350 lines
45 KiB
XML
350 lines
45 KiB
XML
<document id="12C4E0E36991DFDF029E30BE0182C8D7" ID-DOI="10.11646/zootaxa.3946.3.3" ID-GBIF-Dataset="fcec1ba5-a256-4419-a228-ca2b529948bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="245390" ID-ZooBank="8D7575DB-636C-4473-91C3-A58C1E5F9E73" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034695781" checkinUser="plazi" docAuthor="Zakrzewska, Marta & Giłka, Wojciech" docDate="2015" docId="03CD3E7A7115A717FF60AA9FD1A9F76F" docLanguage="en" docName="zt03946p360.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stempellinella electra Gilka et Zakrzewska, sp. nov." docType="treatment" docVersion="11" lastPageNumber="352" masterDocId="FFF446027117A712FFF7AC04D275F355" masterDocTitle="The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk" masterLastPageNumber="360" masterPageNumber="347" pageNumber="349" updateTime="1698574398835" updateUser="plazi">
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<mods:mods id="9FCB415F7261523D40004EE4DE029FA4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CF7D3A4708B64CC88CEED168E626D49C">
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<mods:title id="19A80874E985FAD2E936C4EA1227E5A6">The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk</mods:title>
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</mods:titleInfo>
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<mods:name id="3E8A7E36B8496320048F1A4307BA9610" type="personal">
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<mods:role id="368597F63C412FC6F9833C8A0752A442">
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<mods:roleTerm id="6F59A574CD4DEBC312B5CCE27B166610">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B05655A8C0E81633E089527045B778C3">Zakrzewska, Marta</mods:namePart>
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</mods:name>
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<mods:name id="022AD5BE7D07B75BDC7572E3D86B8220" type="personal">
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<mods:roleTerm id="5C9E77D7F0141787B0E2AF9E57E544A2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="235C49452916427C5C5A40247F1A449C">Giłka, Wojciech</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5C392674868F358AA04794F49CA80EC6">text</mods:typeOfResource>
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<mods:title id="3E23DF9DDA1B41089D79B713A1B3FBA0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="210CDDA54C4D2501AB84A47AC76BC0FA">
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<mods:date id="B9009F47C049F901C806B3E7DF71E216">2015</mods:date>
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<mods:detail id="C6992C44A9DC9C7FA480566139BAB3EA" type="volume">
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<mods:number id="A38CDB7D13F1764E5FF032B8BAC1A68E">3946</mods:number>
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<mods:detail id="78A36F98B750E9DFD1B90FEA34278483" type="issue">
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<mods:number id="6BC06073C76D1F90FAA6037A25D91EE1">3</mods:number>
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<mods:classification id="82240A7A41E7A295CD9FADDBAB3970A4">journal article</mods:classification>
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<mods:identifier id="F62186B22073C307BADFEE987A0E9656" type="DOI">10.11646/zootaxa.3946.3.3</mods:identifier>
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<mods:identifier id="40E07A95B26A91FF1FDD6C14D6601D11" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A8ADA654749B226DFFB47D8CB2B67BFC" type="Zenodo-Dep">245390</mods:identifier>
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<mods:identifier id="68401B175A078907896CD4D611999DF2" type="ZooBank">8D7575DB-636C-4473-91C3-A58C1E5F9E73</mods:identifier>
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<treatment id="03CD3E7A7115A717FF60AA9FD1A9F76F" ID-DOI="http://doi.org/10.5281/zenodo.5616322" ID-GBIF-Taxon="119608921" ID-Zenodo-Dep="5616322" LSID="urn:lsid:plazi:treatment:03CD3E7A7115A717FF60AA9FD1A9F76F" httpUri="http://treatment.plazi.org/id/03CD3E7A7115A717FF60AA9FD1A9F76F" lastPageId="5" lastPageNumber="352" pageId="2" pageNumber="349">
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<subSubSection id="C37EDCE77115A710FF60AA9FD164F5E0" box="[151,785,1691,1717]" pageId="2" pageNumber="349" type="nomenclature">
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<paragraph id="8BDB8F6C7115A710FF60AA9FD164F5E0" blockId="2.[151,785,1691,1717]" box="[151,785,1691,1717]" pageId="2" pageNumber="349">
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<heading id="D09338007115A710FF60AA9FD164F5E0" bold="true" box="[151,785,1691,1717]" fontSize="11" level="1" pageId="2" pageNumber="349" reason="1">
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<emphasis id="B910537E7115A710FF60AA9FD164F5E0" bold="true" box="[151,785,1691,1717]" pageId="2" pageNumber="349">
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<taxonomicName id="4C64F4EF7115A710FF60AA9FD0D2F5E0" ID-CoL="9HM7T" authority="Gilka et Zakrzewska" authorityName="Gilka et Zakrzewska" box="[151,679,1691,1717]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="electra" status="sp. nov.">
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<emphasis id="B910537E7115A710FF60AA9FD3EEF5E0" bold="true" box="[151,411,1691,1717]" italics="true" pageId="2" pageNumber="349">Stempellinella electra</emphasis>
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Giłka
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<emphasis id="B910537E7115A710FE18AA9FD071F5E0" bold="true" box="[495,516,1691,1717]" italics="true" pageId="2" pageNumber="349">et</emphasis>
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Zakrzewska
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</taxonomicName>
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,
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<taxonomicNameLabel id="A223EE057115A710FD43AA9FD164F5E0" box="[692,785,1691,1717]" pageId="2" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37EDCE77115A710FF60AAE5D7FFF417" pageId="2" pageNumber="349" type="materials_examined">
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<paragraph id="8BDB8F6C7115A710FF60AAE5D7FFF417" blockId="2.[151,1436,1761,2038]" pageId="2" pageNumber="349">
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<emphasis id="B910537E7115A710FF60AAE5D330F5AF" bold="true" box="[151,325,1761,1786]" pageId="2" pageNumber="349">
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<typeStatus id="54DF31CE7115A710FF60AAE5D2A7F5AF" box="[151,210,1761,1786]" pageId="2" pageNumber="349">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54DF31CE7115A710FEBAAAE5D3CCF5AF" box="[333,441,1761,1786]" pageId="2" pageNumber="349" type="holotype">Holotype</typeStatus>
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. Adult male, complete specimen with large mite attached under right wing, preserved in
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<date id="FFDAA9AC7115A710FF60AB02D29FF44B" box="[151,234,1798,1822]" pageId="2" pageNumber="349" value="2011-10-14">14 x 11</date>
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x
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<quantity id="4C9C22897115A710FEF1AB02D332F448" box="[262,327,1798,1822]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="349" unit="mm" value="2.0">2 mm</quantity>
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piece of amber (Eocene, ~45–40 Ma, Baltic amber, Gulf of Gdańsk; MAI-4295a;
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<figureCitation id="135F93E97115A710FB15AB01D756F44B" box="[1250,1315,1797,1822]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="2" pageNumber="349">Fig. 3</figureCitation>
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A); animal syninclusions: Acari (2 ind., MAI-4295a),
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<taxonomicName id="4C64F4EF7115A710FD81AB2DD153F417" box="[630,806,1833,1858]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="family">Mycetophilidae</taxonomicName>
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(1 ind., MAI-4295b), Aphidoidea (1 ind., MAI-4295).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37EDCE77115A710FF30AB49D680F433" box="[199,1269,1869,1894]" pageId="2" pageNumber="349" type="etymology">
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<paragraph id="8BDB8F6C7115A710FF30AB49D680F433" blockId="2.[151,1436,1761,2038]" box="[199,1269,1869,1894]" pageId="2" pageNumber="349">
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<emphasis id="B910537E7115A710FF30AB49D3D1F433" bold="true" box="[199,420,1869,1894]" pageId="2" pageNumber="349">Derivatio nominis.</emphasis>
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Adjective in feminine form derived from the Latin noun ‘
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<emphasis id="B910537E7115A710FBD8AB49D6E4F433" box="[1071,1169,1869,1894]" italics="true" pageId="2" pageNumber="349">electrum</emphasis>
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’, amber.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37EDCE77115A710FF30AB75D39EF4A0" pageId="2" pageNumber="349" type="diagnosis">
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<paragraph id="8BDB8F6C7115A710FF30AB75D39EF4A0" blockId="2.[151,1436,1761,2038]" pageId="2" pageNumber="349">
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<emphasis id="B910537E7115A710FF30AB75D334F4DF" bold="true" box="[199,321,1905,1930]" pageId="2" pageNumber="349">Diagnosis.</emphasis>
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Antennal flagellum composed of 13 segments, flagellomeres 11–13 fused in part. Anal point with slender distal section and strong crests. Superior volsella elongated, with round apex, bearing 2 setae. Stem of median volsella club-shaped, with wide pectinate and foliate lamellae. Inferior volsella arcuate, tapering to pointed apex armed with 2 stout setae.
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</paragraph>
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</subSubSection>
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<caption id="DF1BDFE47114A711FF60AB75D330F4B6" httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="3" pageNumber="350" targetBox="[177,1412,193,1863]" targetPageId="3">
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<paragraph id="8BDB8F6C7114A711FF60AB75D330F4B6" blockId="3.[151,1436,1905,2019]" pageId="3" pageNumber="350">
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<emphasis id="B910537E7114A711FF60AB75D363F4D3" bold="true" box="[151,278,1905,1927]" pageId="3" pageNumber="350">FIGURE 3.</emphasis>
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<taxonomicName id="4C64F4EF7114A711FEE8AB75D38AF4D2" box="[287,511,1905,1927]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="electra" status="sp. nov.">
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<emphasis id="B910537E7114A711FEE8AB75D38AF4D2" box="[287,511,1905,1927]" italics="true" pageId="3" pageNumber="350">Stempellinella electra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A223EE057114A711FDFEAB76D029F4D2" box="[521,604,1906,1927]" pageId="3" pageNumber="350" rank="species">
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<emphasis id="B910537E7114A711FDFEAB76D029F4D2" bold="true" box="[521,604,1906,1927]" pageId="3" pageNumber="350">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, adult male, holotype MAI-4295a (Eocene, ~45–40 Ma; Baltic amber, Gulf of Gdańsk).
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<emphasis id="B910537E7114A711FF0DAB94D379F4F0" bold="true" box="[250,268,1936,1957]" pageId="3" pageNumber="350">A</emphasis>
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—inclusion;
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<emphasis id="B910537E7114A711FE66AB94D3D7F4F0" bold="true" box="[401,418,1936,1957]" pageId="3" pageNumber="350">B</emphasis>
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—habitus;
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<emphasis id="B910537E7114A711FDE4AB94D050F4F3" bold="true" box="[531,549,1936,1958]" pageId="3" pageNumber="350">C</emphasis>
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—head;
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<emphasis id="B910537E7114A711FD8AAB94D0FAF4F0" bold="true" box="[637,655,1936,1957]" pageId="3" pageNumber="350">D</emphasis>
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,
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<emphasis id="B910537E7114A711FD6AAB94D0DBF4F0" bold="true" box="[669,686,1936,1957]" pageId="3" pageNumber="350">E</emphasis>
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—antenna photographed in transmitted (
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<emphasis id="B910537E7114A711FBBAAB94D62AF4F0" bold="true" box="[1101,1119,1936,1957]" pageId="3" pageNumber="350">D</emphasis>
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) and reflected light (
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<emphasis id="B910537E7114A711FACDAB94D73EF4F0" bold="true" box="[1338,1355,1936,1957]" pageId="3" pageNumber="350">E</emphasis>
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) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion);
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<emphasis id="B910537E7114A711FB75ABABD6E4F491" bold="true" box="[1154,1169,1967,1988]" pageId="3" pageNumber="350">F</emphasis>
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—wing photographed in transmitted light.
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</paragraph>
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</caption>
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<caption id="DF1BDFE47113A716FF60A851D129F7FD" httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="4" pageNumber="351" targetBox="[151,1407,193,1086]" targetPageId="4">
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<paragraph id="8BDB8F6C7113A716FF60A851D129F7FD" blockId="4.[151,1436,1108,1192]" pageId="4" pageNumber="351">
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<emphasis id="B910537E7113A716FF60A851D363F73F" bold="true" box="[151,278,1109,1131]" pageId="4" pageNumber="351">FIGURE 4.</emphasis>
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<taxonomicName id="4C64F4EF7113A716FEE8A850D38AF73F" box="[287,511,1108,1130]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="electra" status="sp. nov.">
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<emphasis id="B910537E7113A716FEE8A850D38AF73F" box="[287,511,1108,1130]" italics="true" pageId="4" pageNumber="351">Stempellinella electra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A223EE057113A716FDFEA851D029F73F" box="[521,604,1109,1130]" pageId="4" pageNumber="351" rank="species">
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<emphasis id="B910537E7113A716FDFEA851D029F73F" bold="true" box="[521,604,1109,1130]" pageId="4" pageNumber="351">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, adult male, holotype MAI-4295a (Eocene, ~45–40 Ma; Baltic amber, Gulf of Gdańsk).
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<emphasis id="B910537E7113A716FF0BA877D358F7DC" bold="true" box="[252,301,1139,1161]" pageId="4" pageNumber="351">A–C</emphasis>
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—hypopygium, dorsal aspect, photographed in transmitted light (
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<emphasis id="B910537E7113A716FC22A877D192F7DD" bold="true" box="[981,999,1139,1160]" pageId="4" pageNumber="351">A</emphasis>
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), reflected light (
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<emphasis id="B910537E7113A716FB57A870D6C4F7DC" bold="true" box="[1184,1201,1140,1161]" pageId="4" pageNumber="351">B</emphasis>
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), and drawn (
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<emphasis id="B910537E7113A716FAB1A877D72DF7DC" bold="true" box="[1350,1368,1139,1161]" pageId="4" pageNumber="351">C</emphasis>
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);
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<emphasis id="B910537E7113A716FA87A870D7F7F7DC" bold="true" box="[1392,1410,1140,1161]" pageId="4" pageNumber="351">D</emphasis>
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— median volsella (magnified c. 2 times relative to hypopygium drawn).
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</paragraph>
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</caption>
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<subSubSection id="C37EDCE77113A717FF30A8D0D72BF2F9" lastPageId="5" lastPageNumber="352" pageId="4" pageNumber="351" type="description">
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<paragraph id="8BDB8F6C7113A716FF30A8D0D045F7B8" blockId="4.[151,1437,1236,1774]" box="[199,560,1236,1261]" pageId="4" pageNumber="351">
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<emphasis id="B910537E7113A716FF30A8D0D324F7B8" bold="true" box="[199,337,1236,1261]" pageId="4" pageNumber="351">Description</emphasis>
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. Adult male (n = 1)
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</paragraph>
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<paragraph id="8BDB8F6C7113A716FF30A8FCD0A2F644" blockId="4.[151,1437,1236,1774]" box="[199,727,1272,1297]" pageId="4" pageNumber="351">
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Total length c.
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<quantity id="4C9C22897113A716FE99A8FCD3B2F645" box="[366,455,1272,1297]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="351" unit="mm" value="1.5">1.5 mm</quantity>
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; wing length c. 890 µm.
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</paragraph>
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<paragraph id="8BDB8F6C7113A716FF30A918D31FF690" blockId="4.[151,1437,1236,1774]" pageId="4" pageNumber="351">
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<emphasis id="B910537E7113A716FF30A918D371F660" box="[199,260,1308,1333]" italics="true" pageId="4" pageNumber="351">Head</emphasis>
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(
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<figureCitation id="135F93E97113A716FEEFA918D32AF660" box="[280,351,1308,1333]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="4" pageNumber="351">Fig. 3</figureCitation>
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C–E). Eyes bare, small, ovoid, broadly separated. Frontal tubercles conical, c. 10 µm long. Antennal flagellum composed of 13 segments of which 11 are well discernible, flagellomeres 11–13 fused in part, AR 0.76 (when flagellum measured as 11-segmented), AR 0.62 (as 12-segmented), AR 0.50 (as 13-segmented); plume fully developed (
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<figureCitation id="135F93E97113A716FE50A98CD39FF6F4" box="[423,490,1416,1441]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="4" pageNumber="351">Fig. 3</figureCitation>
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D, E). Length of palpomeres 2–5 (in µm): c. 30, 64, 84, 133. Clypeals present, but cannot be counted.
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</paragraph>
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<paragraph id="8BDB8F6C7113A716FF30A9D4D699F6BC" blockId="4.[151,1437,1236,1774]" box="[199,1260,1488,1513]" pageId="4" pageNumber="351">
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<emphasis id="B910537E7113A716FF30A9D4D36DF6BC" box="[199,280,1488,1513]" italics="true" pageId="4" pageNumber="351">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="135F93E97113A716FED0A9D4D31CF6BC" box="[295,361,1488,1513]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="4" pageNumber="351">Fig. 3</figureCitation>
|
||
B). Ac at least 11, Dc at least 7-8 on each side, Pa 1 on each side, Scts at least 4.
|
||
</paragraph>
|
||
<paragraph id="8BDB8F6C7113A716FF30A9F1D14DF52A" blockId="4.[151,1437,1236,1774]" pageId="4" pageNumber="351">
|
||
<emphasis id="B910537E7113A716FF30A9F1D28AF559" box="[199,255,1525,1548]" italics="true" pageId="4" pageNumber="351">Wing</emphasis>
|
||
(
|
||
<figureCitation id="135F93E97113A716FEE6A9F0D321F558" box="[273,340,1524,1549]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="4" pageNumber="351">Fig. 3</figureCitation>
|
||
F). Slender, with anal lobe weak, broadest at 2/3 length; width: 285 µm, length/width ratio: 3.12. Sc ending slightly distal of FCu, R2+3 fading, poorly visible. RM slightly oblique relative to R. FCu placed well distally of RM; VR Cu 1.38. Veins ending as follows (in order from base to tip): An, Sc, Cu1, R1, R4+5 and M3+4, M1+2. Wing covered with dense macrotrichia in distal half at least.
|
||
</paragraph>
|
||
<paragraph id="8BDB8F6C7113A716FF30AA8FD0B6F5BB" blockId="4.[151,1437,1236,1774]" pageId="4" pageNumber="351">
|
||
<emphasis id="B910537E7113A716FF30AA8FD376F5F7" box="[199,259,1675,1698]" italics="true" pageId="4" pageNumber="351">Legs.</emphasis>
|
||
Tibial apices including combs of mid and hind legs weakly visible. Spurs not observed on forelegs. Only one spur visible on mid leg (c. 20 µm long) and hind leg (c. 28 µm long). Sensilla chaetica on ta1 of p2 not observed. Lengths of leg segments and leg ratios in
|
||
<tableCitation id="C6E6BAD77113A716FD9EAAD1D0C8F5BB" box="[617,701,1749,1774]" captionStart="TABLE 1" captionStartId="4.[151,239,1817,1840]" captionTargetBox="[159,1348,1868,2025]" captionTargetPageId="4" captionText="TABLE 1. Leg segment lengths (Μm) and leg ratios of male Stempellinella electra sp. nov." httpUri="http://table.plazi.org/id/DF1BDFE47113A716FF60AB1DD614F465" pageId="4" pageNumber="351" tableUuid="DF1BDFE47113A716FF60AB1DD614F465">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF1BDFE47113A716FF60AB1DD614F465" ID-Table-UUID="DF1BDFE47113A716FF60AB1DD614F465" box="[151,1121,1817,1840]" httpUri="http://table.plazi.org/id/DF1BDFE47113A716FF60AB1DD614F465" pageId="4" pageNumber="351" targetBox="[159,1348,1868,2025]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BDB8F6C7113A716FF60AB1DD614F465" blockId="4.[151,1121,1817,1840]" box="[151,1121,1817,1840]" pageId="4" pageNumber="351">
|
||
<emphasis id="B910537E7113A716FF60AB1DD37FF465" bold="true" box="[151,266,1817,1840]" pageId="4" pageNumber="351">TABLE 1.</emphasis>
|
||
Leg segment lengths (Μm) and leg ratios of male
|
||
<taxonomicName id="4C64F4EF7113A716FCEAAB1DD670F465" box="[797,1029,1817,1840]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="electra" status="sp. nov.">
|
||
<emphasis id="B910537E7113A716FCEAAB1DD670F465" box="[797,1029,1817,1840]" italics="true" pageId="4" pageNumber="351">Stempellinella electra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223EE057113A716FBFBAB1DD614F465" box="[1036,1121,1817,1840]" pageId="4" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB8F6C7113A716FED9AB48D731F4B0" pageId="4" pageNumber="351">
|
||
<table id="F9647DCC711358EDFF68AB48D731F4BC" box="[159,1348,1868,2025]" gridcols="9" gridrows="4" pageId="4" pageNumber="351">
|
||
<tr id="35548D2E711358EDFF68AB48D731F430" box="[159,1348,1868,1893]" gridrow="0" pageId="4" pageNumber="351" rowspan-0="1">
|
||
<th id="7685E452711358EDFED9AB48D321F430" box="[302,340,1868,1893]" gridcol="1" gridrow="0" pageId="4" pageNumber="351">fe</th>
|
||
<th id="7685E452711358EDFE4AAB48D396F430" box="[445,483,1868,1893]" gridcol="2" gridrow="0" pageId="4" pageNumber="351">ti</th>
|
||
<th id="7685E452711358EDFDBBAB48D007F430" box="[588,626,1868,1893]" gridcol="3" gridrow="0" pageId="4" pageNumber="351">ta1</th>
|
||
<th id="7685E452711358EDFD2CAB48D174F430" box="[731,769,1868,1893]" gridcol="4" gridrow="0" pageId="4" pageNumber="351">ta2</th>
|
||
<th id="7685E452711358EDFC9DAB48D1E5F430" box="[874,912,1868,1893]" gridcol="5" gridrow="0" pageId="4" pageNumber="351">ta3</th>
|
||
<th id="7685E452711358EDFC0EAB48D66AF430" box="[1017,1055,1868,1893]" gridcol="6" gridrow="0" pageId="4" pageNumber="351">ta4</th>
|
||
<th id="7685E452711358EDFB7EAB48D6D6F430" box="[1161,1187,1868,1893]" gridcol="7" gridrow="0" pageId="4" pageNumber="351">ta5</th>
|
||
<th id="7685E452711358EDFAEFAB48D731F430" box="[1304,1348,1868,1893]" gridcol="8" gridrow="0" pageId="4" pageNumber="351">LR</th>
|
||
</tr>
|
||
<tr id="35548D2E711358EDFF68AB73D731F4C4" box="[159,1348,1911,1937]" gridrow="1" pageId="4" pageNumber="351">
|
||
<th id="7685E452711358EDFF68AB73D2C6F4C4" box="[159,179,1911,1937]" gridcol="0" gridrow="1" pageId="4" pageNumber="351">p1</th>
|
||
<td id="7685E452711358EDFED9AB73D321F4C4" box="[302,340,1911,1937]" gridcol="1" gridrow="1" pageId="4" pageNumber="351">430</td>
|
||
<td id="7685E452711358EDFE4AAB73D396F4C4" box="[445,483,1911,1937]" gridcol="2" gridrow="1" pageId="4" pageNumber="351">235</td>
|
||
<td id="7685E452711358EDFDBBAB73D007F4C4" box="[588,626,1911,1937]" gridcol="3" gridrow="1" pageId="4" pageNumber="351">500</td>
|
||
<td id="7685E452711358EDFD2CAB73D174F4C4" box="[731,769,1911,1937]" gridcol="4" gridrow="1" pageId="4" pageNumber="351">275</td>
|
||
<td id="7685E452711358EDFC9DAB73D1E5F4C4" box="[874,912,1911,1937]" gridcol="5" gridrow="1" pageId="4" pageNumber="351">210</td>
|
||
<td id="7685E452711358EDFC0EAB73D66AF4C4" box="[1017,1055,1911,1937]" gridcol="6" gridrow="1" pageId="4" pageNumber="351">140</td>
|
||
<td id="7685E452711358EDFB7EAB73D6D6F4C4" box="[1161,1187,1911,1937]" gridcol="7" gridrow="1" pageId="4" pageNumber="351">65</td>
|
||
<td id="7685E452711358EDFAEFAB73D731F4C4" box="[1304,1348,1911,1937]" gridcol="8" gridrow="1" pageId="4" pageNumber="351">2.13</td>
|
||
</tr>
|
||
<tr id="35548D2E711358EDFF68ABA7D731F4E8" box="[159,1348,1955,1981]" gridrow="2" pageId="4" pageNumber="351">
|
||
<th id="7685E452711358EDFF68ABA7D2C6F4E8" box="[159,179,1955,1981]" gridcol="0" gridrow="2" pageId="4" pageNumber="351">p2</th>
|
||
<td id="7685E452711358EDFED9ABA7D321F4E8" box="[302,340,1955,1981]" gridcol="1" gridrow="2" pageId="4" pageNumber="351">485</td>
|
||
<td id="7685E452711358EDFE4AABA7D396F4E8" box="[445,483,1955,1981]" gridcol="2" gridrow="2" pageId="4" pageNumber="351">365</td>
|
||
<td id="7685E452711358EDFDBBABA7D007F4E8" box="[588,626,1955,1981]" gridcol="3" gridrow="2" pageId="4" pageNumber="351">240</td>
|
||
<td id="7685E452711358EDFD2CABA7D174F4E8" box="[731,769,1955,1981]" gridcol="4" gridrow="2" pageId="4" pageNumber="351">115</td>
|
||
<td id="7685E452711358EDFC9DABA7D1E5F4E8" box="[874,912,1955,1981]" gridcol="5" gridrow="2" pageId="4" pageNumber="351">85</td>
|
||
<td id="7685E452711358EDFC0EABA7D66AF4E8" box="[1017,1055,1955,1981]" gridcol="6" gridrow="2" pageId="4" pageNumber="351">65</td>
|
||
<td id="7685E452711358EDFB7EABA7D6D6F4E8" box="[1161,1187,1955,1981]" gridcol="7" gridrow="2" pageId="4" pageNumber="351">45</td>
|
||
<td id="7685E452711358EDFAEFABA7D731F4E8" box="[1304,1348,1955,1981]" gridcol="8" gridrow="2" pageId="4" pageNumber="351">0.66</td>
|
||
</tr>
|
||
<tr id="35548D2E711358EDFF68ABCBD731F4BC" box="[159,1348,1999,2025]" gridrow="3" pageId="4" pageNumber="351">
|
||
<th id="7685E452711358EDFF68ABCBD2C6F4BC" box="[159,179,1999,2025]" gridcol="0" gridrow="3" pageId="4" pageNumber="351">p3</th>
|
||
<td id="7685E452711358EDFED9ABCBD321F4BC" box="[302,340,1999,2025]" gridcol="1" gridrow="3" pageId="4" pageNumber="351">?</td>
|
||
<td id="7685E452711358EDFE4AABCBD396F4BC" box="[445,483,1999,2025]" gridcol="2" gridrow="3" pageId="4" pageNumber="351">415</td>
|
||
<td id="7685E452711358EDFDBBABCBD007F4BC" box="[588,626,1999,2025]" gridcol="3" gridrow="3" pageId="4" pageNumber="351">295</td>
|
||
<td id="7685E452711358EDFD2CABCBD174F4BC" box="[731,769,1999,2025]" gridcol="4" gridrow="3" pageId="4" pageNumber="351">165</td>
|
||
<td id="7685E452711358EDFC9DABCBD1E5F4BC" box="[874,912,1999,2025]" gridcol="5" gridrow="3" pageId="4" pageNumber="351">150</td>
|
||
<td id="7685E452711358EDFC0EABCBD66AF4BC" box="[1017,1055,1999,2025]" gridcol="6" gridrow="3" pageId="4" pageNumber="351">90</td>
|
||
<td id="7685E452711358EDFB7EABCBD6D6F4BC" box="[1161,1187,1999,2025]" gridcol="7" gridrow="3" pageId="4" pageNumber="351">50</td>
|
||
<td id="7685E452711358EDFAEFABCBD731F4BC" box="[1304,1348,1999,2025]" gridcol="8" gridrow="3" pageId="4" pageNumber="351">0.71</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDB8F6C7112A717FF30AC9DD72BF2F9" blockId="5.[151,1436,151,1082]" pageId="5" pageNumber="352">
|
||
<emphasis id="B910537E7112A717FF30AC9DD323F3E5" box="[199,342,153,176]" italics="true" pageId="5" pageNumber="352">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="135F93E97112A717FE92AC93D3D2F3E5" box="[357,423,151,176]" captionStart="FIGURE 4" captionStartId="4.[151,250,1109,1131]" captionTargetBox="[151,1407,193,1086]" captionTargetId="figure@4.[151,1436,193,1087]" captionTargetPageId="4" captionText="FIGURE 4. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, dorsal aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — median volsella (magnified c. 2 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="5" pageNumber="352">Fig. 4</figureCitation>
|
||
A–D). Gonostylus c. 50 Μm long, shorter than gonocoxite, broadest at mid-length, tapering to blunt apex armed with strong apical setae. Anal tergite subtriangular, with several median setae and at least 7 posterolateral setae on each side of anal point (poorly visible on photographs). Anal point broad at base, distinctly narrowed distally, with slender distal section and strong crests tapering towards anal point tip. Superior volsella elongated, with broadened round apex, bearing 2 setae on median margin. Digitus absent. Stem of median volsella club-shaped, slightly broadened apically, c. 20 µm long, with several setiform and 4–5 wide pectinate and foliate lamellae (
|
||
<figureCitation id="135F93E97112A717FEF2AD6BD33DF2DD" box="[261,328,367,392]" captionStart="FIGURE 4" captionStartId="4.[151,250,1109,1131]" captionTargetBox="[151,1407,193,1086]" captionTargetId="figure@4.[151,1436,193,1087]" captionTargetPageId="4" captionText="FIGURE 4. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, dorsal aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — median volsella (magnified c. 2 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="5" pageNumber="352">Fig. 4</figureCitation>
|
||
A, B, D). Inferior volsella reaching 2/3 length of gonostylus at most, somewhat arcuate, tapering to pointed posteromedially directed apex, with several strong setae including 2 stout setae on apex (
|
||
<figureCitation id="135F93E97112A717FB21AD90D76CF2F9" box="[1238,1305,404,429]" captionStart="FIGURE 4" captionStartId="4.[151,250,1109,1131]" captionTargetBox="[151,1407,193,1086]" captionTargetId="figure@4.[151,1436,193,1087]" captionTargetPageId="4" captionText="FIGURE 4. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, dorsal aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — median volsella (magnified c. 2 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="5" pageNumber="352">Fig. 4</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37EDCE77112A717FF30ADB3D1A9F76F" pageId="5" pageNumber="352" type="discussion">
|
||
<paragraph id="8BDB8F6C7112A717FF30ADB3D0E7F0D3" blockId="5.[151,1436,151,1082]" pageId="5" pageNumber="352">
|
||
<emphasis id="B910537E7112A717FF30ADB3D34EF285" bold="true" box="[199,315,439,464]" pageId="5" pageNumber="352">Remarks.</emphasis>
|
||
<taxonomicName id="4C64F4EF7112A717FEB4ADB3D04DF285" box="[323,568,439,464]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="electra">
|
||
<emphasis id="B910537E7112A717FEB4ADB3D04DF285" box="[323,568,439,464]" italics="true" pageId="5" pageNumber="352">Stempellinella electra</emphasis>
|
||
</taxonomicName>
|
||
is the third fossil species of the genus found in Eocene amber, along with
|
||
<taxonomicName id="4C64F4EF7112A717FA70ADBCD049F2A1" authority="Seredszus et Wichard, 2007" authorityName="Seredszus et Wichard" authorityYear="2007" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="bicorna">
|
||
<emphasis id="B910537E7112A717FA70ADBCD29BF2A0" italics="true" pageId="5" pageNumber="352">S. bicorna</emphasis>
|
||
<bibRefCitation id="EFF5F29D7112A717FF0EADD8D049F2A1" author="Seredszus" box="[249,572,476,501]" pageId="5" pageNumber="352" refString="Seredszus, F. & Wichard, W. (2007) Fossil chironomids (Insecta, Diptera) in Baltic amber. Palaeontographica, Beitrage zur Naturgeschichte der Vorzeit, Abteilung A: Palaozoologie - Stratigraphie, 279, 49 - 91." type="journal article" year="2007">
|
||
Seredszus
|
||
<emphasis id="B910537E7112A717FE83ADD9D3FDF2A1" box="[372,392,477,500]" italics="true" pageId="5" pageNumber="352">et</emphasis>
|
||
Wichard, 2007
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C64F4EF7112A717FD8DADD9D636F2A1" authority="Gilka et Zakrzewska, 2014" authorityName="Gilka et Zakrzewska" authorityYear="2014" box="[634,1091,476,501]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="ivanovae">
|
||
<emphasis id="B910537E7112A717FD8DADD9D08AF2A1" box="[634,767,477,500]" italics="true" pageId="5" pageNumber="352">S. ivanovae</emphasis>
|
||
<bibRefCitation id="EFF5F29D7112A717FCFDADD8D636F2A1" author="Gilka" box="[778,1091,476,501]" pageId="5" pageNumber="352" refString="Zakrzewska, M. & Gilka, W. (2014) The oldest known chironomids of the tribe Tanytarsini (Diptera: Chironomidae) indicate plesiomorphic character states. Geobios, 47, 335 - 343." type="journal article" year="2014">
|
||
Giłka
|
||
<emphasis id="B910537E7112A717FCA4ADD9D112F2A1" box="[851,871,477,500]" italics="true" pageId="5" pageNumber="352">et</emphasis>
|
||
Zakrzewska, 2014
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF5F29D7112A717FBA1ADD8D7EDF2A1" author="Seredszus" box="[1110,1432,476,501]" pageId="5" pageNumber="352" refString="Seredszus, F. & Wichard, W. (2007) Fossil chironomids (Insecta, Diptera) in Baltic amber. Palaeontographica, Beitrage zur Naturgeschichte der Vorzeit, Abteilung A: Palaozoologie - Stratigraphie, 279, 49 - 91." type="journal article" year="2007">Seredszus & Wichard 2007</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5F29D7112A717FF60ADFBD3B0F14D" author="Zakrzewska" box="[151,453,511,536]" pageId="5" pageNumber="352" refString="Zakrzewska, M. & Gilka, W. (2014) The oldest known chironomids of the tribe Tanytarsini (Diptera: Chironomidae) indicate plesiomorphic character states. Geobios, 47, 335 - 343." type="journal article" year="2014">Zakrzewska & Giłka 2014</bibRefCitation>
|
||
). The new species fits well the emended generic diagnosis for
|
||
<taxonomicName id="4C64F4EF7112A717FB65ADFBD305F168" authority="Brundin, 1947 (Ekrem 2007)" authorityName="Brundin, 1947 (Ekrem" authorityYear="2007" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910537E7112A717FB65ADFBD744F14D" box="[1170,1329,511,536]" italics="true" pageId="5" pageNumber="352">Stempellinella</emphasis>
|
||
Brundin, 1947 (
|
||
<bibRefCitation id="EFF5F29D7112A717FF2AAE20D31DF168" author="Ekrem" box="[221,360,548,573]" pageId="5" pageNumber="352" refString="Ekrem, T. (2007) A taxonomic revision of the genus Stempellinella (Diptera: Chironomidae). Journal of Natural History, 41, 1367 - 1465. http: // dx. doi. org / 10.1080 / 00222930701437360" type="journal article" year="2007">Ekrem 2007</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—the adult male has bare ovoid and broadly separated eyes (
|
||
<figureCitation id="135F93E97112A717FBE1AE20D622F169" box="[1046,1111,548,573]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="5" pageNumber="352">Fig. 3</figureCitation>
|
||
C), the wing vein R4+5 ending opposite to M3+4 (
|
||
<figureCitation id="135F93E97112A717FEADAE4ED3EEF136" box="[346,411,586,611]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="5" pageNumber="352">Fig. 3</figureCitation>
|
||
F), the broadened superior volsella, and the gonostylus shorter than the gonocoxite (
|
||
<figureCitation id="135F93E97112A717FABDAE4ED7FAF136" box="[1354,1423,586,611]" captionStart="FIGURE 4" captionStartId="4.[151,250,1109,1131]" captionTargetBox="[151,1407,193,1086]" captionTargetId="figure@4.[151,1436,193,1087]" captionTargetPageId="4" captionText="FIGURE 4. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, dorsal aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — median volsella (magnified c. 2 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="5" pageNumber="352">Fig. 4</figureCitation>
|
||
). Significant differences in the hypopygium structure have been observed between the three fossil
|
||
<taxonomicName id="4C64F4EF7112A717FB0BAE75D7EEF1DF" box="[1276,1435,625,650]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910537E7112A717FB0BAE75D7EEF1DF" box="[1276,1435,625,650]" italics="true" pageId="5" pageNumber="352">Stempellinella</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="4C64F4EF7112A717FF0FAE92D311F1FB" box="[248,356,661,686]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="electra">
|
||
<emphasis id="B910537E7112A717FF0FAE92D311F1FB" box="[248,356,661,686]" italics="true" pageId="5" pageNumber="352">S. electra</emphasis>
|
||
</taxonomicName>
|
||
is distinct in having a long anal point, broad at base and narrowed distally (in contrast to the short anal point in
|
||
<taxonomicName id="4C64F4EF7112A717FE8AAEBED383F187" box="[381,502,697,722]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="bicorna">
|
||
<emphasis id="B910537E7112A717FE8AAEBED383F187" box="[381,502,697,722]" italics="true" pageId="5" pageNumber="352">S. bicorna</emphasis>
|
||
</taxonomicName>
|
||
) bearing strong anal point crests (
|
||
<figureCitation id="135F93E97112A717FC65AEBDD1A9F187" box="[914,988,697,722]" captionStart="FIGURE 4" captionStartId="4.[151,250,1109,1131]" captionTargetBox="[151,1407,193,1086]" captionTargetId="figure@4.[151,1436,193,1087]" captionTargetPageId="4" captionText="FIGURE 4. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, dorsal aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — median volsella (magnified c. 2 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="5" pageNumber="352">Fig. 4</figureCitation>
|
||
B, C)—not observed in the two fossil relatives, but resembling those known from several extant species (
|
||
<emphasis id="B910537E7112A717FC70AEDAD1EEF1A3" box="[903,923,734,758]" italics="true" pageId="5" pageNumber="352">cf</emphasis>
|
||
.
|
||
<bibRefCitation id="EFF5F29D7112A717FC5EAED9D64DF1A3" author="Ekrem" box="[937,1080,733,758]" pageId="5" pageNumber="352" refString="Ekrem, T. (2007) A taxonomic revision of the genus Stempellinella (Diptera: Chironomidae). Journal of Natural History, 41, 1367 - 1465. http: // dx. doi. org / 10.1080 / 00222930701437360" type="journal article" year="2007">Ekrem 2007</bibRefCitation>
|
||
). The best diagnostic characters for
|
||
<taxonomicName id="4C64F4EF7112A717FF37AF06D359F04F" box="[192,300,769,794]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="electra">
|
||
<emphasis id="B910537E7112A717FF37AF06D359F04F" box="[192,300,769,794]" italics="true" pageId="5" pageNumber="352">S. electra</emphasis>
|
||
</taxonomicName>
|
||
is the superior volsella elongated and broadened apically, the inferior volsella arcuate and pointed, and the median volsella bearing several setiform and 4–5 wide pectinate and foliate lamellae (
|
||
<figureCitation id="135F93E97112A717FB4DAF21D68BF06B" box="[1210,1278,805,830]" captionStart="FIGURE 4" captionStartId="4.[151,250,1109,1131]" captionTargetBox="[151,1407,193,1086]" captionTargetId="figure@4.[151,1436,193,1087]" captionTargetPageId="4" captionText="FIGURE 4. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, dorsal aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — median volsella (magnified c. 2 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245394/files/figure.png" pageId="5" pageNumber="352">Fig. 4</figureCitation>
|
||
A, B, D)—the combination of shapes not recorded neither in fossil or extant
|
||
<taxonomicName id="4C64F4EF7112A717FCB1AF4DD190F037" box="[838,997,841,866]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910537E7112A717FCB1AF4DD190F037" box="[838,997,841,866]" italics="true" pageId="5" pageNumber="352">Stempellinella</emphasis>
|
||
</taxonomicName>
|
||
(the pectinate lamellae are known only from
|
||
<taxonomicName id="4C64F4EF7112A717FF23AF6AD0F0F0D3" authority="Casas et Vilchez-Quero, 1991" authorityName="Casas et Vilchez-Quero" authorityYear="1991" box="[212,645,877,902]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="reissi">
|
||
<emphasis id="B910537E7112A717FF23AF6AD359F0D0" box="[212,300,878,901]" italics="true" pageId="5" pageNumber="352">S. reissi</emphasis>
|
||
Casas
|
||
<emphasis id="B910537E7112A717FE8CAF6AD3FAF0D0" box="[379,399,878,901]" italics="true" pageId="5" pageNumber="352">et</emphasis>
|
||
Vilchez-Quero, 1991
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB8F6C7112A717FF30AF96D1A9F76F" blockId="5.[151,1436,151,1082]" pageId="5" pageNumber="352">
|
||
It is worth noting that we observed a tendency to formation of a fully-segmented antennal flagellum in
|
||
<taxonomicName id="4C64F4EF7112A717FF60AFB1D3F9F09B" box="[151,396,949,974]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="electra">
|
||
<emphasis id="B910537E7112A717FF60AFB1D3F9F09B" box="[151,396,949,974]" italics="true" pageId="5" pageNumber="352">Stempellinella electra</emphasis>
|
||
</taxonomicName>
|
||
, similar to that known from
|
||
<taxonomicName id="4C64F4EF7112A717FD20AFB2D12CF098" box="[727,857,950,973]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="ivanovae">
|
||
<emphasis id="B910537E7112A717FD20AFB2D12CF098" box="[727,857,950,973]" italics="true" pageId="5" pageNumber="352">S. ivanovae</emphasis>
|
||
</taxonomicName>
|
||
. However, when comparing this character in these two species, the higher number of incompletely fused flagellomeres may indicate more advanced state in
|
||
<taxonomicName id="4C64F4EF7112A717FAC5AFDED7E9F0A7" box="[1330,1436,985,1010]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="electra">
|
||
<emphasis id="B910537E7112A717FAC5AFDED7E9F0A7" box="[1330,1436,985,1010]" italics="true" pageId="5" pageNumber="352">S. electra</emphasis>
|
||
</taxonomicName>
|
||
(flagellomeres 11–13 fused in part,
|
||
<figureCitation id="135F93E97112A717FDB7AFF9D0F0F743" box="[576,645,1021,1046]" captionStart="FIGURE 3" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[177,1412,193,1863]" captionTargetId="figure@3.[177,1413,193,1865]" captionTargetPageId="3" captionText="FIGURE 3. Stempellinella electra sp. nov., adult male, holotype MAI- 4295 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D, E — antenna photographed in transmitted (D) and reflected light (E) (white arrows: borders between well discernible flagellomeres; grey arrows: incomplete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245393/files/figure.png" pageId="5" pageNumber="352">Fig. 3</figureCitation>
|
||
D, E) than that plesiomorphic state known from
|
||
<taxonomicName id="4C64F4EF7112A717FB39AFFAD721F740" box="[1230,1364,1022,1045]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="ivanovae">
|
||
<emphasis id="B910537E7112A717FB39AFFAD721F740" box="[1230,1364,1022,1045]" italics="true" pageId="5" pageNumber="352">S. ivanovae</emphasis>
|
||
</taxonomicName>
|
||
(only flagellomeres 12 and 13 fused in part;
|
||
<bibRefCitation id="EFF5F29D7112A717FDB1A825D101F76F" author="Zakrzewska" box="[582,884,1057,1082]" pageId="5" pageNumber="352" refString="Zakrzewska, M. & Gilka, W. (2014) The oldest known chironomids of the tribe Tanytarsini (Diptera: Chironomidae) indicate plesiomorphic character states. Geobios, 47, 335 - 343." type="journal article" year="2014">Zakrzewska & Giłka 2014</bibRefCitation>
|
||
: fig. 3C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |