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<document id="ACCD8B3712DEF56D28E28AAA6B4EC95C" ID-DOI="10.11646/zootaxa.3872.2.1" ID-GBIF-Dataset="e9b8d555-d661-4f1a-86e8-f2a0acc38798" ID-ISSN="1175-5326" ID-Zenodo-Dep="253224" ID-ZooBank="F9F81CC8-E033-46B7-B73B-9FB777DF4116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460926510372" checkinUser="plazi" docAuthor="Ponder, Winston F., Fukuda, Hiroshi &amp; Hallan, Anders" docDate="2014" docId="03CDAD6557723C29FF05F8BCBEADAECA" docLanguage="en" docName="zt03872p153.pdf" docOrigin="Zootaxa 3872 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clenchiella victoriae Abbott 1948" docType="treatment" docVersion="9" lastPageNumber="114" masterDocId="FFF4D51D57783C24FF92FFB2BB3EA911" masterDocTitle="A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)" masterLastPageNumber="153" masterPageNumber="101" pageNumber="111" updateTime="1698362634141" updateUser="plazi">
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<mods:titleInfo id="CFEB1C36430F08F4368C14D4240C3C8C">
<mods:title id="2B96B4A4073D36F9DADE96A46A8288AC">A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)</mods:title>
</mods:titleInfo>
<mods:name id="6ADC326F242A2DB44DC1F95FD7B4100F" type="personal">
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<mods:namePart id="4BC6713061F0FF7292BE80938AB22AF1">Ponder, Winston F.</mods:namePart>
</mods:name>
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<mods:namePart id="5E337AFE5DAF050B44BA2D21E5EA75AF">Fukuda, Hiroshi</mods:namePart>
</mods:name>
<mods:name id="AE4CB77343A0121CC3B54D6ACD3E9A5D" type="personal">
<mods:role id="38F66799F0170763F30947B4BF908137">
<mods:roleTerm id="147C3FAFD7D03BFCA85A0602FC9B0110">Author</mods:roleTerm>
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<mods:namePart id="DF19F4617CF2350822601329F0F1EDA1">Hallan, Anders</mods:namePart>
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<mods:typeOfResource id="E8245E6080DDF2B774A0D9CF5D81C4C6">text</mods:typeOfResource>
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<mods:title id="8F4159404B6A8523DF225196CFE15514">Zootaxa</mods:title>
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<mods:part id="A9805F1B13CAF1573E300C52317BDECC">
<mods:date id="EEBCEE018E618FF946960C8E71195DC8">2014</mods:date>
<mods:detail id="22944B55A9143AC0AEDE3FA2E96E904E" type="volume">
<mods:number id="D5F7D289518AA14EF7F087FEB85233C6">3872</mods:number>
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<mods:start id="A8082AA19515A279CFD2602DB087F7D0">101</mods:start>
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<mods:identifier id="E506241D2DFA1AEA2839F2239EDDBDB5" type="DOI">10.11646/zootaxa.3872.2.1</mods:identifier>
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<treatment id="03CDAD6557723C29FF05F8BCBEADAECA" ID-DOI="http://doi.org/10.5281/zenodo.5631009" ID-GBIF-Taxon="119589585" ID-Zenodo-Dep="5631009" LSID="urn:lsid:plazi:treatment:03CDAD6557723C29FF05F8BCBEADAECA" httpUri="http://treatment.plazi.org/id/03CDAD6557723C29FF05F8BCBEADAECA" lastPageId="13" lastPageNumber="114" pageId="10" pageNumber="111">
<subSubSection id="C37E4FF857723C2EFF05F8BCBA15AE5B" pageId="10" pageNumber="111" type="nomenclature">
<paragraph id="8BDB1C7357723C2EFF05F8BCB907AE39" blockId="10.[151,569,1806,1866]" box="[151,569,1806,1832]" pageId="10" pageNumber="111">
<heading id="D093AB1F57723C2EFF05F8BCB907AE39" bold="true" box="[151,569,1806,1832]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
<taxonomicName id="4C6467F057723C2EFF05F8BCB907AE39" authority="Abbott, 1948" authorityName="Abbott" authorityYear="1948" box="[151,569,1806,1832]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="victoriae">
<emphasis id="B910C06157723C2EFF05F8BCB907AE39" bold="true" box="[151,569,1806,1832]" pageId="10" pageNumber="111">
<emphasis id="B910C06157723C2EFF05F8BCBAAFAE39" bold="true" box="[151,401,1806,1832]" italics="true" pageId="10" pageNumber="111">Clenchiella victoriae</emphasis>
<bibRefCitation id="EFF5618257723C2EFE0AF8BCB907AE39" author="Abbott" box="[408,569,1806,1832]" pageId="10" pageNumber="111" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott, 1948</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDB1C7357723C2EFF05F883BA15AE5B" blockId="10.[151,569,1806,1866]" box="[151,299,1841,1866]" pageId="10" pageNumber="111">
<figureCitation id="135F00F657723C2EFF05F883BA3EAE5B" box="[151,256,1841,1866]" captionStart="FIGURE 1" captionStartId="11.[151,250,1702,1724]" captionTargetBox="[163,1428,289,1686]" captionTargetId="figure@11.[163,1428,289,1687]" captionTargetPageId="11" captionText="FIGURE 1. Holotypes of species of Clenchiella with distinct keels (from left towards right: dorsal, lateral and ventral view respectively). A C, Clenchiella victoriae (USNM 488534; shell diameter 2.1 mm); D F, Clenchiella bicingulata n. sp. (AMS C. 462959; shell diameter 1.97 mm); G I, Clenchiella varicosa n. sp. (AMS C. 460738; shell diameter 1.96 mm); J L, Clenchiella iriomotensis n. sp. (AMS C. 445413; shell diameter 1.94 mm)." httpUri="https://zenodo.org/record/253225/files/figure.png" pageId="10" pageNumber="111">Figures 1</figureCitation>
,
<figureCitation id="135F00F657723C2EFE9CF880BA15AE5B" box="[270,299,1842,1866]" captionStart="FIGURE 14" captionStartId="24.[151,250,1920,1942]" captionTargetBox="[265,1335,833,1900]" captionTargetId="figure@24.[265,1336,833,1900]" captionTargetPageId="24" captionText="FIGURE 14. Type localities of Clenchiella victoriae, Cl. bicingulata, Cl. varicosa, Cl. iriomotensis, and all recorded localities of Cl. minutissima (type locality Vietnam)." httpUri="https://zenodo.org/record/253238/files/figure.png" pageId="10" pageNumber="111">14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857723C2FFF05F8CBBA3FA9E6" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111" type="reference_group">
<paragraph id="8BDB1C7357723C2EFF05F8CBB989AE9E" blockId="10.[151,695,1913,1935]" box="[151,695,1913,1935]" pageId="10" pageNumber="111">
<treatmentCitationGroup id="AB743B5D57723C2EFF05F8CBB989AE9E" box="[151,695,1913,1935]" pageId="10" pageNumber="111">
<taxonomicName id="4C6467F057723C2EFF05F8CBB921AE9E" authority="Abbott, 1948: 76" authorityName="Abbott" authorityPageNumber="76" authorityYear="1948" box="[151,543,1913,1935]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="victoriae">
<emphasis id="B910C06157723C2EFF05F8CBBA52AE9E" box="[151,364,1913,1935]" italics="true" pageId="10" pageNumber="111">Clenchiella victoriae</emphasis>
<treatmentCitation id="0AC53A6257723C2EFEE0F8CBB921AE9E" author="Abbott" box="[370,543,1913,1935]" page="76" pageId="10" pageNumber="111" year="1948">
<bibRefCitation id="EFF5618257723C2EFEE0F8CBBAC5AE9E" author="Abbott" box="[370,507,1913,1935]" pageId="10" pageNumber="111" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott, 1948</bibRefCitation>
: 76
</treatmentCitation>
</taxonomicName>
78, pl. 5, 17.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB1C7357723C2FFF05F804BA3FA9E6" blockId="10.[151,1436,1974,2035]" lastBlockId="11.[151,1436,151,247]" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111">
<emphasis id="B910C06157723C2EFF05F804BA8CAEDE" bold="true" box="[151,434,1974,1999]" pageId="10" pageNumber="111">
<typeStatus id="54DFA2D157723C2EFF05F804BBE2AEDE" box="[151,220,1974,1999]" pageId="10" pageNumber="111">Types</typeStatus>
and
<typeStatus id="54DFA2D157723C2EFE88F804BA73AEDE" box="[282,333,1974,1999]" pageId="10" pageNumber="111">type</typeStatus>
locality.
</emphasis>
<typeStatus id="54DFA2D157723C2EFE28F804B918AEDE" box="[442,550,1974,1999]" pageId="10" pageNumber="111" type="holotype">Holotype</typeStatus>
: San Joaquin River,
<quantity id="4C9CB19657723C2EFC9DF805B85CAEDE" box="[783,866,1974,1999]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="10" pageNumber="111" unit="mi" value="2.0">2 miles</quantity>
(
<quantity id="4C9CB19657723C2EFCE0F805B8FAAEDE" box="[882,964,1974,1999]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="111" unit="km" value="3.2">3.2 km</quantity>
) inland, north of Tanauan, eastern side of Leyte Island,
<collectingCountry id="F3735CE357723C2EFEA2F868B958AEE2" box="[304,614,2010,2035]" name="Philippines" pageId="10" pageNumber="111">Republic of the Philippines</collectingCountry>
. Brackish water, living in black ooze under rotting nipa palm fronds, on the banks of a sluggish river,
<date id="FFDA3AB357733C2FFE60FF2AB9AEA9A1" box="[498,656,151,176]" pageId="11" pageNumber="112" value="1945-06-29">29 June 1945</date>
. Coll. R.T. Abbott (USNM 488534).
<typeStatus id="54DFA2D157733C2FFBD1FF2ABF8AA9A1" box="[1091,1204,152,176]" pageId="11" pageNumber="112" type="paratype">Paratypes</typeStatus>
: Same data (USNM 488535; ANSP 183548). Other
<typeStatus id="54DFA2D157733C2FFD82FF0FB945A9C4" box="[528,635,189,213]" pageId="11" pageNumber="112" type="paratype">paratypes</typeStatus>
were deposited in MCZ (Harvard), MCZ (Michigan) and Carnegie Museum.
</paragraph>
</subSubSection>
<caption id="DF1B4CFB57733C2FFF05F914B8B7AE09" httpUri="https://zenodo.org/record/253225/files/figure.png" pageId="11" pageNumber="112" targetBox="[163,1428,289,1686]" targetPageId="11">
<paragraph id="8BDB1C7357733C2FFF05F914B8B7AE09" blockId="11.[151,1436,1702,1816]" pageId="11" pageNumber="112">
<emphasis id="B910C06157733C2FFF05F914BA2BAFAD" bold="true" box="[151,277,1702,1724]" pageId="11" pageNumber="112">FIGURE 1.</emphasis>
Holotypes of species of
<taxonomicName id="4C6467F057733C2FFD8AF914B9B3AFAD" box="[536,653,1702,1724]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="112" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157733C2FFD8AF914B9B3AFAD" box="[536,653,1702,1724]" italics="true" pageId="11" pageNumber="112">Clenchiella</emphasis>
</taxonomicName>
with distinct keels (from left towards right: dorsal, lateral and ventral view respectively).
<emphasis id="B910C06157733C2FFEB5F976BA69AFCB" bold="true" box="[295,343,1732,1754]" pageId="11" pageNumber="112">AC</emphasis>
,
<taxonomicName id="4C6467F057733C2FFEF7F976B904AFCA" box="[357,570,1732,1755]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="victoriae">
<emphasis id="B910C06157733C2FFEF7F976B904AFCA" box="[357,570,1732,1755]" italics="true" pageId="11" pageNumber="112">Clenchiella victoriae</emphasis>
</taxonomicName>
(USNM 488534; shell diameter 2.1 mm);
<emphasis id="B910C06157733C2FFC7FF976BF27AFCB" bold="true" box="[1005,1049,1732,1754]" pageId="11" pageNumber="112">DF</emphasis>
,
<taxonomicName id="4C6467F057733C2FFBB5F976BE2BAFCB" box="[1063,1301,1732,1754]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="bicingulata" status="sp. nov.">
<emphasis id="B910C06157733C2FFBB5F976BE2BAFCB" box="[1063,1301,1732,1754]" italics="true" pageId="11" pageNumber="112">Clenchiella bicingulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57733C2FFA8EF976BE68AFCB" box="[1308,1366,1732,1754]" pageId="11" pageNumber="112" rank="species">
<emphasis id="B910C06157733C2FFA8EF976BE68AFCB" bold="true" box="[1308,1366,1732,1754]" pageId="11" pageNumber="112">n. sp.</emphasis>
</taxonomicNameLabel>
(AMS C.462959; shell diameter 1.97 mm);
<emphasis id="B910C06157733C2FFDB5F951B96FAFE8" bold="true" box="[551,593,1763,1785]" pageId="11" pageNumber="112">GI</emphasis>
,
<taxonomicName id="4C6467F057733C2FFDF1F951B804AFE8" box="[611,826,1763,1785]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="varicosa" status="sp. nov.">
<emphasis id="B910C06157733C2FFDF1F951B804AFE8" box="[611,826,1763,1785]" italics="true" pageId="11" pageNumber="112">Clenchiella varicosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57733C2FFCD4F951B8BBAFE8" box="[838,901,1763,1785]" pageId="11" pageNumber="112" rank="species">
<emphasis id="B910C06157733C2FFCD4F951B8BBAFE8" bold="true" box="[838,901,1763,1785]" pageId="11" pageNumber="112">n. sp.</emphasis>
</taxonomicNameLabel>
(AMS C.460738; shell diameter 1.96 mm);
<emphasis id="B910C06157733C2FFAF9F951BEABAFE8" bold="true" box="[1387,1429,1763,1785]" pageId="11" pageNumber="112">JL</emphasis>
,
<taxonomicName id="4C6467F057733C2FFF05F8B0BAB1AE09" box="[151,399,1794,1816]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="iriomotensis" status="sp. nov.">
<emphasis id="B910C06157733C2FFF05F8B0BAB1AE09" box="[151,399,1794,1816]" italics="true" pageId="11" pageNumber="112">Clenchiella iriomotensis</emphasis>
</taxonomicName>
<emphasis id="B910C06157733C2FFE07F8B0BAF0AE09" bold="true" box="[405,462,1794,1816]" pageId="11" pageNumber="112">
<taxonomicNameLabel id="A2237D1A57733C2FFE07F8B0BAF0AE09" box="[405,462,1794,1816]" pageId="11" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(AMS C.445413; shell diameter 1.94 mm).
</paragraph>
</caption>
<subSubSection id="C37E4FF857733C2FFF55F8F6B8E3AEB4" pageId="11" pageNumber="112" type="materials_examined">
<paragraph id="8BDB1C7357733C2FFF55F8F6B8E3AEB4" blockId="11.[151,1436,1860,2029]" pageId="11" pageNumber="112">
<emphasis id="B910C06157733C2FFF55F8F6BA8AAE4C" bold="true" box="[199,436,1860,1885]" pageId="11" pageNumber="112">Material examined.</emphasis>
<typeStatus id="54DFA2D157733C2FFE2DF8F7B90EAE4C" box="[447,560,1861,1885]" pageId="11" pageNumber="112" type="paratype">Paratypes</typeStatus>
: Estuary SE of Abuyog, eastern side of Leyte Island,
<collectingCountry id="F3735CE357733C2FFB3CF8F6BE11AE4C" box="[1198,1327,1860,1885]" name="Philippines" pageId="11" pageNumber="112">Philippines</collectingCountry>
,
<date id="FFDA3AB357733C2FFAACF8F7BBECAE90" pageId="11" pageNumber="112" value="1945-08-14">14 Aug. 1945</date>
. Coll. R. T. Abbott (
<collectionCode id="ED7584B657733C2FFE56F8DBB929AE90" LSID="urn:lsid:biocol.org:col:34871" box="[452,535,1897,1921]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="112">USNM</collectionCode>
488536).
<typeStatus id="54DFA2D157733C2FFD1DF8DBB83EAE90" box="[655,768,1897,1921]" pageId="11" pageNumber="112" type="paratype">Paratypes</typeStatus>
: San Joaquin estuary, 3 kms N of Tanauan, eastern Leyte Island,
<collectingCountry id="F3735CE357733C2FFF7BF83EBA59AEB4" box="[233,359,1932,1957]" name="Philippines" pageId="11" pageNumber="112">Philippines</collectingCountry>
(Museum of Zoology, University of Michigan 169007).
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857733C2FFF55F802BAD5AEFC" pageId="11" pageNumber="112" type="distribution">
<paragraph id="8BDB1C7357733C2FFF55F802BAD5AEFC" blockId="11.[151,1436,1860,2029]" pageId="11" pageNumber="112">
<emphasis id="B910C06157733C2FFF55F802BA5EAED8" bold="true" box="[199,352,1968,1993]" pageId="11" pageNumber="112">Distribution.</emphasis>
Known only from brackish swamps on Leyte Island,
<collectingCountry id="F3735CE357733C2FFC44F802BF69AED8" box="[982,1111,1968,1993]" name="Philippines" pageId="11" pageNumber="112">Philippines</collectingCountry>
, but probably has a broader distribution in the
<collectingCountry id="F3735CE357733C2FFEF7F866BAD9AEFC" box="[357,487,2004,2029]" name="Philippines" pageId="11" pageNumber="112">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857743C29FF55FF25B848A874" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113" type="description">
<paragraph id="8BDB1C7357743C28FF55FF25BF9FA9A1" blockId="12.[151,1437,151,896]" box="[199,1185,151,176]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF55FF25BA67A9A1" bold="true" box="[199,345,151,176]" pageId="12" pageNumber="113">Description.</emphasis>
(Based on
<bibRefCitation id="EFF5618257743C28FE45FF25B95AA9A1" author="Abbott" box="[471,612,151,176]" pageId="12" pageNumber="113" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott 1948</bibRefCitation>
and examination of shells using light microscope).
</paragraph>
<paragraph id="8BDB1C7357743C28FF55FF0EBA59AB09" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF55FF0EBBC1A9C4" box="[199,255,188,213]" italics="true" pageId="12" pageNumber="113">Shell</emphasis>
. Small (up to
<quantity id="4C9CB19657743C28FE37FF0EBAC3A9C5" box="[421,509,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="113" unit="mm" value="2.3">2.3 mm</quantity>
in maximum diameter; see
<tableCitation id="C6E629C857743C28FCAFFF0EB8AFA9C5" box="[829,913,188,213]" captionStart="TABLE 3" captionStartId="13.[151,239,656,679]" captionText="TABLE 3. Shell dimensions of type specimens of species of Clenchiellidae." pageId="12" pageNumber="113">Table 3</tableCitation>
for shell dimensions of all species), spire flat (
<figureCitation id="135F00F657743C28FF0DFF6DBBE0A9E9" box="[159,222,223,248]" captionStart="FIGURE 1" captionStartId="11.[151,250,1702,1724]" captionTargetBox="[163,1428,289,1686]" captionTargetId="figure@11.[163,1428,289,1687]" captionTargetPageId="11" captionText="FIGURE 1. Holotypes of species of Clenchiella with distinct keels (from left towards right: dorsal, lateral and ventral view respectively). A C, Clenchiella victoriae (USNM 488534; shell diameter 2.1 mm); D F, Clenchiella bicingulata n. sp. (AMS C. 462959; shell diameter 1.97 mm); G I, Clenchiella varicosa n. sp. (AMS C. 460738; shell diameter 1.96 mm); J L, Clenchiella iriomotensis n. sp. (AMS C. 445413; shell diameter 1.94 mm)." httpUri="https://zenodo.org/record/253225/files/figure.png" pageId="12" pageNumber="113">Fig. 1</figureCitation>
AC). Protoconch not elevated above spire, of 1.5 smooth whorls, last whorl descends slightly just behind aperture. Teleoconch of 2.53.0 whorls, rounded except for angulations from two prominent carinae, one middorsal, one mid-basal; whole surface sculptured with fine spiral threads, interspaces about equal to width of threads; finer commarginal growth lines present, becoming more prominent towards thickened external varix. Periphery evenly convex, spiral lirae similar in strength on peripheral area as on dorsal and basal surfaces. Umbilicus wide (over half width of base), spiral sculpture continuous within. Sutures impressed. Aperture near circular, with peristome slightly thickened internally by slight rim, external varix moderately thickened, just behind apertural rim. Periostracum moderately thick, bright translucent red in life (
<bibRefCitation id="EFF5618257743C28FC64FE6EBFB7A8E5" author="Abbott" box="[1014,1161,476,501]" pageId="12" pageNumber="113" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott 1948</bibRefCitation>
), brackish-brown in dry shells; shell white.
</paragraph>
<paragraph id="8BDB1C7357743C28FF55FD96B93CAB71" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF55FD96BA7AAB2C" box="[199,324,548,573]" italics="true" pageId="12" pageNumber="113">Operculum</emphasis>
. Horny, near circular, of 4 slowly increasing whorls. Interior with small pimple-like projection in middle part (from
<bibRefCitation id="EFF5618257743C28FEF7FDF5BAC8AB71" author="Abbott" box="[357,502,583,608]" pageId="12" pageNumber="113" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott 1948</bibRefCitation>
).
</paragraph>
<paragraph id="8BDB1C7357743C28FF55FDDEBAF9AA91" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF55FDDEBA08AB94" bold="true" box="[199,310,620,645]" pageId="12" pageNumber="113">Anatomy</emphasis>
(Based on the description and figures of
<bibRefCitation id="EFF5618257743C28FC82FDDEB89CAB95" author="Abbott" box="[784,930,620,645]" pageId="12" pageNumber="113" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott 1948</bibRefCitation>
).
<emphasis id="B910C06157743C28FC25FDDEBF17AB94" box="[951,1065,620,645]" italics="true" pageId="12" pageNumber="113">Head-foot</emphasis>
. Snout bilobed distally, cephalic tentacles long and slender, with 67 cilial tufts on small raised areas (described as serrations by
<bibRefCitation id="EFF5618257743C28FB72FD3DBE51ABB9" author="Abbott" box="[1248,1391,655,680]" pageId="12" pageNumber="113" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott 1948</bibRefCitation>
) on outer edge of left tentacle. Foot with rounded anterior end, laterally extended, with weak indentation, tapering posteriorly. Mucous slit at anterior edge, anterior mucous gland typical of family. Tentacles with black band near distal end. Light orange cells embedded in tentacles and just above right eye. Snout black other than anterior margin and broken clear streak mid-dorsally. Dorsal head asymmetrically coloured with dark grey to black on left side. Patches of grey on right side of neck and anterior lateral dorsal foot. Sides of foot translucent whitish with few yellow or orange cells.
</paragraph>
<caption id="DF1B4CFB57743C28FF05F916BE3DAFEF" httpUri="https://zenodo.org/record/253226/files/figure.png" pageId="12" pageNumber="113" targetBox="[150,1436,985,1678]" targetPageId="12">
<paragraph id="8BDB1C7357743C28FF05F916BE3DAFEF" blockId="12.[151,1436,1700,1790]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF05F916BA2CAFAB" bold="true" box="[151,274,1700,1722]" pageId="12" pageNumber="113">FIGURE 2.</emphasis>
Type specimens of
<taxonomicName id="4C6467F057743C28FE4EF916B9ECAFAB" box="[476,722,1700,1722]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="minutissima">
<emphasis id="B910C06157743C28FE4EF916B9ECAFAB" box="[476,722,1700,1722]" italics="true" pageId="12" pageNumber="113">Clenchiella minutissima</emphasis>
</taxonomicName>
(from left towards right: dorsal, lateral and ventral view respectively).
<emphasis id="B910C06157743C28FF05F977BBF6AFCA" bold="true" box="[151,200,1733,1755]" pageId="12" pageNumber="113">AC</emphasis>
, holotype of
<taxonomicName id="4C6467F057743C28FEC1F977B90BAFCD" box="[339,565,1733,1756]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="papuensis">
<emphasis id="B910C06157743C28FEC1F977B90BAFCD" box="[339,565,1733,1756]" italics="true" pageId="12" pageNumber="113">Clenchiella papuensis</emphasis>
</taxonomicName>
, synonymised with
<taxonomicName id="4C6467F057743C28FC97F977B8C2AFCD" box="[773,1020,1733,1756]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="minutissima">
<emphasis id="B910C06157743C28FC97F977B8C2AFCD" box="[773,1020,1733,1756]" italics="true" pageId="12" pageNumber="113">Clenchiella minutissima</emphasis>
</taxonomicName>
herein (NHML, registration number not known; shell diameter 1.69 mm);
<emphasis id="B910C06157743C28FE7FF95AB91EAFEF" bold="true" box="[493,544,1768,1790]" pageId="12" pageNumber="113">DF,</emphasis>
syntype of
<taxonomicName id="4C6467F057743C28FD05F95AB8B2AFEF" box="[663,908,1768,1790]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="minutissima">
<emphasis id="B910C06157743C28FD05F95AB8B2AFEF" box="[663,908,1768,1790]" italics="true" pageId="12" pageNumber="113">Clenchiella minutissima</emphasis>
</taxonomicName>
(MNHNP; shell diameter 1.70 mm).
</paragraph>
</caption>
<paragraph id="8BDB1C7357743C28FF55F899B96DAE55" blockId="12.[151,1436,1835,2004]" box="[199,595,1835,1860]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF55F899BA7DAE55" box="[199,323,1835,1860]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="4C6467F057743C28FF55F899BA7EAE55" box="[199,320,1835,1860]" class="Bryopsida" family="Hypnaceae" genus="Ctenidium" kingdom="Plantae" order="Hypnales" pageId="12" pageNumber="113" phylum="Bryophyta" rank="genus">Ctenidium</taxonomicName>
.
</emphasis>
About 2527 filaments.
</paragraph>
<paragraph id="8BDB1C7357743C28FF55F8FCB98AAEC2" blockId="12.[151,1436,1835,2004]" pageId="12" pageNumber="113">
<emphasis id="B910C06157743C28FF55F8FCBA21AE76" box="[199,287,1870,1895]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="4C6467F057743C28FF55F8FCBA22AE76" box="[199,284,1870,1895]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="12" pageNumber="113" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
.
</emphasis>
Typical of family. Cusp formulae 4+1+4, 2+1+3, 18, 20, median cusps of central and lateral teeth a little longer than adjacent cusps, elongate, pointed; other cusps on all teeth triangular, pointed. [Note: Abbotts figure of the
<taxonomicName id="4C6467F057743C28FEB9F824BA4FAEBE" box="[299,369,1942,1967]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="12" pageNumber="147" phylum="Bryophyta" rank="genus">radula</taxonomicName>
(his fig. 7) shows the marginal teeth flattened so that they appear to be broader than the SEM images of those of other species of
<taxonomicName id="4C6467F057743C28FDB1F809B99AAEC5" box="[547,676,1979,2004]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157743C28FDB1F809B99AAEC5" box="[547,676,1979,2004]" italics="true" pageId="12" pageNumber="113">Clenchiella</emphasis>
</taxonomicName>
].
</paragraph>
<paragraph id="8BDB1C7357753C29FF55FF2AB9DCA9A1" blockId="13.[151,1436,151,608]" box="[199,738,151,176]" pageId="13" pageNumber="114">
<emphasis id="B910C06157753C29FF55FF2ABBC6A9BE" box="[199,248,152,175]" italics="true" pageId="13" pageNumber="114">Gut.</emphasis>
Anterior oesophagus and rectum unknown.
</paragraph>
<paragraph id="8BDB1C7357753C29FF55FF0FB851A851" blockId="13.[151,1436,151,608]" pageId="13" pageNumber="114">
<emphasis id="B910C06157753C29FF55FF0FBA3BA9C5" box="[199,261,189,212]" italics="true" pageId="13" pageNumber="114">Penis</emphasis>
. Well behind right tentacle, coiled for half turn when retracted. Long, narrow (in life), expanded distally with two glandular lobes: one adjacent to short distal part of penis, and larger oval outer lobe (
<bibRefCitation id="EFF5618257753C29FB27FF6DBEB0A9E9" author="Abbott" box="[1205,1422,223,248]" pageId="13" pageNumber="114" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott 1948, fig. 5</bibRefCitation>
). Tip narrow, bluntly pointed. Penial duct straight to slightly undulating, narrow throughout. Translucent and unpigmented except for few white cells in largest glandular lobe.
</paragraph>
<paragraph id="8BDB1C7357753C29FF55FEFEB848A874" blockId="13.[151,1436,151,608]" box="[199,886,332,357]" pageId="13" pageNumber="114">
<emphasis id="B910C06157753C29FF55FEFEB9D4A875" box="[199,746,332,357]" italics="true" pageId="13" pageNumber="114">Female reproductive system and nervous system.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857753C29FF55FEDDBEADAECA" pageId="13" pageNumber="114" type="discussion">
<paragraph id="8BDB1C7357753C29FF55FEDDB9EEA8C1" blockId="13.[151,1436,151,608]" pageId="13" pageNumber="114">
<emphasis id="B910C06157753C29FF55FEDDBA05A899" bold="true" box="[199,315,367,392]" pageId="13" pageNumber="114">Remarks.</emphasis>
This species is readily distinguished by its relatively thick shell which bears a very heavy, sharp keel on the base and a moderately strong keel on the shoulder. Based on Abbotts description and figure the penis has only two glandular lobes, both located distally.
</paragraph>
<paragraph id="8BDB1C7357753C29FF05FE6EBA3EAB71" blockId="13.[151,1436,151,608]" pageId="13" pageNumber="114">
Abbott figured the aperture as somewhat horizontally elongated and with a very thin peristome, whereas the material inspected in this study appears to possess apertures that are close to perfectly circular and with a slightly thickened peristome. His dimensions are also incorrect giving a size (
<quantity id="4C9CB19657753C29FC28FD96BF3EAB2D" box="[954,1024,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="114" unit="mm" value="4.0">4 mm</quantity>
) twice that of the actual size of the
<typeStatus id="54DFA2D157753C29FF05FDF5BBC3AB71" box="[151,253,583,608]" pageId="13" pageNumber="114" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF1B4CFB57753C29FF05FD22B8FAABB6" box="[151,964,656,679]" pageId="13" pageNumber="114">
<paragraph id="8BDB1C7357753C29FF05FD22B8FAABB6" blockId="13.[151,964,656,679]" box="[151,964,656,679]" pageId="13" pageNumber="114">
<emphasis id="B910C06157753C29FF05FD22BA37ABB6" bold="true" box="[151,265,656,679]" pageId="13" pageNumber="114">TABLE 3.</emphasis>
Shell dimensions of type specimens of species of
<taxonomicName id="4C6467F057753C29FCB0FD22B881ABB6" box="[802,959,656,679]" class="Gastropoda" family="Clenchiellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="114" phylum="Mollusca" rank="family">Clenchiellidae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BDB1C7357753C29FD0EFD76BE62ABCB" blockId="13.[159,1427,708,2011]" box="[668,1372,708,730]" pageId="13" pageNumber="114">SW SH AH UW TW PW TWN</paragraph>
<paragraph id="8BDB1C7357753C29FF0DFD5FBA4AAA12" blockId="13.[159,1427,708,2011]" box="[159,372,749,771]" pageId="13" pageNumber="114">
<taxonomicName id="4C6467F057753C29FF0DFD5FBA4AAA12" box="[159,372,749,771]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="victoriae">
<emphasis id="B910C06157753C29FF0DFD5FBA4AAA12" box="[159,372,749,771]" italics="true" pageId="13" pageNumber="114">Clenchiella victoriae</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BDB1C7357753C29FF0DFCA4BAF3AAC3" blockId="13.[159,1427,708,2011]" pageId="13" pageNumber="114">
<typeStatus id="54DFA2D157753C29FF0DFCA4BBC3AA3D" box="[159,253,790,812]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
2.10 0.86 0.66 1.18 1.75 1.25 3.00
<typeStatus id="54DFA2D157753C29FF0DFCF3BBC6AA47" box="[159,248,833,854]" pageId="13" pageNumber="114" type="paratype">Paratype</typeStatus>
2.17 0.95 0.61 1.17 1.75 - -
<typeStatus id="54DFA2D157753C29FF0DFCD8BBC6AA6E" box="[159,248,874,895]" pageId="13" pageNumber="114" type="paratype">Paratype</typeStatus>
(USNM 488535) 2.08 0.86 0.56 1.24 - - -
<typeStatus id="54DFA2D157753C29FF0DFC21BBC6AAB9" box="[159,248,915,936]" pageId="13" pageNumber="114" type="paratype">Paratype</typeStatus>
(USNM 488535) 2.01 0.81 0.62 1.09 1.75 1.00 2.75
<taxonomicName id="4C6467F057753C29FF0DFC0EBAB3AAC3" box="[159,397,956,978]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="bicingulata" status="sp. nov.">
<emphasis id="B910C06157753C29FF0DFC0EBAB3AAC3" box="[159,397,956,978]" italics="true" pageId="13" pageNumber="114">Clenchiella bicingulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57753C29FE06FC0EBAF3AAC3" box="[404,461,956,978]" pageId="13" pageNumber="114" rank="species">
<emphasis id="B910C06157753C29FE06FC0EBAF3AAC3" bold="true" box="[404,461,956,978]" pageId="13" pageNumber="114">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDB1C7357753C29FF0DFC57BA8EADB1" blockId="13.[159,1427,708,2011]" pageId="13" pageNumber="114">
<typeStatus id="54DFA2D157753C29FF0DFC57BBC3AAEA" box="[159,253,997,1019]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
1.97 0.74 0.47 0.87 1.5 1.25 2.75
<typeStatus id="54DFA2D157753C29FF0DFBBDBA3FAD35" box="[159,257,1039,1060]" pageId="13" pageNumber="114" type="paratype">Paratypes</typeStatus>
2.16 0.90 0.57 0.95 1.75 1.25 3 2.10 0.79 0.58 1.04 1.5 1.5 3 1.96 0.94 0.57 1.05 1.75 0 0
<taxonomicName id="4C6467F057753C29FF0DFB38BA4FADB1" box="[159,369,1162,1184]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="varicosa" status="sp. nov.">
<emphasis id="B910C06157753C29FF0DFB38BA4FADB1" box="[159,369,1162,1184]" italics="true" pageId="13" pageNumber="114">Clenchiella varicosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57753C29FEE5FB38BA8EADB1" box="[375,432,1162,1184]" pageId="13" pageNumber="114" rank="species">
<emphasis id="B910C06157753C29FEE5FB38BA8EADB1" bold="true" box="[375,432,1162,1184]" pageId="13" pageNumber="114">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDB1C7357753C29FF0DFB06BAE8AC57" blockId="13.[159,1427,708,2011]" pageId="13" pageNumber="114">
<typeStatus id="54DFA2D157753C29FF0DFB06BBC3ADDB" box="[159,253,1204,1226]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
1.96 0.70 0.57 0.89 1.5 1.25 2.75
<typeStatus id="54DFA2D157753C29FF0DFB6CBBC6ADE2" box="[159,248,1246,1267]" pageId="13" pageNumber="114" type="paratype">Paratype</typeStatus>
(immature) 1.54 0.56 0.53 0.75 1.75 0 0
<typeStatus id="54DFA2D157753C29FF0DFAB5BBC6AC0D" box="[159,248,1287,1308]" pageId="13" pageNumber="114" type="paratype">Paratype</typeStatus>
(immature) 1.59 0.65 0.00 0.74 1.75 0 0
<taxonomicName id="4C6467F057753C29FF0DFA82BAA9AC57" box="[159,407,1328,1350]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="iriomotensis" status="sp. nov.">
<emphasis id="B910C06157753C29FF0DFA82BAA9AC57" box="[159,407,1328,1350]" italics="true" pageId="13" pageNumber="114">Clenchiella iriomotensis</emphasis>
</taxonomicName>
<emphasis id="B910C06157753C29FE0FFA82BAE8AC57" bold="true" box="[413,470,1328,1350]" pageId="13" pageNumber="114">
<taxonomicNameLabel id="A2237D1A57753C29FE0FFA82BAE8AC57" box="[413,470,1328,1350]" pageId="13" pageNumber="114" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDB1C7357753C29FF0DFAEBBAAAAF04" blockId="13.[159,1427,708,2011]" pageId="13" pageNumber="114">
<typeStatus id="54DFA2D157753C29FF0DFAEBBBC3AC7E" box="[159,253,1369,1391]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
1.94 0.82 0.53 1.06 2 1.25 3.25
<typeStatus id="54DFA2D157753C29FF0DFA31BA3FAC89" box="[159,257,1411,1432]" pageId="13" pageNumber="114" type="paratype">Paratypes</typeStatus>
2.32 1.00 0.68 1.23 1.75 0 0 2.05 0.84 0.68 1.03 1.65 1.25 2.9 2.11 0.87 0.65 1.11 2 1.25 3.25
<taxonomicName id="4C6467F057753C29FF0DFA4CBAAAAF04" box="[159,404,1534,1557]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="minutissima">
<emphasis id="B910C06157753C29FF0DFA4CBAAAAF04" box="[159,404,1534,1557]" italics="true" pageId="13" pageNumber="114">Clenchiella minutissima</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BDB1C7357753C29FF0DF99ABEADAECA" blockId="13.[159,1427,708,2011]" pageId="13" pageNumber="114">
<typeStatus id="54DFA2D157753C29FF0DF99ABBCCAF2F" box="[159,242,1576,1598]" pageId="13" pageNumber="114" type="syntype">Syntype</typeStatus>
1.70 0.75 0.60 0.83 1.5 0 0 ANSP 319936 1.77 0.81 0.57 0.74 0 0 0 1.87 0.83 0.61 0.79 0 0 0 1.84 0.91 0.58 0.74 0 0 0 AMS C.412278 (Darwin) 1.29 0.47 0.40 0.59 1.5 1.25 2.75 1.35 0.56 0.40 0.70 1.75 1.12 2.85 1.23 0.49 0.38 0.57 1.5 1.25 2.75 C.412277 (Magnetic Is.) 1.38 0.51 0.42 0.64 1.5 1.35 2.85 1.66 0.56 0.51 0.77 1.5 1.12 2.62 1.52 0.56 0.49 0.82 1.5 1.25 2.75
<emphasis id="B910C06157753C29FBCFF877BEADAECA" box="[1117,1427,1989,2011]" italics="true" pageId="13" pageNumber="114">......continued on the next page</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>