496 lines
56 KiB
XML
496 lines
56 KiB
XML
<document id="F09CCBC7A2E7AE19D4C3CC495C1A2721" ID-DOI="10.5281/zenodo.5181879" ID-ISSN="1942-1354" ID-Zenodo-Dep="5181879" ID-ZooBank="FADE26B1-F110-443D-A4FA-45A6AF2ACC07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628297584887" checkinUser="felipe" docAuthor="Skelley, Paul E. & Warner, William B." docDate="2015" docId="03CD87BF0F06E76A9EA7FBBFB785FED8" docLanguage="en" docName="InsectaMundi.0416.1-6.pdf" docOrigin="Insecta Mundi 2015 (416)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Trichonotuloides aphoderrans Skelley & Warner 2015, n. sp." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFF4FFC70F05E76D9E67FF9EB225FFE1" masterDocTitle="Trichonotuloides aphoderrans, n. sp. (Coleoptera: Scarabaeidae: Aphodiinae), a new genus and species for the United States" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1699041429326" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="4CAC15566B4F1560C66F99F417D69691" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9C48B585B01226150C43994DE5D4150A">
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<mods:title id="7527505BCA99D7A259A938AE456D5D87">Trichonotuloides aphoderrans, n. sp. (Coleoptera: Scarabaeidae: Aphodiinae), a new genus and species for the United States</mods:title>
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<mods:name id="516FA4ECC3AE4E61CC210E76EEC0CE9E" type="personal">
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<mods:roleTerm id="3EB92214E01297C99B7EDC88770E07EA">Author</mods:roleTerm>
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<mods:namePart id="6D2E565BA851D05E31C38CBF11B38409">Skelley, Paul E.</mods:namePart>
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</mods:name>
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<mods:name id="E012DB174802C1C070CA31DC8FC7ACC0" type="personal">
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<mods:roleTerm id="C19593FA6B95271DC1A7F49A9A73DB0E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="401AB5286250708AEC1E48F78E8F1AA5">Warner, William B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1172F1A4BED55B306AE51296404770D0">text</mods:typeOfResource>
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<mods:titleInfo id="B80820E8A3A2799FCDEFCB95DDF03500">
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<mods:title id="A5A94850DE088CE8B4713CF5D779D1BD">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="EB33AAC8E3F5620168AE56F8B0EF6490">
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<mods:date id="4CCF6FDB9DB1DCCF518F70A37FC52A7E">2015</mods:date>
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<mods:detail id="71D006C7797458456CE585C85386AC66" type="pubDate">
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<mods:number id="CD372613C16D661465046419D2B05C88">2015-04-03</mods:number>
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</mods:detail>
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<mods:detail id="0C72BC709F55C2D61BF54229543FD95A" type="volume">
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<mods:number id="63F214B7191C0B4FB2731C44A1225942">2015</mods:number>
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</mods:detail>
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<mods:detail id="B84D24F76B9BFF85EC220D2AF603C73A" type="issue">
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<mods:number id="F2093EDF295428D3D59BBB23B33A6480">416</mods:number>
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</mods:detail>
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<mods:start id="B6189EB53E008A94DB23C50C799B6EE7">1</mods:start>
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</mods:relatedItem>
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<mods:classification id="DC96E5D794CAFF66E75E8EDEFFDB1DDB">journal article</mods:classification>
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<mods:identifier id="A37B5F1F97ED0430E615B1FB164F909F" type="DOI">10.5281/zenodo.5181879</mods:identifier>
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<mods:identifier id="FBDC51543C5B626D5001D6138D01E9F6" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="7A2985B683505287C7FF3AC65854ADDB" type="Zenodo-Dep">5181879</mods:identifier>
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<mods:identifier id="D0D2C254857233DDCDC8C3B89F12FCF7" type="ZooBank">FADE26B1-F110-443D-A4FA-45A6AF2ACC07</mods:identifier>
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</mods:mods>
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<treatment id="03CD87BF0F06E76A9EA7FBBFB785FED8" LSID="urn:lsid:plazi:treatment:03CD87BF0F06E76A9EA7FBBFB785FED8" httpUri="http://treatment.plazi.org/id/03CD87BF0F06E76A9EA7FBBFB785FED8" lastPageId="7" lastPageNumber="6" pageId="3" pageNumber="2">
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<subSubSection id="C37E65220F06E76E9EA7FBBFB09FFBD8" box="[192,698,1057,1082]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8BDB36A90F06E76E9EA7FBBFB09FFBD8" blockId="3.[192,698,1057,1114]" box="[192,698,1057,1082]" pageId="3" pageNumber="2">
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<heading id="D09381C50F06E76E9EA7FBBFB09FFBD8" bold="true" box="[192,698,1057,1082]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="B910EABB0F06E76E9EA7FBBFB04FFBDB" bold="true" box="[192,618,1057,1082]" pageId="3" pageNumber="2">
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<taxonomicName id="4C644D2A0F06E76E9EA7FBBFB046FBDB" authority="Skelley & Warner, 2015" authorityName="Skelley & Warner" authorityYear="2015" box="[192,611,1057,1082]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="aphoderrans" status="sp. nov.">
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<emphasis id="B910EABB0F06E76E9EA7FBBFB046FBDB" bold="true" box="[192,611,1057,1082]" italics="true" pageId="3" pageNumber="2">Trichonotuloides aphoderrans</emphasis>
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</taxonomicName>
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,
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||
</emphasis>
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<taxonomicNameLabel id="A22357C00F06E76E9C15FBBCB09FFBD8" box="[626,698,1058,1082]" pageId="3" pageNumber="2" rank="species">
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<emphasis id="B910EABB0F06E76E9C15FBBCB096FBDB" bold="true" box="[626,691,1058,1082]" pageId="3" pageNumber="2">n. sp</emphasis>
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||
.
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||
</taxonomicNameLabel>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37E65220F06E76E9EA7FBDCB344FBBB" box="[192,353,1090,1114]" pageId="3" pageNumber="2" type="description">
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<paragraph id="8BDB36A90F06E76E9EA7FBDCB344FBBB" blockId="3.[192,698,1057,1114]" box="[192,353,1090,1114]" pageId="3" pageNumber="2">
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<figureCitation id="135F2A2C0F06E76E9EA7FBDCB344FBBB" box="[192,353,1090,1114]" captionStart-0="Figures 1–8" captionStart-1="Figures 9–14" captionStartId-0="4.[192,288,1846,1868]" captionStartId-1="6.[192,288,1846,1868]" captionTargetBox-0="[192,1440,280,1825]" captionTargetBox-1="[208,1425,738,1836]" captionTargetId-0="figure-68@4.[192,1440,280,1825]" captionTargetId-1="figure-243@6.[208,1425,738,1836]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." captionText-1="Figures 9–14. Trichonotuloides aphoderrans n. sp. 9) Male holotype, ventral view. 10–11) Female allotype, dorsal and ventral view. 12) Male holotype head and pronotum dorso-lateral view. 13) Female allotype protibia. 14) Epipharynx, paratype male." figureDoi-0="http://doi.org/10.5281/zenodo.5181881" figureDoi-1="http://doi.org/10.5281/zenodo.5181883" httpUri-0="https://zenodo.org/record/5181881/files/figure.png" httpUri-1="https://zenodo.org/record/5181883/files/figure.png" pageId="3" pageNumber="2">Figures 4–14</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37E65220F06E76E9EA7FB1CB784FABB" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="8BDB36A90F06E76E9EA7FB1CB6C2FB3B" blockId="3.[192,1441,1153,1370]" pageId="3" pageNumber="2">
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<emphasis id="B910EABB0F06E76E9EA7FB1CB3A3FB78" bold="true" box="[192,390,1153,1178]" pageId="3" pageNumber="2">Type material</emphasis>
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.
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<materialsCitation id="3B0C3CF40F06E76E9FFEFB1CB6C2FB3B" collectingDate="2014-08-13" collectionCode="FSCA" country="United States of America" county="Coconino Co." location="Coconino Co." longitude="-34.32" pageId="3" pageNumber="2" specimenCount="2" typeStatus="holotype">
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<emphasis id="B910EABB0F06E76E9FFEFB1CB032FB7B" bold="true" box="[409,535,1154,1178]" pageId="3" pageNumber="2">
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<typeStatus id="54DF880B0F06E76E9FFEFB1CB032FB7B" box="[409,535,1154,1178]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
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</emphasis>
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male and
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<emphasis id="B910EABB0F06E76E9CC3FB1CB131FB7B" bold="true" box="[676,788,1154,1178]" pageId="3" pageNumber="2">
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<typeStatus id="54DF880B0F06E76E9CC3FB1CB131FB7B" box="[676,788,1154,1178]" pageId="3" pageNumber="2" type="allotype">allotype</typeStatus>
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||
</emphasis>
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||
female appropriately labeled as such, deposited in
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||
<collectionCode id="ED75AE6C0F06E76E9EA7FB3CB329FB5B" box="[192,268,1186,1210]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
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, with the following collection label: “
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<collectingCountry id="F37376390F06E76E9CA2FB3CB127FB5B" box="[709,770,1186,1210]" name="United States of America" pageId="3" pageNumber="2">USA</collectingCountry>
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: AZ:
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<collectingCounty id="62BA4E250F06E76E9D5BFB3CB1F8FB5B" box="[828,989,1186,1210]" pageId="3" pageNumber="2">Coconino Co.</collectingCounty>
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,
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<quantity id="4C9C9B4C0F06E76E9D8FFB3CB611FB5B" box="[1000,1076,1186,1210]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="2" unit="km" value="0.6">0.6km</quantity>
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E jct FS300 &
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<geoCoordinate id="EE50506E0F06E76E9A8FFB3CB71DFB4E" box="[1256,1336,1184,1210]" degrees="34, 32" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="555" value="-34.32">
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34, 32
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<superScript id="7C119BE10F06E76E9B57FB3EB71DFB4E" attach="none" box="[1328,1336,1184,1199]" fontSize="6" pageId="3" pageNumber="2">o</superScript>
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</geoCoordinate>
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24’41”N, 119
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<superScript id="7C119BE10F06E76E9E8BFB5EB2D1FB2E" attach="none" box="[236,244,1216,1231]" fontSize="6" pageId="3" pageNumber="2">o</superScript>
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58’44”[W],
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<date id="FFDA10690F06E76E9F1DFB5CB00FFB3B" box="[378,554,1218,1242]" pageId="3" pageNumber="2" value="2014-08-13">
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<collectingDate id="EF9EE9810F06E76E9F1DFB5CB00FFB3B" box="[378,554,1218,1242]" pageId="3" pageNumber="2" value="2014-08-13">AUG 13, 2014</collectingDate>
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</date>
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||
; clinging to bottom of fresh elk dung; 7730’; WBWarner”.
|
||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8BDB36A90F06E76E9E97FB7CB784FABB" blockId="3.[192,1441,1153,1370]" pageId="3" pageNumber="2">
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<materialsCitation id="3B0C3CF40F06E76E9E97FB7CB1A5FB1B" box="[240,896,1249,1274]" collectingDate="2014-08-13" collectionCode="FSCA" country="United States of America" county="Coconino Co." location="Same" longitude="-34.32" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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<emphasis id="B910EABB0F06E76E9E97FB7CB358FB1B" bold="true" box="[240,381,1250,1274]" pageId="3" pageNumber="2">
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<typeStatus id="54DF880B0F06E76E9E97FB7CB358FB1B" box="[240,381,1250,1274]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
(54 exx.):
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||
<location id="8EBB60720F06E76E9F9DFB7CB01AFB1B" LSID="urn:lsid:plazi:treatment:03CD87BF0F06E76A9EA7FBBFB785FED8:8EBB60720F06E76E9F9DFB7CB01AFB1B" box="[506,575,1250,1274]" name="Same" pageId="3" pageNumber="2">Same</location>
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||
data as holotype (25 exx.)
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</materialsCitation>
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;
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||
<materialsCitation id="3B0C3CF40F06E76E9DEBFB7CB0DFFAFB" collectingDate="2014-08-24" country="United States of America" county="Coconino Co." latitude="32.411392" location="Coconino Co." longLatPrecision="469" longitude="-34.32" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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“
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<collectingCountry id="F37376390F06E76E9DF0FB7CB1F1FB1B" box="[919,980,1250,1274]" name="United States of America" pageId="3" pageNumber="2">USA</collectingCountry>
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: AZ:
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<collectingCounty id="62BA4E250F06E76E9A76FB7CB697FB1B" box="[1041,1202,1250,1274]" pageId="3" pageNumber="2">Coconino Co.</collectingCounty>
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,
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<quantity id="4C9C9B4C0F06E76E9AD9FB7CB72FFB1B" box="[1214,1290,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="2" unit="km" value="0.6">0.6km</quantity>
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E jct FS300 &
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<geoCoordinate id="EE50506E0F06E76E9EBCFA9CB30EFAEE" box="[219,299,1280,1306]" degrees="34, 32" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="555" value="-34.32">
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34,
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<geoCoordinate id="EE50506E0F06E76E9F62FA9CB30EFAEE" box="[261,299,1280,1306]" degrees="32" direction="north" minutes="24" orientation="latitude" pageId="3" pageNumber="2" precision="15" seconds="41" value="32.411392">
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32
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<superScript id="7C119BE10F06E76E9F44FA9EB30EFAEE" attach="right" box="[291,299,1280,1295]" fontSize="6" pageId="3" pageNumber="2">o</superScript>
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</geoCoordinate>
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</geoCoordinate>
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24’41”N, 119
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<superScript id="7C119BE10F06E76E9FA3FA9EB3E9FAEE" attach="none" box="[452,460,1280,1295]" fontSize="6" pageId="3" pageNumber="2">o</superScript>
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58’44”[W],
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<date id="FFDA10690F06E76E9C28FA9CB0DFFAFB" box="[591,762,1282,1306]" pageId="3" pageNumber="2" value="2014-08-24">
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<collectingDate id="EF9EE9810F06E76E9C28FA9CB0DFFAFB" box="[591,762,1282,1306]" pageId="3" pageNumber="2" value="2014-08-24">AUG 24, 2014</collectingDate>
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</date>
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||
</materialsCitation>
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||
; clinging to bottom of moist elk dung; 7730’;
|
||
<materialsCitation id="3B0C3CF40F06E76E9B6CFA9CB73CFADB" collectorName="White Mtn. & Ariz." country="United States of America" county="Apache Co." location="Apache Co." pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
WBWarner” (28 exx.); [AZ:
|
||
<collectingCounty id="62BA4E250F06E76E9F16FABCB3D8FADB" box="[369,509,1314,1338]" pageId="3" pageNumber="2">Apache Co.</collectingCounty>
|
||
, hand written] “
|
||
<collectorName id="2691537F0F06E76E9CA3FABCB176FAD8" box="[708,851,1314,1338]" pageId="3" pageNumber="2">White Mtn.</collectorName>
|
||
,
|
||
<collectorName id="2691537F0F06E76E9D38FABCB1B8FADB" box="[863,925,1314,1338]" pageId="3" pageNumber="2">Ariz.</collectorName>
|
||
, VIII-28-32” (1 damaged male)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54DF880B0F06E76E9B44FABCB785FADB" box="[1315,1440,1314,1338]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
|
||
deposited in
|
||
<collectionCode id="ED75AE6C0F06E76E9F3FFADCB383FABB" box="[344,422,1346,1370]" country="USA" httpUri="http://grbio.org/cool/jxy6-3sw7" name="Frank M. Hasbrouck Insect Collection" pageId="3" pageNumber="2">ASUT</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9FD7FADCB02DFABB" box="[432,520,1346,1370]" pageId="3" pageNumber="2">CMNC</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9C75FADCB07AFABB" box="[530,607,1346,1370]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="3" pageNumber="2" type="Museum">CSCA</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9C0EFADCB096FABB" box="[617,691,1346,1370]" pageId="3" pageNumber="2">DCGI</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9CDAFADCB12CFABB" box="[701,777,1346,1370]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9D74FADCB143FABB" box="[787,870,1346,1370]" country="USA" httpUri="http://grbio.org/cool/ktis-ep89" name="Northern Arizona University" pageId="3" pageNumber="2" type="University or college">NAUF</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9D17FADCB1E5FABB" box="[880,960,1346,1370]" pageId="3" pageNumber="2">RACC</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9DAEFADCB607FABB" box="[969,1058,1346,1370]" pageId="3" pageNumber="2">RHMC</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9A4BFADCB65AFABB" box="[1068,1151,1346,1370]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="3" pageNumber="2" type="Museum">SEMC</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9AEEFADCB6F6FABB" box="[1161,1235,1346,1370]" country="0" httpUri="http://grbio.org/cool/tizb-nd41" name="University of Alabama, Ichthyological Collection" pageId="3" pageNumber="2">UAIC</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9ABBFADCB711FABB" box="[1244,1332,1346,1370]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="3" pageNumber="2">UNSM</collectionCode>
|
||
,
|
||
<collectionCode id="ED75AE6C0F06E76E9B59FADCB7B8FABB" box="[1342,1437,1346,1370]" pageId="3" pageNumber="2">WBWC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F06E76E9EA7FA1FB785F9F8" pageId="3" pageNumber="2" type="diagnosis">
|
||
<paragraph id="8BDB36A90F06E76E9EA7FA1FB785F9F8" blockId="3.[192,1441,1409,1562]" pageId="3" pageNumber="2">
|
||
<emphasis id="B910EABB0F06E76E9EA7FA1FB36DFA78" bold="true" box="[192,328,1409,1433]" pageId="3" pageNumber="2">Diagnosis</emphasis>
|
||
. Besides being the only species of the genus known from the
|
||
<collectingCountry id="F37376390F06E76E9A4AFA1CB6FFFA7B" box="[1069,1242,1410,1434]" name="United States of America" pageId="3" pageNumber="2">United States</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C644D2A0F06E76E9A83FA1CB785FA7B" authorityName="Skelley & Warner" authorityYear="2015" box="[1252,1440,1410,1434]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F06E76E9A83FA1CB785FA7B" box="[1252,1440,1410,1434]" italics="true" pageId="3" pageNumber="2">T. aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished from other species in the genus by the following combination of characters: head with genae angulate, angle approximately 90
|
||
<superScript id="7C119BE10F06E76E9C8EFA5EB0D4FA2E" attach="none" box="[745,753,1472,1487]" fontSize="6" pageId="3" pageNumber="2">o</superScript>
|
||
, pronotal punctures uniformly coarse, elytral punctures distinct, elytral surface between punctures flattened and almost glossy (at most only vaguely vermiculose), male protibial spur weakly hooked inward at apex, and mesotarsomere I shorter than upper spur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F06E7689EA7F9DFB3E3FB7B" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="description">
|
||
<paragraph id="8BDB36A90F06E7689EA7F9DFB318FE5B" blockId="3.[192,1440,1601,1914]" lastBlockId="5.[192,1441,194,730]" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
|
||
<emphasis id="B910EABB0F06E76E9EA7F9DFB34EF9B8" bold="true" box="[192,363,1601,1625]" pageId="3" pageNumber="2">Description.</emphasis>
|
||
Length 3.7–5.0 mm; oval-elongate, convex, almost glossy beneath punctation and pubescence; each elytral puncture with short, erect silver-grey seta. Blackish or piceous black; head, pronotum and elytra with margins dark reddish. Head with epistome convex at center, coarsely and densely punctured, punctures sparser on disc, denser on sides; clypeus feebly sinuate at middle, rounded or obtusely angulate at each side, bordered, edge imperceptibly bristled laterally; genae angulate, ~90
|
||
<superScript id="7C119BE10F06E76E9BF6F95EB7BCF92E" attach="none" box="[1425,1433,1728,1743]" fontSize="6" pageId="3" pageNumber="2">o</superScript>
|
||
, rather shortly ciliate, distinctly protruding from the eyes; frontal suture nearly obsolete at middle, moderately raised laterally; front coarsely and rather densely punctured. Epipharynx transverse (
|
||
<figureCitation id="135F2A2C0F06E76E9B5CF89CB7B7F8FB" box="[1339,1426,1794,1818]" captionStart="Figures 9–14" captionStartId="6.[192,288,1846,1868]" captionTargetBox="[208,1425,738,1836]" captionTargetId="figure-243@6.[208,1425,738,1836]" captionTargetPageId="6" captionText="Figures 9–14. Trichonotuloides aphoderrans n. sp. 9) Male holotype, ventral view. 10–11) Female allotype, dorsal and ventral view. 12) Male holotype head and pronotum dorso-lateral view. 13) Female allotype protibia. 14) Epipharynx, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181883" httpUri="https://zenodo.org/record/5181883/files/figure.png" pageId="3" pageNumber="2">Fig. 14</figureCitation>
|
||
). Pronotum transverse, moderately convex medially, narrowly flattened on sides, strongly and densely punctured with nearly uniformly sized punctures; sides of disc shortly pubescent; lateral margins with border minutely bristled, slightly inwardly sinuate before hind angles; hind angle obtusely rounded; base slightly bisinuate and distinctly bordered. Scutellum with curved sides, coarsely and confusedly punctured. Elytra elongate-oval, moderately convex, not denticulate at shoulder; epipleural carina very distinct at humeral callus and minutely bristled; striae wide, glossy, superficially punctured and subcrenulate discally, indistinctly punctured and canaliculate toward apex; intervals flat with coarse punctation distinct, separated by almost glossy surface between punctures [surface best studied under diffused light]. Upper spur of mesotibia longer than first tarsal segment. Upper spur of hind tibiae shorter than first tarsal segment; latter as long as or slightly longer than the following three segments combined.
|
||
</paragraph>
|
||
<caption id="DF1B66210F01E7699EA7F8A8B62DF867" ID-DOI="http://doi.org/10.5281/zenodo.5181881" ID-Zenodo-Dep="5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,288,1846,1868]" targetBox="[192,1440,280,1825]" targetPageId="4">
|
||
<paragraph id="8BDB36A90F01E7699EA7F8A8B62DF867" blockId="4.[192,1441,1846,1926]" pageId="4" pageNumber="3">
|
||
<emphasis id="B910EABB0F01E7699EA7F8A8B36BF8AA" bold="true" box="[192,334,1846,1868]" pageId="4" pageNumber="3">Figures 1–8</emphasis>
|
||
.
|
||
<taxonomicName id="4C644D2A0F01E7699F3BF8A8B07AF8AD" box="[348,607,1846,1868]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="undefined-1">
|
||
<emphasis id="B910EABB0F01E7699F3BF8A8B033F8AA" box="[348,534,1846,1867]" italics="true" pageId="4" pageNumber="3">Trichonotuloides</emphasis>
|
||
spp.
|
||
<emphasis id="B910EABB0F01E7699C35F8A8B07AF8AD" bold="true" box="[594,607,1846,1868]" pageId="4" pageNumber="3">1</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B910EABB0F01E7699C39F8A8B05AF8AD" bold="true" box="[606,639,1846,1868]" pageId="4" pageNumber="3">-3)</emphasis>
|
||
<taxonomicName id="4C644D2A0F01E7699CE1F8A9B16EF8AD" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[646,843,1846,1868]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="glyptus">
|
||
<emphasis id="B910EABB0F01E7699CE1F8A9B0D1F8AA" box="[646,756,1846,1868]" italics="true" pageId="4" pageNumber="3">T. glyptus</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
male.
|
||
<emphasis id="B910EABB0F01E7699DF1F8A8B189F8AD" bold="true" box="[918,940,1846,1868]" pageId="4" pageNumber="3">1)</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="B910EABB0F01E7699A01F8A8B658F8AD" bold="true" box="[1126,1149,1846,1868]" pageId="4" pageNumber="3">2)</emphasis>
|
||
Elytra and pronotal base.
|
||
<emphasis id="B910EABB0F01E7699EA7F8CDB2F2F888" bold="true" box="[192,215,1875,1897]" pageId="4" pageNumber="3">3)</emphasis>
|
||
Protibia, dorsal view.
|
||
<emphasis id="B910EABB0F01E7699FA9F8CDB3DFF888" bold="true" box="[462,506,1875,1897]" pageId="4" pageNumber="3">4-6)</emphasis>
|
||
<taxonomicName id="4C644D2A0F01E7699C67F8CAB08CF889" authority="Skelley & Warner, 2015" authorityName="Skelley & Warner" authorityYear="2015" box="[512,681,1875,1897]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="aphoderrans" status="sp. nov.">
|
||
<emphasis id="B910EABB0F01E7699C67F8CAB08CF889" box="[512,681,1875,1897]" italics="true" pageId="4" pageNumber="3">T. aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22357C00F01E7699CC8F8CDB0C9F889" box="[687,748,1875,1896]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype male.
|
||
<emphasis id="B910EABB0F01E7699DF9F8CDB191F888" bold="true" box="[926,948,1875,1897]" pageId="4" pageNumber="3">4)</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="B910EABB0F01E7699A0DF8CDB6A4F888" bold="true" box="[1130,1153,1875,1897]" pageId="4" pageNumber="3">5)</emphasis>
|
||
Elytra and pronotal base.
|
||
<emphasis id="B910EABB0F01E7699EA7F8F1B2F2F864" bold="true" box="[192,215,1903,1925]" pageId="4" pageNumber="3">6)</emphasis>
|
||
Protibia, dorsal view.
|
||
<emphasis id="B910EABB0F01E7699FB6F8F1B3D8F864" bold="true" box="[465,509,1903,1925]" pageId="4" pageNumber="3">7-8)</emphasis>
|
||
<taxonomicName id="4C644D2A0F01E7699C63F8EEB08BF864" authorityName="Skelley & Warner" authorityYear="2015" box="[516,686,1904,1925]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F01E7699C63F8EEB08BF864" box="[516,686,1904,1925]" italics="true" pageId="4" pageNumber="3">T. aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
male genitalia, paratype male.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB36A90F00E7689E97FE5CB324FD9B" blockId="5.[192,1441,194,730]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689E97FE5CB317FE3B" bold="true" box="[240,306,450,474]" pageId="5" pageNumber="4">Male</emphasis>
|
||
. Clypeal angles rounded; pronotum relatively more transverse; fore tibial spur stout, spatulate, and weakly hooked inwardly at apex; fore tibia more elongate, narrowed, apical half ventrally curved; metasternal plate spoon-shaped, punctured and shortly pubescent distally; setae on basal half of hind femur dense and long; aedeagus (
|
||
<figureCitation id="135F2A2C0F00E7689C3FFDBCB09EFDDB" box="[600,699,546,570]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="5" pageNumber="4">Fig. 7–8</figureCitation>
|
||
) with parameres gradually narrowing to dorsoventrally flattened apex, paramere apices not contiguous at apex, parameres bent ventrally at apical third in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F00E7689E97FD1CB785FD3B" blockId="5.[192,1441,194,730]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689E97FD1CB370FD7B" bold="true" box="[240,341,642,666]" pageId="5" pageNumber="4">Female</emphasis>
|
||
. Clypeal angles often angulate; pronotum relatively narrower toward front; fore tibial spur slender and regularly acuminate apically; fore tibia normal, flattened, lacking ventral curvature on apical half; metasternal plate nearly flat, glabrous; setae on basal half of hind femur sparse and short.
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F00E7689EA7FC9FB3E3FB7B" blockId="5.[192,1441,769,1178]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689EA7FC9FB020FCF8" bold="true" box="[192,517,769,793]" pageId="5" pageNumber="4">Biological observations</emphasis>
|
||
. Nearly all specimens of the
|
||
<typeStatus id="54DF880B0F00E7689D0FFC9CB1B8FCFB" box="[872,925,770,794]" pageId="5" pageNumber="4">type</typeStatus>
|
||
series were collected from beneath aggregated pellets or piles of fresh elk dung to which they mostly clung upside-down to the bottom surface. Some specimens were also found in cracks and surfaces between the coagulated pellets. No specimens were found burrowing into the interior of the dung, but many had excavated shallow channels in the surface as much as half the body depth, and appeared to be “grazing” on the surface of the dung. This is in sharp contrast to the many specimens of
|
||
<taxonomicName id="4C644D2A0F00E7689C9AFC3CB663FC5B" authority="(Fall)" baseAuthorityName="Fall" baseAuthorityYear="1907" box="[765,1094,930,954]" class="Insecta" family="Aphodiidae" genus="Agoliinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="plutonicus">
|
||
<emphasis id="B910EABB0F00E7689C9AFC3CB1DFFC5B" box="[765,1018,930,954]" italics="true" pageId="5" pageNumber="4">Agoliinus plutonicus</emphasis>
|
||
(Fall)
|
||
</taxonomicName>
|
||
and few specimens of
|
||
<taxonomicName id="4C644D2A0F00E7689B3AFC3CB002FC3B" authority="(Horn)" baseAuthorityName="Horn" baseAuthorityYear="1870" class="Insecta" family="Aphodiidae" genus="Pseudagolius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="coloradensis">
|
||
<emphasis id="B910EABB0F00E7689B3AFC3CB3E9FC3B" italics="true" pageId="5" pageNumber="4">Pseudagolius coloradensis</emphasis>
|
||
(Horn)
|
||
</taxonomicName>
|
||
found burrowing completely into the same dung (some on the bottom surface as well). Specimens of
|
||
<taxonomicName id="4C644D2A0F00E7689C52FC7CB0D4FC1B" authorityName="Skelley & Warner" authorityYear="2015" box="[565,753,994,1018]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F00E7689C52FC7CB062FC18" box="[565,583,994,1017]" italics="true" pageId="5" pageNumber="4">T</emphasis>
|
||
.
|
||
<emphasis id="B910EABB0F00E7689C31FC7CB0D4FC1B" box="[598,753,994,1018]" italics="true" pageId="5" pageNumber="4">aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
were densely packed beneath the freshest dung piles in an open, narrow, grassy canyon bottom in ponderosa pine/Douglas fir/aspen woods (
|
||
<figureCitation id="135F2A2C0F00E7689AACFB9CB707FBF8" box="[1227,1314,1025,1050]" captionStart="Figure 15" captionStartId="7.[192,276,1902,1924]" captionTargetBox="[192,1440,1182,1886]" captionTargetId="figure-26@7.[192,1440,1182,1886]" captionTargetPageId="7" captionText="Figure 15. Type locality for Trichonotuloides aphoderrans n. sp." figureDoi="http://doi.org/10.5281/zenodo.5181885" httpUri="https://zenodo.org/record/5181885/files/figure.png" pageId="5" pageNumber="4">Fig. 15</figureCitation>
|
||
); but very few specimens were found beneath dung that had dried to a hard crust on its exposed surfaces. No specimens were found beneath logs or rocks in the area, nor under older dung piles even though the ground beneath was damp. Based on these observations, the species appears to be a true dung feeder and not a detritivore.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F00E7689EA7FB5CB169FA98" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="8BDB36A90F00E7689EA7FB5CB169FA98" blockId="5.[192,1441,1218,1402]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689EA7FB5CB319FB3B" bold="true" box="[192,316,1218,1242]" pageId="5" pageNumber="4">Remarks</emphasis>
|
||
. In general,
|
||
<taxonomicName id="4C644D2A0F00E7689FBBFB5CB0BFFB3B" authorityName="Skelley & Warner" authorityYear="2015" box="[476,666,1218,1242]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F00E7689FBBFB5CB0BFFB3B" box="[476,666,1218,1242]" italics="true" pageId="5" pageNumber="4">T. aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
is most similar in general morphological characters, genitalia and sexual dimorphisms to
|
||
<taxonomicName id="4C644D2A0F00E7689C47FB7CB0B8FB1B" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[544,669,1250,1274]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glyptus">
|
||
<emphasis id="B910EABB0F00E7689C47FB7CB0B8FB1B" box="[544,669,1250,1274]" italics="true" pageId="5" pageNumber="4">T. glyptus</emphasis>
|
||
</taxonomicName>
|
||
, the most commonly collected and widespread species in the genus.
|
||
<taxonomicName id="4C644D2A0F00E7689F7FFA9CB062FAFB" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[280,583,1282,1306]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glyptus">
|
||
<emphasis id="B910EABB0F00E7689F7FFA9CB062FAFB" box="[280,583,1282,1306]" italics="true" pageId="5" pageNumber="4">Trichonotuloides glyptus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by having the genae obtusely rounded, pronotal punctures coarse and moderate in size (
|
||
<figureCitation id="135F2A2C0F00E7689CCBFABCB137FADB" box="[684,786,1314,1338]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="5" pageNumber="4">Fig. 1–2</figureCitation>
|
||
), elytral punctures nearly coalescing making surface distinctly rugose-vermiculose (
|
||
<figureCitation id="135F2A2C0F00E7689C5CFADCB0A6FABB" box="[571,643,1346,1370]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="5" pageNumber="4">Fig. 2</figureCitation>
|
||
), male fore tibial spur distinctly hooked inward at apex (
|
||
<figureCitation id="135F2A2C0F00E7689B2EFADCB7B5FABB" box="[1353,1424,1346,1370]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
|
||
), and middle tarsomere I a bit longer than upper spur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F00E7689EA7FA3CB375F9F8" pageId="5" pageNumber="4" type="etymology">
|
||
<paragraph id="8BDB36A90F00E7689EA7FA3CB375F9F8" blockId="5.[192,1440,1442,1562]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689EA7FA3CB371FA5B" bold="true" box="[192,340,1442,1466]" pageId="5" pageNumber="4">Etymology</emphasis>
|
||
. The specific epithet “
|
||
<taxonomicName id="4C644D2A0F00E7689C05FA3CB0D8FA5B" authorityName="Skelley & Warner" authorityYear="2015" box="[610,765,1442,1466]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F00E7689C05FA3CB0D8FA5B" box="[610,765,1442,1466]" italics="true" pageId="5" pageNumber="4">aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
” was chosen for the fact that the
|
||
<typeStatus id="54DF880B0F00E7689AFFFA3CB6E8FA5B" box="[1176,1229,1442,1466]" pageId="5" pageNumber="4">type</typeStatus>
|
||
series was found walking on the dung, not burrowing into it. It is derived from ‘
|
||
<emphasis id="B910EABB0F00E7689DDAFA5CB606FA3B" box="[957,1059,1474,1498]" italics="true" pageId="5" pageNumber="4">aphodos</emphasis>
|
||
’ which is Greek meaning dung and ‘
|
||
<emphasis id="B910EABB0F00E7689E99FA7CB368FA18" box="[254,333,1506,1529]" italics="true" pageId="5" pageNumber="4">errans</emphasis>
|
||
’ which is Latin meaning wanderer, and is an intentional combination of the Greek and Latin roots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F00E7689EA7F9DCB785F8F8" pageId="5" pageNumber="4" type="key">
|
||
<paragraph id="8BDB36A90F00E7689EA7F9DCB648F97B" blockId="5.[192,1440,1602,1690]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689EA7F9DCB3B6F9BB" bold="true" box="[192,403,1602,1626]" pageId="5" pageNumber="4">Key Placement</emphasis>
|
||
.
|
||
<taxonomicName id="4C644D2A0F00E7689FC2F9DCB13CF9BB" authorityName="Skelley & Warner" authorityYear="2015" box="[421,793,1602,1626]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F00E7689FC2F9DCB13CF9BB" box="[421,793,1602,1626]" italics="true" pageId="5" pageNumber="4">Trichonotuloides aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
, is most similar to
|
||
<taxonomicName id="4C644D2A0F00E7689A72F9DCB6DCF9BB" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[1045,1273,1602,1626]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glyptus">
|
||
<emphasis id="B910EABB0F00E7689A72F9DCB6B6F9BB" box="[1045,1171,1602,1626]" italics="true" pageId="5" pageNumber="4">T. glyptus</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C644D2A0F00E7689BE1F9DCB37EF99B" authorityName="Skelley & Warner" authorityYear="2015" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="aphoderrans">
|
||
<emphasis id="B910EABB0F00E7689BE1F9DCB37EF99B" italics="true" pageId="5" pageNumber="4">T. aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
does not easily fall to that species in the key of Dellacasa et al. (2014). To help resolve this problem, we here reproduce and modify their key to include the new species.
|
||
</paragraph>
|
||
<keyStep id="30902E0C0F00E7689EA7F95CB2FEF8D8" pageId="5" pageNumber="4">
|
||
<key id="FFF561980F00E7689EA7F95CB2FEF8D8" pageId="5" pageNumber="4">
|
||
<paragraph id="8BDB36A90F00E7689EA7F95CB2F3F93B" blockId="5.[192,214,1730,1754]" box="[192,214,1730,1754]" pageId="5" pageNumber="4">
|
||
<keyLead id="3095959C0F00E7689EA7F95CB2F3F93B" box="[192,214,1730,1754]" pageId="5" pageNumber="4">1.</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F00E7689EA7F8BCB2FEF8D8" blockId="5.[192,219,1826,1849]" box="[192,219,1826,1849]" pageId="5" pageNumber="4">
|
||
<keyLead id="3095959C0F00E7689EA7F8BCB2FEF8D8" box="[192,219,1826,1849]" pageId="5" pageNumber="4">—</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</keyStep>
|
||
<paragraph id="8BDB36A90F00E7689F47F95CB785F8F8" blockId="5.[288,1441,1730,1882]" pageId="5" pageNumber="4">
|
||
Elytra strongly denticulate at shoulder; striae almost as broad as intervals; latter coarsely, densely, evenly punctured; pronotum dually punctured with hind angles truncate. Dark brownish. Length 4.0–
|
||
<quantity id="4C9C9B4C0F00E7689C29F89FB089F8F8" box="[590,684,1793,1817]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="4" unit="mm" value="4.5">4.5 mm</quantity>
|
||
.
|
||
<collectingCountry id="F37376390F00E7689CD0F89CB166F8FB" box="[695,835,1794,1818]" name="Guatemala" pageId="5" pageNumber="4">Guatemala</collectingCountry>
|
||
,
|
||
<collectingCountry id="F37376390F00E7689D28F89CB180F8FB" box="[847,933,1794,1818]" name="Mexico" pageId="5" pageNumber="4">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A0F84B0F00E7689DD1F89CB63AF8FB" box="[950,1055,1794,1818]" country="Mexico" name="Chiapas" pageId="5" pageNumber="4">Chiapas</collectingRegion>
|
||
). .......
|
||
<taxonomicName id="4C644D2A0F00E7689A00F89CB785F8F8" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[1127,1440,1793,1818]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="latecrenatus">
|
||
<emphasis id="B910EABB0F00E7689A00F89CB785F8F8" bold="true" box="[1127,1440,1793,1818]" pageId="5" pageNumber="4">
|
||
<emphasis id="B910EABB0F00E7689A00F89CB712F8FB" bold="true" box="[1127,1335,1793,1818]" italics="true" pageId="5" pageNumber="4">T. latecrenatus</emphasis>
|
||
(Bates)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F00E7689F47F8BCB785F8B8" pageId="5" pageNumber="4" type="description">
|
||
<paragraph id="8BDB36A90F00E7689F47F8BCB785F8B8" blockId="5.[288,1441,1730,1882]" pageId="5" pageNumber="4">
|
||
Elytra not denticulate at shoulder, at most with distinct epipleural carina; striae not so broad; pronotum irregularly punctured, with hind angles obtusely rounded. ...................................
|
||
<emphasis id="B910EABB0F00E7689BF6F8DFB785F8B8" bold="true" box="[1425,1440,1857,1881]" pageId="5" pageNumber="4">2</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F03E76B9EA7FF5CB785FEF8" pageId="6" pageNumber="5" type="key">
|
||
<key id="FFF561980F03E76B9EA7FF5CB2D2FDDB" pageId="6" pageNumber="5">
|
||
<keyStep id="30902E0C0F03E76B9EA7FF5CB2FEFED8" pageId="6" pageNumber="5">
|
||
<paragraph id="8BDB36A90F03E76B9EA7FF5CB2D3FF3B" blockId="6.[192,246,194,218]" box="[192,246,194,218]" pageId="6" pageNumber="5">
|
||
<keyLead id="3095959C0F03E76B9EA7FF5CB2D3FF3B" box="[192,246,194,218]" pageId="6" pageNumber="5">2(1).</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F03E76B9EA7FEBCB2FEFED8" blockId="6.[192,219,290,313]" box="[192,219,290,313]" pageId="6" pageNumber="5">
|
||
<keyLead id="3095959C0F03E76B9EA7FEBCB2FEFED8" box="[192,219,290,313]" pageId="6" pageNumber="5">—</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30902E0C0F03E76B9EA7FE1CB2FEFE18" pageId="6" pageNumber="5">
|
||
<paragraph id="8BDB36A90F03E76B9EA7FE1CB2D3FE7B" blockId="6.[192,246,386,410]" box="[192,246,386,410]" pageId="6" pageNumber="5">
|
||
<keyLead id="3095959C0F03E76B9EA7FE1CB2D3FE7B" box="[192,246,386,410]" pageId="6" pageNumber="5">3(2).</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F03E76B9EA7FE7CB2FEFE18" blockId="6.[192,219,482,505]" box="[192,219,482,505]" pageId="6" pageNumber="5">
|
||
<keyLead id="3095959C0F03E76B9EA7FE7CB2FEFE18" box="[192,219,482,505]" pageId="6" pageNumber="5">—</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8BDB36A90F03E76B9EA7FDBCB2D2FDDB" blockId="6.[192,247,546,570]" box="[192,247,546,570]" pageId="6" pageNumber="5">
|
||
<keyLead id="3095959C0F03E76B9EA7FDBCB2D2FDDB" box="[192,247,546,570]" pageId="6" pageNumber="5">
|
||
<keyStep id="30902E0C0F03E76B9EA7FDBCB2D2FDDB" box="[192,247,546,570]" pageId="6" pageNumber="5">4(3).</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
<paragraph id="8BDB36A90F03E76B9F47FF5CB785FEF8" blockId="6.[288,1441,194,698]" pageId="6" pageNumber="5">
|
||
Clypeus widely rounded at sides of median sinuosity; genae rounded, feebly protruding from the eyes; fore tibiae spur almost straight and acuminate in both sexes. Blackish. Length 4.5–6.0 mm.
|
||
<collectingCountry id="F37376390F03E76B9F11FE9CB3E9FEFB" box="[374,460,258,282]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A0F84B0F03E76B9FBBFE9CB06EFEFB" box="[476,587,258,282]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="5">Veracruz</collectingRegion>
|
||
).............................
|
||
<taxonomicName id="4C644D2A0F03E76B9D49FE9CB785FEF8" authority="Dellacasa, Dellacasa and Gordon" authorityName="Dellacasa, Dellacasa and Gordon" box="[814,1440,257,282]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="hansferyi">
|
||
<emphasis id="B910EABB0F03E76B9D49FE9CB785FEF8" bold="true" box="[814,1440,257,282]" pageId="6" pageNumber="5">
|
||
<emphasis id="B910EABB0F03E76B9D49FE9CB1F7FEFB" bold="true" box="[814,978,257,282]" italics="true" pageId="6" pageNumber="5">T. hansferyi</emphasis>
|
||
Dellacasa, Dellacasa and Gordon
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E65220F03E76A9F47FEBCB785FED8" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="description">
|
||
<paragraph id="8BDB36A90F03E76B9F47FEBCB785FE98" blockId="6.[288,1441,194,698]" pageId="6" pageNumber="5">
|
||
Clypeus subangulate at sides of median sinuosity; genae obtusely to angularly rounded, strongly protruding from the eyes; fore tibiae spur in males strongly curved or hooked inwardly at apex..............................................................................................................................................
|
||
<emphasis id="B910EABB0F03E76B9BF6FEFFB785FE98" bold="true" box="[1425,1440,353,377]" pageId="6" pageNumber="5">3</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F03E76B9F47FE1CB785FE3B" blockId="6.[288,1441,194,698]" pageId="6" pageNumber="5">
|
||
Elytral intervals densely, coarsely, evenly punctured, surface smooth between punctures (
|
||
<figureCitation id="135F2A2C0F03E76B9B2AFE1CB7B7FE78" box="[1357,1426,385,410]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="6" pageNumber="5">Fig. 5</figureCitation>
|
||
); genae angularly rounded (
|
||
<figureCitation id="135F2A2C0F03E76B9C1CFE3CB0E6FE5B" box="[635,707,418,442]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="6" pageNumber="5">Fig. 4</figureCitation>
|
||
); upper mesotibial spur longer than first tarsomere. Length 3.7–5.0 mm.
|
||
<collectingCountry id="F37376390F03E76B9FB0FE5CB034FE3B" box="[471,529,450,474]" name="United States of America" pageId="6" pageNumber="5">USA</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A0F84B0F03E76B9C47FE5CB05AFE3B" box="[544,639,450,474]" country="United States of America" name="Arizona" pageId="6" pageNumber="5">Arizona</collectingRegion>
|
||
)...................................................................
|
||
<emphasis id="B910EABB0F03E76B9A1AFE5CB785FE3B" bold="true" box="[1149,1440,449,474]" pageId="6" pageNumber="5">
|
||
<taxonomicName id="4C644D2A0F03E76B9A1AFE5CB774FE3B" authority="Skelley & Warner, 2015" authorityName="Skelley & Warner" authorityYear="2015" box="[1149,1361,449,474]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="aphoderrans" status="sp. nov.">
|
||
<emphasis id="B910EABB0F03E76B9A1AFE5CB774FE3B" bold="true" box="[1149,1361,449,474]" italics="true" pageId="6" pageNumber="5">T. aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22357C00F03E76B9B3FFE5CB785FE3B" box="[1368,1440,450,474]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F03E76B9F47FE7CB785FDF8" blockId="6.[288,1441,194,698]" pageId="6" pageNumber="5">
|
||
Elytral intervals with coarse punctation irregularly vermiculose (
|
||
<figureCitation id="135F2A2C0F03E76B9A4FFE7CB64EFE1B" box="[1064,1131,482,506]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="6" pageNumber="5">Fig. 2</figureCitation>
|
||
); genae obtusely rounded (
|
||
<figureCitation id="135F2A2C0F03E76B9F26FD9CB3A0FDFB" box="[321,389,514,538]" captionStart="Figures 1–8" captionStartId="4.[192,288,1846,1868]" captionTargetBox="[192,1440,280,1825]" captionTargetId="figure-68@4.[192,1440,280,1825]" captionTargetPageId="4" captionText="Figures 1–8. Trichonotuloides spp. 1-3) T. glyptus (Bates) male. 1) Dorsal habitus. 2) Elytra and pronotal base. 3) Protibia, dorsal view. 4-6) T. aphoderrans n. sp., holotype male. 4) Dorsal habitus. 5) Elytra and pronotal base. 6) Protibia, dorsal view. 7-8) T. aphoderrans male genitalia, paratype male." figureDoi="http://doi.org/10.5281/zenodo.5181881" httpUri="https://zenodo.org/record/5181881/files/figure.png" pageId="6" pageNumber="5">Fig. 1</figureCitation>
|
||
); upper mesotibial spur shorter than first tarsomere.
|
||
<collectingCountry id="F37376390F03E76B9A7AFD9CB653FDFB" box="[1053,1142,514,538]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
|
||
......................................
|
||
<emphasis id="B910EABB0F03E76B9BF6FD9FB785FDF8" bold="true" box="[1425,1440,513,537]" pageId="6" pageNumber="5">4</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDB36A90F03E76B9F47FDBCB785FD58" blockId="6.[288,1441,194,698]" pageId="6" pageNumber="5">
|
||
Epistome subshiny, coarsely, distinctly almost evenly punctured; clypeus relatively more feebly sinuate at middle;
|
||
<materialsCitation id="3B0C3CF40F03E76B9C43FDDCB785FD58" country="Mexico" location="Colima" pageId="6" pageNumber="5" specimenCount="1" stateProvince="Colima">
|
||
spur of fore tibiae, in males, spatulate and apically hooked. Blackish or piceous black, margins of head and pronotum reddish. Species relatively larger: length 4.5–5.0 mm.
|
||
<collectingCountry id="F37376390F03E76B9F11FD1CB3E9FD7B" box="[374,460,642,666]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A0F84B0F03E76B9FBBFD1CB01DFD7B" box="[476,568,642,666]" country="Mexico" name="Colima" pageId="6" pageNumber="5">Colima</collectingRegion>
|
||
,
|
||
<collectingRegion id="49A0F84B0F03E76B9C23FD1CB096FD7B" box="[580,691,642,666]" country="Mexico" name="Durango" pageId="6" pageNumber="5">Durango</collectingRegion>
|
||
,
|
||
<collectingRegion id="49A0F84B0F03E76B9CD9FD1CB107FD7B" box="[702,802,642,666]" country="Mexico" name="Hidalgo" pageId="6" pageNumber="5">Hidalgo</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37376390F03E76B9D49FD1CB1ADFD7B" box="[814,904,642,666]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="49A0F84B0F03E76B9DF4FD1CB1D4FD7B" box="[915,1009,642,666]" country="Mexico" name="Oaxaca" pageId="6" pageNumber="5">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingRegion id="49A0F84B0F03E76B9D9AFD1CB670FD7B" box="[1021,1109,642,666]" country="Mexico" name="Puebla" pageId="6" pageNumber="5">Puebla</collectingRegion>
|
||
,
|
||
<collectingRegion id="49A0F84B0F03E76B9A05FD1CB6F4FD7B" box="[1122,1233,642,666]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="5">Veracruz</collectingRegion>
|
||
).......................... .......................................................................................................................
|
||
<taxonomicName id="4C644D2A0F03E76B9AD7FD3CB785FD58" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[1200,1440,673,698]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="glyptus">
|
||
<emphasis id="B910EABB0F03E76B9AD7FD3CB785FD58" bold="true" box="[1200,1440,673,698]" pageId="6" pageNumber="5">
|
||
<emphasis id="B910EABB0F03E76B9AD7FD3CB712FD5B" bold="true" box="[1200,1335,673,698]" italics="true" pageId="6" pageNumber="5">T. glyptus</emphasis>
|
||
(Bates)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF1B66210F03E76B9EA7F8A8B3DDF867" ID-DOI="http://doi.org/10.5281/zenodo.5181883" ID-Zenodo-Dep="5181883" httpUri="https://zenodo.org/record/5181883/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,288,1846,1868]" targetBox="[208,1425,738,1836]" targetPageId="6">
|
||
<paragraph id="8BDB36A90F03E76B9EA7F8A8B3DDF867" blockId="6.[192,1440,1846,1926]" pageId="6" pageNumber="5">
|
||
<emphasis id="B910EABB0F03E76B9EA7F8A8B344F8AA" bold="true" box="[192,353,1846,1868]" pageId="6" pageNumber="5">Figures 9–14.</emphasis>
|
||
<taxonomicName id="4C644D2A0F03E76B9F01F8A8B097F8AA" authority="Skelley & Warner, 2015" authorityName="Skelley & Warner" authorityYear="2015" box="[358,690,1846,1867]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="aphoderrans" status="sp. nov.">
|
||
<emphasis id="B910EABB0F03E76B9F01F8A8B097F8AA" box="[358,690,1846,1867]" italics="true" pageId="6" pageNumber="5">Trichonotuloides aphoderrans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22357C00F03E76B9CD0F8A8B0D4F8AA" box="[695,753,1846,1867]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
<emphasis id="B910EABB0F03E76B9C91F8A8B128F8AD" bold="true" box="[758,781,1846,1868]" pageId="6" pageNumber="5">9)</emphasis>
|
||
Male holotype, ventral view.
|
||
<emphasis id="B910EABB0F03E76B9A29F8A8B6BEF8AD" bold="true" box="[1102,1179,1846,1868]" pageId="6" pageNumber="5">10–11)</emphasis>
|
||
Female allotype, dorsal and ventral view.
|
||
<emphasis id="B910EABB0F03E76B9FF7F8CCB390F889" bold="true" box="[400,437,1874,1896]" pageId="6" pageNumber="5">12)</emphasis>
|
||
Male holotype head and pronotum dorso-lateral view.
|
||
<emphasis id="B910EABB0F03E76B9A4FF8CCB668F889" bold="true" box="[1064,1101,1874,1896]" pageId="6" pageNumber="5">13)</emphasis>
|
||
Female allotype protibia.
|
||
<emphasis id="B910EABB0F03E76B9B1CF8CCB785F889" bold="true" box="[1403,1440,1874,1896]" pageId="6" pageNumber="5">14)</emphasis>
|
||
Epipharynx, paratype male.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB36A90F02E76A9EA7FF5CB2FEFF38" blockId="7.[192,219,194,217]" box="[192,219,194,217]" pageId="7" pageNumber="6">—</paragraph>
|
||
<paragraph id="8BDB36A90F02E76A9F47FF5CB785FED8" blockId="7.[288,1441,194,314]" pageId="7" pageNumber="6">
|
||
Epistome dull, confusedly subrugosely punctured; clypeus relatively more deeply sinuate at middle, fore tibiae spur in males stout, subcylindrical, acuminate and strongly curved inwardly apically. Blackish. Species relatively smaller: length 3.5–4.0 mm.
|
||
<collectingCountry id="F37376390F02E76A9A05FE9CB69DFEFB" box="[1122,1208,258,282]" name="Mexico" pageId="7" pageNumber="6">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A0F84B0F02E76A9AAFFE9CB71FFEFB" box="[1224,1338,258,282]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="7" pageNumber="6">Veracruz</collectingRegion>
|
||
)............ ...............................................................
|
||
<taxonomicName id="4C644D2A0F02E76A9D78FEBCB785FED8" authority="Dellacasa, Dellacasa and Gordon" authorityName="Dellacasa, Dellacasa and Gordon" box="[799,1440,289,314]" class="Insecta" family="Aphodiidae" genus="Trichonotuloides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="alfonsinae">
|
||
<emphasis id="B910EABB0F02E76A9D78FEBCB785FED8" bold="true" box="[799,1440,289,314]" pageId="7" pageNumber="6">
|
||
<emphasis id="B910EABB0F02E76A9D78FEBCB1F7FEDB" bold="true" box="[799,978,289,314]" italics="true" pageId="7" pageNumber="6">T. alfonsinae</emphasis>
|
||
Dellacasa, Dellacasa and Gordon
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |