date>
, Hefti, Tomka &amp; Zurwerra leg. [
<collectionCode id="ED75D883FFC6961176F244F9FBD3F9BE" box="[960,1025,1521,1544]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="20" pageNumber="379">MZL</collectionCode>
]
</paragraph>
</subSubSection>
</treatment>
</document>