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<document id="D69B18EF7B09E7BB86ADAE0181AB2F9B" ID-DOI="10.1080/00222933.2012.655797" ID-ISSN="1464-5262" ID-Zenodo-Dep="5200053" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628905690664" checkinUser="felipe" docAuthor="Takahashi, Yoshie, Kajihara, Hiroshi &amp; Mawatari, Shunsuke F." docDate="2012" docId="03CD87A8FF836504DB9EFBFB995BFC4D" docLanguage="en" docName="JNatHist.46.21-22.1337-1358.pdf" docOrigin="Journal of Natural History 46 (21 - 22)" docSource="http://dx.doi.org/10.1080/00222933.2012.655797" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Nymphon nagannuense Takahashi &amp; Kajihara &amp; Mawatari 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1349" masterDocId="FFF4FFD0FF896509DA5AFE519D7AFFE8" masterDocTitle="Sea spiders of the genus Nymphon (Arthropoda: Pycnogonida) from waters around the Nansei Islands, Japan" masterLastPageNumber="1358" masterPageNumber="1337" pageNumber="1346" updateTime="1699055937477" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="E2E2DBC58A79F45E43279B64B73C1F30" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BF850A18732D78C9D6E09F49DBFD1218">
<mods:title id="7A94323B3332056DE43DE5139121399E">Sea spiders of the genus Nymphon (Arthropoda: Pycnogonida) from waters around the Nansei Islands, Japan</mods:title>
</mods:titleInfo>
<mods:name id="1B897A55C5DE26BCE97DE2B4D37E69A4" type="personal">
<mods:role id="E4A8E0EBA9CE7181B9830B188B523C74">
<mods:roleTerm id="3AF5AA839EE4EF03BBDB9E6ED660D96D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2D2729A425AFE2D189DF0D0BA2D72C9">Takahashi, Yoshie</mods:namePart>
</mods:name>
<mods:name id="367C1C7918478EEC200F69D3294D22E7" type="personal">
<mods:role id="1B052EA663213571914471F7EC00ADCE">
<mods:roleTerm id="BF8BF9F3E337F314DE107D09099CCFD9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1B64B6A3912BAF22E0362DCC5712EBE">Kajihara, Hiroshi</mods:namePart>
</mods:name>
<mods:name id="E9CE4EA42F41A89FAD5642061F2EB668" type="personal">
<mods:role id="EFE3B43BA106976EA7EE3A710BFD274C">
<mods:roleTerm id="A2EB26690CD949320BE667F34ACFE031">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9C2940E22D0937D3247A742CB8D799D">Mawatari, Shunsuke F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CD70E0A0C83C1AB337D7D9F11E379D59">text</mods:typeOfResource>
<mods:relatedItem id="5B2C15C61B2DFA8DC32490504416A0BD" type="host">
<mods:titleInfo id="33F549A6F99E5BC9D0493A4E8A94CCB1">
<mods:title id="CEBB0B1F71785F2DFDF842D2CC58EAA1">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="C9744B7647B4F625E2D1C31092B6A1CD">
<mods:date id="64976B3E7643398E251CCDD716A76CA5">2012</mods:date>
<mods:detail id="EBD0466D19908F896A58ACEA0DBD722F" type="pubDate">
<mods:number id="CDFAD9EEC7B77955B749CCCC84D35B95">2012-06-30</mods:number>
</mods:detail>
<mods:detail id="9D986331B4CC329C846D7D9BC76E5269" type="volume">
<mods:number id="526EA87258CB63AA9E0321E87816B032">46</mods:number>
</mods:detail>
<mods:detail id="EBF36EC1850E0879E20EDEA28B0B8C4C" type="issue">
<mods:number id="03B10109F859568C11FC617BCA264143">21 - 22</mods:number>
</mods:detail>
<mods:extent id="F5CE11A089E3DC0438E2DD68E5CE3717" unit="page">
<mods:start id="055C2E0EF4F44A699A69EF22DC4DBC27">1337</mods:start>
<mods:end id="492E732C27E4600AB9E9B5C0ADEB7AA9">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3D352DBCBAA82AFECC4A0DDBE99C24E8">
<mods:url id="8BC69A16DA38711AD2A087CF9B2D46B9">http://dx.doi.org/10.1080/00222933.2012.655797</mods:url>
</mods:location>
<mods:classification id="0137039C87776071AEA964048933C6D5">journal article</mods:classification>
<mods:identifier id="A3B41AFE8165CC2ABD979A5F7E2B8713" type="DOI">10.1080/00222933.2012.655797</mods:identifier>
<mods:identifier id="984D789093C221BEE8CDB4398D98BEA4" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="D57BE6314C79F2FEA77E0A89E3041DD7" type="Zenodo-Dep">5200053</mods:identifier>
</mods:mods>
<treatment id="03CD87A8FF836504DB9EFBFB995BFC4D" LSID="urn:lsid:plazi:treatment:03CD87A8FF836504DB9EFBFB995BFC4D" httpUri="http://treatment.plazi.org/id/03CD87A8FF836504DB9EFBFB995BFC4D" lastPageId="13" lastPageNumber="1349" pageId="10" pageNumber="1346">
<subSubSection id="C37E6535FF836503DB9EFBFB9E66FA2A" box="[452,796,1450,1475]" pageId="10" pageNumber="1346" type="nomenclature">
<paragraph id="8BDB36BEFF836503DB9EFBFB9E66FA2A" blockId="10.[452,796,1450,1506]" box="[452,796,1450,1475]" pageId="10" pageNumber="1346">
<heading id="D09381D2FF836503DB9EFBFB9E66FA2A" box="[452,796,1450,1475]" centered="true" fontSize="36" level="2" pageId="10" pageNumber="1346" reason="7">
<taxonomicName id="4C644D3DFF836503DB9EFBFB9FBBFA2A" ID-CoL="486Y9" authority="Takahashi &amp; Kajihara &amp; Mawatari, 2012" authorityName="Takahashi &amp; Kajihara &amp; Mawatari" authorityYear="2012" box="[452,705,1450,1475]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="10" pageNumber="1346" phylum="Arthropoda" rank="species" species="nagannuense" status="sp. nov.">
<emphasis id="B910EAACFF836503DB9EFBFB9FBBFA2A" bold="true" box="[452,705,1450,1475]" italics="true" pageId="10" pageNumber="1346">Nymphon nagannuense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357D7FF836503D89DFBFA9E66FA2A" box="[711,796,1451,1474]" pageId="10" pageNumber="1346" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E6535FF836504D86DFB9B995BFC4D" lastPageId="13" lastPageNumber="1349" pageId="10" pageNumber="1346" type="description">
<paragraph id="8BDB36BEFF836503D86DFB9B9FD2FA0A" blockId="10.[452,796,1450,1506]" box="[567,680,1482,1506]" pageId="10" pageNumber="1346">
(
<figureCitation id="135F2A3BFF836503D865FB9B9FE5FA0A" box="[575,671,1482,1506]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="10" pageNumber="1346">Figure 5</figureCitation>
)
</paragraph>
<paragraph id="8BDB36BEFF836503DAD5F8439C1BF9C2" blockId="10.[143,353,1554,1578]" box="[143,353,1554,1578]" pageId="10" pageNumber="1346">
<emphasis id="B910EAACFF836503DAD5F8439C1BF9C2" box="[143,353,1554,1578]" italics="true" pageId="10" pageNumber="1346">Material examined</emphasis>
</paragraph>
<paragraph id="8BDB36BEFF836503DAD5F8139FD7F992" blockId="10.[143,1103,1598,1659]" pageId="10" pageNumber="1346">
<materialsCitation id="3B0C3CE3FF836503DAD5F8139FD7F992" collectingDate="2005-05-22" collectionCode="ZIHU" collectorName="Y. Takahashi." latitude="26.244" location="Nagannu Island" longLatPrecision="12" longitude="127.51683" pageId="10" pageNumber="1346" specimenCode="ZIHU 3329" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="B910EAACFF836503DAD5F8139D87F9B2" box="[143,253,1602,1626]" italics="true" pageId="10" pageNumber="1346">
<typeStatus id="54DF881CFF836503DAD5F8139D82F9B2" box="[143,248,1602,1626]" pageId="10" pageNumber="1346" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBC29EC5FF836503DB5CF8129CF5F9B2" box="[262,399,1602,1626]" collectionCode="ZIHU" country="Japan" httpUri="http://grbio.org/cool/mc14-1jj5" name="Zoological Institute, Hokkaido University" pageId="10" pageNumber="1346">ZIHU 3329</specimenCode>
, male,
<geoCoordinate id="EE505079FF836503DBB8F8139F11F9B2" box="[482,619,1598,1626]" degrees="26" direction="north" minutes="14.64" orientation="latitude" pageId="10" pageNumber="1346" precision="9" value="26.244">
26
<superScript id="7C119BF6FF836503DBA7F86F9F7CF9A7" attach="none" box="[509,518,1598,1615]" fontSize="7" pageId="10" pageNumber="1346">◦</superScript>
14.64
<superScript id="7C119BF6FF836503D81EF86F9F33F9A7" attach="left" box="[580,585,1598,1615]" fontSize="7" pageId="10" pageNumber="1346">′</superScript>
N
</geoCoordinate>
,
<geoCoordinate id="EE505079FF836503D823F8139E76F9B2" box="[633,780,1598,1626]" degrees="127" direction="east" minutes="31.01" orientation="longitude" pageId="10" pageNumber="1346" precision="9" value="127.51683">
127
<superScript id="7C119BF6FF836503D8FBF86F9FD0F9A7" attach="none" box="[673,682,1598,1615]" fontSize="7" pageId="10" pageNumber="1346">◦</superScript>
31.01
<superScript id="7C119BF6FF836503D8B2F86F9F97F9A7" attach="left" box="[744,749,1598,1615]" fontSize="7" pageId="10" pageNumber="1346">′</superScript>
E
</geoCoordinate>
,
<location id="8EBB6065FF836503D941F8129EAFF9B2" LSID="urn:lsid:plazi:treatment:03CD87A8FF836504DB9EFBFB995BFC4D:8EBB6065FF836503D941F8129EAFF9B2" box="[795,981,1602,1627]" latitude="26.244" longLatPrecision="12" longitude="127.51683" name="Nagannu Island" pageId="10" pageNumber="1346" stateProvince="Okinawa">Nagannu Island</location>
,
<collectingRegion id="49A0F85CFF836503D9BEF8139936F9B2" box="[996,1100,1602,1626]" country="Japan" name="Okinawa" pageId="10" pageNumber="1346">Okinawa</collectingRegion>
,
<date id="FFDA107EFF836503DAD5F8339C59F992" box="[143,291,1634,1659]" pageId="10" pageNumber="1346" value="2005-05-22">
<collectingDate id="EF9EE996FF836503DAD5F8339C59F992" box="[143,291,1634,1659]" pageId="10" pageNumber="1346" value="2005-05-22">22 May 2005</collectingDate>
</date>
, 57 m depth, dredge,
<collectorName id="26915368FF836503D848F8329FD7F992" box="[530,685,1634,1658]" pageId="10" pageNumber="1346">Y. Takahashi.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BDB36BEFF826502DA89FEE29E34FF02" blockId="11.[211,1171,174,234]" pageId="11" pageNumber="1347">
<materialsCitation id="3B0C3CE3FF826502DA89FEE29E30FF02" collectingDate="1994-11-11" collectionCode="ZIHU" collectorName="S. Ohtsuka" latitude="27.9805" location="Uke Island" longLatPrecision="12" longitude="129.28" pageId="11" pageNumber="1347" specimenCode="ZIHU 3330" specimenCount="1" typeStatus="paratype">
<emphasis id="B910EAACFF826502DA89FEE29C47FF22" box="[211,317,179,202]" italics="true" pageId="11" pageNumber="1347">
<typeStatus id="54DF881CFF826502DA89FEE29C43FF22" box="[211,313,179,202]" pageId="11" pageNumber="1347" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="DBC29EC5FF826502DB12FEE29CA9FF22" box="[328,467,178,202]" collectionCode="ZIHU" country="Japan" httpUri="http://grbio.org/cool/mc14-1jj5" name="Zoological Institute, Hokkaido University" pageId="11" pageNumber="1347">ZIHU 3330</specimenCode>
, male,
<geoCoordinate id="EE505079FF826502D871FEE39FCCFF22" box="[555,694,174,202]" degrees="27" direction="north" minutes="58.83" orientation="latitude" pageId="11" pageNumber="1347" precision="9" value="27.9805">
27
<superScript id="7C119BF6FF826502D81CFEFF9F35FF57" attach="none" box="[582,591,174,191]" fontSize="7" pageId="11" pageNumber="1347">◦</superScript>
58.83
<superScript id="7C119BF6FF826502D8D6FEFF9FEBFF57" attach="left" box="[652,657,174,191]" fontSize="7" pageId="11" pageNumber="1347">′</superScript>
N
</geoCoordinate>
,
<geoCoordinate id="EE505079FF826502D89CFEE39E21FF22" box="[710,859,174,202]" degrees="129" direction="east" minutes="16.80" orientation="longitude" pageId="11" pageNumber="1347" precision="9" value="129.28">
129
<superScript id="7C119BF6FF826502D8B4FEFF9F8DFF57" attach="none" box="[750,759,174,191]" fontSize="7" pageId="11" pageNumber="1347">◦</superScript>
16.80
<superScript id="7C119BF6FF826502D96FFEFF9E40FF57" attach="left" box="[821,826,174,191]" fontSize="7" pageId="11" pageNumber="1347">′</superScript>
E
</geoCoordinate>
, southwest of
<location id="8EBB6065FF826502DE51FEE399F5FF22" LSID="urn:lsid:plazi:treatment:03CD87A8FF836504DB9EFBFB995BFC4D:8EBB6065FF826502DE51FEE399F5FF22" box="[1035,1167,178,202]" latitude="27.9805" longLatPrecision="12" longitude="129.28" name="Uke Island" pageId="11" pageNumber="1347">Uke Island</location>
,
<date id="FFDA107EFF826502DA89FE839CD0FF02" box="[211,426,210,234]" pageId="11" pageNumber="1347" value="1994-11-11">
<collectingDate id="EF9EE996FF826502DA89FE839CD0FF02" box="[211,426,210,234]" pageId="11" pageNumber="1347" value="1994-11-11">
11 November
<quantity id="4C9C9B5BFF826502DB28FE839CD0FF02" box="[370,426,210,234]" metricMagnitude="6" metricUnit="m" metricValue="1.994128" pageId="11" pageNumber="1347" unit="m" value="1994128.0">1994</quantity>
</collectingDate>
</date>
, 128 m depth,
<collectingMethod id="52254EA9FF826502D814FE839FC5FF02" box="[590,703,210,234]" pageId="11" pageNumber="1347">sledge net</collectingMethod>
,
<collectorName id="26915368FF826502D890FE839E30FF02" box="[714,842,210,234]" pageId="11" pageNumber="1347">S. Ohtsuka</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDB36BEFF826502DA89FF7A9FEAFEAA" blockId="11.[211,656,298,322]" box="[211,656,298,322]" pageId="11" pageNumber="1347">
<emphasis id="B910EAACFF826502DA89FF7A9FEAFEAA" box="[211,656,298,322]" italics="true" pageId="11" pageNumber="1347">
Measurements of
<typeStatus id="54DF881CFF826502DBCDFF7B9C8FFEAA" box="[407,501,298,322]" pageId="11" pageNumber="1347" type="holotype">holotype</typeStatus>
(millimetres)
</emphasis>
</paragraph>
<paragraph id="8BDB36BEFF826502DA89FF08995FFE19" blockId="11.[211,1172,345,497]" pageId="11" pageNumber="1347">Trunk length, 2.3; body width, 1.13; proboscis length, 0.60; length of abdomen, 0.25; third leg (L3) coxa 1, 0.20; L3 coxa 2, 1.04; L3 coxa 3, 0.25; L3 femur, 2.04; L3 tibia 1, 2.60; L3 tibia 2, 3.16; L3 tarsus, 0.82; L3 propodus, 0.77; L3 main claw, 0.19; L3 auxiliary claws, 0.19; oviger first article (O1), 0.15; O2, 0.23; O3, 0.23; O4, 0.82; O5, 1.07; O6, 0.35; O7, 0.21; O8, 0.19; O9, 0.16; O10, 0.15; oviger terminal claw, 0.11.</paragraph>
<paragraph id="8BDB36BEFF826502DA89FC609C31FDA1" blockId="11.[211,331,561,585]" box="[211,331,561,585]" pageId="11" pageNumber="1347">
<emphasis id="B910EAACFF826502DA89FC609C31FDA1" box="[211,331,561,585]" italics="true" pageId="11" pageNumber="1347">Etymology</emphasis>
</paragraph>
<paragraph id="8BDB36BEFF826502DA89FC319951FD90" blockId="11.[211,1067,608,633]" box="[211,1067,608,633]" pageId="11" pageNumber="1347">
<materialsCitation id="3B0C3CE3FF826502DA89FC31995DFD90" box="[211,1063,608,633]" location="Nagannu Island" pageId="11" pageNumber="1347" specimenCount="1">
The specific epithet, an adjective, refers to the
<typeStatus id="54DF881CFF826502D882FC309E72FD91" box="[728,776,609,633]" pageId="11" pageNumber="1347">type</typeStatus>
locality,
<location id="8EBB6065FF826502D934FC30995DFD90" LSID="urn:lsid:plazi:treatment:03CD87A8FF836504DB9EFBFB995BFC4D:8EBB6065FF826502D934FC30995DFD90" box="[878,1063,608,633]" name="Nagannu Island" pageId="11" pageNumber="1347">Nagannu Island</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDB36BEFF826502DA89FCE89C28FD38" blockId="11.[211,338,697,720]" box="[211,338,697,720]" pageId="11" pageNumber="1347">
<emphasis id="B910EAACFF826502DA89FCE89C28FD38" box="[211,338,697,720]" italics="true" pageId="11" pageNumber="1347">Description</emphasis>
</paragraph>
<paragraph id="8BDB36BEFF826502DA89FCB99C09FBF7" blockId="11.[211,1172,744,1660]" pageId="11" pageNumber="1347">
Trunk (
<figureCitation id="135F2A3BFF826502DB72FCB99CCCFCE8" box="[296,438,744,768]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="11" pageNumber="1347">Figure 5A,B</figureCitation>
) elongate, completely segmented. Lateral processes slightly more than twice as long as their diameter and separated by twice their diameter, without tubercles or setae. Neck moderately long, at least half as narrow as trunk diameter, widening distally; crop 3.6 times as wide as the minimum width of neck. Ocular tubercle in midline at level of anterior extent of first lateral processes, almost as tall as its basal diameter, rounded, with two small lateral projections. Eyes large, lightly pigmented. Proboscis cylindrical, rounded distally, slightly constricted laterally at onequarter the length from the tip, 2.6 times as long as basal diameter. Abdomen robust, twice as long as basal diameter, slightly tapering distally, with several short setae dorsolaterally.
</paragraph>
<paragraph id="8BDB36BEFF826502DAA4FA769F8CFB36" blockId="11.[211,1172,744,1660]" pageId="11" pageNumber="1347">
Chelifores (
<figureCitation id="135F2A3BFF826502DBDAFA769F75FBD7" box="[384,527,1063,1087]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="11" pageNumber="1347">Figure 5A,B</figureCitation>
) very slender; scapes slightly curved endally, six times as long as its diameter, with many tiny setae laterally and some longer setae dorsally and dorsodistally. Chela palms (
<figureCitation id="135F2A3BFF826502D85FFA379F0DFB96" box="[517,631,1126,1150]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="11" pageNumber="1347">Figure 5C</figureCitation>
) almost twice as long as wide, with several dorsal setae. Fingers very long; movable finger straight, with 16 sharp teeth endally, without setae; immovable finger slightly curved endally, with 18 sharp teeth endally and with slight inflation bearing several long setae at base.
</paragraph>
<paragraph id="8BDB36BEFF826502DAA4FAB79E10FA75" blockId="11.[211,1172,744,1660]" pageId="11" pageNumber="1347">
Palp (
<figureCitation id="135F2A3BFF826502DB67FAB79CC8FB16" box="[317,434,1254,1278]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="11" pageNumber="1347">Figure 5D</figureCitation>
) first segment as long as wide; second segment longest, 1.2 times as long as third segment, armed with low dorsal tubercle distally; third segment 1.5 times as long as fifth segment, with several ventral setae; fourth segment two-thirds length of fifth segment, with a few lateral setae and many ventral setae; fifth segment with many lateral and ventral setae and longer distal setae. Second and third segments each with several short setae ventrodistally (not visible in
<figureCitation id="135F2A3BFF826502D8BCFBD49E27FA75" box="[742,861,1413,1437]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="11" pageNumber="1347">Figure 5D</figureCitation>
).
</paragraph>
<paragraph id="8BDB36BEFF826502DAA4FBF49C2DF994" blockId="11.[211,1172,744,1660]" pageId="11" pageNumber="1347">
Ovigers (
<figureCitation id="135F2A3BFF826502DB3DFBF49C8FFA55" box="[359,501,1445,1469]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="11" pageNumber="1347">Figure 5E,F</figureCitation>
) long, 10-segmented; first segment very short, as long as wide; second segment equal in length to third segment; fourth segment about 0.8 times as long as fifth; sixth segment 0.3 times as long as fifth segment; fifth and sixth segments each with ectal row of short setae. Strigilis with several ectal setae, with endal compound spines in the formula 7: 6: 5: 6 :: 6. Spines with three denticles per side. Terminal claw 0.7 times the length of terminal segment, slightly curved, with six sharp endal teeth.
</paragraph>
<caption id="DF1B6636FF856505DAD5F8349FCBF93A" ID-DOI="http://doi.org/10.5281/zenodo.5200063" ID-Zenodo-Dep="5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="12" pageNumber="1348" startId="12.[143,210,1637,1659]" targetBox="[143,1101,163,1604]" targetPageId="12">
<paragraph id="8BDB36BEFF856505DAD5F8349FCBF93A" blockId="12.[143,1104,1637,1747]" pageId="12" pageNumber="1348">
Figure 5.
<taxonomicName id="4C644D3DFF856505DAADF8349CA1F993" authority="Takahashi &amp; Kajihara &amp; Mawatari, 2012" authorityName="Takahashi &amp; Kajihara &amp; Mawatari" authorityYear="2012" box="[247,475,1637,1659]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="1348" phylum="Arthropoda" rank="species" species="nagannuense" status="sp. nov.">
<emphasis id="B910EAACFF856505DAADF8349CA1F993" box="[247,475,1637,1659]" italics="true" pageId="12" pageNumber="1348">Nymphon nagannuense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357D7FF856505DBB8F8379F4BF993" box="[482,561,1638,1659]" pageId="12" pageNumber="1348" rank="species">sp. nov.</taxonomicNameLabel>
, ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BDB36BEFF846504DAA4FEF99EBCFE56" blockId="13.[211,1172,167,446]" pageId="13" pageNumber="1349">
Legs (
<figureCitation id="135F2A3BFF846504DB1BFEF69CA9FF57" box="[321,467,167,191]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="13" pageNumber="1349">Figure 5G,H</figureCitation>
) slender, with many minute and some longer setae. Coxae with several tiny setae ventrally and laterally; first coxa short, 1.5 times as long as wide; second coxa long, more than five times as long as first coxa; third coxa 0.4 times length of second coxa. Femur sparsely setose; first tibia 1.3 times as long as femur, with some sparse short setae on ventral side; second tibia 1.5 times as long as femur, setose on ventral side especially, half the thickness of femur; tarsus (
<figureCitation id="135F2A3BFF846504D930FF169E9BFEB7" box="[874,993,327,351]" captionStart="Figure 5" captionStartId="12.[143,210,1637,1659]" captionTargetBox="[143,1101,163,1604]" captionTargetId="graphics-5@12.[143,1101,164,1604]" captionTargetPageId="12" captionText="Figure 5. Nymphon nagannuense sp. nov., ZIHU 3329, holotype, male. (A) Trunk, dorsal view; (B) trunk, lateral view; (C) chelifore; (D) palp; (E) oviger; (E) third leg; (F) enlargement of distal segments of oviger; (G) third leg; (H) enlargement of distal segments of third leg. Scale bars: G, 1 mm; A, B, E, H, 0.5 mm; D, F, 0.25 mm; C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5200063" httpUri="https://zenodo.org/record/5200063/files/figure.png" pageId="13" pageNumber="1349">Figure 5H</figureCitation>
) slightly longer than propodus, both without heel or large heel spines, sole with row of very short spines over entire length; claw slightly curved, about one-quarter as long as propodus; auxiliary claws same length as main claw. Cement gland not visible.
</paragraph>
<paragraph id="8BDB36BEFF846504DA89FFAF9C4FFDFE" blockId="13.[211,309,510,534]" box="[211,309,510,534]" pageId="13" pageNumber="1349">
<emphasis id="B910EAACFF846504DA89FFAF9C4FFDFE" box="[211,309,510,534]" italics="true" pageId="13" pageNumber="1349">Remarks</emphasis>
</paragraph>
<paragraph id="8BDB36BEFF846504DA89FC7F995BFC4D" blockId="13.[211,1172,558,933]" pageId="13" pageNumber="1349">
This species is similar to
<taxonomicName id="4C644D3DFF846504DBB9FC7F9971FDAE" authority="Nakamura and Child, 1991" authorityName="Nakamura and Child" authorityYear="1991" box="[483,1035,558,582]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="improcerum">
<emphasis id="B910EAACFF846504DBB9FC7F9FACFDAE" box="[483,726,558,582]" italics="true" pageId="13" pageNumber="1349">Nymphon improcerum</emphasis>
<bibRefCitation id="EFF54B4FFF846504D886FC7F9971FDAE" author="Nakamura K &amp; Child CA" box="[732,1035,558,582]" pageId="13" pageNumber="1349" pagination="1 - 74" refId="ref10149" refString="Nakamura K, Child CA. 1991. Pycnogonida from waters adjacent to Japan. Smithson Contrib Zool. 512: 1 - 74." type="journal article" year="1991">
Nakamura and Child,
<quantity id="4C9C9B5BFF846504D98CFC7F9971FDAE" box="[982,1035,558,582]" metricMagnitude="1" metricUnit="m" metricValue="5.05714" pageId="13" pageNumber="1349" unit="in" value="1991.0">1991</quantity>
</bibRefCitation>
</taxonomicName>
in the trunk configuration, the moderately long neck, the short chela palm and long chela fingers with many endal teeth, and the short main claw, which is almost the same length as the auxiliary claws. However, it differs from
<taxonomicName id="4C644D3DFF846504D8C6FCDE9E3EFD4E" authorityName="Nakamura and Child" authorityYear="1991" box="[668,836,654,678]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="improcerum">
<emphasis id="B910EAACFF846504D8C6FCDE9FCAFD4E" box="[668,688,655,678]" italics="true" pageId="13" pageNumber="1349">N</emphasis>
.
<emphasis id="B910EAACFF846504D89BFCDE9E3EFD4E" box="[705,836,655,678]" italics="true" pageId="13" pageNumber="1349">improcerum</emphasis>
</taxonomicName>
in having no tubercles on the lateral processes or anterodistally on the cephalic segment, and in having the third segment of the palp short, almost the same length as the fifth segment. In
<collectingCountry id="F373762EFF846504D9B6FC9F994BFD0D" box="[1004,1073,718,741]" name="Japan" pageId="13" pageNumber="1349">Japan</collectingCountry>
, another short-palmed
<taxonomicName id="4C644D3DFF846504DB2EFCBC9F4DFCED" box="[372,567,749,773]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B910EAACFF846504DB2EFCBC9CA5FCED" box="[372,479,749,773]" italics="true" pageId="13" pageNumber="1349">Nymphon</emphasis>
species
</taxonomicName>
is
<taxonomicName id="4C644D3DFF846504D807FCBF9960FCED" authority="Nakamura and Child, 1983" authorityName="Nakamura and Child" authorityYear="1983" box="[605,1050,749,773]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="akanei">
<emphasis id="B910EAACFF846504D807FCBF9F0BFCED" box="[605,625,750,773]" italics="true" pageId="13" pageNumber="1349">N</emphasis>
.
<emphasis id="B910EAACFF846504D8DEFCBC9FB7FCED" box="[644,717,749,773]" italics="true" pageId="13" pageNumber="1349">akanei</emphasis>
<bibRefCitation id="EFF54B4FFF846504D882FCBC9960FCED" author="Nakamura K &amp; Child CA" box="[728,1050,749,773]" pageId="13" pageNumber="1349" pagination="1 - 71" refId="ref10059" refString="Nakamura K, Child CA. 1983. Shallow-water Pycnogonida from the Izu Peninsura, Japan. Smithson Contrib Zool. 386: 1 - 71." type="journal article" year="1983">Nakamura and Child, 1983</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C644D3DFF846504DE72FCBC9C24FCCD" authorityName="Takahashi &amp; Kajihara &amp; Mawatari" authorityYear="2012" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="nagannuense">
<emphasis id="B910EAACFF846504DE72FCBC9C24FCCD" italics="true" pageId="13" pageNumber="1349">Nymphon nagannuense</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C644D3DFF846504DBB4FD5F9F20FCCD" authorityName="Nakamura and Child" authorityYear="1983" box="[494,602,781,805]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="akanei">
<emphasis id="B910EAACFF846504DBB4FD5F9F78FCCD" box="[494,514,782,805]" italics="true" pageId="13" pageNumber="1349">N</emphasis>
.
<emphasis id="B910EAACFF846504D84BFD5C9F20FCCD" box="[529,602,781,805]" italics="true" pageId="13" pageNumber="1349">akanei</emphasis>
</taxonomicName>
in having slender trunk and neck; short and robust claws; more chela teeth (
<quantity id="4C9C9B5BFF846504DBA8FD7C9F2FFCAD" box="[498,597,813,837]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="13" pageNumber="1349" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
<taxonomicName id="4C644D3DFF846504D805FD7F9FAAFCAD" authorityName="Nakamura and Child" authorityYear="1983" box="[607,720,813,837]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="akanei">
<emphasis id="B910EAACFF846504D805FD7F9F09FCAD" box="[607,627,814,837]" italics="true" pageId="13" pageNumber="1349">N</emphasis>
.
<emphasis id="B910EAACFF846504D8DDFD7C9FAAFCAD" box="[647,720,813,837]" italics="true" pageId="13" pageNumber="1349">akanei</emphasis>
</taxonomicName>
;
<quantity id="4C9C9B5BFF846504D8BBFD7C9E3EFCAD" box="[737,836,813,837]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="13" pageNumber="1349" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
<taxonomicName id="4C644D3DFF846504D917FD7F997AFCAD" authorityName="Takahashi &amp; Kajihara &amp; Mawatari" authorityYear="2012" box="[845,1024,814,837]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="nagannuense">
<emphasis id="B910EAACFF846504D917FD7F9E1BFCAD" box="[845,865,814,837]" italics="true" pageId="13" pageNumber="1349">N</emphasis>
.
<emphasis id="B910EAACFF846504D92FFD7F997AFCAD" box="[885,1024,814,837]" italics="true" pageId="13" pageNumber="1349">nagannuense</emphasis>
</taxonomicName>
); and different ratios of leg-segment lengths.
<taxonomicName id="4C644D3DFF846504D83AFD1C9E18FC8D" authorityName="Takahashi &amp; Kajihara &amp; Mawatari" authorityYear="2012" box="[608,866,845,869]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="13" pageNumber="1349" phylum="Arthropoda" rank="species" species="nagannuense">
<emphasis id="B910EAACFF846504D83AFD1C9E18FC8D" box="[608,866,845,869]" italics="true" pageId="13" pageNumber="1349">Nymphon nagannuense</emphasis>
</taxonomicName>
is unique in having some pigmented spots (reddish-brown in ethanol-preserved material) at the base of each chelifore scape and each first coxa, and on the anterior two trunk segments.
</paragraph>
</subSubSection>
</treatment>
</document>
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