treatments-xml/data/03/CD/BE/03CDBE749322B50FCEB4FF17FDCAF813.xml

<document id="AB987F16B415E8C056EA3D43BBFDB4BA" ID-DOI="10.5252/zoosystema2023v45a9" ID-ISSN="1638-9387" ID-Zenodo-Dep="8071253" ID-ZooBank="urn:lsid:zoobank.org:pub:69C34731-8C25-4A1E-B336-B222CD3CBAC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686747199180" checkinUser="felipe" docAuthor="Guinot, Danièle" docDate="2023" docId="03CDBE749322B50FCEB4FF17FDCAF813" docLanguage="en" docName="Zoosystema.45.9.225-372.pdf" docOrigin="Zoosystema 45 (9)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.5:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="5" docTitle="Dorippoidinae Guinot 2023, n. subfam." docType="treatment" docVersion="4" lastPageNumber="266" masterDocId="FFF4C60C9304B524CD15FFC0FF9AFFF8" masterDocTitle="A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system" masterLastPageNumber="372" masterPageNumber="225" pageNumber="261" updateTime="1687453794845" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CD095E4C1CDA1AA1DC6EE38AAF4D1FEE">A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system</mods:title>
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<mods:namePart id="D69A64309B323C89FC70AB8E5BD7750F">Guinot, Danièle</mods:namePart>
<mods:affiliation id="48EBA81E74524BA5DAB68FD3958F209C">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) daniele. guinot @ mnhn. fr Dedicated to the memory of my colleague and dearest friend Ngan Kee NG (1966 - 2022)</mods:affiliation>
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<mods:date id="BB1E19A2202F502A7C5650BC72EAC3EB">2023</mods:date>
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<subSubSection id="C37E5CE99322B502CEB4FF17FAA1FF09" box="[929,1339,215,242]" pageId="38" pageNumber="261" type="nomenclature">
<paragraph id="8BDB0F629322B502CEB4FF17FAA1FF09" blockId="38.[929,1339,215,242]" box="[929,1339,215,242]" pageId="38" pageNumber="261">
<heading id="D093B80E9322B502CEB4FF17FAA1FF09" box="[929,1339,215,242]" centered="true" fontSize="36" level="2" pageId="38" pageNumber="261" reason="6">
Subfamily 
<taxonomicName id="4C6474E19322B502C903FF18FB5AFF08" authority="Guinot, 2023" authorityName="Guinot" authorityYear="2023" box="[1046,1216,216,242]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="subFamily" status="n. subfam." subFamily="Dorippoidinae">
<smallCapsWord id="8D3D99BE9322B502C903FF18FB5AFF08" baselines="235,235" box="[1046,1216,216,242]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dorippoidinae" pageId="38" pageNumber="261">DORIPPOIDINAE</smallCapsWord>
</taxonomicName>
<taxonomicNameLabel id="A2236E0B9322B502C9DDFF18FAA1FF09" box="[1224,1339,215,242]" pageId="38" pageNumber="261" rank="subFamily">n. subfam.</taxonomicNameLabel>
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<typeStatus id="54DFB1C09322B502CE38FED1FCC7FEDE" box="[813,861,273,296]" pageId="38" pageNumber="261">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D3D99BE9322B502CE71FED6FC3EFEDF" baselines="291" box="[868,932,278,295]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="genus" pageId="38" pageNumber="261">GENUS</smallCapsWord>
. — 
<taxonomicName id="4C6474E19322B502CEDAFED0FAE2FED0" authority="Serene &amp; Romimohtarto, 1969" authorityName="Serene &amp; Romimohtarto" authorityYear="1969" box="[975,1400,272,296]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709322B502CEDAFED0FBA5FED0" box="[975,1087,272,296]" italics="true" pageId="38" pageNumber="261">Dorippoides</emphasis>
<bibRefCitation id="EFF572939322B502C950FED0FAE2FED0" author="Serene &amp; Romimohtarto" box="[1093,1400,272,296]" firstAuthor="Serene" pageId="38" pageNumber="261" pagination="1 - 35" refId="ref141809" refString="SERENE R. &amp; ROMIMOHTARTO K. 1969. - ̿ bservations on the species of Dorippe from the Indo-Malayan Region. Marine Research in Indonesia (Penelitian Laut di Indonesia) 9: 1 - 35." type="journal article" year="1969">Serène &amp; Romimohtarto, 1969</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54DFB1C09322B502C891FED0FA35FED0" box="[1412,1455,272,296]" pageId="38" pageNumber="261">type</typeStatus>
species by original designation and monotypy: 
<taxonomicName id="4C6474E19322B502C807FEEBFC2FFEA5" authority="Herbst, 1785" authorityName="Herbst" authorityYear="1785" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709322B502C807FEEBFA35FEBB" box="[1298,1455,299,323]" italics="true" pageId="38" pageNumber="261">Cancer facchino</emphasis>
Herbst, 1785
</taxonomicName>
). Other included species: 
<taxonomicName id="4C6474E19322B502C9A0FE85FB8EFE8F" authority="Manning &amp; Holthuis, 1986" authorityName=": Manning &amp; Holthuis" authorityYear="1986" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709322B502C9A0FE85FAE9FEA5" box="[1205,1395,325,349]" italics="true" pageId="38" pageNumber="261">Dorippoides nudipes</emphasis>
<bibRefCitation id="EFF572939322B502C86FFE86FB8EFE8F" author="Manning &amp; Holthuis" firstAuthor="Manning" pageId="38" pageNumber="261" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis, 1986</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE99322B50CCE38FE57FD5FF9B5" lastPageId="40" lastPageNumber="263" pageId="38" pageNumber="261" type="description">
<paragraph id="8BDB0F629322B502CE38FE57FC27FE57" blockId="38.[811,1457,407,1677]" box="[813,957,407,433]" pageId="38" pageNumber="261">
<smallCapsWord id="8D3D99BE9322B502CE38FE57FC27FE57" baselines="426,426" box="[813,957,407,433]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="38" pageNumber="261">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BDB0F629322B502CE38FE77FC6AFE29" blockId="38.[811,1457,407,1677]" box="[813,1008,438,465]" pageId="38" pageNumber="261">
<emphasis id="B910D3709322B502CE38FE77FC6AFE29" box="[813,1008,438,465]" italics="true" pageId="38" pageNumber="261">
Carapace (
<figureCitation id="135F13E79322B502CE8EFE77FC7DFE28" box="[923,999,438,465]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="38" pageNumber="261">Fig. 16</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629322B502CE38FE16FA36FAF6" blockId="38.[811,1457,407,1677]" pageId="38" pageNumber="261">
Carapace wider than long, convex posteriorly, flattened anteriorly, appearing flatter in large individuals. Dorsal surface weakly sculptured, rather smooth, covered with pubescence; ‘human face’ distinctly delineated. Only a few grooves: precervical groove continuous, distinct in small specimens, with extreme lateral part behind orbit vaguely distinct, or not indicated at all medially in adult males; indistinct, medially interrupted in females; cervical and branchiocardiac grooves deep. Median area (urogastric region) circular or oval, flanked by large, rounded, convex branchial lobes. Meso-metagastric region with two small, oblique submedian gastric pits at the base. Front consisting of two distinct triangular teeth directed forward, or of blunter teeth directed outward, separated by rather deep emargination. Inner orbital angle triangular or blunt. Outer orbital tooth sharp, pointed, reaching as far forwards as or slightly beyond frontal teeth. Antero- and posterolateral margins devoid of spine or tooth, only demarcated by very faint angle. Exposed pleurites 5-7 as granulous sclerites; exposed pleurite 6 rather narrow, granular, separated from P3 coxa by thick, whitish membrane. Carapace posterior rim not extending laterally at all along inflated posterolateral margin and lined posteriorly by nearly rectangular strip, thinner medially in male 
<taxonomicName id="4C6474E19322B502C944FB54FAA9FB56" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[1105,1331,1172,1198]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709322B502C944FB54FAA9FB56" box="[1105,1331,1172,1198]" italics="true" pageId="38" pageNumber="261">Dorippoides facchino</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502C857FB54FA32FB56" box="[1346,1448,1172,1198]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="38" pageNumber="261">Fig. 16A</figureCitation>
) and 
<taxonomicName id="4C6474E19322B502CE48FB75FC55FB36" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[861,975,1204,1230]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709322B502CE48FB75FC55FB36" box="[861,975,1204,1230]" italics="true" pageId="38" pageNumber="261">D. nudipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CECAFB74FBD8FB36" box="[991,1090,1204,1230]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="38" pageNumber="261">Fig. 16C</figureCitation>
); strip with two developed lateral extensions and thus appearing much more hollowed medially in female 
<taxonomicName id="4C6474E19322B502CE80FB34FB97FAF5" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[917,1037,1267,1294]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709322B502CE80FB34FC30FAF5" box="[917,938,1268,1293]" italics="true" pageId="38" pageNumber="261">D</emphasis>
. 
<emphasis id="B910D3709322B502CEADFB33FB97FAF5" box="[952,1037,1267,1293]" italics="true" pageId="38" pageNumber="261">facchino</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502C909FB33FBE1FAF6" box="[1052,1147,1267,1294]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="38" pageNumber="261">Fig. 16B</figureCitation>
) and 
<taxonomicName id="4C6474E19322B502C9ACFB34FAB1FAF5" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[1209,1323,1267,1293]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709322B502C9ACFB34FAB1FAF5" box="[1209,1323,1267,1293]" italics="true" pageId="38" pageNumber="261">D. nudipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502C82CFB33FA04FAF6" box="[1337,1438,1267,1294]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="38" pageNumber="261">Fig. 16D</figureCitation>
).
</paragraph>
<paragraph id="8BDB0F629322B502CE56FAD3FC45F974" blockId="38.[811,1457,407,1677]" pageId="38" pageNumber="261">
Illustrations: 
<taxonomicName id="4C6474E19322B502CECDFAD3FACFFAD6" authority=": Herbst 1785" authorityName="Herbst" authorityYear="1785" box="[984,1365,1299,1326]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709322B502CECDFAD3FB2CFAD5" box="[984,1206,1299,1325]" italics="true" pageId="38" pageNumber="261">Dorippoides facchino</emphasis>
: Herbst 1785
</taxonomicName>
: pl. 11, fig. 68, as 
<taxonomicName id="4C6474E19322B502CE8FFAF4FBDAFAB5" authorityName="Herbst" authorityYear="1785" box="[922,1088,1331,1358]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709322B502CE8FFAF4FBDAFAB5" box="[922,1088,1331,1358]" italics="true" pageId="38" pageNumber="261">Cancer facchino</emphasis>
</taxonomicName>
(reproduced by Holthuis &amp; Manning 1990: fig. 23); H. Milne Edwards 1837: pl. 20, fig. 11, as 
<taxonomicName id="4C6474E19322B502CE52FAB4FC4BFA75" authorityName="H. Milne Edwards" authorityYear="1837" box="[839,977,1396,1421]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="sima">
<emphasis id="B910D3709322B502CE52FAB4FC4BFA75" box="[839,977,1396,1421]" italics="true" pageId="38" pageNumber="261">Dorippe sima</emphasis>
</taxonomicName>
(reproduced by Holthuis &amp; Manning 1990: fig. 24); 
<bibRefCitation id="EFF572939322B502CE90FA53FB95FA55" author="Verrill" box="[901,1039,1427,1453]" firstAuthor="Verrill" pageId="38" pageNumber="261" pagination="51 - 104" refId="ref147502" refString="VERRILL A. E. 1869 a. - Actiniaria, with Supplement and Geographical Lists. Synopses of the Polyps and Corals of the North Pacific ‰ xploring ‰ xpedition, under Commodore C. Ringgold and Captain John Rodgers, U. S. N., from 1853 to 1856, Collected by Dr. William Stimpson, Naturalist to the ‰ xpedition, Part IV. Proceedings of the Essex Institute 6: 51 - 104, pls 1 - 2." type="journal article" year="1869" yearSuffix="a">Verrill 1869a</bibRefCitation>
: pl. 2, fig. 1, as 
<taxonomicName id="4C6474E19322B502C9A4FA54FAC5FA55" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[1201,1375,1427,1453]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709322B502C9A4FA54FAC5FA55" box="[1201,1375,1427,1453]" italics="true" pageId="38" pageNumber="261">Dorippe facchino</emphasis>
</taxonomicName>
(reproduced by Holthuis &amp; Manning 1990: fig. 25a); Holthuis &amp; Manning 1990: figs 19a-d, 22 (male 
<typeStatus id="54DFB1C09322B502C9CDFA13FAB4FA15" box="[1240,1326,1491,1517]" pageId="38" pageNumber="261">syntype</typeStatus>
of 
<taxonomicName id="4C6474E19322B502C842FA14FB87F9F5" authority="Fabricius, 1785" authorityName="Fabricius" authorityYear="1785" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="astuta">
<emphasis id="B910D3709322B502C842FA14FCF6F9F4" italics="true" pageId="38" pageNumber="261">Dorippe astuta</emphasis>
Fabricius, 1785
</taxonomicName>
), 19b (reproduced by 
<bibRefCitation id="EFF572939322B502C819FA33FA32F9F5" author="Sin" box="[1292,1448,1523,1549]" etAl="et al." firstAuthor="Sin" pageId="38" pageNumber="261" pagination="223 - 230" refId="ref142785" refString="SIN Y. W., LAI J. C. Y., NG P. K. L. &amp; CHU K. H. 2009. - Phylogeny of Dorippoidea (Crustacea: Decapoda: Brachyura) inferred from three mitochondrial genes. Invertebrate Systematics 23 (3): 223 - 230. https: // doi. org / 10.1071 / IS 09020" type="journal article" year="2009">
Sin 
<emphasis id="B910D3709322B502C823FA33FAF0F9F5" box="[1334,1386,1523,1549]" italics="true" pageId="38" pageNumber="261">et al.</emphasis>
2009
</bibRefCitation>
: fig. 3B), 25b; 
<bibRefCitation id="EFF572939322B502CEDBF9D2FB38F9D5" author="Chen &amp; Sun" box="[974,1186,1554,1581]" firstAuthor="Chen" pageId="38" pageNumber="261" refId="ref112964" refString="CHEN H. &amp; SUN H. 2002. - Brachyura. Marine primitive crabs, in Fauna Sinica, Invertebrata, vol. 30. Arthropoda, Crustacea. Academia Sinica, Science Press, Beijing, China, xiii + 597 p. (In Chinese with ‰ nglish abstract). https: // www. biodiversitylibrary. org / part / 119053" type="book" year="2002">Chen &amp; Sun 2002</bibRefCitation>
: fig. 92.1; 
<bibRefCitation id="EFF572939322B502C837F9D3FCFFF9B5" author="Naruse" etAl="et al." firstAuthor="Naruse" pageId="38" pageNumber="261" pagination="24 - 36" refId="ref134433" refString="NARUSE T., YEO D. C. J. &amp; OSAWA M. 2014. - Notes on a collection of stomatopod and decapod crustaceans from Cambodia. Cambodian Journal of Natural History 1: 24 - 36." type="journal article" year="2014">
Naruse 
<emphasis id="B910D3709322B502C86CF9D3FA2AF9D4" box="[1401,1456,1554,1580]" italics="true" pageId="38" pageNumber="261">et al.</emphasis>
2014
</bibRefCitation>
: fig. 2d; 
<bibRefCitation id="EFF572939322B502CED7F9F3FBE2F9B5" author="Wong" box="[962,1144,1586,1613]" etAl="et al." firstAuthor="Wong" pageId="38" pageNumber="261" pagination="102013" refId="ref148990" refString="WONG K. J. H., TAO L. S. R. &amp; LEUNG K. M. Y. 2021. - Subtidal crabs of Hong Kong: Brachyura (Crustacea: Decapoda) from benthic trawl surveys conducted by the University of Hong Kong, 2012 to 2018. Regional Studies in Marine Science 48: 102013. https: // doi. org / https: // doi. org / 10.1016 / j. rsma. 2021.102013" type="journal article" year="2021">
Wong 
<emphasis id="B910D3709322B502C91CF9F3FBA1F9B4" box="[1033,1083,1586,1612]" italics="true" pageId="38" pageNumber="261">et al.</emphasis>
2021
</bibRefCitation>
: fig. 10a, pl. 2D. 
<taxonomicName id="4C6474E19322B502C827F9F2FB9BF994" authority=": Chen 1988" authorityName=": Chen" authorityYear="1988" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709322B502C827F9F2FCE0F994" italics="true" pageId="38" pageNumber="261">Dorippoides nudipes</emphasis>
: 
<bibRefCitation id="EFF572939322B502CE9DF992FB9BF994" author="Chen" box="[904,1025,1618,1644]" firstAuthor="Chen" pageId="38" pageNumber="261" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen 1988</bibRefCitation>
</taxonomicName>
: fig. 1a, pl. 1B, C; Holthuis &amp; Manning 1990: fig. 26a-c.
</paragraph>
<paragraph id="8BDB0F629322B502CE38F972FBF9F934" blockId="38.[813,1455,1714,1899]" box="[813,1123,1714,1740]" pageId="38" pageNumber="261">
<emphasis id="B910D3709322B502CE38F972FBF9F934" box="[813,1123,1714,1740]" italics="true" pageId="38" pageNumber="261">
Cephalic structures (
<figureCitation id="135F13E79322B502CEE8F972FBC1F934" box="[1021,1115,1714,1740]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="38" pageNumber="261">Fig. 17A</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629322B502CE38F912FA39F893" blockId="38.[813,1455,1714,1899]" pageId="38" pageNumber="261">Eyestalk inclined, well protected in rather deep orbital hollow and along outer orbital tooth. Antennule almost entirely folded into fossa. Antenna: basal article exposed; articles 2+3 very developed, moveable, with salient setose external part; other articles directed forward; flagellum curved outwards.</paragraph>
<paragraph id="8BDB0F629322B502CE38F852FBAAF853" blockId="38.[813,1455,1937,2028]" box="[813,1072,1937,1964]" pageId="38" pageNumber="261">
<emphasis id="B910D3709322B502CE38F852FBAAF853" box="[813,1072,1937,1964]" italics="true" pageId="38" pageNumber="261">Oxystomatous disposition</emphasis>
</paragraph>
<paragraph id="8BDB0F629322B503CE38F871FE53FF0A" blockId="38.[813,1455,1937,2028]" lastBlockId="39.[130,777,215,975]" lastPageId="39" lastPageNumber="262" pageId="38" pageNumber="261">
Opening of exhalant channels perceptible in dorsal view between rostral teeth, hardly visible in 
<taxonomicName id="4C6474E19322B502C9C5F811FA3BF813" authorityName=": Chen" authorityYear="1988" box="[1232,1441,2001,2027]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709322B502C9C5F811FA3BF813" box="[1232,1441,2001,2027]" italics="true" pageId="38" pageNumber="261">Dorippoides nudipes</emphasis>
</taxonomicName>
. 
<emphasis id="B910D3709323B503CD91FF17FE53FF0A" box="[132,457,215,242]" italics="true" pageId="39" pageNumber="262">
Pereiopods (
<figureCitation id="135F13E79323B503CDE8FF18FED0FF09" box="[253,330,215,242]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="39" pageNumber="262">Figs 16</figureCitation>
; 
<figureCitation id="135F13E79323B503CC4DFF17FE1FFF0A" box="[344,389,215,242]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="39" pageNumber="262">17A</figureCitation>
; 
<figureCitation id="135F13E79323B503CC86FF17FE58FF0A" box="[403,450,215,242]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">18C</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CD91FF37FC9DFE89" blockId="39.[130,777,215,975]" pageId="39" pageNumber="262">Chelipeds of females and most males of same size and shape; heterochely only in large adult males: major chela swollen, short, with short fingers; minor chela with slender and downwardly curved fingers, forming slight angle with axis of palm.</paragraph>
<paragraph id="8BDB0F629323B503CD8CFEB7FEBDFE08" blockId="39.[130,777,215,975]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CC38FEB7FF24FE49" authority=": Holthuis &amp; Manning 1990" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CC38FEB7FD92FE69" box="[301,520,375,401]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: Holthuis &amp; Manning 1990
</taxonomicName>
: fig. 19e; 
<bibRefCitation id="EFF572939323B503CC39FE56FE62FE49" author="Chen &amp; Sun" box="[300,504,406,433]" firstAuthor="Chen" pageId="39" pageNumber="262" refId="ref112964" refString="CHEN H. &amp; SUN H. 2002. - Brachyura. Marine primitive crabs, in Fauna Sinica, Invertebrata, vol. 30. Arthropoda, Crustacea. Academia Sinica, Science Press, Beijing, China, xiii + 597 p. (In Chinese with ‰ nglish abstract). https: // www. biodiversitylibrary. org / part / 119053" type="book" year="2002">Chen &amp; Sun 2002</bibRefCitation>
: fig. 92.2, 3; 
<bibRefCitation id="EFF572939323B503CF9FFE57FF26FE29" author="Wong" etAl="et al." firstAuthor="Wong" pageId="39" pageNumber="262" pagination="102013" refId="ref148990" refString="WONG K. J. H., TAO L. S. R. &amp; LEUNG K. M. Y. 2021. - Subtidal crabs of Hong Kong: Brachyura (Crustacea: Decapoda) from benthic trawl surveys conducted by the University of Hong Kong, 2012 to 2018. Regional Studies in Marine Science 48: 102013. https: // doi. org / https: // doi. org / 10.1016 / j. rsma. 2021.102013" type="journal article" year="2021">
Wong 
<emphasis id="B910D3709323B503CFC6FE57FC9DFE48" box="[723,775,406,432]" italics="true" pageId="39" pageNumber="262">et al.</emphasis>
2021
</bibRefCitation>
: fig. 10b. 
<taxonomicName id="4C6474E19323B503CC27FE76FF24FE08" authority=": Holthuis &amp; Manning 1990" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CC27FE76FD9DFE28" box="[306,519,438,464]" italics="true" pageId="39" pageNumber="262">Dorippoides nudipes</emphasis>
: Holthuis &amp; Manning 1990
</taxonomicName>
: fig. 26d.
</paragraph>
<paragraph id="8BDB0F629323B503CD8EFE36FC92FCD7" blockId="39.[130,777,215,975]" pageId="39" pageNumber="262">
P2 and P3 not very long, P3 longest, all articles rather short and wide, flattened, unarmed. Meri without spines on dorsal margins; sexual dimorphism of setation in 
<taxonomicName id="4C6474E19323B503CF85FDF7FC9DFDA8" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[656,775,566,593]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CF85FDF7FD3FFDA8" box="[656,677,567,592]" italics="true" pageId="39" pageNumber="262">D</emphasis>
. 
<emphasis id="B910D3709323B503CFA7FDF6FC9DFDA8" box="[690,775,566,592]" italics="true" pageId="39" pageNumber="262">facchino</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79323B503CD99FD96FE91FD88" box="[140,267,598,624]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="39" pageNumber="262">Fig. 16A, B</figureCitation>
): males with dense setae only on posterior margins and females with entirely naked margins; dimorphism not marked in 
<taxonomicName id="4C6474E19323B503CC31FD56FE0FFD48" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[292,405,662,688]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CC31FD56FE0FFD48" box="[292,405,662,688]" italics="true" pageId="39" pageNumber="262">D. nudipes</emphasis>
</taxonomicName>
, with naked margins in both sexes (
<figureCitation id="135F13E79323B503CD99FD75FE89FD28" box="[140,275,693,720]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="39" pageNumber="262">Fig. 16C, D</figureCitation>
). Dactyli flattened, twisted, without fringes of hair. P3 ischium of females with spur-like process on anterior margin; process may be present on ischium of P2 but smaller (
<figureCitation id="135F13E79323B503CD9FFCD5FF47FCD7" box="[138,221,789,815]" captionStart="FIG" captionStartId="26.[132,143,1214,1232]" captionTargetBox="[140,1456,215,1172]" captionTargetId="figure-502@26.[140,800,691,1172]" captionTargetPageId="26" captionText="FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area." figureDoi="http://doi.org/10.5281/zenodo.8071277" httpUri="https://zenodo.org/record/8071277/files/figure.png" pageId="39" pageNumber="262">Figs 9A</figureCitation>
; 
<figureCitation id="135F13E79323B503CDFDFCD5FEAEFCD7" box="[232,308,789,815]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">18A, B</figureCitation>
); these processes completely absent in males.
</paragraph>
<paragraph id="8BDB0F629323B503CD8CFCF5FEC6FC37" blockId="39.[130,777,215,975]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CC38FCF5FF25FC97" authority=": Holthuis &amp; Manning 1990" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CC38FCF5FD92FCB7" box="[301,520,821,847]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: Holthuis &amp; Manning 1990
</taxonomicName>
: fig. 20. 
<taxonomicName id="4C6474E19323B503CC3FFC95FF24FC77" authority=": Manning &amp; Holthuis 1986" authorityName=": Manning &amp; Holthuis" authorityYear="1986" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CC3FFC95FD9EFC97" box="[298,516,853,879]" italics="true" pageId="39" pageNumber="262">Dorippoides nudipes</emphasis>
: 
<bibRefCitation id="EFF572939323B503CF06FC95FF24FC77" author="Manning &amp; Holthuis" firstAuthor="Manning" pageId="39" pageNumber="262" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis 1986</bibRefCitation>
</taxonomicName>
: fig. 1c; 
<bibRefCitation id="EFF572939323B503CC34FCB5FE07FC77" author="Chen" box="[289,413,885,911]" firstAuthor="Chen" pageId="39" pageNumber="262" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen 1988</bibRefCitation>
: fig. 1d, e; Holthuis &amp; Manning 1990: figs 26e, f, 27f; 
<bibRefCitation id="EFF572939323B503CC7AFC55FDACFC57" author="Chen &amp; Sun" box="[367,566,917,943]" firstAuthor="Chen" pageId="39" pageNumber="262" refId="ref112964" refString="CHEN H. &amp; SUN H. 2002. - Brachyura. Marine primitive crabs, in Fauna Sinica, Invertebrata, vol. 30. Arthropoda, Crustacea. Academia Sinica, Science Press, Beijing, China, xiii + 597 p. (In Chinese with ‰ nglish abstract). https: // www. biodiversitylibrary. org / part / 119053" type="book" year="2002">Chen &amp; Sun 2002</bibRefCitation>
: fig. 92.2, 3; 
<bibRefCitation id="EFF572939323B503CFD0FC55FF6CFC37" author="Wong" etAl="et al." firstAuthor="Wong" pageId="39" pageNumber="262" pagination="102013" refId="ref148990" refString="WONG K. J. H., TAO L. S. R. &amp; LEUNG K. M. Y. 2021. - Subtidal crabs of Hong Kong: Brachyura (Crustacea: Decapoda) from benthic trawl surveys conducted by the University of Hong Kong, 2012 to 2018. Regional Studies in Marine Science 48: 102013. https: // doi. org / https: // doi. org / 10.1016 / j. rsma. 2021.102013" type="journal article" year="2021">
Wong 
<emphasis id="B910D3709323B503CD91FC75FF22FC37" box="[132,184,949,975]" italics="true" pageId="39" pageNumber="262">et al.</emphasis>
2021
</bibRefCitation>
: fig. 10c.
</paragraph>
<paragraph id="8BDB0F629323B503CD91FC34FE5EFBF7" blockId="39.[130,776,1012,1421]" box="[132,452,1012,1039]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CD91FC34FE5EFBF7" box="[132,452,1012,1039]" italics="true" pageId="39" pageNumber="262">
Thoracic sternum (
<figureCitation id="135F13E79323B503CC51FC35FE0BFBF7" box="[324,401,1013,1039]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="39" pageNumber="262">Figs 17</figureCitation>
; 
<figureCitation id="135F13E79323B503CC8AFC35FE21FBF7" box="[415,443,1013,1039]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">18</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CD91FBD4FEAEFA75" blockId="39.[130,776,1012,1421]" pageId="39" pageNumber="262">Thoracic sternum almost smooth in males, minutely granular and eroded in females.Sternite 1 pointed; sternite 2 pentagonal, high, separated from sternite 3 by broad depression; sternite 3 large, extended laterally; sternite 4 with two raised submedian prominences protruding from pubescence; sternites 4 and 5 lacking sharp ridges, only with blunt, naked transverse ridge. Suture 3/4 rather long, ending in deep depression in males. In males, interruption points of sutures very close to each other at bottom of deep sterno-pleonal cavity, which may give the false impression of a median line. Female thoracic sternum extremely tilted backwards at level of ridge running through entire sternite 6.</paragraph>
<paragraph id="8BDB0F629323B503CD91FA73FE7AFA35" blockId="39.[130,777,1459,2027]" box="[132,480,1459,1485]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CD91FA73FE7AFA35" box="[132,480,1459,1485]" italics="true" pageId="39" pageNumber="262">
Pleon and telson (
<figureCitation id="135F13E79323B503CC2EFA73FE03FA35" box="[315,409,1459,1485]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="39" pageNumber="262">Figs 17B</figureCitation>
; 
<figureCitation id="135F13E79323B503CCB3FA73FE4DFA35" box="[422,471,1459,1485]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">18D</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CD91FA13FD7BF8D4" blockId="39.[130,777,1459,2027]" pageId="39" pageNumber="262">
Sterno-pleonal cavity very narrow and deep. On pleon no tubercles or spines, only low, blunt transverse elevations or indistinct grooves. 
<collectingRegion id="49A0C1809323B503CC49F9D2FE0EF9D4" box="[348,404,1554,1580]" country="Maldives" name="Maale" pageId="39" pageNumber="262">Male</collectingRegion>
pleon with articular membranes situated between all somites and across entire breadth of each somite. In males, somite 1 long, trapezoidal, widening posteriorly, with posterior margin deeply concave in middle; somite 2 long, widening posteriorly, lacking erect tubercles or spines; somite 4 short, wide; somite 5 distinctly narrowing; somite 6 elongate, very narrow, with concave lateral margins and produced posterolateral angles; telson triangular, with constricted base, its tip far exceeding level of suture 5/6.
</paragraph>
<paragraph id="8BDB0F629323B503CD8EF8F1FC9DF894" blockId="39.[130,777,1459,2027]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CC38F8F1FF24F894" authority=": Holthuis &amp; Manning 1990" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CC38F8F1FD92F8B3" box="[301,520,1841,1867]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: Holthuis &amp; Manning 1990
</taxonomicName>
: fig. 
<quantity id="4C9CA2879323B503CDE1F892FE85F894" box="[244,287,1874,1900]" metricMagnitude="-2" metricUnit="kg" metricValue="2.1" pageId="39" pageNumber="262" unit="g" value="21.0">21g</quantity>
, f. 
<taxonomicName id="4C6474E19323B503CC2AF891FD0CF894" authority=": Chen 1988" authorityName=": Chen" authorityYear="1988" box="[319,662,1873,1900]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CC2AF891FD8AF893" box="[319,528,1873,1899]" italics="true" pageId="39" pageNumber="262">Dorippoides nudipes</emphasis>
: 
<bibRefCitation id="EFF572939323B503CF0EF891FD0CF894" author="Chen" box="[539,662,1873,1900]" firstAuthor="Chen" pageId="39" pageNumber="262" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen 1988</bibRefCitation>
</taxonomicName>
: fig. 1b, c.
</paragraph>
<paragraph id="8BDB0F629323B503CD8EF8B1FDDDF813" blockId="39.[130,777,1459,2027]" pageId="39" pageNumber="262">
Female pleon (
<figureCitation id="135F13E79323B503CC55F8B1FE5EF873" box="[320,452,1905,1932]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">Fig. 18A, E</figureCitation>
) wide and rounded, without tubercles or teeth; somites 2-5 with blunt but distinct transverse carina; somite 5 widest; somites 5 and 6 longest; telson small, with semicircular posterior margin.
</paragraph>
<paragraph id="8BDB0F629323B503CE38FF17FC3CFEE9" blockId="39.[813,1425,215,306]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CE38FF17FC3CFEE9" italics="true" pageId="39" pageNumber="262">
Pleonal-locking mechanism by press-button (
<figureCitation id="135F13E79323B503C9E6FF18FAEFFF09" box="[1267,1397,215,242]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="39" pageNumber="262">Figs 17C, D</figureCitation>
; 
<figureCitation id="135F13E79323B503CE38FF37FC00FEE9" box="[813,922,247,273]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">18B, C, F</figureCitation>
),
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CE38FED7FA0BFECA" blockId="39.[813,1425,215,306]" box="[813,1425,279,306]" pageId="39" pageNumber="262">Press-button in deep curve of thoracic sternal suture 5/6.</paragraph>
<paragraph id="8BDB0F629323B503CE38FE97FA8FFE89" blockId="39.[811,1462,343,593]" box="[813,1301,343,369]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CE38FE97FA8FFE89" box="[813,1301,343,369]" italics="true" pageId="39" pageNumber="262">Additional female pleonal-retention mechanism</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CE38FEB7FA2AFDA8" blockId="39.[811,1462,343,593]" pageId="39" pageNumber="262">
In females of 
<taxonomicName id="4C6474E19323B503CEA8FEB7FBA8FE69" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[957,1074,375,401]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CEA8FEB7FBA8FE69" box="[957,1074,375,401]" italics="true" pageId="39" pageNumber="262">D. facchino</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79323B503C954FEB7FB3AFE69" box="[1089,1184,375,401]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="39" pageNumber="262">Fig. 16B</figureCitation>
), dorsally exposed part of sternite 8 with very small prominence that does not overhang pleonal somite 2 and seems not functional; in females of 
<taxonomicName id="4C6474E19323B503CE38FE17FC0CFE08" authorityName=": Chen" authorityYear="1988" box="[813,918,470,496]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CE38FE17FC0CFE08" box="[813,918,470,496]" italics="true" pageId="39" pageNumber="262">D. nudipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79323B503CEB7FE16FC66FE08" box="[930,1020,470,496]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="39" pageNumber="262">Fig. 16C</figureCitation>
), the process of sternite 8 is quite developed but seems too distant to be able to overhang pleonal somite 2, thus also nonfunctional. Small telson engaged between raised slopes of sterno-pleonal cavity at level of sternite 5 (
<figureCitation id="135F13E79323B503C83EFDF6FA39FDA8" box="[1323,1443,566,592]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">Fig. 18A, E</figureCitation>
).
</paragraph>
<paragraph id="8BDB0F629323B503CE3EFDB6FBB0FD68" blockId="39.[811,1457,630,848]" box="[811,1066,630,656]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CE3EFDB6FBB0FD68" box="[811,1066,630,656]" italics="true" pageId="39" pageNumber="262">
<collectingRegion id="49A0C1809323B503CE3EFDB6FCFAFD68" box="[811,864,630,656]" country="Maldives" name="Maale" pageId="39" pageNumber="262">Male</collectingRegion>
gonopore and penis
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CE38FD56FAADFCF7" blockId="39.[811,1457,630,848]" pageId="39" pageNumber="262">
Gonopore coxal; coxo-sternal condition with penial tube consisting of inclined portion, then vertical portion, without visible membrane between the two (sternites 7 and 
<quantity id="4C9CA2879323B503C821FD16FAC6FD17" box="[1332,1372,725,752]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="39" pageNumber="262" unit="in" value="8.0">8 in</quantity>
contact over very short distance); bulb more or less long.
</paragraph>
<paragraph id="8BDB0F629323B503CE56FCD5FA2AFCA8" blockId="39.[811,1457,630,848]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CEC9FCD5FA32FCD7" authority=": Guinot et al. 2013" authorityName="Guinot" authorityYear="2013" box="[988,1448,789,816]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CEC9FCD5FB5AFCD7" box="[988,1216,789,815]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: 
<bibRefCitation id="EFF572939323B503C9C4FCD5FA32FCD7" author="Guinot" box="[1233,1448,789,815]" etAl="et al." firstAuthor="Guinot" pageId="39" pageNumber="262" pagination="1 - 414" refId="ref122703" refString="GUINOT D., TAVARES M. &amp; CASTRO P. 2013. - Significance of sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zootaxa 3665 (1): 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot 
<emphasis id="B910D3709323B503C838FCD6FAFFFCD7" box="[1325,1381,789,815]" italics="true" pageId="39" pageNumber="262">et al.</emphasis>
2013
</bibRefCitation>
</taxonomicName>
: fig.18A, B. 
<taxonomicName id="4C6474E19323B503CEB0FCF5FAAAFCB7" authority=": Guinot et al. 2013" authorityName="Guinot" authorityYear="2013" box="[933,1328,821,848]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CEB0FCF5FBF1FCB7" box="[933,1131,821,847]" italics="true" pageId="39" pageNumber="262">Dorippoides nudipes</emphasis>
: 
<bibRefCitation id="EFF572939323B503C963FCF5FAAAFCB7" author="Guinot" box="[1142,1328,821,847]" etAl="et al." firstAuthor="Guinot" pageId="39" pageNumber="262" pagination="1 - 414" refId="ref122703" refString="GUINOT D., TAVARES M. &amp; CASTRO P. 2013. - Significance of sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zootaxa 3665 (1): 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot 
<emphasis id="B910D3709323B503C9D2FCF6FB6CFCB7" box="[1223,1270,821,847]" italics="true" pageId="39" pageNumber="262">et al.</emphasis>
2013
</bibRefCitation>
</taxonomicName>
: fig. 19A, B.
</paragraph>
<paragraph id="8BDB0F629323B503CE3FFCB5FBA5FC77" blockId="39.[810,1456,885,1454]" box="[810,1087,885,911]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CE3FFCB5FBA5FC77" box="[810,1087,885,911]" italics="true" pageId="39" pageNumber="262">
Gonopods (
<figureCitation id="135F13E79323B503CE88FCB5FC66FC77" box="[925,1020,885,911]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="39" pageNumber="262">Figs 17C</figureCitation>
; 
<figureCitation id="135F13E79323B503C91FFCB5FBADFC77" box="[1034,1079,885,911]" captionStart="FIG" captionStartId="83.[132,143,1939,1957]" captionTargetBox="[294,1300,228,1817]" captionTargetId="figure-13@83.[250,1338,183,1836]" captionTargetPageId="83" captionText="FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis &amp; Manning 1990; I, after Manning &amp; Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured." figureDoi="http://doi.org/10.5281/zenodo.8071329" httpUri="https://zenodo.org/record/8071329/files/figure.png" pageId="39" pageNumber="262">31B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CE38FC55FBDCFBB7" blockId="39.[810,1456,885,1454]" pageId="39" pageNumber="262">
G1 short, stubby, with very elongated coxa and well-developed basis encircling most of endopodite. Apical process twisted, ending in slender simple point, rolled-up in a spiral, triangular (
<taxonomicName id="4C6474E19323B503CE20FC35FC32FBF6" box="[821,936,1012,1038]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CE20FC35FC32FBF6" box="[821,936,1012,1038]" italics="true" pageId="39" pageNumber="262">D. facchino</emphasis>
</taxonomicName>
) or produced into long, thin whip-like appendage (
<taxonomicName id="4C6474E19323B503CE21FBD5FC3DFBD6" box="[820,935,1044,1070]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CE21FBD5FC3DFBD6" box="[820,935,1044,1070]" italics="true" pageId="39" pageNumber="262">D. nudipes</emphasis>
</taxonomicName>
); subdistal cluster of setae. Basal lobe rounded, covered with several setae.
</paragraph>
<paragraph id="8BDB0F629323B503CE56FB94FAA7FAD5" blockId="39.[810,1456,885,1454]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CEC7FB94FAA7FB96" authority=": Chen 1986 a" authorityName="Chen" authorityYear="1986" box="[978,1341,1108,1135]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CEC7FB94FB33FB96" box="[978,1193,1108,1134]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: 
<bibRefCitation id="EFF572939323B503C9A2FB94FAA7FB96" author="Chen" box="[1207,1341,1108,1134]" firstAuthor="Chen" pageId="39" pageNumber="262" pagination="179 - 203" refId="ref112757" refString="CHEN H. 1986 a. - Decapod Crustacea: Dorippidae, in CROSNIER A. (ed.), Resultats des Campagnes MUS ̿ RST ̿ M I et II. Philippines (1976, 1980), vol. 2. (Memoires du Museum national d'Histoire naturelle Serie A - Zoologie; 133 (5) [1985]): 179 - 203, 2 pls." type="journal article" year="1986" yearSuffix="a">Chen 1986a</bibRefCitation>
</taxonomicName>
: fig. 4a, b; Holthuis &amp; Manning 1990: fig. 21a-e (reproduced by 
<bibRefCitation id="EFF572939323B503C84DFBB4FCFFFB56" author="Sin" etAl="et al." firstAuthor="Sin" pageId="39" pageNumber="262" pagination="223 - 230" refId="ref142785" refString="SIN Y. W., LAI J. C. Y., NG P. K. L. &amp; CHU K. H. 2009. - Phylogeny of Dorippoidea (Crustacea: Decapoda: Brachyura) inferred from three mitochondrial genes. Invertebrate Systematics 23 (3): 223 - 230. https: // doi. org / 10.1071 / IS 09020" type="journal article" year="2009">
Sin 
<emphasis id="B910D3709323B503C895FBB5FA2AFB76" box="[1408,1456,1140,1166]" italics="true" pageId="39" pageNumber="262">et al.</emphasis>
2009
</bibRefCitation>
: fig. 4B); 
<bibRefCitation id="EFF572939323B503CEDAFB54FB17FB56" author="Dai &amp; Yang" box="[975,1165,1172,1198]" firstAuthor="Dai" pageId="39" pageNumber="262" pagination="1 - 74" refId="ref114658" refString="DAI A. &amp; YANG S. 1991. - Crabs of the China Seas. China ̿ cean Press, Beijing and Springer-Verlag, Berlin Heidelberg New York Tokyo, ‰ nglish edition. (Translation from Chinese original 1986): 1 - 682, figs 1 - 295, pls 1 - 74." type="journal article" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: fig. 22, as 
<taxonomicName id="4C6474E19323B503C810FB55FC4AFB36" baseAuthorityName="Herbst" baseAuthorityYear="1785" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino" subGenus="Dorippoides">
<emphasis id="B910D3709323B503C810FB55FAC3FB56" box="[1285,1369,1173,1198]" italics="true" pageId="39" pageNumber="262">Dorippe</emphasis>
(
<emphasis id="B910D3709323B503C87DFB55FCF1FB36" italics="true" pageId="39" pageNumber="262">Dorippoides</emphasis>
) 
<emphasis id="B910D3709323B503CE6EFB74FC4AFB36" box="[891,976,1204,1230]" italics="true" pageId="39" pageNumber="262">facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF572939323B503CECAFB74FB36FB36" author="Chen &amp; Sun" box="[991,1196,1204,1230]" firstAuthor="Chen" pageId="39" pageNumber="262" refId="ref112964" refString="CHEN H. &amp; SUN H. 2002. - Brachyura. Marine primitive crabs, in Fauna Sinica, Invertebrata, vol. 30. Arthropoda, Crustacea. Academia Sinica, Science Press, Beijing, China, xiii + 597 p. (In Chinese with ‰ nglish abstract). https: // www. biodiversitylibrary. org / part / 119053" type="book" year="2002">Chen &amp; Sun 2002</bibRefCitation>
: fig. 92.4; 
<bibRefCitation id="EFF572939323B503C831FB74FCFFFB16" author="Guinot" etAl="et al." firstAuthor="Guinot" pageId="39" pageNumber="262" pagination="1 - 414" refId="ref122703" refString="GUINOT D., TAVARES M. &amp; CASTRO P. 2013. - Significance of sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zootaxa 3665 (1): 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot 
<emphasis id="B910D3709323B503C86EFB75FA2AFB36" box="[1403,1456,1204,1230]" italics="true" pageId="39" pageNumber="262">et al.</emphasis>
2013
</bibRefCitation>
: fig. 18C, D; 
<bibRefCitation id="EFF572939323B503CEE8FB14FBE6FB16" author="Vehof" box="[1021,1148,1236,1262]" firstAuthor="Vehof" pageId="39" pageNumber="262" refId="ref147044" refString="VEHOF J. 2020. - Evolution of sperm storage organs in the early diverging lineage of eubrachyuran Dorippidae (Brachyura, Decapoda). Unpublished Doctoral thesis, Lebenswissenschaftlichen Fakultat der Humboldt-Universitat zu Berlin, 109 p." type="book" year="2020">Vehof 2020</bibRefCitation>
: fig.12. 
<taxonomicName id="4C6474E19323B503C9C2FB14FC7FFAF5" authority=": Stephensen 1946" authorityName=": Stephensen" authorityYear="1946" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503C9C2FB14FA33FB16" box="[1239,1449,1236,1262]" italics="true" pageId="39" pageNumber="262">Dorippoides nudipes</emphasis>
: 
<bibRefCitation id="EFF572939323B503CE38FB33FC7FFAF5" author="Stephensen" box="[813,997,1267,1293]" firstAuthor="Stephensen" pageId="39" pageNumber="262" pagination="1 - 237" refId="ref143509" refString="STEPHENSEN K. 1946. - The Brachyura of the Iranian Gulf with an appendix: the male pleopods of the Brachyura, in JESSEN K. &amp; SPARCK R. (eds), Danish Scientific Investigations in Iran, part 4 [1945]. Copenhagen, Munksgaard: 1 - 237." type="book chapter" year="1946">Stephensen 1946</bibRefCitation>
</taxonomicName>
: fig. 4C, as 
<taxonomicName id="4C6474E19323B503C972FB34FA8DFAF5" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[1127,1303,1267,1293]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503C972FB34FA8DFAF5" box="[1127,1303,1267,1293]" italics="true" pageId="39" pageNumber="262">Dorippe facchino</emphasis>
</taxonomicName>
?; 
<bibRefCitation id="EFF572939323B503C83BFB33FA32FAF6" author="Chen" box="[1326,1448,1267,1294]" firstAuthor="Chen" pageId="39" pageNumber="262" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen 1988</bibRefCitation>
: fig. 1f-h; Holthuis &amp; Manning 1990: fig. 
<quantity id="4C9CA2879323B503C9E4FAD3FA86FAD5" box="[1265,1308,1299,1325]" metricMagnitude="-2" metricUnit="kg" metricValue="2.6" pageId="39" pageNumber="262" unit="g" value="26.0">26g</quantity>
, h.
</paragraph>
<paragraph id="8BDB0F629323B503CE56FAF3FA03FAB5" blockId="39.[810,1456,885,1454]" box="[835,1433,1331,1357]" pageId="39" pageNumber="262">G2 shorter than G1, straight, with indistinct flagellum.</paragraph>
<paragraph id="8BDB0F629323B503CE56FA93FB8DFA55" blockId="39.[810,1456,885,1454]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CED9FA93FACEFA95" authority=": Stephensen 1946" authorityName=": Stephensen" authorityYear="1946" box="[972,1364,1363,1390]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D3709323B503CED9FA93FB0FFA95" box="[972,1173,1363,1389]" italics="true" pageId="39" pageNumber="262">Dorippoides nudipes</emphasis>
: 
<bibRefCitation id="EFF572939323B503C9B4FA93FACEFA95" author="Stephensen" box="[1185,1364,1363,1389]" firstAuthor="Stephensen" pageId="39" pageNumber="262" pagination="1 - 237" refId="ref143509" refString="STEPHENSEN K. 1946. - The Brachyura of the Iranian Gulf with an appendix: the male pleopods of the Brachyura, in JESSEN K. &amp; SPARCK R. (eds), Danish Scientific Investigations in Iran, part 4 [1945]. Copenhagen, Munksgaard: 1 - 237." type="book chapter" year="1946">Stephensen 1946</bibRefCitation>
</taxonomicName>
: fig. 4D, as 
<taxonomicName id="4C6474E19323B503CE53FAB4FC68FA75" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[838,1010,1395,1421]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CE53FAB4FC68FA75" box="[838,1010,1395,1421]" italics="true" pageId="39" pageNumber="262">Dorippe facchino</emphasis>
</taxonomicName>
?; 
<bibRefCitation id="EFF572939323B503C91DFAB3FB1AFA75" author="Chen" box="[1032,1152,1395,1421]" firstAuthor="Chen" pageId="39" pageNumber="262" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen 1988</bibRefCitation>
: fig. 1i; 
<taxonomicName id="4C6474E19323B503C9C1FAB3FC31FA55" authority=": Vehof 2020" authorityName="Vehof" authorityYear="2020" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503C9C1FAB3FA32FA75" box="[1236,1448,1395,1421]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: 
<bibRefCitation id="EFF572939323B503CE38FA53FC31FA55" author="Vehof" box="[813,939,1427,1453]" firstAuthor="Vehof" pageId="39" pageNumber="262" refId="ref147044" refString="VEHOF J. 2020. - Evolution of sperm storage organs in the early diverging lineage of eubrachyuran Dorippidae (Brachyura, Decapoda). Unpublished Doctoral thesis, Lebenswissenschaftlichen Fakultat der Humboldt-Universitat zu Berlin, 109 p." type="book" year="2020">Vehof 2020</bibRefCitation>
</taxonomicName>
: fig. 12A.
</paragraph>
<paragraph id="8BDB0F629323B503CE38FA13FB07FA15" blockId="39.[813,1457,1491,1836]" box="[813,1181,1491,1517]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CE38FA13FB07FA15" box="[813,1181,1491,1517]" italics="true" pageId="39" pageNumber="262">
Vulvae (
<figureCitation id="135F13E79323B503CE94FA13FC79FA15" box="[897,995,1491,1517]" captionStart="FIG" captionStartId="41.[133,144,1271,1289]" captionTargetBox="[132,1455,215,1224]" captionTargetId="figure-460@41.[128,1457,215,1233]" captionTargetPageId="41" captionText="FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="39" pageNumber="262">Figs 17D</figureCitation>
; 
<figureCitation id="135F13E79323B503CEE4FA13FBC1FA15" box="[1009,1115,1491,1517]" captionStart="FIG" captionStartId="42.[132,143,1690,1708]" captionTargetBox="[137,1443,215,1647]" captionTargetId="figure-118@42.[137,1444,215,1647]" captionTargetPageId="42" captionText="FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning &amp; Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B." figureDoi="http://doi.org/10.5281/zenodo.8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="39" pageNumber="262">18B, C, F</figureCitation>
; 
<figureCitation id="135F13E79323B503C97DFA13FB0CFA15" box="[1128,1174,1491,1517]" captionStart="FIG" captionStartId="84.[132,143,1111,1129]" captionTargetBox="[145,994,236,1052]" captionTargetId="figure-415@84.[153,572,508,768]" captionTargetPageId="84" captionText="FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis &amp; Manning (1990)." figureDoi="http://doi.org/10.5281/zenodo.8071331" httpUri="https://zenodo.org/record/8071331/files/figure.png" pageId="39" pageNumber="262">32B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CE38FA33FA80F914" blockId="39.[813,1457,1491,1836]" pageId="39" pageNumber="262">
Vulvae on clearly raised, well delimited, globose, papillaelike whitish prominences on sternite 6, each very close to the other, almost joining on median axis, in prolongation of raised oblique setose ridge; opening relatively large, not recessed, near external margin of prominence. In the diagnosis of 
<taxonomicName id="4C6474E19323B503CE91F952FB9AF954" authorityName="Serene &amp; Romimohtarto" authorityYear="1969" box="[900,1024,1682,1708]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709323B503CE91F952FB9AF954" box="[900,1024,1682,1708]" italics="true" pageId="39" pageNumber="262">Dorippoides</emphasis>
</taxonomicName>
by Holthuis &amp; Manning (1990: 48) the statement that the ‘female gonopore’ is on the third sternite is erroneous (a confusion with sternite of P3).
</paragraph>
<paragraph id="8BDB0F629323B503CE56F932FB78F8D4" blockId="39.[813,1457,1491,1836]" pageId="39" pageNumber="262">
Illustrations: 
<taxonomicName id="4C6474E19323B503CEC0F932FCFCF8D4" authority=": Holthuis &amp; Manning 1990" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CEC0F932FB2BF8F4" box="[981,1201,1778,1804]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
: Holthuis &amp; Manning 1990
</taxonomicName>
: fig. 21h, i; 
<bibRefCitation id="EFF572939323B503CEFCF8D2FBFDF8D4" author="Vehof" box="[1001,1127,1810,1836]" firstAuthor="Vehof" pageId="39" pageNumber="262" refId="ref147044" refString="VEHOF J. 2020. - Evolution of sperm storage organs in the early diverging lineage of eubrachyuran Dorippidae (Brachyura, Decapoda). Unpublished Doctoral thesis, Lebenswissenschaftlichen Fakultat der Humboldt-Universitat zu Berlin, 109 p." type="book" year="2020">Vehof 2020</bibRefCitation>
: fig. 7A-C.
</paragraph>
<paragraph id="8BDB0F629323B503CE38F891FBDAF893" blockId="39.[811,1456,1873,2027]" box="[813,1088,1873,1899]" pageId="39" pageNumber="262">
<emphasis id="B910D3709323B503CE38F891FBDAF893" box="[813,1088,1873,1899]" italics="true" pageId="39" pageNumber="262">Female reproductive system</emphasis>
</paragraph>
<paragraph id="8BDB0F629323B503CE38F8B1FC5DF813" blockId="39.[811,1456,1873,2027]" pageId="39" pageNumber="262">
Studied in 
<taxonomicName id="4C6474E19323B503CE88F8B1FBF7F873" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[925,1133,1905,1931]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D3709323B503CE88F8B1FBF7F873" box="[925,1133,1905,1931]" italics="true" pageId="39" pageNumber="262">Dorippoides facchino</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFF572939323B503C985F8B1FAA6F874" author="Vehof" box="[1168,1340,1905,1932]" firstAuthor="Vehof" page="55" pageId="39" pageNumber="262" refId="ref147044" refString="VEHOF J. 2020. - Evolution of sperm storage organs in the early diverging lineage of eubrachyuran Dorippidae (Brachyura, Decapoda). Unpublished Doctoral thesis, Lebenswissenschaftlichen Fakultat der Humboldt-Universitat zu Berlin, 109 p." type="book" year="2020">Vehof (2020: 55</bibRefCitation>
, figs 7E-G, 17, 20, 22). See 
<figureCitation id="135F13E79323B503CEF6F851FBD4F853" box="[995,1102,1937,1963]" captionStart="FIG" captionStartId="95.[132,143,1805,1823]" captionTargetBox="[132,696,217,1773]" captionTargetId="figure-516@95.[191,696,194,1763]" captionTargetPageId="95" captionText="FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22)." figureDoi="http://doi.org/10.5281/zenodo.8071341" httpUri="https://zenodo.org/record/8071341/files/figure.png" pageId="39" pageNumber="262">Figure 37</figureCitation>
and below, 
<emphasis id="B910D3709323B503C9C1F851FC5DF813" italics="true" pageId="39" pageNumber="262">
The female reproductive system in 
<taxonomicName id="4C6474E19323B503CED4F871FBB4F833" authorityName="Latreille" authorityYear="1802" box="[961,1070,1969,1995]" class="Reptilia" family="Colubridae" genus="Brachyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="262" phylum="Chordata" rank="genus">Brachyura</taxonomicName>
, its evolution and unique disposition in 
<taxonomicName id="4C6474E19323B503CE5CF811FC58F813" authorityName="H. Milne Edwards" authorityYear="1837" box="[841,962,2001,2027]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="262" phylum="Arthropoda" rank="family">Dorippidae</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="DF1B5FEA932CB50CCD90FB37FCADFACF" ID-DOI="http://doi.org/10.5281/zenodo.8071293" ID-Zenodo-Dep="8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="40" pageNumber="263" startId="40.[133,144,1271,1289]" targetBox="[135,1455,216,1229]" targetPageId="40" targetType="figure">
<paragraph id="8BDB0F62932CB50CCD90FB37FCADFACF" blockId="40.[133,1457,1271,1337]" pageId="40" pageNumber="263">
<smallCapsWord id="8D3D99BE932CB50CCD90FB37FF07FAF0" baselines="1284,1285" box="[133,157,1271,1289]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="40" pageNumber="263">FIG</smallCapsWord>
. 16. — 
<taxonomicName id="4C6474E1932CB50CCDCBFB37FEC9FAF1" authority="Guinot, 2023" authorityName="Guinot" authorityYear="2023" box="[222,339,1271,1289]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="subFamily" status="n. subfam." subFamily="Dorippoidinae">Dorippoidinae</taxonomicName>
<taxonomicNameLabel id="A2236E0B932CB50CCC4DFB37FE2EFAF1" box="[344,436,1271,1289]" pageId="40" pageNumber="263" rank="subFamily">n. subfam.</taxonomicNameLabel>
: 
<emphasis id="B910D370932CB50CCCACFB37FE5CFAF1" bold="true" box="[441,454,1271,1289]" pageId="40" pageNumber="263">A</emphasis>
, 
<emphasis id="B910D370932CB50CCCC5FB37FE47FAF1" bold="true" box="[464,477,1271,1289]" pageId="40" pageNumber="263">B</emphasis>
, 
<taxonomicName id="4C6474E1932CB50CCCF2FB37FC8FFAF1" authority="(Herbst, 1785)" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[487,789,1271,1289]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932CB50CCCF2FB37FD02FAF1" box="[487,664,1271,1289]" italics="true" pageId="40" pageNumber="263">Dorippoides facchino</emphasis>
(Herbst, 1785)
</taxonomicName>
: 
<emphasis id="B910D370932CB50CCE0EFB37FCB2FAF1" bold="true" box="[795,808,1271,1289]" pageId="40" pageNumber="263">A</emphasis>
, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; 
<emphasis id="B910D370932CB50CC820FB37FAD8FAF1" bold="true" box="[1333,1346,1271,1289]" pageId="40" pageNumber="263">B</emphasis>
, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; 
<emphasis id="B910D370932CB50CCF9CFACEFD0DFAD8" bold="true" box="[649,663,1294,1312]" pageId="40" pageNumber="263">C</emphasis>
, 
<emphasis id="B910D370932CB50CCFB4FACEFD35FAD8" bold="true" box="[673,687,1294,1312]" pageId="40" pageNumber="263">D</emphasis>
, 
<taxonomicName id="4C6474E1932CB50CCFADFACEFBDBFAD8" authority="Manning &amp; Holthuis, 1986" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[696,1089,1294,1312]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CCFADFACEFCFEFAD8" box="[696,868,1294,1312]" italics="true" pageId="40" pageNumber="263">Dorippoides nudipes</emphasis>
<bibRefCitation id="EFF57293932CB50CCE7CFACEFBDBFAD8" author="Manning &amp; Holthuis" box="[873,1089,1294,1312]" firstAuthor="Manning" pageId="40" pageNumber="263" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis,1986</bibRefCitation>
</taxonomicName>
: 
<emphasis id="B910D370932CB50CC95DFACEFBCCFAD8" bold="true" box="[1096,1110,1294,1312]" pageId="40" pageNumber="263">C</emphasis>
, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; 
<emphasis id="B910D370932CB50CCD90FAE5FF09FACF" bold="true" box="[133,147,1317,1335]" pageId="40" pageNumber="263">D</emphasis>
, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.
</paragraph>
</caption>
<paragraph id="8BDB0F62932CB50CCD91FAB3FF71FA75" blockId="40.[130,775,1395,1613]" box="[132,235,1395,1421]" pageId="40" pageNumber="263">
<emphasis id="B910D370932CB50CCD91FAB3FF71FA75" box="[132,235,1395,1421]" italics="true" pageId="40" pageNumber="263">Callosities</emphasis>
</paragraph>
<paragraph id="8BDB0F62932CB50CCD91FA54FD5FF9B5" blockId="40.[130,775,1395,1613]" pageId="40" pageNumber="263">
In both sexes of 
<taxonomicName id="4C6474E1932CB50CCC3FFA53FE61FA55" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[298,507,1427,1453]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932CB50CCC3FFA53FE61FA55" box="[298,507,1427,1453]" italics="true" pageId="40" pageNumber="263">Dorippoides facchino</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E7932CB50CCF1DFA53FDFCFA55" box="[520,614,1427,1453]" captionStart="FIG" captionStartId="87.[132,143,1829,1847]" captionTargetBox="[347,1239,216,1787]" captionTargetId="figure-107@87.[790,881,1000,1790]" captionTargetPageId="87" captionText="FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning &amp; Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8." figureDoi="http://doi.org/10.5281/zenodo.8071333" httpUri="https://zenodo.org/record/8071333/files/figure.png" pageId="40" pageNumber="263">Fig. 33A</figureCitation>
), on dorsal part of P3, between the coxa and the widely exposed pleurite 6, a large thick, movable, whitish membrane lined on pleural side by narrow calcified strip: structure considered here as a ‘simple’ callosity. A similar structure observed in females of 
<taxonomicName id="4C6474E1932CB50CCD91F9F3FF6CF9B4" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[132,246,1586,1612]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CCD91F9F3FF6CF9B4" box="[132,246,1586,1612]" italics="true" pageId="40" pageNumber="263">D. nudipes</emphasis>
</taxonomicName>
but seemingly lacking in males (
<figureCitation id="135F13E7932CB50CCF42F9F2FD22F9B5" box="[599,696,1586,1613]" captionStart="FIG" captionStartId="87.[132,143,1829,1847]" captionTargetBox="[347,1239,216,1787]" captionTargetId="figure-107@87.[790,881,1000,1790]" captionTargetPageId="87" captionText="FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning &amp; Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8." figureDoi="http://doi.org/10.5281/zenodo.8071333" httpUri="https://zenodo.org/record/8071333/files/figure.png" pageId="40" pageNumber="263">Fig. 33B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9932CB50CCD91F9B3FE2DF973" box="[132,439,1651,1677]" pageId="40" pageNumber="263" type="distribution">
<paragraph id="8BDB0F62932CB50CCD91F9B3FE2DF973" blockId="40.[131,775,1651,2028]" box="[132,439,1651,1677]" pageId="40" pageNumber="263">
<smallCapsWord id="8D3D99BE932CB50CCD91F9B3FEBAF973" baselines="1670,1670" box="[132,288,1651,1677]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Distribution" pageId="40" pageNumber="263">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D3D99BE932CB50CCC32F9B8FECFF973" baselines="1670" box="[295,341,1656,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="40" pageNumber="263">AND</smallCapsWord>
<smallCapsWord id="8D3D99BE932CB50CCC49F9B8FE2DF973" baselines="1670" box="[348,439,1656,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="habitat" pageId="40" pageNumber="263">HABITAT</smallCapsWord>
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9932CB50DCD91F952FEB3FA35" lastPageId="41" lastPageNumber="264" pageId="40" pageNumber="263" type="discussion">
<paragraph id="8BDB0F62932CB50CCD91F952FD3CF813" blockId="40.[131,775,1651,2028]" pageId="40" pageNumber="263">
<taxonomicName id="4C6474E1932CB50CCD91F952FED1F954" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[132,331,1682,1708]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932CB50CCD91F952FED1F954" box="[132,331,1682,1708]" italics="true" pageId="40" pageNumber="263">Dorippoides facchino</emphasis>
</taxonomicName>
is known from 
<collectingCountry id="F3734FF2932CB50CCCF6F952FDDBF954" box="[483,577,1682,1708]" name="Sri Lanka" pageId="40" pageNumber="263">Sri Lanka</collectingCountry>
and 
<collectingCountry id="F3734FF2932CB50CCF65F952FD3FF954" box="[624,677,1682,1708]" name="India" pageId="40" pageNumber="263">India</collectingCountry>
(? 
<bibRefCitation id="EFF57293932CB50CCFAFF952FF74F934" author="Pillai &amp; Nair" firstAuthor="Pillai" pageId="40" pageNumber="263" pagination="384 - 386" refId="ref137483" refString="PILLAI K. K. &amp; NAIR N. B. 1970. - ̿ bservations on the reproductive cycles of some crabs from the South-West coast of India. Journal of the Marine Biological Association of India 10 (2): 384 - 386." type="journal article" year="1970">Pillai &amp; Nair 1970</bibRefCitation>
, 
<bibRefCitation id="EFF57293932CB50CCDEFF972FEAAF934" author="Pillai &amp; Nair" box="[250,304,1714,1740]" firstAuthor="Pillai" pageId="40" pageNumber="263" pagination="754 - 770" refId="ref137530" refString="PILLAI K. K &amp; NAIR N. B. 1976. - ̿ bservations on the breeding biology of some crabs from the Southwest coast of India. Journal of the Marine Biological Association of India 15 (2) [1973]: 754 - 770." type="journal article" year="1976">1976</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932CB50CCC41F973FE7CF934" authorityName="Fabricius" authorityYear="1785" box="[340,486,1715,1740]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="astuta">
<emphasis id="B910D370932CB50CCC41F973FE7CF934" box="[340,486,1715,1740]" italics="true" pageId="40" pageNumber="263">Dorippe astuta</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932CB50CCCE4F973FD1CF934" author="DEV ROY M. K." box="[497,646,1714,1741]" pageId="40" pageNumber="263" pagination="1 - 212" refId="ref116553" refString="DEV ROY M. K. 2008. - An annotated checklist of mangrove and coral reef inhabiting Brachyuran crabs of India. Records of Zoological Survey of India ̿ ccasional Paper, 289: 1 - 212." type="journal article" year="2008">Dev Roy 2008</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCF87F973FF65F914" author="Dev Roy &amp; Nandi" firstAuthor="Dev Roy" pageId="40" pageNumber="263" pagination="9 - 23" refId="ref116654" refString="DEV ROY M. K. &amp; NANDI N. C. 2001. - Crustacean biodiversity in mangrove ecosystems of Sundarbans and Bay Islands. Bulletin of the National Institute of Ecology 11: 9 - 23." type="journal article" year="2001">Dev Roy &amp; Nandi 2001</bibRefCitation>
, 
<bibRefCitation id="EFF57293932CB50CCC19F912FED8F914" author="DEV ROY M. K. &amp; NANDI N. C." box="[268,322,1746,1772]" pageId="40" pageNumber="263" pagination="191 - 214" refId="ref116731" refString="DEV ROY M. K. &amp; NANDI N. C. 2008. - Checklist and distribution of Brachyuran crabs of West Bengal. Journal of Environment and Sociobiology 5 (2): 191 - 214." type="journal article" year="2008">2008</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCC58F912FDCBF914" author="Venkataraman" box="[333,593,1746,1772]" etAl="et al." firstAuthor="Venkataraman" pageId="40" pageNumber="263" pagination="1 - 468" refId="ref147407" refString="VENKATARAMAN K., JEYABASKARAN R., RAGHURAM K. P &amp; ALFRED J. R. B. 2004. - Bibliography and checklist of corals and coral reef associated organisms of India. Records of the Zoological Survey of India, ̿ cc. Paper 226: 1 - 468." type="journal article" year="2004">
Venkataraman 
<emphasis id="B910D370932CB50CCCF2F913FD8DF914" box="[487,535,1746,1772]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2004
</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCF49F912FEDEF8F4" author="Ravichandran &amp; Kannupandi" firstAuthor="Ravichandran" pageId="40" pageNumber="263" pagination="331 - 340" refId="ref138519" refString="RAVICHANDRAN S. &amp; KANNUPANDI T. 2007. - Biodiversity of crabs in Pichavaram mangrove environment. ZooIogical Survey of India, National Symposium on Conservation and Valuation of Marine Biodiversity: 331 - 340." type="proceedings paper" year="2007">Ravichandran &amp; Kannupandi 2007</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCC44F932FD88F8F4" author="Vidhya" box="[337,530,1778,1804]" etAl="et al." firstAuthor="Vidhya" pageId="40" pageNumber="263" pagination="2331 - 2336" refId="ref147740" refString="VIDHYA V., JAWAHAR P. &amp; KARUPPASAMY K. 2017. - Annotated check list of the brachyuran crabs (Crustacea: Decapoda) from Gulf of Mannar region, south east coast of India. Journal of Entomology and Zoology Studies 5 (6): 2331 - 2336." type="journal article" year="2017">
Vidhya 
<emphasis id="B910D370932CB50CCCB6F933FE4CF8F4" box="[419,470,1778,1804]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2017
</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCF0AF932FD47F8F4" author="Trivedi" box="[543,733,1778,1804]" etAl="et al." firstAuthor="Trivedi" pageId="40" pageNumber="263" pagination="1 - 83" refId="ref145269" refString="TRIVEDI J. N., TRIVEDI D. J., VACHRAJANI K. D. &amp; NG P. K. L. 2018. - An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa 4502 (1): 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
Trivedi 
<emphasis id="B910D370932CB50CCF7AF933FD3BF8F4" box="[623,673,1778,1804]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2018
</bibRefCitation>
: table 1; 
<bibRefCitation id="EFF57293932CB50CCDD4F8D2FEFBF8D4" author="Bhat" box="[193,353,1810,1836]" etAl="et al." firstAuthor="Bhat" pageId="40" pageNumber="263" pagination="26 - 44" refId="ref110784" refString="BHAT M., TRIVEDI J. &amp; RIVONKER C. 2021. - An annotated checklist of marine brachyuran crabs (Crustacea, Decapoda, Brachyura) from Goa, west coast of India with ten new records. Proceedings of the &quot; Marine Biology Research Symposium - MBRS 2021 &quot; ̿ rganized by Sathyabama Institute of Science and Technology, Chennai, in association with ICAR-National Bureau of Fish Genetic Resources, Lucknow. Taxonomy, Biodiversity and Conservation, TBC- 111 - 0 R: 26 - 44." type="proceedings paper" year="2021">
Bhat 
<emphasis id="B910D370932CB50CCDE2F8D2FEBCF8D4" box="[247,294,1810,1836]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2021
</bibRefCitation>
: table 1) eastward to 
<collectingCountry id="F3734FF2932CB50CCF23F8D2FD09F8D4" box="[566,659,1810,1836]" name="Malaysia" pageId="40" pageNumber="263">Malaysia</collectingCountry>
, 
<collectingCountry id="F3734FF2932CB50CCF89F8D2FC99F8D4" box="[668,771,1810,1836]" name="Indonesia" pageId="40" pageNumber="263">Indonesia</collectingCountry>
, 
<collectingCountry id="F3734FF2932CB50CCD96F8F1FF47F8B3" box="[131,221,1841,1867]" name="Vietnam" pageId="40" pageNumber="263">Vietnam</collectingCountry>
(
<bibRefCitation id="EFF57293932CB50CCDFEF8F1FE3AF8B3" author="Andre" box="[235,416,1841,1868]" firstAuthor="Andre" page="639" pageId="40" pageNumber="263" pagination="638 - 650" refId="ref108978" refString="ANDRE M. 1931. - Crustaces Decapodes provenant de I'lnstitut ̿ ceanographique de Nha-Trang (Annam). Bulletin du Museum national d'Histoire naturelle 2 e serie 3 (7): 638 - 650." type="journal article" year="1931">André 1931: 639</bibRefCitation>
), 
<collectingCountry id="F3734FF2932CB50CCCA4F8F1FD8AF8B3" box="[433,528,1841,1867]" name="Thailand" pageId="40" pageNumber="263">Thailand</collectingCountry>
, southern 
<collectingCountry id="F3734FF2932CB50CCF6EF8F1FD24F8B3" box="[635,702,1841,1867]" name="China" pageId="40" pageNumber="263">China</collectingCountry>
, 
<collectingCountry id="F3734FF2932CB50CCFDCF8F2FF27F894" name="Hong Kong" pageId="40" pageNumber="263">Hong Kong</collectingCountry>
(
<bibRefCitation id="EFF57293932CB50CCDDFF891FE1FF894" author="Chen &amp; Sun" box="[202,389,1873,1900]" firstAuthor="Chen" pageId="40" pageNumber="263" refId="ref112964" refString="CHEN H. &amp; SUN H. 2002. - Brachyura. Marine primitive crabs, in Fauna Sinica, Invertebrata, vol. 30. Arthropoda, Crustacea. Academia Sinica, Science Press, Beijing, China, xiii + 597 p. (In Chinese with ‰ nglish abstract). https: // www. biodiversitylibrary. org / part / 119053" type="book" year="2002">Chen &amp; Sun 2002</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCC85F892FDA7F894" author="Wong" box="[400,573,1873,1900]" etAl="et al." firstAuthor="Wong" pageId="40" pageNumber="263" pagination="102013" refId="ref148990" refString="WONG K. J. H., TAO L. S. R. &amp; LEUNG K. M. Y. 2021. - Subtidal crabs of Hong Kong: Brachyura (Crustacea: Decapoda) from benthic trawl surveys conducted by the University of Hong Kong, 2012 to 2018. Regional Studies in Marine Science 48: 102013. https: // doi. org / https: // doi. org / 10.1016 / j. rsma. 2021.102013" type="journal article" year="2021">
Wong 
<emphasis id="B910D370932CB50CCCC1F892FD99F893" box="[468,515,1873,1899]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2021
</bibRefCitation>
), at depths between 2 and 
<quantity id="4C9CA287932CB50CCDD7F8B1FF60F874" box="[194,250,1905,1932]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="40" pageNumber="263" unit="m" value="80.0">80 m</quantity>
, most often between 10 and 
<quantity id="4C9CA287932CB50CCF35F8B1FDC2F874" box="[544,600,1905,1932]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="40" pageNumber="263" unit="m" value="30.0">30 m</quantity>
, usually in sandy mud, but also in soft clay, soft gray mud, and in stones and sand (Holthuis &amp; Manning 1990; 
<bibRefCitation id="EFF57293932CB50CCCA2F872FDECF833" author="Ng &amp; Davie" box="[439,630,1969,1996]" firstAuthor="Ng" pageId="40" pageNumber="263" pagination="313 - 339" refId="ref115345" refString="DAVIE P. J. F., NG P. K. L., KAENPHET A. &amp; AUNGTONYA C. 2002. - Annotated checklist of Brachyura (Crustacea: Decapoda) principally obtained during the BI ̿ SH ‰ LF survey off western Thailand from 1996 - 1998. Phuket Marine Biological Center Research Bulletin Spec. Publ., 23: 313 - 339." type="journal article" year="2002">Ng &amp; Davie 2002</bibRefCitation>
); there are no definite records from either 
<collectingCountry id="F3734FF2932CB50CCCB0F812FE45F814" box="[421,479,2002,2028]" name="Japan" pageId="40" pageNumber="263">Japan</collectingCountry>
or the 
<collectingCountry id="F3734FF2932CB50CCF3FF811FD38F813" box="[554,674,2001,2027]" name="Philippines" pageId="40" pageNumber="263">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BDB0F62932CB50DCE56FAB3FEB3FA35" blockId="40.[810,1456,1395,2028]" lastBlockId="41.[130,775,1363,1485]" lastPageId="41" lastPageNumber="264" pageId="40" pageNumber="263">
<taxonomicName id="4C6474E1932CB50CCE56FAB3FB93FA75" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[835,1033,1395,1421]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CCE56FAB3FB93FA75" box="[835,1033,1395,1421]" italics="true" pageId="40" pageNumber="263">Dorippoides nudipes</emphasis>
</taxonomicName>
is known from various parts of the eastern Indian Ocean, including the southern Red Sea, the Persian Gulf (
<bibRefCitation id="EFF57293932CB50CCE7BFA73FC45FA35" author="Apel" box="[878,991,1459,1485]" firstAuthor="Apel" pageId="40" pageNumber="263" refId="ref109195" refString="APEL M. 2001. - Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch- Arabischen Golfes. Doctoral thesis, Johann Wolfgang Goethe- Universitat, Frankfurt am Main, 260 p." type="book" year="2001">Apel 2001</bibRefCitation>
), the Gulf of 
<collectingCountry id="F3734FF2932CB50CC961FA73FB20FA35" box="[1140,1210,1459,1485]" name="Oman" pageId="40" pageNumber="263">Oman</collectingCountry>
(
<bibRefCitation id="EFF57293932CB50CC9DFFA73FA32FA36" author="Naderloo" box="[1226,1448,1459,1486]" etAl="et al." firstAuthor="Naderloo" pageId="40" pageNumber="263" pagination="397 - 412" refId="ref134135" refString="NADERLOO R., EBRAHIMNEZHAD S. &amp; SARI A. 2015. - Annotated checklist of the decapod crustaceans of the Gulf of ̿ man, northwestern Indian ̿ cean. Zootaxa 4028 (3): 397 - 412. https: // doi. org / 10.11646 / zootaxa. 4028.3.5" type="journal article" year="2015">
Naderloo 
<emphasis id="B910D370932CB50CC823FA74FAF0FA35" box="[1334,1386,1459,1485]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2015
</bibRefCitation>
: table 2), 
<collectingCountry id="F3734FF2932CB50CCE99FA13FC22FA15" box="[908,952,1491,1517]" name="Iran" pageId="40" pageNumber="263">Iran</collectingCountry>
(
<bibRefCitation id="EFF57293932CB50CCED2FA13FB1AFA15" author="Stephensen" box="[967,1152,1491,1517]" firstAuthor="Stephensen" pageId="40" pageNumber="263" pagination="1 - 237" refId="ref143509" refString="STEPHENSEN K. 1946. - The Brachyura of the Iranian Gulf with an appendix: the male pleopods of the Brachyura, in JESSEN K. &amp; SPARCK R. (eds), Danish Scientific Investigations in Iran, part 4 [1945]. Copenhagen, Munksgaard: 1 - 237." type="book chapter" year="1946">Stephensen 1946</bibRefCitation>
), southern 
<collectingCountry id="F3734FF2932CB50CC9E9FA13FAE6FA15" box="[1276,1404,1491,1517]" name="Madagascar" pageId="40" pageNumber="263">Madagascar</collectingCountry>
, and probably reaches 
<collectingCountry id="F3734FF2932CB50CCEF2FA33FBEAF9F5" box="[999,1136,1523,1549]" name="South Africa" pageId="40" pageNumber="263">South Africa</collectingCountry>
(Holthuis &amp; Manning 1990; 
<bibRefCitation id="EFF57293932CB50CCE38F9D2FC3CF9D5" author="Chen" box="[813,934,1554,1581]" firstAuthor="Chen" pageId="40" pageNumber="263" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen 1988</bibRefCitation>
). And also from the western Indian Ocean: 
<collectingCountry id="F3734FF2932CB50CC863F9D2FA35F9D4" box="[1398,1455,1554,1580]" name="India" pageId="40" pageNumber="263">India</collectingCountry>
(
<bibRefCitation id="EFF57293932CB50CCE23F9F2FB8CF9B5" author="Devi &amp; Kumar" box="[822,1046,1586,1613]" firstAuthor="Devi" pageId="40" pageNumber="263" pagination="625 - 630" refId="ref116809" refString="DEVI S. S. &amp; KUMAR A. B. 2017. - ̿ n two species of crabs (Decapoda, Brachyura): Cycloes marisrubri Galil &amp; Clark, 1996 (Calappidae) and Dorippoides nudipes Manning &amp; Holthuis, 1986 (Dorippidae) from India. Crustaceana 90 (5): 625 - 630. https: // doi. org / 10.1163 / 15685403 - 00003670" type="journal article" year="2017">Devi &amp; Kumar 2017</bibRefCitation>
: fig. 1E, F; 
<bibRefCitation id="EFF57293932CB50CC984F9F2FAC9F9B5" author="Trivedi" box="[1169,1363,1586,1613]" etAl="et al." firstAuthor="Trivedi" pageId="40" pageNumber="263" pagination="1 - 83" refId="ref145269" refString="TRIVEDI J. N., TRIVEDI D. J., VACHRAJANI K. D. &amp; NG P. K. L. 2018. - An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa 4502 (1): 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
Trivedi 
<emphasis id="B910D370932CB50CC9F6F9F3FA8CF9B4" box="[1251,1302,1586,1612]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2018
</bibRefCitation>
: table 1; 
<bibRefCitation id="EFF57293932CB50CCE39F992FC73F994" author="Gosavi" box="[812,1001,1618,1644]" etAl="et al." firstAuthor="Gosavi" pageId="40" pageNumber="263" pagination="45 - 66" refId="ref120837" refString="GOSAVI S., PUROHIT B., MITRA S., PATEL K., VACHHRAJANI K. &amp; TRIVEDI J. 2021. - Annotated checklist of marine decapods (Crustacea: Decapoda) of Gujarat state with three new records. Proceedings of the &quot; Marine Biology Research Symposium - MBRS 2021 &quot; ̿ rganized by Sathyabama Institute of Science and Technology, Chennai, in association with ICAR-National Bureau of Fish Genetic Resources, Lucknow. Taxonomy, Biodiversity and Conservation, TBC - 115 - P ̿: 45 - 66." type="book chapter" year="2021">
Gosavi 
<emphasis id="B910D370932CB50CCE6EF993FC37F994" box="[891,941,1618,1644]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2021
</bibRefCitation>
: table 3) and western 
<collectingCountry id="F3734FF2932CB50CC9D8F992FAB6F994" box="[1229,1324,1618,1644]" name="Thailand" pageId="40" pageNumber="263">Thailand</collectingCountry>
(
<bibRefCitation id="EFF57293932CB50CC82FF992FCF8F974" author="Davie" etAl="et al." firstAuthor="Davie" pageId="40" pageNumber="263" pagination="313 - 339" refId="ref115345" refString="DAVIE P. J. F., NG P. K. L., KAENPHET A. &amp; AUNGTONYA C. 2002. - Annotated checklist of Brachyura (Crustacea: Decapoda) principally obtained during the BI ̿ SH ‰ LF survey off western Thailand from 1996 - 1998. Phuket Marine Biological Center Research Bulletin Spec. Publ., 23: 313 - 339." type="journal article" year="2002">
Davie 
<emphasis id="B910D370932CB50CC86BF993FA2AF994" box="[1406,1456,1618,1644]" italics="true" pageId="40" pageNumber="263">et al.</emphasis>
2002
</bibRefCitation>
; 
<bibRefCitation id="EFF57293932CB50CCE78F9B3FBBEF974" author="Ng &amp; Davie" box="[877,1060,1650,1677]" firstAuthor="Ng" pageId="40" pageNumber="263" pagination="313 - 339" refId="ref115345" refString="DAVIE P. J. F., NG P. K. L., KAENPHET A. &amp; AUNGTONYA C. 2002. - Annotated checklist of Brachyura (Crustacea: Decapoda) principally obtained during the BI ̿ SH ‰ LF survey off western Thailand from 1996 - 1998. Phuket Marine Biological Center Research Bulletin Spec. Publ., 23: 313 - 339." type="journal article" year="2002">Ng &amp; Davie 2002</bibRefCitation>
). Recent records of 
<taxonomicName id="4C6474E1932CB50CC9FEF9B2FA2AF974" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[1259,1456,1650,1676]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CC9FEF9B2FA2AF974" box="[1259,1456,1650,1676]" italics="true" pageId="40" pageNumber="263">Dorippoides nudipes</emphasis>
</taxonomicName>
in the Middle East need to be reviewed. In 
<bibRefCitation id="EFF57293932CB50CC9EAF952FA35F954" author="Naderloo's" box="[1279,1455,1682,1708]" firstAuthor="Naderloo's" pageId="40" pageNumber="263" refId="ref134094" refString="NADERLOO R. 2017. - Atlas of Crabs of the Persian Gulf. Springer, Switzerland, xi + 444 p. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo’s 2017</bibRefCitation>
<emphasis id="B910D370932CB50CCE38F972FBE8F934" box="[813,1138,1714,1740]" italics="true" pageId="40" pageNumber="263">
<taxonomicName id="4C6474E1932CB50CCE38F972FCFAF934" box="[813,864,1714,1740]" class="Insecta" family="Cetoniidae" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="263" phylum="Arthropoda" rank="genus">Atlas</taxonomicName>
of crabs of the Persian Gulf
</emphasis>
, although the keys to 
<taxonomicName id="4C6474E1932CB50CC84EF973FC06F914" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D370932CB50CC84EF973FC06F914" italics="true" pageId="40" pageNumber="263">Dorippe quadridens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6474E1932CB50CCED9F912FB0FF914" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[972,1173,1746,1772]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CCED9F912FB0FF914" box="[972,1173,1746,1772]" italics="true" pageId="40" pageNumber="263">Dorippoides nudipes</emphasis>
</taxonomicName>
, their figures of the G1 and distribution maps (
<bibRefCitation id="EFF57293932CB50CCEE8F932FB55F8F4" author="Naderloo" box="[1021,1231,1778,1804]" firstAuthor="Naderloo" page="47" pageId="40" pageNumber="263" refId="ref134094" refString="NADERLOO R. 2017. - Atlas of Crabs of the Persian Gulf. Springer, Switzerland, xi + 444 p. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017: 47</bibRefCitation>
, fig. 4.2.e and 4.2.f, respectively) are correct, the carapace assumed to be that of 
<taxonomicName id="4C6474E1932CB50CCE38F8F1FC67F8B3" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[813,1021,1841,1867]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CCE38F8F1FC67F8B3" box="[813,1021,1841,1867]" italics="true" pageId="40" pageNumber="263">Dorippoides nudipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF57293932CB50CC919F8F1FB35F8B4" author="Naderloo" box="[1036,1199,1841,1868]" firstAuthor="Naderloo" pageId="40" pageNumber="263" refId="ref134094" refString="NADERLOO R. 2017. - Atlas of Crabs of the Persian Gulf. Springer, Switzerland, xi + 444 p. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017</bibRefCitation>
: fig. 7.3) is actually that of 
<taxonomicName id="4C6474E1932CB50CCE5DF892FB89F893" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[840,1043,1873,1899]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D370932CB50CCE5DF892FB89F893" box="[840,1043,1873,1899]" italics="true" pageId="40" pageNumber="263">Dorippe quadridens</emphasis>
</taxonomicName>
. In our opinion, the figures for the two species have been accidentally mixed. The correct representation of the carapace of 
<taxonomicName id="4C6474E1932CB50CC939F851FA9AF853" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[1068,1280,1937,1963]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932CB50CC939F851FA9AF853" box="[1068,1280,1937,1963]" italics="true" pageId="40" pageNumber="263">Dorippoides nudipes</emphasis>
</taxonomicName>
is shown under 
<taxonomicName id="4C6474E1932CB50CCE38F872FC63F833" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[813,1017,1969,1995]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="263" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D370932CB50CCE38F872FC63F833" box="[813,1017,1969,1995]" italics="true" pageId="40" pageNumber="263">Dorippe quadridens</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF57293932CB50CC91CF871FB36F833" author="Naderloo" box="[1033,1196,1969,1995]" firstAuthor="Naderloo" pageId="40" pageNumber="263" refId="ref134094" refString="NADERLOO R. 2017. - Atlas of Crabs of the Persian Gulf. Springer, Switzerland, xi + 444 p. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017</bibRefCitation>
: fig. 7.2). Subsequently, crabs from the northwestern Persian Gulf, 
<collectingCountry id="F3734FF2932CB50CC9E0F812FAB9F814" box="[1269,1315,2002,2028]" name="Iraq" pageId="40" pageNumber="263">Iraq</collectingCountry>
, identified as 
<taxonomicName id="4C6474E1932DB50DCD91FA94FEC8FA95" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[132,338,1363,1389]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D370932DB50DCD91FA94FEC8FA95" box="[132,338,1363,1389]" italics="true" pageId="41" pageNumber="264">Dorippe quadridens</emphasis>
</taxonomicName>
by the records of 
<bibRefCitation id="EFF57293932DB50DCF01FA94FD65FA95" author="Yasser &amp; Naser" box="[532,767,1363,1390]" firstAuthor="Yasser" pageId="41" pageNumber="264" pagination="1 - 3" refId="ref150095" refString="YASSER A. &amp; NASER M. 2019. - A new record of Dorippe quadridens (Fabricius, 1793) (Decapoda, Brachyura, Dorippidae) from the north western Persian-Arabian Gulf, Iraq. Journal of Biological Studies 2 (1): 1 - 3." type="journal article" year="2019">Yasser &amp; Naser (2019</bibRefCitation>
: fig. 2) and 
<bibRefCitation id="EFF57293932DB50DCDE0FAB3FE7AFA75" author="Al-Khafaji" box="[245,480,1395,1421]" etAl="et al." firstAuthor="Al-Khafaji" pageId="41" pageNumber="264" pagination="118 - 126" refId="ref108801" refString="AL-KHAFAJI K., AL-MALIKY T. H. Y. &amp; AL-MALIKY A. M. J. 2019. - First record of the crab, Droippe [sic] quadridens (Fabricius, 1793) (Brachyura: Dorippidae), from the Iraqi coastal waters of the NWArabian Gulf, with notes on the occurrence of seven species of crabs in the region. Arthropods 8 (4): 118 - 126." type="journal article" year="2019">
Al-Khafaji 
<emphasis id="B910D370932DB50DCC7CFAB4FE01FA75" box="[361,411,1395,1421]" italics="true" pageId="41" pageNumber="264">et al.</emphasis>
(2019
</bibRefCitation>
: fig. 2a, table 2) are likewise 
<taxonomicName id="4C6474E1932DB50DCD97FA53FECCFA55" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[130,342,1427,1453]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932DB50DCD97FA53FECCFA55" box="[130,342,1427,1453]" italics="true" pageId="41" pageNumber="264">Dorippoides nudipes</emphasis>
</taxonomicName>
and not 
<taxonomicName id="4C6474E1932DB50DCCA8FA54FD11FA55" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[445,651,1427,1453]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D370932DB50DCCA8FA54FD11FA55" box="[445,651,1427,1453]" italics="true" pageId="41" pageNumber="264">Dorippe quadridens</emphasis>
</taxonomicName>
(see under 
<taxonomicName id="4C6474E1932DB50DCD91FA74FE83FA35" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[132,281,1459,1485]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D370932DB50DCD91FA74FE83FA35" box="[132,281,1459,1485]" italics="true" pageId="41" pageNumber="264">D. quadridens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF1B5FEA932DB50DCD90FB37FAE9FAD8" ID-DOI="http://doi.org/10.5281/zenodo.8071297" ID-Zenodo-Dep="8071297" httpUri="https://zenodo.org/record/8071297/files/figure.png" pageId="41" pageNumber="264" startId="41.[133,144,1271,1289]" targetBox="[132,1455,215,1224]" targetPageId="41" targetType="figure">
<paragraph id="8BDB0F62932DB50DCD90FB37FAE9FAD8" blockId="41.[133,1455,1271,1314]" pageId="41" pageNumber="264">
<smallCapsWord id="8D3D99BE932DB50DCD90FB37FF07FAF0" baselines="1284,1285" box="[133,157,1271,1289]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="41" pageNumber="264">FIG</smallCapsWord>
. 17. — 
<taxonomicName id="4C6474E1932DB50DCDCBFB37FEC9FAF1" authority="Guinot, 2023" authorityName="Guinot" authorityYear="2023" box="[222,339,1271,1289]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="subFamily" status="n. subfam." subFamily="Dorippoidinae">Dorippoidinae</taxonomicName>
<taxonomicNameLabel id="A2236E0B932DB50DCC4DFB37FE28FAF1" box="[344,434,1271,1289]" pageId="41" pageNumber="264" rank="subFamily">n. subfam.</taxonomicNameLabel>
<taxonomicName id="4C6474E1932DB50DCCA0FB37FD7EFAF1" authority="(Herbst, 1785)" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[437,740,1271,1289]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCCA0FB37FDFCFAF1" box="[437,614,1271,1289]" italics="true" pageId="41" pageNumber="264">Dorippoides facchino</emphasis>
(Herbst, 1785)
</taxonomicName>
: 
<emphasis id="B910D370932DB50DCFFFFB37FC96FAF1" bold="true" box="[746,780,1271,1289]" pageId="41" pageNumber="264">A -C</emphasis>
, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: 
<emphasis id="B910D370932DB50DC809FB37FAB3FAF1" bold="true" box="[1308,1321,1271,1289]" pageId="41" pageNumber="264">A</emphasis>
, anterior ventral view; 
<emphasis id="B910D370932DB50DCDA0FACEFF58FAD8" bold="true" box="[181,194,1294,1312]" pageId="41" pageNumber="264">B</emphasis>
, thoracic sternum with pleon; 
<emphasis id="B910D370932DB50DCCAAFACDFE57FAE7" bold="true" box="[447,461,1293,1311]" pageId="41" pageNumber="264">C</emphasis>
, without pleon. 
<emphasis id="B910D370932DB50DCF47FACEFDFAFAD8" bold="true" box="[594,608,1294,1312]" pageId="41" pageNumber="264">D</emphasis>
, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae.
</paragraph>
</caption>
<subSubSection id="C37E5CE9932DB50DCD91FA33FA35F8B4" pageId="41" pageNumber="264" type="biology_ecology">
<paragraph id="8BDB0F62932DB50DCD91FA33FEEEF9F3" blockId="41.[132,776,1523,2028]" box="[132,372,1523,1549]" pageId="41" pageNumber="264">
<smallCapsWord id="8D3D99BE932DB50DCD91FA33FF69F9F3" baselines="1542,1542" box="[132,243,1523,1549]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Carrying" pageId="41" pageNumber="264">CARRYING</smallCapsWord>
<smallCapsWord id="8D3D99BE932DB50DCDEFFA38FEEEF9F3" baselines="1542" box="[250,372,1528,1547]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="behaviour" pageId="41" pageNumber="264">BEHAVIOUR</smallCapsWord>
</paragraph>
<paragraph id="8BDB0F62932DB50DCD91F9D2FB7DF934" blockId="41.[132,776,1523,2028]" lastBlockId="41.[813,1456,1363,1868]" pageId="41" pageNumber="264">
The association of 
<taxonomicName id="4C6474E1932DB50DCC44F9D2FDB0F9D4" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[337,554,1554,1580]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCC44F9D2FDB0F9D4" box="[337,554,1554,1580]" italics="true" pageId="41" pageNumber="264">Dorippoides facchino</emphasis>
</taxonomicName>
, the ‘porter crab’ or ‘anemone-carrying crab’, with another organism, in fact with a sea anemone, has long been recognised (Herbst 1796: 215, as 
<taxonomicName id="4C6474E1932DB50DCDB4F9B3FEDFF974" authorityName="Herbst" authorityYear="1785" box="[161,325,1650,1677]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCDB4F9B3FEDFF974" box="[161,325,1650,1677]" italics="true" pageId="41" pageNumber="264">Cancer facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCC40F9B2FDBDF974" author="Stimpson" box="[341,551,1650,1677]" firstAuthor="Stimpson" page="37" pageId="41" pageNumber="264" pagination="375 - 384" refId="ref143899" refString="STIMPSON W. 1855. - Descriptions of some of the new marine Invertebrata from the Chinese and Japanese Seas. Proceedings of the Academy of Natural Sciences of Philadelphia 7: 375 - 384. https: // www. biodiversitylibrary. org / bibliography / 51444" type="journal article" year="1855">Stimpson 1855: 37</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCF45F9B3FC9AF974" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[592,768,1650,1676]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCF45F9B3FC9AF974" box="[592,768,1650,1676]" italics="true" pageId="41" pageNumber="264">Dorippe facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCD91F952FEC5F955" author="Verrill" box="[132,351,1682,1709]" firstAuthor="Verrill" page="58 - 60" pageId="41" pageNumber="264" pagination="51 - 104" refId="ref147502" refString="VERRILL A. E. 1869 a. - Actiniaria, with Supplement and Geographical Lists. Synopses of the Polyps and Corals of the North Pacific ‰ xploring ‰ xpedition, under Commodore C. Ringgold and Captain John Rodgers, U. S. N., from 1853 to 1856, Collected by Dr. William Stimpson, Naturalist to the ‰ xpedition, Part IV. Proceedings of the Essex Institute 6: 51 - 104, pls 1 - 2." type="journal article" year="1869" yearSuffix="a">Verrill 1869a: 58-60</bibRefCitation>
; 
<bibRefCitation id="EFF57293932DB50DCC7FF952FD81F955" author="Verrill" box="[362,539,1682,1709]" firstAuthor="Verrill" page="249 - 250" pageId="41" pageNumber="264" pagination="239 - 250" refId="ref147588" refString="VERRILL A. E. 1869 b. - ̿ n the parasitic habits of Crustacea. American Naturalist 3: 239 - 250. https: // doi. org / 10.1086 / 270420" type="journal article" year="1869" yearSuffix="b">1869b: 249-250</bibRefCitation>
, pl. 2, fig. 1, as 
<taxonomicName id="4C6474E1932DB50DCFAAF953FF21F934" baseAuthorityName="Herbst" baseAuthorityYear="1785" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCFAAF953FF21F934" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
[reproduced by Holthuis &amp; Manning 1990: fig. 25a]; 
<bibRefCitation id="EFF57293932DB50DCD91F912FEEFF915" author="Henderson" box="[132,373,1746,1773]" firstAuthor="Henderson" page="405" pageId="41" pageNumber="264" pagination="325 - 458" refId="ref123912" refString="HENDERSON J. R. 1893. - A Contribution to Indian Carcinology. Transactions of the Linnean Society of London (2) 5: 325 - 458." type="journal article" year="1893">Henderson 1893: 405</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCC88F913FD8EF914" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[413,532,1746,1772]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCC88F913FD8EF914" box="[413,532,1746,1772]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCF36F912FD7CF914" author="ALCOCK A." box="[547,742,1746,1772]" pageId="41" pageNumber="264" pagination="134 - 296" refId="ref108587" refString="ALCOCK A. 1896. - The Brachyura ̿ xystoma: materials for a carcinological fauna of India, No. 2. Journal of the Asiatic Society of Bengal 65 (2): 134 - 296." type="journal article" year="1896">Alcock 1896: 279</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCD91F933FE9DF8F4" baseAuthorityName="De Haan" baseAuthorityYear="1841" box="[132,263,1778,1804]" class="Malacostraca" family="Dorippidae" genus="Paradorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="B910D370932DB50DCD91F933FE9DF8F4" box="[132,263,1778,1804]" italics="true" pageId="41" pageNumber="264">D. granulata</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCC01F932FE66F8F4" author="Lanchester" box="[276,508,1778,1804]" firstAuthor="Lanchester" page="769" pageId="41" pageNumber="264" pagination="719 - 770" refId="ref129223" refString="LANCHESTER W. F. 1900. - ̿ n a collection of Crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London 1900: 719 - 770. https: // www. biodiversitylibrary. org / part / 49623" type="journal article" year="1900">Lanchester 1900: 769</bibRefCitation>
; 
<bibRefCitation id="EFF57293932DB50DCF13F932FDFDF8F5" author="Lanchester" box="[518,615,1778,1805]" firstAuthor="Lanchester" page="55" pageId="41" pageNumber="264" pagination="33" refId="ref129274" refString="LANCHESTER W. F. 1902. - ̿ n the Crustacea collected during the &quot; Skeat &quot; ‰ xpedition to the Malay Penainsula, together with a note on the genus Actaeopsis, Part 1. Brachyura, Stomatopoda, and Macrura. Proceedings of the Zoological Society of London (2) (1901): 534 - 574, pls 33, 34." type="journal article" year="1902">1902: 55</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCF9EF933FC9AF8F4" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[651,768,1778,1804]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCF9EF933FC9AF8F4" box="[651,768,1778,1804]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCD91F8D2FE13F8D4" author="Shelford" box="[132,393,1810,1836]" firstAuthor="Shelford" page="299 - 300" pageId="41" pageNumber="264" refId="ref141930" refString="SHELFORD R. W. C. 1916. - A Naturalist in Borneo. London, xxviii + 331 p., 32 pls. [A facsimile reprint edition was published in 1985 by ̿ xford University Press]. https: // www. biodiversitylibrary. org / bibliography / 32180" type="book" year="1916">Shelford 1916: 299-300</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCCA5F8D2FDBCF8D4" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[432,550,1810,1836]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCCA5F8D2FDBCF8D4" box="[432,550,1810,1836]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCF21F8D2FF2BF8B4" author="Hornell" firstAuthor="Hornell" page="934 - 935" pageId="41" pageNumber="264" pagination="926 - 936" refId="ref124703" refString="HORNELL J. S. 1922. - Some commensals of Indian Alcyonarians and Crabs. Journal of the Bombay Natural History Society 28 (4): 926 - 936. https: // www. biodiversitylibrary. org / part / 154427" type="journal article" year="1922">Hornell 1922: 934- 935</bibRefCitation>
, fig. 5, as 
<taxonomicName id="4C6474E1932DB50DCC30F8F2FE02F8B3" authorityName=", Manning" authorityYear="1984" baseAuthorityName="Linnaeus" baseAuthorityYear="1764" box="[293,408,1841,1867]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="dorsipes">
<emphasis id="B910D370932DB50DCC30F8F2FE02F8B3" box="[293,408,1841,1867]" italics="true" pageId="41" pageNumber="264">D. dorsipes</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCCBDF8F1FDC0F8B3" author="Verrill" box="[424,602,1841,1868]" firstAuthor="Verrill" page="16" pageId="41" pageNumber="264" pagination="1 - 30" refId="ref147623" refString="VERRILL A. E. 1928. - Havaiian shallow water Anthozoa. Bernice P. Bishop Museum Bulletin 49: 1 - 30." type="journal article" year="1928">Verrill 1928: 16</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCF90F8F2FD65F8B3" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[645,767,1841,1867]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCF90F8F2FD65F8B3" box="[645,767,1841,1867]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCD91F892FEB1F894" author="Hose" box="[132,299,1874,1900]" firstAuthor="Hose" page="31" pageId="41" pageNumber="264" refId="ref124747" refString="HOSE C. 1929. - The Field-Book of a Jungle-Wallah: being a Description of Shore, River and Forest Life in Sarawak. London, H. F. &amp; G. Witherby, viii + 216 p., 14 pls." type="book" year="1929">Hose 1929: 31</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCC43F892FE55F893" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[342,463,1873,1899]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCC43F892FE55F893" box="[342,463,1873,1899]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCCCAF891FD0BF894" author="Shen" box="[479,657,1873,1900]" firstAuthor="Shen" page="101" pageId="41" pageNumber="264" pagination="92 - 110" refId="ref142035" refString="SHEN C. - J. 1931. - The Crabs of Hong Kong. Part I. HongKong Naturalist 2 (2): 92 - 110." type="journal article" year="1931">Shen 1931: 101</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCFA9F892FF27F873" baseAuthorityName="Herbst" baseAuthorityYear="1785" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCFA9F892FF27F873" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCDDBF8B1FE09F873" author="Gravely" box="[206,403,1905,1932]" firstAuthor="Gravely" page="81" pageId="41" pageNumber="264" pagination="1 - 112" refId="ref120934" refString="GRAVELY F. H. 1941. - Shells and other animals remains found on the Madras beach. I. Groups other than snails, etc. (Mollusca Gastropoda). Bulletin of the Madras Government Museum new series (Natural History) 5 (1): 1 - 112." type="journal article" year="1941">Gravely 1941: 81</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCCAAF8B2FDA1F873" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[447,571,1905,1931]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCCAAF8B2FDA1F873" box="[447,571,1905,1931]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCF59F8B1FD78F874" author="Chopra" box="[588,738,1905,1932]" firstAuthor="Chopra" pageId="41" pageNumber="264" pagination="25 - 52" refId="ref113246" refString="CHOPRA B. N. 1935. - Further Notes on Crustacea Decapoda in the Indian Museum. III. ̿ n the Decapod Crustacea collected by the Bengal Pilot Service off the Mouth of the River Hughli. Dromiacea and ̿ xystomata. Records of the Indian Museum 35 (1): 25 - 52." type="journal article" year="1935">Chopra 1935</bibRefCitation>
, as 
<taxonomicName id="4C6474E1932DB50DCD91F852FF66F853" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[132,252,1937,1963]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCD91F852FF66F853" box="[132,252,1937,1963]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCC19F851FD1CF853" author="Serene &amp; Romimohtarto" box="[268,646,1937,1964]" firstAuthor="Serene" page="11" pageId="41" pageNumber="264" pagination="1 - 35" refId="ref141809" refString="SERENE R. &amp; ROMIMOHTARTO K. 1969. - ̿ bservations on the species of Dorippe from the Indo-Malayan Region. Marine Research in Indonesia (Penelitian Laut di Indonesia) 9: 1 - 35." type="journal article" year="1969">Serène &amp; Romimohtarto 1969: 11</bibRefCitation>
; 
<bibRefCitation id="EFF57293932DB50DCF81F852FEC9F833" author="Morton &amp; Morton" firstAuthor="Morton" page="187" pageId="41" pageNumber="264" refId="ref133689" refString="MORTON B. &amp; MORTON J. 1983. - The Sea Shore Ecology of Hong Kong. Hong Kong University Press, xiv + 350 p. https: // doi. org / 10.1017 / S 0025315400059816" type="book" year="1983">Morton &amp; Morton 1983: 187</bibRefCitation>
, fig. 10.4, 7, as 
<taxonomicName id="4C6474E1932DB50DCF14F872FD1DF833" baseAuthorityName="De Haan" baseAuthorityYear="1841" box="[513,647,1969,1995]" class="Malacostraca" family="Dorippidae" genus="Paradorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="B910D370932DB50DCF14F872FD1DF833" box="[513,647,1969,1995]" italics="true" pageId="41" pageNumber="264">D. granulata</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF57293932DB50DCF80F872FF6DF813" author="Tan &amp; Ng" firstAuthor="Tan" page="149" pageId="41" pageNumber="264" refId="ref144682" refString="TAN S. H. &amp; NG P. K. L. 1988. - A guide to Seashore Life. First edition. Singapore, Singapore Science Centre, 160 p." type="book" year="1988">Tan &amp; Ng 1988: 149</bibRefCitation>
, unnumbered fig.; 1992: 149; Holthuis &amp; Manning 1990: 60, 63, fig. 25a, b; 
<bibRefCitation id="EFF57293932DB50DC97EFA93FADEFA95" author="Manning" box="[1131,1348,1363,1390]" firstAuthor="Manning" page="114" pageId="41" pageNumber="264" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning 1993: 114</bibRefCitation>
, fig. 3a, b; 
<bibRefCitation id="EFF57293932DB50DCE38FAB3FC77FA76" author="GUINOT D. &amp; RICHER DE FORGES B." box="[813,1005,1395,1422]" pageId="41" pageNumber="264" pagination="283 - 517" refId="ref121946" refString="GUINOT D. &amp; RICHER DE FORGES B. 1995. - Crustacea, Decapoda, Brachyura: revision de la famille des Homolidae de Haan, 1841, in CROSNIER A. (ed.), Resultats des Campagnes MUS ̿ RST ̿ M. Vol. 13. Museum national d'Histoire naturelle, Paris: 283 - 517 (Memoires du Museum national d'Histoire naturelle; 163). https: // biodiversitylibrary. org / page / 58815176" type="journal article" year="1995">
Guinot 
<emphasis id="B910D370932DB50DCE95FAB4FC2BFA75" box="[896,945,1395,1421]" italics="true" pageId="41" pageNumber="264">et al.</emphasis>
1995
</bibRefCitation>
: fig. 4A, pl. 1B; 
<bibRefCitation id="EFF57293932DB50DC98DFAB3FCC3FA56" author="Guinot &amp; Wicksten" firstAuthor="Guinot" page="599" pageId="41" pageNumber="264" pagination="583 - 638" refId="ref122397" refString="GUINOT D. &amp; WICKSTEN M. K. 2015. - Camouflage: carrying behaviour, decoration behaviour, and other modalities of concealment in Brachyura (Chapter 71 - 11), in CASTRO P., DAVIE P. J. F., GUINOT D., SCHRAM F. R. &amp; VON VAUPEL KLEIN J. C. (eds), Decapoda: Brachyura. Treatise on Zoology - Anatomy, Taxonomy, Biology, vol. 9, Part C, 2 vols, Crustacea. Brill, Leiden and Boston: 583 - 638. https: // doi. org / 10.1163 / 9789004190832 _ 013" type="book chapter" year="2015">Guinot &amp; Wicksten 2015: 599</bibRefCitation>
, fig. 71-11.8B; 
<bibRefCitation id="EFF57293932DB50DCEEFFA53FB40FA55" author="Fautin" box="[1018,1242,1427,1454]" etAl="et al." firstAuthor="Fautin" page="47" pageId="41" pageNumber="264" pagination="44 - 59" refId="ref118220" refString="FAUTIN D. G., TAN R., YAP W. L. N., TAN S. H., CROWTHER A., GOODWILL R., SANPANICH K. &amp; TAY Y. C. 2015. - Sea anemones (Cnidaria: Actiniaria) of Singapore: shallow-water species known also from the Indian subcontinent. Raffles Bulletin of Zoology Supplement 31: 44 - 59." type="journal article" year="2015">
Fautin 
<emphasis id="B910D370932DB50DC956FA54FBEEFA55" box="[1091,1140,1427,1453]" italics="true" pageId="41" pageNumber="264">et al.</emphasis>
2015: 47
</bibRefCitation>
, fig. 5). The symbiosis between 
<taxonomicName id="4C6474E1932DB50DCEBBFA73FB1FFA35" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[942,1157,1459,1485]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCEBBFA73FB1FFA35" box="[942,1157,1459,1485]" italics="true" pageId="41" pageNumber="264">Dorippoides facchino</emphasis>
</taxonomicName>
and the actinid 
<taxonomicName id="4C6474E1932DB50DC82CFA73FBB1FA16" authority="Stimpson, 1856" authorityName="Stimpson" authorityYear="1856" class="Anthozoa" family="Sagartiidae" genus="Cancrisocia" kingdom="Animalia" order="Actiniaria" pageId="41" pageNumber="264" phylum="Cnidaria" rank="species" species="expansa">
<emphasis id="B910D370932DB50DC82CFA73FCE6FA15" italics="true" pageId="41" pageNumber="264">Cancrisocia expansa</emphasis>
Stimpson, 1856
</taxonomicName>
is very close and obligatory, with the young crab carrying a very small shell and holding with its P4 and P5 dactyli an ovoid to kidney-shaped platform on which the actinian is sitting and will grow. Most platforms consist of an eccentrically positioned bivalve shell around which the anemone had secreted chitinous material as it grows; in some cases, the shell extends the edge of the platform and gives the crab greater coverage (
<bibRefCitation id="EFF57293932DB50DC90FF972FB4DF935" author="Fautin" box="[1050,1239,1714,1741]" etAl="et al." firstAuthor="Fautin" pageId="41" pageNumber="264" pagination="44 - 59" refId="ref118220" refString="FAUTIN D. G., TAN R., YAP W. L. N., TAN S. H., CROWTHER A., GOODWILL R., SANPANICH K. &amp; TAY Y. C. 2015. - Sea anemones (Cnidaria: Actiniaria) of Singapore: shallow-water species known also from the Indian subcontinent. Raffles Bulletin of Zoology Supplement 31: 44 - 59." type="journal article" year="2015">
Fautin 
<emphasis id="B910D370932DB50DC973F973FB00F934" box="[1126,1178,1714,1740]" italics="true" pageId="41" pageNumber="264">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
<paragraph id="8BDB0F62932DB50DCE56F912FA35F8B4" blockId="41.[813,1456,1363,1868]" pageId="41" pageNumber="264">
The carrying behaviour of 
<taxonomicName id="4C6474E1932DB50DC977F913FB4EF914" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[1122,1236,1746,1772]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932DB50DC977F913FB4EF914" box="[1122,1236,1746,1772]" italics="true" pageId="41" pageNumber="264">D. nudipes</emphasis>
</taxonomicName>
, a species closely related to 
<taxonomicName id="4C6474E1932DB50DCE91F933FC61F8F4" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[900,1019,1778,1804]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932DB50DCE91F933FC61F8F4" box="[900,1019,1778,1804]" italics="true" pageId="41" pageNumber="264">D. facchino</emphasis>
</taxonomicName>
, has not been documented (
<bibRefCitation id="EFF57293932DB50DC838F932FB8AF8D4" author="Manning &amp; Holthuis" firstAuthor="Manning" page="364" pageId="41" pageNumber="264" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis 1986: 364</bibRefCitation>
, fig. 1c; Holthuis &amp; Manning 1990: 70; 
<bibRefCitation id="EFF57293932DB50DCE43F8F1FBDFF8B4" author="Davie" box="[854,1093,1841,1868]" etAl="et al." firstAuthor="Davie" page="315" pageId="41" pageNumber="264" pagination="313 - 339" refId="ref115345" refString="DAVIE P. J. F., NG P. K. L., KAENPHET A. &amp; AUNGTONYA C. 2002. - Annotated checklist of Brachyura (Crustacea: Decapoda) principally obtained during the BI ̿ SH ‰ LF survey off western Thailand from 1996 - 1998. Phuket Marine Biological Center Research Bulletin Spec. Publ., 23: 313 - 339." type="journal article" year="2002">
Davie 
<emphasis id="B910D370932DB50DCE8EF8F2FC54F8B3" box="[923,974,1841,1867]" italics="true" pageId="41" pageNumber="264">et al.</emphasis>
2002: 315
</bibRefCitation>
; 
<bibRefCitation id="EFF57293932DB50DC95AF8F1FAAAF8B4" author="Devi &amp; Kumar" box="[1103,1328,1841,1868]" firstAuthor="Devi" pageId="41" pageNumber="264" pagination="625 - 630" refId="ref116809" refString="DEVI S. S. &amp; KUMAR A. B. 2017. - ̿ n two species of crabs (Decapoda, Brachyura): Cycloes marisrubri Galil &amp; Clark, 1996 (Calappidae) and Dorippoides nudipes Manning &amp; Holthuis, 1986 (Dorippidae) from India. Crustaceana 90 (5): 625 - 630. https: // doi. org / 10.1163 / 15685403 - 00003670" type="journal article" year="2017">Devi &amp; Kumar 2017</bibRefCitation>
: fig. 1E, F).
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9932DB50FCE38F8B2FDCAF813" lastPageId="43" lastPageNumber="266" pageId="41" pageNumber="264" type="discussion">
<paragraph id="8BDB0F62932DB50DCE38F8B2FC0BF872" blockId="41.[813,1457,1906,2028]" box="[813,913,1906,1932]" pageId="41" pageNumber="264">
<smallCapsWord id="8D3D99BE932DB50DCE38F8B2FC0BF872" baselines="1925,1925" box="[813,913,1906,1932]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="41" pageNumber="264">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BDB0F62932DB50FCE38F851FC9DFCF7" blockId="41.[813,1457,1906,2028]" lastBlockId="43.[130,777,215,2028]" lastPageId="43" lastPageNumber="266" pageId="41" pageNumber="264">
The species known as 
<taxonomicName id="4C6474E1932DB50DC90AF851FB42F853" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1055,1240,1937,1963]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="264" phylum="Arthropoda" rank="species" species="lanata">
<emphasis id="B910D370932DB50DC90AF851FB42F853" box="[1055,1240,1937,1963]" italics="true" pageId="41" pageNumber="264">Medorippe lanata</emphasis>
</taxonomicName>
in the Adriatic was actually described by James Plancus (Latinisation of Simon Giovanni Bianchi, 1693-1775), who gave it the Italian vernacular name ‘facchino’, which means ‘porter’, to the inhabitants of his hometown, Rimini, a famous city on the Adriatic coast of northern 
<collectingCountry id="F3734FF2932EB50ECC5CF851FEE0F853" box="[329,378,1937,1963]" name="Italy" pageId="42" pageNumber="265">Italy</collectingCountry>
. According to Holthuis &amp; Manning (1990: 52), Plancus only referred to the fact that the carapace of this crab resembles an ugly human face, as supposedly often found among members of the porter profession, and without mentioning the possibility that the crabs can carry objects with their last two pairs of legs. The point is that, in originally establishing and figuring 
<taxonomicName id="4C6474E1932EB50EC9B1F872FADCF833" authorityName="Herbst" authorityYear="1785" box="[1188,1350,1969,1996]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="265" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932EB50EC9B1F872FADCF833" box="[1188,1350,1969,1996]" italics="true" pageId="42" pageNumber="265">Cancer facchino</emphasis>
</taxonomicName>
for material from the East 
<collectingCountry id="F3734FF2932EB50ECEFEF811FBBCF813" box="[1003,1062,2001,2027]" name="India" pageId="42" pageNumber="265">India</collectingCountry>
, Herbst (1785) erroneously included in the description a reference to the description and figure by Plancus (1739, 1760) of the Mediterranean crab now known as 
<taxonomicName id="4C6474E1932FB50FCDB4FED7FEC1FEC9" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[161,347,279,305]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="lanata">
<emphasis id="B910D370932FB50FCDB4FED7FEC1FEC9" box="[161,347,279,305]" italics="true" pageId="43" pageNumber="266">Medorippe lanata</emphasis>
</taxonomicName>
, thus incorrectly assuming that 
<taxonomicName id="4C6474E1932FB50FCFABFED7FF43FEA9" authorityName="Herbst" authorityYear="1785" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCFABFED7FF43FEA9" italics="true" pageId="43" pageNumber="266">Cancer facchino</emphasis>
</taxonomicName>
occurred in the Mediterranean. The specific name 
<taxonomicName id="4C6474E1932FB50FCD93FE97FF40FE89" authorityName="Herbst" authorityYear="1785" box="[134,218,343,369]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCD93FE97FF40FE89" box="[134,218,343,369]" italics="true" pageId="43" pageNumber="266">facchino</emphasis>
</taxonomicName>
is thus the result of the misidentification by Herbst (1785) of the pre-Linnean ‘ 
<taxonomicName id="4C6474E1932FB50FCCB1FEB7FDD4FE69" box="[420,590,375,401]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="hirsutus">Cancer hirsutus</taxonomicName>
personatus maris Superi, vulgo 
<taxonomicName id="4C6474E1932FB50FCC1BFE56FD9FFE48" authority="Ariminensibus" authorityName="Ariminensibus" box="[270,517,406,432]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">Facchino Ariminensibus</taxonomicName>
dictus’ by Plancus (1739; 1760: 36-38, pl. 5, fig. 1) with 
<taxonomicName id="4C6474E1932FB50FCCDBFE76FD38FE28" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[462,674,438,464]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCCDBFE76FD38FE28" box="[462,674,438,464]" italics="true" pageId="43" pageNumber="266">Dorippoides facchino</emphasis>
</taxonomicName>
. Plancus’ specimen thus becomes a 
<typeStatus id="54DFB1C0932FB50FCC8DFE17FE73FE09" box="[408,489,471,497]" pageId="43" pageNumber="266">syntype</typeStatus>
of the Herbst’s 
<taxonomicName id="4C6474E1932FB50FCF87FE17FF26FDE8" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCF87FE17FF26FDE8" italics="true" pageId="43" pageNumber="266">Cancer facchino</emphasis>
</taxonomicName>
. As Herbst’s 
<taxonomicName id="4C6474E1932FB50FCC5BFE37FE6FFDE8" box="[334,501,502,529]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCC5BFE37FE6FFDE8" box="[334,501,502,529]" italics="true" pageId="43" pageNumber="266">Cancer facchino</emphasis>
</taxonomicName>
was a composite species, 
<bibRefCitation id="EFF57293932FB50FCD93FDD6FD9AFDC8" author="Serene &amp; Romimohtarto" box="[134,512,534,561]" firstAuthor="Serene" page="4" pageId="43" pageNumber="266" pagination="1 - 35" refId="ref141809" refString="SERENE R. &amp; ROMIMOHTARTO K. 1969. - ̿ bservations on the species of Dorippe from the Indo-Malayan Region. Marine Research in Indonesia (Penelitian Laut di Indonesia) 9: 1 - 35." type="journal article" year="1969">Serène &amp; Romimohtarto (1969: 4)</bibRefCitation>
selected as the 
<typeStatus id="54DFB1C0932FB50FCFBDFDD6FC93FDC8" box="[680,777,534,560]" pageId="43" pageNumber="266">lectotype</typeStatus>
the specimen represented by Herbst (1785: pl. 11, fig. 68) (reproduced by Holthuis &amp; Manning 1990: fig. 23), thereby fixing the identity of Herbst’s species and making 
<taxonomicName id="4C6474E1932FB50FCFA5FDB6FC9CFD68" box="[688,774,630,656]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCFA5FDB6FC9CFD68" box="[688,774,630,656]" italics="true" pageId="43" pageNumber="266">facchino</emphasis>
</taxonomicName>
its valid name. Through this 
<typeStatus id="54DFB1C0932FB50FCCA9FD56FD87FD48" box="[444,541,662,688]" pageId="43" pageNumber="266">lectotype</typeStatus>
selection, the specific name 
<taxonomicName id="4C6474E1932FB50FCDD0FD75FE83FD37" box="[197,281,693,719]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCDD0FD75FE83FD37" box="[197,281,693,719]" italics="true" pageId="43" pageNumber="266">facchino</emphasis>
</taxonomicName>
is now definitively linked to the Indo-West Pacific species of 
<taxonomicName id="4C6474E1932FB50FCC36FD15FE04FD17" authorityName="Serene &amp; Romimohtarto" authorityYear="1969" box="[291,414,725,751]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="B910D370932FB50FCC36FD15FE04FD17" box="[291,414,725,751]" italics="true" pageId="43" pageNumber="266">Dorippoides</emphasis>
</taxonomicName>
rather than to the Mediterranean 
<taxonomicName id="4C6474E1932FB50FCD90FD35FEADFCF7" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[133,311,757,783]" class="Malacostraca" family="Dorippidae" genus="Medorippe" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="lanata">
<emphasis id="B910D370932FB50FCD90FD35FEADFCF7" box="[133,311,757,783]" italics="true" pageId="43" pageNumber="266">Medorippe lanata</emphasis>
</taxonomicName>
(Holthuis &amp; Manning 1990: 51-52, 63-64).
</paragraph>
<caption id="DF1B5FEA932EB50ECD91F95AFD92F8FF" ID-DOI="http://doi.org/10.5281/zenodo.8071299" ID-Zenodo-Dep="8071299" httpUri="https://zenodo.org/record/8071299/files/figure.png" pageId="42" pageNumber="265" startId="42.[132,143,1690,1708]" targetBox="[137,1443,215,1647]" targetPageId="42" targetType="figure">
<paragraph id="8BDB0F62932EB50ECD91F95AFD92F8FF" blockId="42.[132,1457,1690,1799]" pageId="42" pageNumber="265">
<smallCapsWord id="8D3D99BE932EB50ECD91F95AFF07F953" baselines="1703,1704" box="[132,157,1690,1708]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="42" pageNumber="265">FIG</smallCapsWord>
. 18. — 
<taxonomicName id="4C6474E1932EB50ECDF4F95AFECCF954" authority="Guinot, 2023" authorityName="Guinot" authorityYear="2023" box="[225,342,1690,1708]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="265" phylum="Arthropoda" rank="subFamily" status="n. subfam." subFamily="Dorippoidinae">Dorippoidinae</taxonomicName>
<taxonomicNameLabel id="A2236E0B932EB50ECC48F95AFE26F954" box="[349,444,1690,1708]" pageId="42" pageNumber="265" rank="subFamily">n. subfam.</taxonomicNameLabel>
: 
<emphasis id="B910D370932EB50ECCD6F95AFE7FF954" bold="true" box="[451,485,1690,1708]" pageId="42" pageNumber="265">A -C</emphasis>
, 
<taxonomicName id="4C6474E1932EB50ECCE4F95AFCBCF954" authority="(Herbst, 1785)" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[497,806,1690,1708]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="265" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932EB50ECCE4F95AFD3EF954" box="[497,676,1690,1708]" italics="true" pageId="42" pageNumber="265">Dorippoides facchino</emphasis>
(Herbst, 1785)
</taxonomicName>
, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: 
<emphasis id="B910D370932EB50ECD91F971FF0BF93B" bold="true" box="[132,145,1713,1731]" pageId="42" pageNumber="265">A</emphasis>
, thoracic sternum with pleon; 
<emphasis id="B910D370932EB50ECC80F971FE38F93B" bold="true" box="[405,418,1713,1731]" pageId="42" pageNumber="265">B</emphasis>
, without pleon, vulvae. 
<emphasis id="B910D370932EB50ECF7BF970FDE6F93A" bold="true" box="[622,636,1712,1730]" pageId="42" pageNumber="265">C</emphasis>
, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802). 
<emphasis id="B910D370932EB50ECC3DF907FED3F921" bold="true" box="[296,329,1735,1753]" pageId="42" pageNumber="265">D -F</emphasis>
, 
<taxonomicName id="4C6474E1932EB50ECC46F907FD42F921" authority="Manning &amp; Holthuis 1986" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[339,728,1735,1753]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="265" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932EB50ECC46F907FE65F921" box="[339,511,1735,1753]" italics="true" pageId="42" pageNumber="265">Dorippoides nudipes</emphasis>
<bibRefCitation id="EFF57293932EB50ECF11F907FD42F921" author="Manning &amp; Holthuis" box="[516,728,1735,1753]" firstAuthor="Manning" pageId="42" pageNumber="265" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis 1986</bibRefCitation>
</taxonomicName>
: 
<emphasis id="B910D370932EB50ECFCBF907FD76F921" bold="true" box="[734,748,1735,1753]" pageId="42" pageNumber="265">D</emphasis>
, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; 
<emphasis id="B910D370932EB50EC82BF907FAD0F921" bold="true" box="[1342,1354,1735,1753]" pageId="42" pageNumber="265">E</emphasis>
, 
<emphasis id="B910D370932EB50EC841F907FAC5F921" bold="true" box="[1364,1375,1735,1753]" pageId="42" pageNumber="265">F</emphasis>
, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: 
<emphasis id="B910D370932EB50ECFC1F91EFD7AF908" bold="true" box="[724,736,1758,1776]" pageId="42" pageNumber="265">E</emphasis>
, thoracic sternum with pleon; 
<emphasis id="B910D370932EB50ECEC8F91EFC72F908" bold="true" box="[989,1000,1758,1776]" pageId="42" pageNumber="265">F</emphasis>
, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on 
<emphasis id="B910D370932EB50ECCCBF935FE71F8FF" bold="true" box="[478,491,1781,1799]" pageId="42" pageNumber="265">A</emphasis>
, 
<emphasis id="B910D370932EB50ECCE0F935FD98F8FF" bold="true" box="[501,514,1781,1799]" pageId="42" pageNumber="265">B</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BDB0F62932FB50FCD8CFCD6FDC7FCA8" blockId="43.[130,777,215,2028]" pageId="43" pageNumber="266">
<taxonomicName id="4C6474E1932FB50FCD8CFCD6FE42FCD7" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[153,472,789,815]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="astuta">
<emphasis id="B910D370932FB50FCD8CFCD6FEB4FCD7" box="[153,302,790,815]" italics="true" pageId="43" pageNumber="266">Dorippe astuta</emphasis>
<bibRefCitation id="EFF57293932FB50FCC26FCD5FE42FCD7" author="Fabricius" box="[307,472,789,815]" firstAuthor="Fabricius" pageId="43" pageNumber="266" refId="ref118022" refString="FABRICIUS J. C. 1798. - Supplementum Entomologiae Systematicae. Hafniae, 572 p. https: // www. biodiversitylibrary. org / bibliography / 123559" type="book" year="1798">Fabricius, 1798</bibRefCitation>
</taxonomicName>
, is actually a junior synonym of 
<taxonomicName id="4C6474E1932FB50FCD8AFCF5FDC3FCA8" authority="(Ng et al. 2008: 59)" baseAuthorityName="Ng" baseAuthorityPageNumber="59" baseAuthorityYear="2008" box="[159,601,821,848]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCD8AFCF5FEEDFCB7" box="[159,375,821,847]" italics="true" pageId="43" pageNumber="266">Dorippoides facchino</emphasis>
(
<bibRefCitation id="EFF57293932FB50FCC92FCF6FDD5FCA8" author="NG P. K. L. &amp; GUINOT D. &amp; DAVIE P. J. F." box="[391,591,821,848]" pageId="43" pageNumber="266" pagination="1 - 286" refId="ref135161" refString="NG P. K. L., GUINOT D. &amp; DAVIE P. J. F. 2008 a. - Systema Brachyurorum: Part 1. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, S] upplement 17: 1 - 286. [updates at http: // rmbr. nus. edu. sg / research / cotw / supplement 17 php]" type="journal article" year="2008">
Ng 
<emphasis id="B910D370932FB50FCCA5FCF6FE7EFCB7" box="[432,484,821,847]" italics="true" pageId="43" pageNumber="266">et al.</emphasis>
2008: 59
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB0F62932FB50FCD8EFC95FDE0F9F5" blockId="43.[130,777,215,2028]" pageId="43" pageNumber="266">
<taxonomicName id="4C6474E1932FB50FCD8EFC95FEFEFC97" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[155,356,853,879]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCD8EFC95FEFEFC97" box="[155,356,853,879]" italics="true" pageId="43" pageNumber="266">Dorippoides nudipes</emphasis>
</taxonomicName>
was established by 
<bibRefCitation id="EFF57293932FB50FCF3DFC95FF66FC77" author="Manning &amp; Holthuis" firstAuthor="Manning" page="364" pageId="43" pageNumber="266" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis (1986: 364</bibRefCitation>
, fig. 1c) on a single male specimen, the 
<typeStatus id="54DFB1C0932FB50FCFBCFCB5FC9FFC77" box="[681,773,885,911]" pageId="43" pageNumber="266">holotype</typeStatus>
17.0 × 19.0 mm, from Massawa, 
<collectingCountry id="F3734FF2932FB50FCCCDFC55FDAEFC57" box="[472,564,917,943]" name="Ethiopia" pageId="43" pageNumber="266">Ethiopia</collectingCountry>
, Red Sea, and on the basis of the more granular carapace and the dactyli of P2 and P3 broadest in the distal fourth rather than at midlength, in contrast to the only other species of the genus, 
<taxonomicName id="4C6474E1932FB50FCF41FC35FD59FBF6" box="[596,707,1012,1038]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCF41FC35FD59FBF6" box="[596,707,1012,1038]" italics="true" pageId="43" pageNumber="266">D. facchino</emphasis>
</taxonomicName>
. Later, Holthuis &amp; Manning (1990: 66-68) designated as 
<typeStatus id="54DFB1C0932FB50FCFB6FBD5FC9FFBD7" box="[675,773,1045,1071]" pageId="43" pageNumber="266">paratypes</typeStatus>
several specimens from the Red Sea and various samples from the Gulf of Aden, Gulf of 
<collectingCountry id="F3734FF2932FB50FCC9FFB94FE4AFB96" box="[394,464,1108,1134]" name="Oman" pageId="43" pageNumber="266">Oman</collectingCountry>
, Persian Gulf and 
<collectingCountry id="F3734FF2932FB50FCF9DFB94FC99FB96" box="[648,771,1108,1134]" name="Madagascar" pageId="43" pageNumber="266">Madagascar</collectingCountry>
, deposited in various institutions. This was an unjustified act, and these 
<typeStatus id="54DFB1C0932FB50FCDFEFB54FED7FB56" box="[235,333,1172,1198]" pageId="43" pageNumber="266">paratypes</typeStatus>
are invalid (see above, 
<emphasis id="B910D370932FB50FCF23FB54FEE6FB36" italics="true" pageId="43" pageNumber="266">
Subsequent erroneous designation of 
<typeStatus id="54DFB1C0932FB50FCC1FFB75FEF8FB36" box="[266,354,1205,1230]" pageId="43" pageNumber="266">paratypes</typeStatus>
by
</emphasis>
Holthuis &amp; Manning 1990). Although Holthuis &amp; Manning (1990: 49 key, 69, 70, fig. 26) have listed the characters distinguishing 
<taxonomicName id="4C6474E1932FB50FCCA7FB34FD85FAF5" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[434,543,1267,1293]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCCA7FB34FD85FAF5" box="[434,543,1267,1293]" italics="true" pageId="43" pageNumber="266">D. nudipes</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C6474E1932FB50FCF4AFB34FD49FAF5" box="[607,723,1267,1293]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCF4AFB34FD49FAF5" box="[607,723,1267,1293]" italics="true" pageId="43" pageNumber="266">D. facchino</emphasis>
</taxonomicName>
, it is useful to restate them here. In 
<taxonomicName id="4C6474E1932FB50FCCAAFAD4FDB6FAD5" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[447,556,1299,1325]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCCAAFAD4FDB6FAD5" box="[447,556,1299,1325]" italics="true" pageId="43" pageNumber="266">D. nudipes</emphasis>
</taxonomicName>
, the general granulation is more marked, as for example on the anterolateral margin behind the outer orbital tooth; the fingers of chelae are shorter in young males as well in males with inflated palm; the P2 and P3 are cylindrical, with longer and slender articles with naked margins, and with paddle-like dactyli (in 
<taxonomicName id="4C6474E1932FB50FCF33FA74FD0DFA35" box="[550,663,1459,1485]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCF33FA74FD0DFA35" box="[550,663,1459,1485]" italics="true" pageId="43" pageNumber="266">D. facchino</emphasis>
</taxonomicName>
merus and propodus stout and wider, with a fringe of setae on posterior border in adult males, dactylus tapering distally.
</paragraph>
<paragraph id="8BDB0F62932FB50FCD8EF9D2FD8BF853" blockId="43.[130,777,215,2028]" pageId="43" pageNumber="266">
When 
<bibRefCitation id="EFF57293932FB50FCDF3F9D2FE3CF9D4" author="Chen" box="[230,422,1554,1581]" firstAuthor="Chen" page="678" pageId="43" pageNumber="266" pagination="677 - 693" refId="ref112869" refString="CHEN H. 1988. - Dorippidae (Crustacea Decapoda Brachyura) collected in Madagascar waters. Bulletin du Museum national d'Histoire naturelle 4 e serie 9 A (3) [1987]: 677 - 693." type="journal article" year="1988">Chen (1988: 678</bibRefCitation>
, fig. 1, pl. 1B, C) assigned some specimens from 
<collectingCountry id="F3734FF2932FB50FCC2EF9F2FE21F9B4" box="[315,443,1586,1612]" name="Madagascar" pageId="43" pageNumber="266">Madagascar</collectingCountry>
to 
<taxonomicName id="4C6474E1932FB50FCCF7F9F3FDCFF9B4" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[482,597,1586,1612]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCCF7F9F3FDCFF9B4" box="[482,597,1586,1612]" italics="true" pageId="43" pageNumber="266">D. nudipes</emphasis>
</taxonomicName>
, she found that the main differences between the two were related to the length and setation of P2 and P3. According to 
<bibRefCitation id="EFF57293932FB50FCF90F9B2FEAFF954" author="Devi &amp; Kumar" firstAuthor="Devi" page="626" pageId="43" pageNumber="266" pagination="625 - 630" refId="ref116809" refString="DEVI S. S. &amp; KUMAR A. B. 2017. - ̿ n two species of crabs (Decapoda, Brachyura): Cycloes marisrubri Galil &amp; Clark, 1996 (Calappidae) and Dorippoides nudipes Manning &amp; Holthuis, 1986 (Dorippidae) from India. Crustaceana 90 (5): 625 - 630. https: // doi. org / 10.1163 / 15685403 - 00003670" type="journal article" year="2017">Devi &amp; Kumar (2017: 626</bibRefCitation>
, fig. 1E, F), their specimen from southern 
<collectingCountry id="F3734FF2932FB50FCD91F972FF27F934" box="[132,189,1714,1740]" name="India" pageId="43" pageNumber="266">India</collectingCountry>
fits well with the characters previously mentioned for 
<taxonomicName id="4C6474E1932FB50FCD91F913FF6FF914" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[132,245,1746,1772]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCD91F913FF6FF914" box="[132,245,1746,1772]" italics="true" pageId="43" pageNumber="266">D. nudipes</emphasis>
</taxonomicName>
, and some of the older records of 
<taxonomicName id="4C6474E1932FB50FCF48F913FD49F914" box="[605,723,1746,1772]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCF48F913FD49F914" box="[605,723,1746,1772]" italics="true" pageId="43" pageNumber="266">D. facchino</emphasis>
</taxonomicName>
may well refer to 
<taxonomicName id="4C6474E1932FB50FCC16F933FEEAF8F4" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[259,368,1778,1804]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCC16F933FEEAF8F4" box="[259,368,1778,1804]" italics="true" pageId="43" pageNumber="266">D. nudipes</emphasis>
</taxonomicName>
due to their close similarity. The sexual dimorphism of the setation of P2 and P3 meri is marked in 
<taxonomicName id="4C6474E1932FB50FCD91F8F2FF66F8B3" box="[132,252,1841,1867]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="facchino">
<emphasis id="B910D370932FB50FCD91F8F2FF66F8B3" box="[132,252,1841,1867]" italics="true" pageId="43" pageNumber="266">D. facchino</emphasis>
</taxonomicName>
, with thick fringe of setae on the posterior margins in males instead of margins naked in females (
<figureCitation id="135F13E7932FB50FCFB7F891FF0DF874" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="43" pageNumber="266">Fig. 16A, B</figureCitation>
, respectively), but not marked in 
<taxonomicName id="4C6474E1932FB50FCF05F8B2FD19F873" authorityName=": Manning &amp; Holthuis" authorityYear="1986" box="[528,643,1905,1931]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B910D370932FB50FCF05F8B2FD19F873" box="[528,643,1905,1931]" italics="true" pageId="43" pageNumber="266">D. nudipes</emphasis>
</taxonomicName>
with naked margins in both sexes (
<figureCitation id="135F13E7932FB50FCC69F851FD99F853" box="[380,515,1937,1963]" captionStart="FIG" captionStartId="40.[133,144,1271,1289]" captionTargetBox="[135,1455,216,1229]" captionTargetId="figure-454@40.[135,1462,216,1233]" captionTargetPageId="40" captionText="FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning &amp; Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885." figureDoi="http://doi.org/10.5281/zenodo.8071293" httpUri="https://zenodo.org/record/8071293/files/figure.png" pageId="43" pageNumber="266">Fig. 16C, D</figureCitation>
).
</paragraph>
<paragraph id="8BDB0F62932FB50FCD8EF871FDCAF813" blockId="43.[130,777,215,2028]" pageId="43" pageNumber="266">
The 
<taxonomicName id="4C6474E1932FB50FCDDDF871FEFEF833" authority="Guinot, 2023" authorityName="Guinot" authorityYear="2023" box="[200,356,1969,1995]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="subFamily" status="n. subfam." subFamily="Dorippoidinae">Dorippoidinae</taxonomicName>
<taxonomicNameLabel id="A2236E0B932FB50FCC7CF872FE42F833" box="[361,472,1969,1996]" pageId="43" pageNumber="266" rank="subFamily">n. subfam.</taxonomicNameLabel>
is monotypic, with the genus 
<taxonomicName id="4C6474E1932FB50FCD97F811FF64F813" authorityName="Serene &amp; Romimohtarto" authorityYear="1969" box="[130,254,2001,2027]" class="Malacostraca" family="Dorippidae" genus="Dorippoides" kingdom="Animalia" order="Decapoda" pageId="43" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="B910D370932FB50FCD97F811FF64F813" box="[130,254,2001,2027]" italics="true" pageId="43" pageNumber="266">Dorippoides</emphasis>
</taxonomicName>
, only known from two species.
</paragraph>
</subSubSection>
</treatment>
</document>