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<document id="27B265986E6F124B46F41E06AA4B221D" ID-DOI="10.5252/zoosystema2023v45a9" ID-ISSN="1638-9387" ID-Zenodo-Dep="8071253" ID-ZooBank="urn:lsid:zoobank.org:pub:69C34731-8C25-4A1E-B336-B222CD3CBAC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686747199180" checkinUser="felipe" docAuthor="Guinot, Danièle" docDate="2023" docId="03CDBE74938EB5AECEE4FF17FBB8F9B5" docLanguage="en" docName="Zoosystema.45.9.225-372.pdf" docOrigin="Zoosystema 45 (9)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Medorippe margaretha Lőrenthey 1929" docType="treatment" docVersion="7" lastPageNumber="361" masterDocId="FFF4C60C9304B524CD15FFC0FF9AFFF8" masterDocTitle="A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system" masterLastPageNumber="372" masterPageNumber="225" pageNumber="361" updateTime="1706900238514" updateUser="felipe" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DBB0B6F49C8BB39BF5CD147D0B9D55F7">A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system</mods:title>
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<mods:namePart id="7E3F0B05DBF88D91B9B69D4D2C9C123E">Guinot, Danièle</mods:namePart>
<mods:affiliation id="021F4BB24B85E027FA30B1E4F24797D0">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) daniele. guinot @ mnhn. fr Dedicated to the memory of my colleague and dearest friend Ngan Kee NG (1966 - 2022)</mods:affiliation>
<mods:nameIdentifier id="11E16A09BA84233FDA64AC16C4B97A93" type="email">guinot@mnhn.fr</mods:nameIdentifier>
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<mods:title id="A641F1F5CF15BE671385459E9D848C89">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="8D0A7E2636BAE11BA75FC434DFF02AA8">
<mods:date id="277B2159A1D5CD71F9CA9E9C052CB522">2023</mods:date>
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<mods:number id="804AA2B96ED89D52437F97C80A237928">2023-06-05</mods:number>
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<mods:number id="DA1E19B5E2F742F17F4E667809E99A2F">45</mods:number>
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<mods:classification id="E89FF7F901B2B2AD47B6DC8FB6CF1B2A">journal article</mods:classification>
<mods:identifier id="9B7C1A4FF9427140F5D784D0F6A44B26" type="DOI">10.5252/zoosystema2023v45a9</mods:identifier>
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<treatment id="03CDBE74938EB5AECEE4FF17FBB8F9B5" ID-DOI="http://doi.org/10.5281/zenodo.8235625" ID-Zenodo-Dep="8235625" LSID="urn:lsid:plazi:treatment:03CDBE74938EB5AECEE4FF17FBB8F9B5" httpUri="http://treatment.plazi.org/id/03CDBE74938EB5AECEE4FF17FBB8F9B5" lastPageNumber="361" pageId="138" pageNumber="361">
<subSubSection id="C37E5CE9938EB5AECEE4FF17FACDFEE9" pageId="138" pageNumber="361" type="nomenclature">
<paragraph id="8BDB0F62938EB5AECEE4FF17FACDFEE9" blockId="138.[901,1367,215,277]" pageId="138" pageNumber="361">
<heading id="D093B80E938EB5AECEE4FF17FACDFEE9" centered="true" fontSize="11" level="2" pageId="138" pageNumber="361" reason="2">
<taxonomicName id="4C6474E1938EB5AECEE4FF17FAD5FEE9" ID-CoL="f59e52e0-5e5a-461b-bea8-cb1214b8132e" authorityName="Lőrenthey" authorityYear="1929" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="margaretha">
<emphasis id="B910D370938EB5AECEE4FF17FB70FF0A" bold="true" box="[1009,1258,215,242]" italics="true" pageId="138" pageNumber="361">Medorippe margaretha</emphasis>
(Lőrenthey
<emphasis id="B910D370938EB5AEC916FF38FB83FEE9" box="[1027,1049,248,273]" italics="true" pageId="138" pageNumber="361">in</emphasis>
Lőrenthey &amp; Beurlen, 1929
</taxonomicName>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9938EB5AECE38FEF1FA35FD01" pageId="138" pageNumber="361" type="reference_group">
<paragraph id="8BDB0F62938EB5AECE38FEF1FBC3FE49" blockId="138.[811,1455,304,434]" pageId="138" pageNumber="361">
<taxonomicName id="4C6474E1938EB5AECE38FEF1FC7DFEB0" ID-CoL="30d269b7-f4b8-46ba-93d3-b47a3099483f" authorityName="Lorenthey" authorityYear="1929" box="[813,999,304,328]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="margaretha">
<emphasis id="B910D370938EB5AECE38FEF1FC7DFEB0" box="[813,999,304,328]" italics="true" pageId="138" pageNumber="361">Dorippe margaretha</emphasis>
</taxonomicName>
Lőrenthey
<emphasis id="B910D370938EB5AEC941FEF1FBF2FEB0" box="[1108,1128,305,328]" italics="true" pageId="138" pageNumber="361">in</emphasis>
Lőrenthey &amp; Beurlen, 1929: 135, pl. 6, fig. 1a-c. — Lőrenthey 1911: 528 (
<emphasis id="B910D370938EB5AEC9BAFE8BFAACFE9A" box="[1199,1334,330,354]" italics="true" pageId="138" pageNumber="361">nomen nudum</emphasis>
); 1913: 326 (
<emphasis id="B910D370938EB5AECE21FEA6FC20FE85" box="[820,954,357,381]" italics="true" pageId="138" pageNumber="361">nomen nudum</emphasis>
). — Glaessner 1929: 137; 1969: R492, fig. 304.8. — Müller 1978: 279, 287, pl. 13, fig. 4; 1984a: 66, pl. 34, figs 1-5. — Holthuis &amp; Manning 1990: 7.
</paragraph>
<paragraph id="8BDB0F62938EB5AECE26FE09FAE9FE1A" blockId="138.[819,1395,457,482]" box="[819,1395,457,482]" pageId="138" pageNumber="361">
<taxonomicName id="4C6474E1938EB5AECE26FE09FB91FE19" ID-CoL="f59e52e0-5e5a-461b-bea8-cb1214b8132e" authorityName="Lőrenthey" authorityYear="1929" box="[819,1035,457,481]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="margaretha">
<emphasis id="B910D370938EB5AECE26FE09FB91FE19" box="[819,1035,457,481]" italics="true" pageId="138" pageNumber="361">Medorippe Margaretha</emphasis>
</taxonomicName>
– Karasawa 2000: 811, 812, table 1.
</paragraph>
<paragraph id="8BDB0F62938EB5AECE38FDC3FABCFDE2" blockId="138.[813,1318,514,538]" box="[813,1318,514,538]" pageId="138" pageNumber="361">
<taxonomicName id="4C6474E1938EB5AECE38FDC3FBFEFDE2" ID-CoL="30d269b7-f4b8-46ba-93d3-b47a3099483f" authorityName="Lorenthey" authorityYear="1929" box="[813,1124,514,538]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="margaretha" subGenus="Medorippe">
<emphasis id="B910D370938EB5AECE38FDC3FCE3FDE2" box="[813,889,515,538]" italics="true" pageId="138" pageNumber="361">Dorippe</emphasis>
(
<emphasis id="B910D370938EB5AECE92FDC2FC71FDE2" box="[903,1003,514,538]" italics="true" pageId="138" pageNumber="361">Medorippe</emphasis>
)
<emphasis id="B910D370938EB5AECEECFDC2FBFEFDE2" box="[1017,1124,514,538]" italics="true" pageId="138" pageNumber="361">margaretha</emphasis>
</taxonomicName>
– Müller 2006: 42.
</paragraph>
<paragraph id="8BDB0F62938EB5AECE38FDFCFA2AFD95" blockId="138.[813,1456,571,621]" pageId="138" pageNumber="361">
<taxonomicName id="4C6474E1938EB5AECE38FDFCFC70FDAB" ID-CoL="30d269b7-f4b8-46ba-93d3-b47a3099483f" authorityName="Lorenthey" authorityYear="1929" box="[813,1002,571,595]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="margaretha">
<emphasis id="B910D370938EB5AECE38FDFCFC70FDAB" box="[813,1002,571,595]" italics="true" pageId="138" pageNumber="361">Dorippe margaretha</emphasis>
</taxonomicName>
– Schweitzer
<emphasis id="B910D370938EB5AEC966FDFCFB39FDAB" box="[1139,1187,571,595]" italics="true" pageId="138" pageNumber="361">et al.</emphasis>
2010: 79. — Hyžný 2016: table 1. — Sasaki 2019: 7774. — Hyžný &amp; Dulai 2021: fig. 62.1-5.
</paragraph>
<paragraph id="8BDB0F62938EB5AECE3EFD4EFB5BFD39" blockId="138.[811,1455,654,705]" pageId="138" pageNumber="361">
<taxonomicName id="4C6474E1938EB5AECE3EFD4EFC67FD5E" ID-CoL="f59e52e0-5e5a-461b-bea8-cb1214b8132e" authorityName="Lőrenthey" authorityYear="1929" box="[811,1021,654,678]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="margaretha">
<emphasis id="B910D370938EB5AECE3EFD4EFC67FD5E" box="[811,1021,654,678]" italics="true" pageId="138" pageNumber="361">Medorippe margaretha</emphasis>
</taxonomicName>
– Guinot
<emphasis id="B910D370938EB5AEC975FD4FFB17FD5E" box="[1120,1165,654,678]" italics="true" pageId="138" pageNumber="361">et al.</emphasis>
2013: 187. — Jagt
<emphasis id="B910D370938EB5AEC853FD4FFAE8FD5E" box="[1350,1394,654,678]" italics="true" pageId="138" pageNumber="361">et al.</emphasis>
2015: 882. — Van Bakel
<emphasis id="B910D370938EB5AECEF2FD69FB8CFD38" box="[999,1046,680,704]" italics="true" pageId="138" pageNumber="361">et al.</emphasis>
2020: fig. 10.34.
</paragraph>
<paragraph id="8BDB0F62938EB5AECE38FD22FA35FD01" blockId="138.[813,1455,737,761]" box="[813,1455,737,761]" pageId="138" pageNumber="361">
<taxonomicName id="4C6474E1938EB5AECE38FD22FBBDFD01" box="[813,1063,737,761]" class="Malacostraca" family="Dorippidae" genus="Dorippe" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="B910D370938EB5AECE38FD22FCE2FD01" box="[813,888,738,761]" italics="true" pageId="138" pageNumber="361">Dorippe</emphasis>
sp. aff.
<emphasis id="B910D370938EB5AECEABFD21FBBDFD01" box="[958,1063,737,761]" italics="true" pageId="138" pageNumber="361">margaretha</emphasis>
</taxonomicName>
– Polkowsky 2014: 123, pl. 125, figs 1-4.
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9938EB5AECE38FCF6FBB8F9B5" pageId="138" pageNumber="361" type="discussion">
<paragraph id="8BDB0F62938EB5AECE38FCF6FC0BFCB6" blockId="138.[811,1457,822,1613]" box="[813,913,822,848]" pageId="138" pageNumber="361">
<smallCapsWord id="8D3D99BE938EB5AECE38FCF6FC0BFCB6" baselines="841,841" box="[813,913,822,848]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="138" pageNumber="361">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BDB0F62938EB5AECE38FC96FBB8F9B5" blockId="138.[811,1457,822,1613]" pageId="138" pageNumber="361">
By its outline, its anterolateral spine in the figre of Lőrenthey &amp; Beurlen (1929: pl. 6, fig. 1a), its two closely spaced triangular frontal teeth, the carapace of this species, from the Miocene of
<collectingCountry id="F3734FF2938EB5AECE3EFC75FC16FC37" box="[811,908,949,975]" name="Hungary" pageId="138" pageNumber="361">Hungary</collectingCountry>
,
<collectingCountry id="F3734FF2938EB5AECE81FC75FC6BFC37" box="[916,1009,949,975]" name="Portugal" pageId="138" pageNumber="361">Portugal</collectingCountry>
,
<collectingCountry id="F3734FF2938EB5AECEECFC75FBC4FC37" box="[1017,1118,949,975]" name="Germany" pageId="138" pageNumber="361">Germany</collectingCountry>
(Upper Miocene) and
<collectingCountry id="F3734FF2938EB5AEC858FC75FA37FC37" box="[1357,1453,949,975]" name="Hungary" pageId="138" pageNumber="361">Hungary</collectingCountry>
(Middle Miocene), indeed resembles that of extant species
<taxonomicName id="4C6474E1938EB5AECE38FC34FC72FBF6" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[813,1000,1012,1038]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="lanata">
<emphasis id="B910D370938EB5AECE38FC34FC72FBF6" box="[813,1000,1012,1038]" italics="true" pageId="138" pageNumber="361">Medorippe lanata</emphasis>
</taxonomicName>
. In Polkowsky’s (2014: pl. 125, figs 1-4) four good figures of several Schleswig-Holstein specimens, the cardiac region does not show the characteristic Y-shaped ridge of the extant
<taxonomicName id="4C6474E1938EB5AECEEFFB94FB2FFB96" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1018,1205,1108,1134]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="lanata">
<emphasis id="B910D370938EB5AECEEFFB94FB2FFB96" box="[1018,1205,1108,1134]" italics="true" pageId="138" pageNumber="361">Medorippe lanata</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E7938EB5AEC9D1FB94FAD2FB96" box="[1220,1352,1108,1134]" captionStart="FIG" captionStartId="53.[133,144,1771,1789]" captionTargetBox="[132,1455,215,1727]" captionTargetId="figure-84@53.[698,1267,782,1351]" captionTargetPageId="53" captionText="FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon." figureDoi="http://doi.org/10.5281/zenodo.8071309" httpUri="https://zenodo.org/record/8071309/files/figure.png" pageId="138" pageNumber="361">Fig. 22A, B</figureCitation>
), and the strong tripatite branchial ridge is more reminiscent of
<taxonomicName id="4C6474E1938EB5AEC862FBB4FC47FB56" baseAuthorityName="Miers" baseAuthorityYear="1881" class="Malacostraca" family="Dorippidae" genus="Phyllodorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="B910D370938EB5AEC862FBB4FC47FB56" italics="true" pageId="138" pageNumber="361">Phyllodorippe armata</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E7938EB5AECEFCFB54FBFCFB56" box="[1001,1126,1172,1198]" captionStart="FIG" captionStartId="66.[133,144,1812,1830]" captionTargetBox="[137,1452,216,1769]" captionTargetId="figure-83@66.[128,1451,215,1820]" captionTargetPageId="66" captionText="FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8." figureDoi="http://doi.org/10.5281/zenodo.8071325" httpUri="https://zenodo.org/record/8071325/files/figure.png" pageId="138" pageNumber="361">Fig. 29A, B</figureCitation>
); but the shape of the carapace, the strong lobulation, the spaced frontal teeth, the absence of a lateral spine (possibly lost) are not typical of this genus. But most importantly, the valuable figures of the
<typeStatus id="54DFB1C0938EB5AEC85BFB33FA35FAF5" box="[1358,1455,1267,1293]" pageId="138" pageNumber="361" type="holotype">holotype</typeStatus>
and other specimens provided by Hyžný &amp; Dulai (2021: fig. 62.1-5) show the characteristic Y-shaped ridge on the cardiac region, a small spine on the carapace lateral margin, all of which are characters in favour of belonging to
<taxonomicName id="4C6474E1938EB5AEC823FAB3FA3CFA75" baseAuthorityName="Karaswa" baseAuthorityYear="2000" box="[1334,1446,1395,1421]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B910D370938EB5AEC823FAB3FA3CFA75" box="[1334,1446,1395,1421]" italics="true" pageId="138" pageNumber="361">Medorippe</emphasis>
</taxonomicName>
, if not, at least, to the subfamily
<taxonomicName id="4C6474E1938EB5AEC992FA53FAB9FA55" authority="It" authorityName="Guinot" authorityYear="2023" box="[1159,1315,1427,1453]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Medorippinae">Medorippinae</taxonomicName>
<taxonomicNameLabel id="A2236E0B938EB5AEC83CFA53FA02FA55" box="[1321,1432,1427,1453]" pageId="138" pageNumber="361" rank="subFamily">n.subfam.</taxonomicNameLabel>
It should be noted that a second species of
<taxonomicName id="4C6474E1938EB5AEC9FAFA73FAFAFA35" baseAuthorityName="Karaswa" baseAuthorityYear="2000" box="[1263,1376,1459,1485]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B910D370938EB5AEC9FAFA73FAFAFA35" box="[1263,1376,1459,1485]" italics="true" pageId="138" pageNumber="361">Medorippe</emphasis>
</taxonomicName>
, under the name of
<taxonomicName id="4C6474E1938EB5AECEA1FA14FBA8FA15" authorityName="Chen" authorityYear="1988" box="[948,1074,1492,1517]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="crosnieri">
<emphasis id="B910D370938EB5AECEA1FA14FBA8FA15" box="[948,1074,1492,1517]" italics="true" pageId="138" pageNumber="361">M. crosnieri</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E7938EB5AEC955FA13FB78FA15" box="[1088,1250,1491,1517]" captionStart="FIG" captionStartId="54.[132,143,965,983]" captionTargetBox="[132,1455,214,919]" captionTargetId="figure-553@54.[698,1267,211,745]" captionTargetPageId="54" captionText="FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne." figureDoi="http://doi.org/10.5281/zenodo.8071311" httpUri="https://zenodo.org/record/8071311/files/figure.png" pageId="138" pageNumber="361">Fig. 23A, B, D</figureCitation>
), with a similar facies, including the cardiac region, probably exists, but with a still questionable status (see
<emphasis id="B910D370938EB5AEC993F9D2FAB2F9D4" box="[1158,1320,1554,1580]" italics="true" pageId="138" pageNumber="361">Remarks about</emphasis>
<taxonomicName id="4C6474E1938EB5AEC824F9D2FB88F9B5" authority="Chen, 1988" authorityName="Chen" authorityYear="1988" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="138" pageNumber="361" phylum="Arthropoda" rank="species" species="crosnieri">
Medorippe crosnieri
<emphasis id="B910D370938EB5AECE87F9F2FB88F9B5" box="[914,1042,1586,1613]" italics="true" pageId="138" pageNumber="361">Chen, 1988</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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