680 lines
76 KiB
XML
680 lines
76 KiB
XML
<document id="7A90CED482125C20B15F90570539BE35" ID-CLB-Dataset="46497" ID-DOI="10.5281/zenodo.278107" ID-GBIF-Dataset="572f25c8-cd28-425b-a7cc-2df8744fc69e" ID-ISSN="1175-5326" ID-Zenodo-Dep="278107" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460380651600" checkinUser="plazi" docAuthor="Yang, Jian-Huan, Orlov, Nikolai L. & Wang, Ying-Yong" docDate="2011" docId="03CC87A2FFC9FFD8FF173B48FB21F8E0" docLanguage="en" docName="zt02936p068.pdf" docOrigin="Zootaxa 2936" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protobothrops maolanensis Yang, Orlov & Wang, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="FFF5FFDAFFC8FFDEFF803E38FFA3FFAF" masterDocTitle="A new species of pitviper of the genus Protobothrops from China (Squamata: Viperidae)" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698296810739" updateUser="ExternalLinkService">
|
||
<mods:mods id="09A718245153D830E00804EC1AE22B10" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="166459549E564CD419FAA6A70796060D">
|
||
<mods:title id="700EB57448B57FEED5349D8A2CEF71C8">A new species of pitviper of the genus Protobothrops from China (Squamata: Viperidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A1FCDE8B66C8986CF16AF65FDAF40D8D" type="personal">
|
||
<mods:role id="CB3E2B5E1873EAE38312894AD8CB02CE">
|
||
<mods:roleTerm id="56E732BB4C6000F6B11B2FF295CB7F02">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F444617E177D2E29602C83AE778A3491">Yang, Jian-Huan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FF9898F591E3F7180BB3386E00A653EF" type="personal">
|
||
<mods:role id="453F4366A7A58A173A474D4A7B5BCC0A">
|
||
<mods:roleTerm id="AA5074D2EF1F8AB86FFAB7EE6A2887A1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CF139E4C61B57BC35EBAE4554011CB3F">Orlov, Nikolai L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="30E71265C729259259D95E86C94BCDDE" type="personal">
|
||
<mods:role id="AA79198EBEFE70E8BF52403D3C53246C">
|
||
<mods:roleTerm id="72599D7D967E50AE1E6E88BF22C89EC3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E15782E8855F7F3060D3593F2882C2C8">Wang, Ying-Yong</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="5B31A1237E2605BF202F2C8D66BE28BC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="AD7AE77B7AB8BFDEA00084FFF05B99F7" type="host">
|
||
<mods:titleInfo id="19C9A85E10C14D59982537C93B86F9A3">
|
||
<mods:title id="0996329624D930E21A46712D8443AD19">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1B55D93FC6922B206139C93C6ABE7008">
|
||
<mods:date id="D1A84B389ADF3C4C46F04523DF4C3E84">2011</mods:date>
|
||
<mods:detail id="AECEA329BD7218C37298ED1CE5884599" type="volume">
|
||
<mods:number id="EF9EE96AAD0847260DF4B392D6F0991A">2936</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6AA7D966A9C190F1F625C61894FAC3AA" unit="page">
|
||
<mods:start id="515E9C47E7FAE8B0E9530529897A244A">59</mods:start>
|
||
<mods:end id="B25BD2F81D466B2328064BE86BC972E7">68</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="A51BB9D9387FAB055E0C65488027B3AF">journal article</mods:classification>
|
||
<mods:identifier id="F8CC68B2890709506C42085B6E1E572A" type="CLB-Dataset">46497</mods:identifier>
|
||
<mods:identifier id="190231B234B5761BD9A4F9AFD9FC435B" type="DOI">10.5281/zenodo.278107</mods:identifier>
|
||
<mods:identifier id="9C7AEA989F0D988247707BF0A4A50870" type="GBIF-Dataset">572f25c8-cd28-425b-a7cc-2df8744fc69e</mods:identifier>
|
||
<mods:identifier id="9B357B4A4FAE062C41448E3D144C91BB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="7A8E657BF926E557278012EC2FD539F8" type="Zenodo-Dep">278107</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03CC87A2FFC9FFD8FF173B48FB21F8E0" ID-DOI="http://doi.org/10.5281/zenodo.5696976" ID-GBIF-Taxon="119414136" ID-Zenodo-Dep="5696976" LSID="urn:lsid:plazi:treatment:03CC87A2FFC9FFD8FF173B48FB21F8E0" httpUri="http://treatment.plazi.org/id/03CC87A2FFC9FFD8FF173B48FB21F8E0" lastPageId="6" lastPageNumber="65" pageId="1" pageNumber="60">
|
||
<subSubSection id="C37F653FFFC9FFDFFF173B48FED0FA05" pageId="1" pageNumber="60" type="nomenclature">
|
||
<paragraph id="8BDA36B4FFC9FFDFFF173B48FDE4FA25" blockId="1.[151,583,1392,1450]" box="[151,583,1392,1418]" pageId="1" pageNumber="60">
|
||
<heading id="D09281D8FFC9FFDFFF173B48FDE4FA25" bold="true" box="[151,583,1392,1418]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
|
||
<emphasis id="B911EAA6FFC9FFDFFF173B48FDE4FA25" bold="true" box="[151,583,1392,1418]" pageId="1" pageNumber="60">
|
||
<taxonomicName id="4C654D37FFC9FFDFFF173B48FE40FA25" box="[151,483,1392,1418]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFC9FFDFFF173B48FE40FA25" bold="true" box="[151,483,1392,1418]" italics="true" pageId="1" pageNumber="60">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257DDFFC9FFDFFE693B48FDE4FA25" box="[489,583,1392,1418]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFC9FFDFFF173BAAFED0FA05" blockId="1.[151,583,1392,1450]" box="[151,371,1426,1450]" pageId="1" pageNumber="60">
|
||
(
|
||
<figureCitation id="135E2A31FFC9FFDFFF1F3BAAFEA9FA05" box="[159,266,1426,1450]" captionStart-0="FIGURE 1. A – E" captionStart-1="FIGURE 2. A" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1952,1975]" captionStartId-1="3.[151,250,1934,1957]" captionStartId-2="5.[151,250,1955,1978]" captionTargetBox-0="[191,1391,195,1910]" captionTargetBox-1="[195,1392,193,1901]" captionTargetBox-2="[151,1436,1140,1933]" captionTargetId-0="figure@2.[191,1392,195,1910]" captionTargetId-1="figure@3.[195,1392,193,1903]" captionTargetId-2="figure@5.[151,1436,1140,1934]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 1. A – E (Holotype of Protobothrops maolanensis sp. nov., SYSr 000211). A: General aspect in life; B: dorsal views of head; C: ventral views of head; D: lateral views of head; E: dorsal view of midbody (showing thin dark brown bands and keeled scales). F (Paratype, SYS r 000210): left hemipenis in asulcate view. Photographs by J. H. Yang." captionText-1="FIGURE 2. A: paratype of Protobothrops maolanensis sp. nov. (female, SYS r 000277); B: holotype of P. trungkhanhensis (male); C: paratype of P. trungkhanhensis (female); D: karst forest in Maolan National Nature Reserve; E: microhabitats of the type locality. Photographs by Sheng Zheng (A), N. L. Orlov (B, C) and J. H. Yang (D, E)." captionText-2="FIGURE 3. The type locality of Protobothrops maolanensis sp. nov., Maolan National Nature Reserve (circle, “ MNNR ”), Maolan town, Guizhou, China." httpUri-0="https://zenodo.org/record/278108/files/figure.png" httpUri-1="https://zenodo.org/record/278109/files/figure.png" httpUri-2="https://zenodo.org/record/278110/files/figure.png" pageId="1" pageNumber="60">Figs. 1–3</figureCitation>
|
||
,
|
||
<tableCitation id="C6E7030FFFC9FFDFFE963BAAFEC4FA05" box="[278,359,1426,1450]" captionStart="TABLE 1" captionStartId="4.[151,235,368,389]" captionTargetBox="[151,1436,448,1079]" captionTargetPageId="4" captionText="TABLE 1. Measurements and scale characters of the type specimens of Protobothrops maolanensis sp. nov. See text for abbreviations." httpUri="http://table.plazi.org/id/DF1A663CFFCCFFDAFF173F48FF4DFE0B" pageId="1" pageNumber="60" tableUuid="DF1A663CFFCCFFDAFF173F48FF4DFE0B">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F653FFFC9FFDFFF173BE3FE8DF908" pageId="1" pageNumber="60" type="materials_examined">
|
||
<paragraph id="8BDA36B4FFC9FFDFFF173BE3FE97F995" blockId="1.[151,1437,1498,2030]" pageId="1" pageNumber="60">
|
||
<emphasis id="B911EAA6FFC9FFDFFF173BE3FEA8FA5C" bold="true" box="[151,267,1499,1523]" pageId="1" pageNumber="60">
|
||
<typeStatus id="54DE8816FFC9FFDFFF173BE3FEA5FA5C" box="[151,262,1499,1523]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
SYS r000211, adult male (
|
||
<figureCitation id="135E2A31FFC9FFDFFDC43BE2FD25FA5D" box="[580,646,1498,1522]" captionStart="FIGURE 1. A – E" captionStartId="2.[151,250,1952,1975]" captionTargetBox="[191,1391,195,1910]" captionTargetId="figure@2.[191,1392,195,1910]" captionTargetPageId="2" captionText="FIGURE 1. A – E (Holotype of Protobothrops maolanensis sp. nov., SYSr 000211). A: General aspect in life; B: dorsal views of head; C: ventral views of head; D: lateral views of head; E: dorsal view of midbody (showing thin dark brown bands and keeled scales). F (Paratype, SYS r 000210): left hemipenis in asulcate view. Photographs by J. H. Yang." httpUri="https://zenodo.org/record/278108/files/figure.png" pageId="1" pageNumber="60">Fig. 1</figureCitation>
|
||
) from Maolan National Nature Reserve, Maolan town, Libo County, Guizhou,
|
||
<collectingCountry id="F3727624FFC9FFDFFE863BC7FEEAF9B8" box="[262,329,1535,1559]" name="China" pageId="1" pageNumber="60">China</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE515073FFC9FFDFFED83BC7FE58F9B8" box="[344,507,1535,1559]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="1" value="25.21389">25°12'50.0'' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE515073FFC9FFDFFD873BC7FD61F9B8" box="[519,706,1535,1559]" direction="east" orientation="longitude" pageId="1" pageNumber="60" precision="1" value="107.99889">107°59'56.04'' E</geoCoordinate>
|
||
,
|
||
<quantity id="4C9D9B51FFC9FFDFFD4D3BC7FCB6F9B8" box="[717,789,1535,1559]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="60" unit="m" value="560.0">560 m</quantity>
|
||
a.s.l.), collected by Sheng Zheng and Jian-Huan Yang on
|
||
<date id="FFDB1074FFC9FFDFFF17381AFE93F995" box="[151,304,1570,1595]" pageId="1" pageNumber="60" value="2010-05-18">18 May 2010</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFC9FFDFFF45387FFE8DF908" blockId="1.[151,1437,1498,2030]" pageId="1" pageNumber="60">
|
||
<emphasis id="B911EAA6FFC9FFDFFF45387FFEE7F9F0" bold="true" box="[197,324,1607,1631]" pageId="1" pageNumber="60">
|
||
<typeStatus id="54DE8816FFC9FFDFFF45387FFE9CF9F0" box="[197,319,1607,1631]" pageId="1" pageNumber="60" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
SYS r000210, adult male, same data as
|
||
<typeStatus id="54DE8816FFC9FFDFFCA4387FFC29F9F0" box="[804,906,1607,1631]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
. SYS r000276, adult male and SYS r000277, adult female, from the same locality as
|
||
<typeStatus id="54DE8816FFC9FFDFFDEA3852FD6CF92D" box="[618,719,1642,1666]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
, collected by Jian-Huan Yang, Yun Li and Run-Lin Li on
|
||
<date id="FFDB1074FFC9FFDFFA003852FE88F908" pageId="1" pageNumber="60" value="2010-08-10">10 August 2010</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F653FFFC9FFDFFF45388AFF43F841" pageId="1" pageNumber="60" type="diagnosis">
|
||
<paragraph id="8BDA36B4FFC9FFDFFF45388AFF43F841" blockId="1.[151,1437,1498,2030]" pageId="1" pageNumber="60">
|
||
<emphasis id="B911EAA6FFC9FFDFFF45388AFE9CF964" bold="true" box="[197,319,1714,1739]" pageId="1" pageNumber="60">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C654D37FFC9FFDFFEC8388BFDDDF964" box="[328,638,1715,1739]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFC9FFDFFEC8388BFDDDF964" box="[328,638,1715,1739]" italics="true" pageId="1" pageNumber="60">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFC9FFDFFD07388BFD41F964" bold="true" box="[647,738,1715,1739]" pageId="1" pageNumber="60">
|
||
<taxonomicNameLabel id="A22257DDFFC9FFDFFD07388BFD41F964" box="[647,738,1715,1739]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from other species of
|
||
<taxonomicName id="4C654D37FFC9FFDFFBB8388BFB7AF964" box="[1080,1241,1715,1739]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="genus">
|
||
<emphasis id="B911EAA6FFC9FFDFFBB8388BFB7AF964" box="[1080,1241,1715,1739]" italics="true" pageId="1" pageNumber="60">Protobothrops</emphasis>
|
||
</taxonomicName>
|
||
by the combination of the following characters: (1) a relatively small body size (TL
|
||
<quantity id="4C9D9B51FFC9FFDFFC5138EFFBCCF940" box="[977,1135,1751,1775]" metricMagnitude="-1" metricUnit="m" metricValue="7.07" metricValueMax="8.05" metricValueMin="6.09" pageId="1" pageNumber="60" unit="mm" value="707.0" valueMax="805.0" valueMin="609.0">609–805 mm</quantity>
|
||
); (2) dorsal scales keeled throughout, in 23(rarely 25): 21(19): 17(15) rows; (3) ventral scales
|
||
<quantity id="4C9D9B51FFC9FFDFFC2138C2FB82F8BD" box="[929,1057,1786,1810]" metricMagnitude="0" metricUnit="m" metricValue="4.8133" metricValueMax="4.9022" metricValueMin="4.7244" pageId="1" pageNumber="60" unit="in" value="189.5" valueMax="193.0" valueMin="186.0">186–193 in</quantity>
|
||
male,
|
||
<quantity id="4C9D9B51FFC9FFDFFBEF38C2FB1BF8BD" box="[1135,1208,1786,1810]" metricMagnitude="0" metricUnit="m" metricValue="4.826" pageId="1" pageNumber="60" unit="in" value="190.0">190 in</quantity>
|
||
female; (4) subcaudals 78–85 pairs in male,
|
||
<quantity id="4C9D9B51FFC9FFDFFE383927FE52F898" box="[440,497,1823,1847]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" pageId="1" pageNumber="60" unit="in" value="74.0">74 in</quantity>
|
||
female; (5) 7–8 supralabials; (6) 11–12 infralabials; (7) 10–12 scales between midsection of supraoculars in a line; (8) the 4th supralabials separated from suboculars by one row of scales; (9) 68–72 small and thin body bands across body and tail; (10) TaL/SVL ratio
|
||
<quantity id="4C9D9B51FFC9FFDFFC0F3951FB9BF82E" box="[911,1080,1897,1921]" metricMagnitude="-3" metricUnit="m" metricValue="6.438899999999999" metricValueMax="6.832600000000001" metricValueMin="6.0451999999999995" pageId="1" pageNumber="60" unit="in" value="0.2535" valueMax="0.269" valueMin="0.238">0.238–0.269 in</quantity>
|
||
male,
|
||
<quantity id="4C9D9B51FFC9FFDFFB023951FB7DF82E" box="[1154,1246,1897,1921]" metricMagnitude="-3" metricUnit="m" metricValue="4.826" pageId="1" pageNumber="60" unit="in" value="0.19">0.190 in</quantity>
|
||
female; (11) SL/ HL ratio
|
||
<quantity id="4C9D9B51FFC9FFDFFE8039B5FE09F80A" box="[256,426,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="8.0518" metricValueMax="8.2804" metricValueMin="7.8232" pageId="1" pageNumber="60" unit="in" value="0.317" valueMax="0.326" valueMin="0.308">0.308–0.326 in</quantity>
|
||
male,
|
||
<quantity id="4C9D9B51FFC9FFDFFE7739B5FDF7F80A" box="[503,596,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="8.7376" pageId="1" pageNumber="60" unit="in" value="0.344">0.344 in</quantity>
|
||
female; (12) ED/HL ratio
|
||
<quantity id="4C9D9B51FFC9FFDFFC0439B5FB8CF80A" box="[900,1071,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="4.318" metricValueMax="4.4704" metricValueMin="4.1656" pageId="1" pageNumber="60" unit="in" value="0.16999999999999998" valueMax="0.176" valueMin="0.164">0.164–0.176 in</quantity>
|
||
male,
|
||
<quantity id="4C9D9B51FFC9FFDFFBFC39B5FB7AF80A" box="[1148,1241,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="3.8608" pageId="1" pageNumber="60" unit="in" value="0.152">0.152 in</quantity>
|
||
female; and (13) the unique color pattern. See below for differential diagnosis by way of comparison with all other species in the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1A663CFFCAFFDCFF173998FBFEF85B" httpUri="https://zenodo.org/record/278108/files/figure.png" pageId="2" pageNumber="61" targetBox="[191,1391,195,1910]" targetPageId="2">
|
||
<paragraph id="8BDA36B4FFCAFFDCFF173998FBFEF85B" blockId="2.[151,1436,1952,2036]" pageId="2" pageNumber="61">
|
||
<emphasis id="B911EAA6FFCAFFDCFF173998FEB1F819" bold="true" box="[151,274,1952,1975]" pageId="2" pageNumber="61">FIGURE 1.</emphasis>
|
||
A–E (Holotype of
|
||
<taxonomicName id="4C654D37FFCAFFDCFE533999FD49F818" box="[467,746,1953,1975]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCAFFDCFE533999FD49F818" box="[467,746,1953,1975]" italics="true" pageId="2" pageNumber="61">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCAFFDCFD6F3999FC9CF819" bold="true" box="[751,831,1953,1974]" pageId="2" pageNumber="61">
|
||
<taxonomicNameLabel id="A22257DDFFCAFFDCFD6F3999FC9CF819" box="[751,831,1953,1974]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, SYSr000211). A: General aspect in life; B: dorsal views of head; C: ventral views of head; D: lateral views of head; E: dorsal view of midbody (showing thin dark brown bands and keeled scales). F (Paratype, SYS r000210): left hemipenis in asulcate view. Photographs by J.H. Yang.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A663CFFCBFFDDFF1739B6FB87F84D" httpUri="https://zenodo.org/record/278109/files/figure.png" pageId="3" pageNumber="62" targetBox="[195,1392,193,1901]" targetPageId="3">
|
||
<paragraph id="8BDA36B4FFCBFFDDFF1739B6FB87F84D" blockId="3.[151,1436,1934,2018]" pageId="3" pageNumber="62">
|
||
<emphasis id="B911EAA6FFCBFFDDFF1739B6FEB6F80B" bold="true" box="[151,277,1934,1957]" pageId="3" pageNumber="62">FIGURE 2.</emphasis>
|
||
A: paratype of
|
||
<taxonomicName id="4C654D37FFCBFFDDFE3D39B7FD75F80A" box="[445,726,1935,1957]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCBFFDDFE3D39B7FD75F80A" box="[445,726,1935,1957]" italics="true" pageId="3" pageNumber="62">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257DDFFCBFFDDFD5E39B7FC92F80B" box="[734,817,1935,1956]" pageId="3" pageNumber="62" rank="species">
|
||
<emphasis id="B911EAA6FFCBFFDDFD5E39B7FC88F80B" bold="true" box="[734,811,1935,1956]" pageId="3" pageNumber="62">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(female, SYS r000277); B: holotype of
|
||
<taxonomicName id="4C654D37FFCBFFDDFB5939A8FA38F80A" box="[1241,1435,1935,1957]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="trungkhanhensis">
|
||
<emphasis id="B911EAA6FFCBFFDDFB5939A8FA38F80A" box="[1241,1435,1935,1957]" italics="true" pageId="3" pageNumber="62">P. trungkhanhensis</emphasis>
|
||
</taxonomicName>
|
||
(male); C: paratype of
|
||
<taxonomicName id="4C654D37FFCBFFDDFEFD3996FD9DF86C" box="[381,574,1965,1987]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="trungkhanhensis">
|
||
<emphasis id="B911EAA6FFCBFFDDFEFD3996FD9DF86C" box="[381,574,1965,1987]" italics="true" pageId="3" pageNumber="62">P. trungkhanhensis</emphasis>
|
||
</taxonomicName>
|
||
(female); D: karst forest in Maolan National Nature Reserve; E: microhabitats of the type locality. Photographs by Sheng Zheng (A), N. L. Orlov (B, C) and J. H. Yang (D, E).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37F653FFFCCFFDBFF453EAFFA21FB90" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63" type="description">
|
||
<paragraph id="8BDA36B4FFCCFFDAFF453EAFFD27FE90" blockId="4.[151,1436,151,320]" pageId="4" pageNumber="63">
|
||
<emphasis id="B911EAA6FFCCFFDAFF453EAFFE47FF1F" bold="true" box="[197,484,151,176]" pageId="4" pageNumber="63">
|
||
Description of
|
||
<typeStatus id="54DE8816FFCCFFDAFEF63EA0FE7CFF1F" box="[374,479,152,176]" pageId="4" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male with detailed measurements as listed in
|
||
<tableCitation id="C6E7030FFFCCFFDAFBB63EAFFB28FF00" box="[1078,1163,151,175]" captionStart="TABLE 1" captionStartId="4.[151,235,368,389]" captionTargetBox="[151,1436,448,1079]" captionTargetPageId="4" captionText="TABLE 1. Measurements and scale characters of the type specimens of Protobothrops maolanensis sp. nov. See text for abbreviations." httpUri="http://table.plazi.org/id/DF1A663CFFCCFFDAFF173F48FF4DFE0B" pageId="4" pageNumber="63" tableUuid="DF1A663CFFCCFFDAFF173F48FF4DFE0B">Table 1</tableCitation>
|
||
. Body moderately elongate, thin and slightly compressed; head elongate and triangular, distinctly wider than neck, covered with very small, convex and irregular shaped scales, 0.69 times wider than long; upper head scales smooth anteriorly, keeled on occipital, 10 scales between midsection of supraoculars in a transverse line; snout elongate, SL/HL ratio 0.31, nearly twice ED; eye convex, pupil vertical.
|
||
</paragraph>
|
||
<caption id="DF1A663CFFCCFFDAFF173F48FF4DFE0B" ID-Table-UUID="DF1A663CFFCCFFDAFF173F48FF4DFE0B" httpUri="http://table.plazi.org/id/DF1A663CFFCCFFDAFF173F48FF4DFE0B" pageId="4" pageNumber="63" targetBox="[151,1436,448,1079]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BDA36B4FFCCFFDAFF173F48FF4DFE0B" blockId="4.[151,1436,368,420]" pageId="4" pageNumber="63">
|
||
<emphasis id="B911EAA6FFCCFFDAFF173F48FEA7FE2A" bold="true" box="[151,260,368,389]" pageId="4" pageNumber="63">TABLE 1.</emphasis>
|
||
Measurements and scale characters of the type specimens of
|
||
<taxonomicName id="4C654D37FFCCFFDAFCEF3F48FB27FE29" box="[879,1156,368,390]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCCFFDAFCEF3F48FB27FE29" box="[879,1156,368,390]" italics="true" pageId="4" pageNumber="63">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCCFFDAFB0B3F48FB78FE2A" bold="true" box="[1163,1243,368,389]" pageId="4" pageNumber="63">
|
||
<taxonomicNameLabel id="A22257DDFFCCFFDAFB0B3F48FB78FE2A" box="[1163,1243,368,389]" pageId="4" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
See text for abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA36B4FFCCFFDAFD963FF8FB06FB98" pageId="4" pageNumber="63">
|
||
<table id="F965C414FFCC0021FF173FF8FA3FFB98" box="[151,1436,448,1079]" gridcols="5" gridrows="6" pageId="4" pageNumber="63">
|
||
<tr id="355534F6FFCC0021FF173FF8FA3FFE5B" box="[151,1436,448,500]" gridrow="0" pageId="4" pageNumber="63" rowspan-0="1">
|
||
<th id="76845D8AFFCC0021FD963FF8FD3DFE5B" box="[534,670,448,500]" gridcol="1" gridrow="0" pageId="4" pageNumber="63">SYS r000211 Holotype</th>
|
||
<th id="76845D8AFFCC0021FD613FF8FCC9FE5B" box="[737,874,448,500]" gridcol="2" gridrow="0" pageId="4" pageNumber="63">SYS r000210 Paratype</th>
|
||
<th id="76845D8AFFCC0021FC333FF8FB9FFE5B" box="[947,1084,448,500]" gridcol="3" gridrow="0" pageId="4" pageNumber="63">SYS r000276 Paratype</th>
|
||
<th id="76845D8AFFCC0021FB053FF8FA3FFE5B" box="[1157,1436,448,500]" gridcol="4" gridrow="0" pageId="4" pageNumber="63">SYS r000277 Paratype</th>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173C30FA3FFD36" box="[151,1436,520,665]" gridrow="1" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173C30FEB4FD36" box="[151,279,520,665]" gridcol="0" gridrow="1" pageId="4" pageNumber="63">Sex SVL TaL TaL/SVL</th>
|
||
<td id="76845D8AFFCC0021FD963C30FD3DFD36" box="[534,670,520,665]" gridcol="1" gridrow="1" pageId="4" pageNumber="63">male 634 171 0.27</td>
|
||
<td id="76845D8AFFCC0021FD613C30FCC9FD36" box="[737,874,520,665]" gridcol="2" gridrow="1" pageId="4" pageNumber="63">male 638 160 0.25</td>
|
||
<td id="76845D8AFFCC0021FC333C30FB9FFD36" box="[947,1084,520,665]" gridcol="3" gridrow="1" pageId="4" pageNumber="63">male 492 117 0.24</td>
|
||
<td id="76845D8AFFCC0021FB053C30FA3FFD36" box="[1157,1436,520,665]" gridcol="4" gridrow="1" pageId="4" pageNumber="63">female 521 99 0.19</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173C95FA3FFD43" box="[151,1436,685,748]" gridrow="2" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173C95FEB4FD43" box="[151,279,685,748]" gridcol="0" gridrow="2" pageId="4" pageNumber="63">TL HW</th>
|
||
<td id="76845D8AFFCC0021FD963C95FD3DFD43" box="[534,670,685,748]" gridcol="1" gridrow="2" pageId="4" pageNumber="63">805 16.6</td>
|
||
<td id="76845D8AFFCC0021FD613C95FCC9FD43" box="[737,874,685,748]" gridcol="2" gridrow="2" pageId="4" pageNumber="63">798 16.6</td>
|
||
<td id="76845D8AFFCC0021FC333C95FB9FFD43" box="[947,1084,685,748]" gridcol="3" gridrow="2" pageId="4" pageNumber="63">609 13.8</td>
|
||
<td id="76845D8AFFCC0021FB053C95FA3FFD43" box="[1157,1436,685,748]" gridcol="4" gridrow="2" pageId="4" pageNumber="63">620 14.8</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173D38FA3FFC3E" box="[151,1436,768,913]" gridrow="3" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173D38FEB4FC3E" box="[151,279,768,913]" gridcol="0" gridrow="3" pageId="4" pageNumber="63">HL HW/HL SL SL/HL</th>
|
||
<td id="76845D8AFFCC0021FD963D38FD3DFC3E" box="[534,670,768,913]" gridcol="1" gridrow="3" pageId="4" pageNumber="63">24.0 0.69 7.7 0.32</td>
|
||
<td id="76845D8AFFCC0021FD613D38FCC9FC3E" box="[737,874,768,913]" gridcol="2" gridrow="3" pageId="4" pageNumber="63">24.5 0.68 7.5 0.31</td>
|
||
<td id="76845D8AFFCC0021FC333D38FB9FFC3E" box="[947,1084,768,913]" gridcol="3" gridrow="3" pageId="4" pageNumber="63">19.6 0.70 6.4 0.33</td>
|
||
<td id="76845D8AFFCC0021FB053D38FA3FFC3E" box="[1157,1436,768,913]" gridcol="4" gridrow="3" pageId="4" pageNumber="63">21.7 0.68 7.5 0.35</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173D9DFA3FFC4B" box="[151,1436,933,996]" gridrow="4" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173D9DFEB4FC4B" box="[151,279,933,996]" gridcol="0" gridrow="4" pageId="4" pageNumber="63">ED ED/HL</th>
|
||
<td id="76845D8AFFCC0021FD963D9DFD3DFC4B" box="[534,670,933,996]" gridcol="1" gridrow="4" pageId="4" pageNumber="63">4.2 0.18</td>
|
||
<td id="76845D8AFFCC0021FD613D9DFCC9FC4B" box="[737,874,933,996]" gridcol="2" gridrow="4" pageId="4" pageNumber="63">4.3 0.18</td>
|
||
<td id="76845D8AFFCC0021FC333D9DFB9FFC4B" box="[947,1084,933,996]" gridcol="3" gridrow="4" pageId="4" pageNumber="63">3.2 0.16</td>
|
||
<td id="76845D8AFFCC0021FB053D9DFA3FFC4B" box="[1157,1436,933,996]" gridcol="4" gridrow="4" pageId="4" pageNumber="63">3.3 0.15</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173DC0FA3FFB98" box="[151,1436,1016,1079]" gridrow="5" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173DC0FEB4FB98" box="[151,279,1016,1079]" gridcol="0" gridrow="5" pageId="4" pageNumber="63">EN supralabials</th>
|
||
<td id="76845D8AFFCC0021FD963DC0FD3DFB98" box="[534,670,1016,1079]" gridcol="1" gridrow="5" pageId="4" pageNumber="63">4.8 8/8</td>
|
||
<td id="76845D8AFFCC0021FD613DC0FCC9FB98" box="[737,874,1016,1079]" gridcol="2" gridrow="5" pageId="4" pageNumber="63">4.9 7/8</td>
|
||
<td id="76845D8AFFCC0021FC333DC0FB9FFB98" box="[947,1084,1016,1079]" gridcol="3" gridrow="5" pageId="4" pageNumber="63">4.2 8/8</td>
|
||
<td id="76845D8AFFCC0021FB053DC0FA3FFB98" box="[1157,1436,1016,1079]" gridcol="4" gridrow="5" pageId="4" pageNumber="63">4.8 8/8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCCFFDAFF1E3A75FB3DF9B0" pageId="4" pageNumber="63">
|
||
<table id="F965C414FFCC0021FF173A72FA3FF9B0" box="[151,1436,1098,1567]" gridcols="5" gridrows="6" pageId="4" pageNumber="63">
|
||
<tr id="355534F6FFCC0021FF173A72FA3FFB23" box="[151,1436,1098,1164]" gridrow="0" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173A72FE4DFB23" box="[151,494,1098,1164]" gridcol="0" gridrow="0" pageId="4" pageNumber="63">3rd supralabials contact subocular infralabials</th>
|
||
<th id="76845D8AFFCC0021FD963A72FD12FB23" box="[534,689,1098,1164]" gridcol="1" gridrow="0" pageId="4" pageNumber="63">no 11/12</th>
|
||
<th id="76845D8AFFCC0021FD613A72FCDFFB23" box="[737,892,1098,1164]" gridcol="2" gridrow="0" pageId="4" pageNumber="63">yes 11/12</th>
|
||
<th id="76845D8AFFCC0021FC333A72FBEDFB23" box="[947,1102,1098,1164]" gridcol="3" gridrow="0" pageId="4" pageNumber="63">yes 12/12</th>
|
||
<th id="76845D8AFFCC0021FB053A72FA3FFB23" box="[1157,1436,1098,1164]" gridcol="4" gridrow="0" pageId="4" pageNumber="63">no 11/11</th>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173A98FA3FFB1A" box="[151,1436,1184,1205]" gridrow="1" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173A98FE4DFB1A" box="[151,494,1184,1205]" gridcol="0" gridrow="1" pageId="4" pageNumber="63">SBSO</th>
|
||
<td id="76845D8AFFCC0021FD963A98FD12FB1A" box="[534,689,1184,1205]" gridcol="1" gridrow="1" pageId="4" pageNumber="63">10</td>
|
||
<td id="76845D8AFFCC0021FD613A98FCDFFB1A" box="[737,892,1184,1205]" gridcol="2" gridrow="1" pageId="4" pageNumber="63">11</td>
|
||
<td id="76845D8AFFCC0021FC333A98FBEDFB1A" box="[947,1102,1184,1205]" gridcol="3" gridrow="1" pageId="4" pageNumber="63">12</td>
|
||
<td id="76845D8AFFCC0021FB053A98FA3FFB1A" box="[1157,1436,1184,1205]" gridcol="4" gridrow="1" pageId="4" pageNumber="63">10</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173AF2FA3FFAA7" box="[151,1436,1226,1288]" gridrow="2" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173AF2FE4DFAA7" box="[151,494,1226,1288]" gridcol="0" gridrow="2" pageId="4" pageNumber="63">L canthal scales</th>
|
||
<td id="76845D8AFFCC0021FD963AF2FD12FAA7" box="[534,689,1226,1288]" gridcol="1" gridrow="2" pageId="4" pageNumber="63">2 2/2</td>
|
||
<td id="76845D8AFFCC0021FD613AF2FCDFFAA7" box="[737,892,1226,1288]" gridcol="2" gridrow="2" pageId="4" pageNumber="63">2 3/3</td>
|
||
<td id="76845D8AFFCC0021FC333AF2FBEDFAA7" box="[947,1102,1226,1288]" gridcol="3" gridrow="2" pageId="4" pageNumber="63">2 3/3</td>
|
||
<td id="76845D8AFFCC0021FB053AF2FA3FFAA7" box="[1157,1436,1226,1288]" gridcol="4" gridrow="2" pageId="4" pageNumber="63">2 4/4</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173B24FA3FFA02" box="[151,1436,1308,1453]" gridrow="3" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173B24FE4DFA02" box="[151,494,1308,1453]" gridcol="0" gridrow="3" pageId="4" pageNumber="63">pre-V V Scd Sq1: Sq2: Sq3</th>
|
||
<td id="76845D8AFFCC0021FD963B24FD12FA02" box="[534,689,1308,1453]" gridcol="1" gridrow="3" pageId="4" pageNumber="63">2 186 85 25: 21: 17</td>
|
||
<td id="76845D8AFFCC0021FD613B24FCDFFA02" box="[737,892,1308,1453]" gridcol="2" gridrow="3" pageId="4" pageNumber="63">2 188 78 23: 19: 15</td>
|
||
<td id="76845D8AFFCC0021FC333B24FBEDFA02" box="[947,1102,1308,1453]" gridcol="3" gridrow="3" pageId="4" pageNumber="63">1 193 80 23: 21: 17</td>
|
||
<td id="76845D8AFFCC0021FB053B24FA3FFA02" box="[1157,1436,1308,1453]" gridcol="4" gridrow="3" pageId="4" pageNumber="63">2 190 74 23: 21: 17</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173BF9FA3FFA5A" box="[151,1436,1473,1525]" gridrow="4" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173BF9FE4DFA5A" box="[151,494,1473,1525]" gridcol="0" gridrow="4" pageId="4" pageNumber="63">outermost row of dorsal scales</th>
|
||
<td id="76845D8AFFCC0021FD963BF9FD12FA5A" box="[534,689,1473,1525]" gridcol="1" gridrow="4" pageId="4" pageNumber="63">strongly keeled</td>
|
||
<td id="76845D8AFFCC0021FD613BF9FCDFFA5A" box="[737,892,1473,1525]" gridcol="2" gridrow="4" pageId="4" pageNumber="63">strongly keeled</td>
|
||
<td id="76845D8AFFCC0021FC333BF9FBEDFA5A" box="[947,1102,1473,1525]" gridcol="3" gridrow="4" pageId="4" pageNumber="63">strongly keeled</td>
|
||
<td id="76845D8AFFCC0021FB053BF9FA3FFA5A" box="[1157,1436,1473,1525]" gridcol="4" gridrow="4" pageId="4" pageNumber="63">smooth anteriorly, weakly keeled posteriorly</td>
|
||
</tr>
|
||
<tr id="355534F6FFCC0021FF173831FA3FF9B0" box="[151,1436,1545,1567]" gridrow="5" pageId="4" pageNumber="63">
|
||
<th id="76845D8AFFCC0021FF173831FE4DF9B0" box="[151,494,1545,1567]" gridcol="0" gridrow="5" pageId="4" pageNumber="63">body bands</th>
|
||
<td id="76845D8AFFCC0021FD963831FD12F9B0" box="[534,689,1545,1567]" gridcol="1" gridrow="5" pageId="4" pageNumber="63">71</td>
|
||
<td id="76845D8AFFCC0021FD613831FCDFF9B0" box="[737,892,1545,1567]" gridcol="2" gridrow="5" pageId="4" pageNumber="63">72</td>
|
||
<td id="76845D8AFFCC0021FC333831FBEDF9B0" box="[947,1102,1545,1567]" gridcol="3" gridrow="5" pageId="4" pageNumber="63">68</td>
|
||
<td id="76845D8AFFCC0021FB053831FA3FF9B0" box="[1157,1436,1545,1567]" gridcol="4" gridrow="5" pageId="4" pageNumber="63">72</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCCFFDBFF45385CFAABFF52" blockId="4.[151,1437,1636,2022]" lastBlockId="5.[151,1436,151,1088]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
|
||
Rostral triangular, about 1.3 times broader than high, barely visible from above, broader posteriorly with three apical scales bordered laterally by internasals; internasals not in contact with rostral and separated from each other by four small scales; nasal trapezoid, undivided, with a round nostril opening in middle; 2/2 (L and R, hereafter) canthal scales between supraocular and internasal, distinctly larger than adjacent snout scales, anterior one largest, bordering sharp, raised
|
||
<emphasis id="B911EAA6FFCCFFDAFE2738CCFDC5F8A3" box="[423,614,1780,1804]" italics="true" pageId="4" pageNumber="63">canthus rostralis</emphasis>
|
||
; two loreals, approximately equal size; supraocular large and elongate, wider than adjacent head scales, pointed anteriorly and 0.58 times as wide as interspace between supraoculars and surrounded by 13/13 small scales; two elongate upper-preoculars above loreal pit in contact with posterior loreal, lower one longer than upper and forming upper margin of loreal pit; one elongate lower-preocular, forming lower margin of loreal pit; two small postoculars; one elongate, thin, crescent-shaped subocular, separated from lower preocular by one scale; temporals numerous, upper ones keeled, lower ones smooth, those bordering supralabials larger than latter; anterior part of nasal large, turning over
|
||
<emphasis id="B911EAA6FFCCFFDAFCA739F6FC47F849" box="[807,996,1998,2022]" italics="true" pageId="4" pageNumber="63">canthus rostralis</emphasis>
|
||
and visible above; 8/8 supralabials, 2nd forming anterior margin of loreal pit and separated from nasal by one small scale, 3rd largest, 3rd and 4th separated from subocular by a row of scales on both sides; 11/12 infralabials, 1st pair in contact with each other, first three in contact with anterior chin shield; posterior chin shield small, scarcely differentiated from postgenials.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCDFFDBFF453F31FEF9FEC6" blockId="5.[151,1436,151,1088]" pageId="5" pageNumber="64">Dorsal scales narrow, pointed and strongly keeled throughout; in 25 rows on anterior body, 21 rows at midbody, 17 rows at posterior of body; pre-ventral scales two; ventral scales 186; anal scale undivided; subcaudals divided, 85 pairs.</paragraph>
|
||
<paragraph id="8BDA36B4FFCDFFDBFF453F4DFD62FDCA" blockId="5.[151,1436,151,1088]" pageId="5" pageNumber="64">
|
||
<emphasis id="B911EAA6FFCDFFDBFF453F4DFE34FE21" bold="true" box="[197,407,373,398]" pageId="5" pageNumber="64">Coloration in life.</emphasis>
|
||
Grayish brown above, with 71 dark brown transverse bands with discontinuous pale yellow rough edges, across body and tail. The dark brown bands are relatively small and thin, occupy about six dorsal scale rows in width and 0.5–1.5 dorsal scales in length and often broken into spots. The bands become gradually closer towards tail; tip of tail dark brown with scatted pale spots. Underside gray, with a series of pale brown blotches on both sides. Dorsal head gray brown, with indistinctly symmetrical dark streaks. A postorbital thin dark brown stripe with discontinuous pale yellow edge, starting from posterior margin of eye, extending across upper temporals posteriorly, ending above corner of lip.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCDFFDBFF453C49FF5FFCB3" blockId="5.[151,1436,151,1088]" pageId="5" pageNumber="64">
|
||
<emphasis id="B911EAA6FFCDFFDBFF453C49FE9AFD25" bold="true" box="[197,313,625,650]" pageId="5" pageNumber="64">Variation.</emphasis>
|
||
Measurements and scale characters of all the specimens are listed in
|
||
<tableCitation id="C6E7030FFFCDFFDBFBEE3C49FB66FD26" box="[1134,1221,625,649]" captionStart="TABLE 1" captionStartId="4.[151,235,368,389]" captionTargetBox="[151,1436,448,1079]" captionTargetPageId="4" captionText="TABLE 1. Measurements and scale characters of the type specimens of Protobothrops maolanensis sp. nov. See text for abbreviations." httpUri="http://table.plazi.org/id/DF1A663CFFCCFFDAFF173F48FF4DFE0B" pageId="5" pageNumber="64" tableUuid="DF1A663CFFCCFFDAFF173F48FF4DFE0B">Table 1</tableCitation>
|
||
. All the
|
||
<typeStatus id="54DE8816FFCDFFDBFAB13C4AFA3FFD25" box="[1329,1436,626,650]" pageId="5" pageNumber="64" type="paratype">paratypes</typeStatus>
|
||
match the
|
||
<typeStatus id="54DE8816FFCDFFDBFE8F3CADFED2FD02" box="[271,369,661,685]" pageId="5" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
in overall morphology and scalation. Dorsal scales in 23 rows on anterior part of body in all three
|
||
<typeStatus id="54DE8816FFCDFFDBFF593C85FEE4FD7A" box="[217,327,701,725]" pageId="5" pageNumber="64" type="paratype">paratypes</typeStatus>
|
||
, 19 rows on mid-body in SYS r000210, 15 rows on posterior part of body in SYS r000210; 3rd supralabials touching the subocular in SYSr000210; 3/3 canthal scales in SYS r000210 and 0 0 0 276, 4/
|
||
<quantity id="4C9D9B51FFCDFFDBFAB33CD8FAC3FD57" box="[1331,1376,736,760]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="64" unit="in" value="4.0">4 in</quantity>
|
||
SYS r000277.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCDFFDBFF453D10FBE1FC27" blockId="5.[151,1436,151,1088]" pageId="5" pageNumber="64">
|
||
<emphasis id="B911EAA6FFCDFFDBFF453D10FE17FCEE" bold="true" box="[197,436,808,833]" pageId="5" pageNumber="64">Sexual dimorphism.</emphasis>
|
||
No significant sexual dimorphism occurs, other than the female having fewer subcaudals than the males (74 vs. 78–85) and the outermost row of dorsal scales are weakly keeled in the female, nearly invisible at anterior body, while the dorsal scales are strongly keeled throughout in male.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCDFFDBFF453DACFA21FB90" blockId="5.[151,1436,151,1088]" pageId="5" pageNumber="64">
|
||
<emphasis id="B911EAA6FFCDFFDBFF453DACFEE8FC02" bold="true" box="[197,331,916,941]" pageId="5" pageNumber="64">Hemipenis.</emphasis>
|
||
The everted left hemipenis (
|
||
<typeStatus id="54DE8816FFCDFFDBFD103DACFD56FC03" box="[656,757,916,940]" pageId="5" pageNumber="64" type="paratype">paratype</typeStatus>
|
||
, SYS r000210) is
|
||
<quantity id="4C9D9B51FFCDFFDBFC443DACFB8AFC03" box="[964,1065,916,940]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="5" pageNumber="64" unit="mm" value="18.5">18.5 mm</quantity>
|
||
long, robust and forked, spinous proximally and calyculate distally; it bears two lobes covered with spines and microspines, extending to the 8th subcaudal.
|
||
<emphasis id="B911EAA6FFCDFFDBFF6E3DE7FE66FC58" box="[238,453,991,1015]" italics="true" pageId="5" pageNumber="64">Sulcus spermaticus</emphasis>
|
||
forked at the truncus
|
||
<quantity id="4C9D9B51FFCDFFDBFD3B3DE7FCB2FC58" box="[699,785,991,1015]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="64" unit="mm" value="6.5">6.5 mm</quantity>
|
||
from the base of the hemipenis. Strong spines on asulcate side at the base part of each lobe, followed by smaller spines more distally; sulcate side covered with microspines. The hemipenis of
|
||
<taxonomicName id="4C654D37FFCDFFDBFEE23A10FDAAFB90" box="[354,521,1063,1087]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="maolanensis">
|
||
<emphasis id="B911EAA6FFCDFFDBFEE23A10FDAAFB90" box="[354,521,1063,1087]" italics="true" pageId="5" pageNumber="64">P. maolanensis</emphasis>
|
||
</taxonomicName>
|
||
closely resembles the
|
||
<typeStatus id="54DE8816FFCDFFDBFC873A10FC9CFBEF" box="[775,831,1064,1088]" pageId="5" pageNumber="64">Type</typeStatus>
|
||
3 spinose hemipenis of Malhotra & Thorpe (2004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1A663CFFCDFFDBFF17399BFE72F877" httpUri="https://zenodo.org/record/278110/files/figure.png" pageId="5" pageNumber="64" targetBox="[151,1436,1140,1933]" targetPageId="5">
|
||
<paragraph id="8BDA36B4FFCDFFDBFF17399BFE72F877" blockId="5.[151,1436,1955,2008]" pageId="5" pageNumber="64">
|
||
<emphasis id="B911EAA6FFCDFFDBFF17399BFEB6F817" bold="true" box="[151,277,1955,1978]" pageId="5" pageNumber="64">FIGURE 3.</emphasis>
|
||
The type locality of
|
||
<taxonomicName id="4C654D37FFCDFFDBFE76399BFCACF816" box="[502,783,1955,1977]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCDFFDBFE76399BFCACF816" box="[502,783,1955,1977]" italics="true" pageId="5" pageNumber="64">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCDFFDBFC99399CFCCFF816" bold="true" box="[793,876,1956,1977]" pageId="5" pageNumber="64">
|
||
<taxonomicNameLabel id="A22257DDFFCDFFDBFC99399CFCCFF816" box="[793,876,1956,1977]" pageId="5" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Maolan National Nature Reserve (circle, “MNNR”), Maolan town, Guizhou, China.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37F653FFFCEFFD8FF453EAFFEA6FE60" pageId="6" pageNumber="65" type="distribution">
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453EAFFEA6FE60" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453EAFFDF6FF1F" bold="true" box="[197,597,151,176]" pageId="6" pageNumber="65">Distribution and natural history.</emphasis>
|
||
Currently,
|
||
<taxonomicName id="4C654D37FFCEFFD8FD593EA1FC20FF1F" box="[729,899,152,176]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD593EA1FC20FF1F" box="[729,899,152,176]" italics="true" pageId="6" pageNumber="65">P. maolanensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54DE8816FFCEFFD8FB293EA0FB7AFF1F" box="[1193,1241,152,176]" pageId="6" pageNumber="65">type</typeStatus>
|
||
locality, Maolan National Natural Reserve, Maolan town, Guizhou,
|
||
<collectingCountry id="F3727624FFCEFFD8FD513E84FCBBFF7B" box="[721,792,188,212]" name="China" pageId="6" pageNumber="65">China</collectingCountry>
|
||
. All individuals were collected at
|
||
<quantity id="4C9D9B51FFCEFFD8FB143E84FAB1FF7B" box="[1172,1298,188,212]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="6" pageNumber="65" unit="m" value="575.0" valueMax="600.0" valueMin="550.0">550–600 m</quantity>
|
||
elevation in a subtropical karst evergreen forest. They were found on the ground under scrub at night; no individuals were observed during the day. Other species of amphibians and reptiles at the same habitat in the
|
||
<typeStatus id="54DE8816FFCEFFD8FB193F3DFB6AFEB2" box="[1177,1225,261,285]" pageId="6" pageNumber="65">type</typeStatus>
|
||
locality during the survey included
|
||
<taxonomicName id="4C654D37FFCEFFD8FEDB3F10FD33FEEF" box="[347,656,296,320]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B911EAA6FFCEFFD8FEDB3F10FD33FEEF" box="[347,656,296,320]" italics="true" pageId="6" pageNumber="65">Polypedates megacephalus</emphasis>
|
||
</taxonomicName>
|
||
(SYS a000959-962),
|
||
<taxonomicName id="4C654D37FFCEFFD8FC153F10FB01FEEF" box="[917,1186,296,320]" class="Amphibia" family="Rhacophoridae" genus="Kurixalus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="odontotarsus">
|
||
<emphasis id="B911EAA6FFCEFFD8FC153F10FB01FEEF" box="[917,1186,296,320]" italics="true" pageId="6" pageNumber="65">Kurixalus odontotarsus</emphasis>
|
||
</taxonomicName>
|
||
(SYS a000926-931),
|
||
<taxonomicName id="4C654D37FFCEFFD8FF173F74FECEFECB" box="[151,365,332,356]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="fissipes">
|
||
<emphasis id="B911EAA6FFCEFFD8FF173F74FECEFECB" box="[151,365,332,356]" italics="true" pageId="6" pageNumber="65">Microhyla fissipes</emphasis>
|
||
</taxonomicName>
|
||
(SYS a000940),
|
||
<taxonomicName id="4C654D37FFCEFFD8FDC93F74FC95FECB" box="[585,822,332,356]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="heymonsi">
|
||
<emphasis id="B911EAA6FFCEFFD8FDC93F74FC95FECB" box="[585,822,332,356]" italics="true" pageId="6" pageNumber="65">Microhyla heymonsi</emphasis>
|
||
</taxonomicName>
|
||
(SYS a000938-939),
|
||
<taxonomicName id="4C654D37FFCEFFD8FBC63F75FAECFECB" box="[1094,1359,332,356]" class="Reptilia" family="Eublepharidae" genus="Goniurosaurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="luii">
|
||
<emphasis id="B911EAA6FFCEFFD8FBC63F75FB4DFECB" box="[1094,1262,333,356]" italics="true" pageId="6" pageNumber="65">Goniurosaurus</emphasis>
|
||
cf.
|
||
<emphasis id="B911EAA6FFCEFFD8FAAA3F74FAECFECB" box="[1322,1359,332,356]" italics="true" pageId="6" pageNumber="65">luii</emphasis>
|
||
</taxonomicName>
|
||
(SYS r000217-219),
|
||
<taxonomicName id="4C654D37FFCEFFD8FEC63F48FE6EFE27" box="[326,461,368,392]" class="Reptilia" family="Lacertidae" genus="Takydromus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="genus">
|
||
<emphasis id="B911EAA6FFCEFFD8FEC63F48FE6EFE27" box="[326,461,368,392]" italics="true" pageId="6" pageNumber="65">Takydromus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C654D37FFCEFFD8FD913F48FCF2FE27" box="[529,849,367,392]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B911EAA6FFCEFFD8FD913F48FD66FE27" box="[529,709,368,392]" italics="true" pageId="6" pageNumber="65">Sphenomorphus</emphasis>
|
||
cf.
|
||
<emphasis id="B911EAA6FFCEFFD8FC803F48FCF2FE27" box="[768,849,368,392]" italics="true" pageId="6" pageNumber="65">indicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C654D37FFCEFFD8FCE63F48FB76FE27" box="[870,1237,368,392]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mucrosquamatus">
|
||
<emphasis id="B911EAA6FFCEFFD8FCE63F48FB76FE27" box="[870,1237,368,392]" italics="true" pageId="6" pageNumber="65">Protobothrops mucrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
(SYS r000238),
|
||
<taxonomicName id="4C654D37FFCEFFD8FF173FACFE3DFE03" box="[151,414,404,428]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="flavozonatum">
|
||
<emphasis id="B911EAA6FFCEFFD8FF173FACFE3DFE03" box="[151,414,404,428]" italics="true" pageId="6" pageNumber="65">Dinodon flavozonatum</emphasis>
|
||
</taxonomicName>
|
||
(SYS r000274),
|
||
<taxonomicName id="4C654D37FFCEFFD8FDF33FACFCEBFE03" box="[627,840,404,428]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="spinalis">
|
||
<emphasis id="B911EAA6FFCEFFD8FDF33FACFCEBFE03" box="[627,840,404,428]" italics="true" pageId="6" pageNumber="65">Achalinus spinalis</emphasis>
|
||
</taxonomicName>
|
||
(SYS r000273) and
|
||
<taxonomicName id="4C654D37FFCEFFD8FBCD3FADFAF2FE03" box="[1101,1361,404,428]" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="multicinctus">
|
||
<emphasis id="B911EAA6FFCEFFD8FBCD3FADFAF2FE03" box="[1101,1361,404,428]" italics="true" pageId="6" pageNumber="65">Bungarus multicinctus</emphasis>
|
||
</taxonomicName>
|
||
(SYS r000305).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F653FFFCEFFD8FF453FE4FB21F8E0" pageId="6" pageNumber="65" type="etymology">
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453FE4FDB7FDB8" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453FE4FEE8FE5B" bold="true" box="[197,331,476,500]" pageId="6" pageNumber="65">Etymology.</emphasis>
|
||
The specific epithet refers to the
|
||
<typeStatus id="54DE8816FFCEFFD8FD573FE5FCA4FE5A" box="[727,775,477,501]" pageId="6" pageNumber="65">type</typeStatus>
|
||
locality, Maolan Town, Guizhou,
|
||
<collectingCountry id="F3727624FFCEFFD8FB153FE4FB7FFE5B" box="[1173,1244,476,500]" name="China" pageId="6" pageNumber="65">China</collectingCountry>
|
||
. We propose the English name “Mao-lan pitviper”.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453C1BFB53FDF0" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453C1BFEC8FD93" bold="true" box="[197,363,547,572]" pageId="6" pageNumber="65">Comparisons.</emphasis>
|
||
<taxonomicName id="4C654D37FFCEFFD8FEF23C1CFD04FD93" box="[370,679,548,572]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FEF23C1CFD04FD93" box="[370,679,548,572]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FD2F3C1CFCA9FD93" bold="true" box="[687,778,548,572]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FD2F3C1CFCA9FD93" box="[687,778,548,572]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be readily distinguished from
|
||
<taxonomicName id="4C654D37FFCEFFD8FB143C1DFAB1FD93" box="[1172,1298,549,572]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="cornutus">
|
||
<emphasis id="B911EAA6FFCEFFD8FB143C1DFAB1FD93" box="[1172,1298,549,572]" italics="true" pageId="6" pageNumber="65">P. cornutus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D37FFCEFFD8FACB3C1DFF57FDCF" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="sieversorum">
|
||
<emphasis id="B911EAA6FFCEFFD8FACB3C1DFF57FDCF" italics="true" pageId="6" pageNumber="65">P. sieversorum</emphasis>
|
||
</taxonomicName>
|
||
by the absence of horn-shaped projections on the supraocular and a different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453C54FE8EFD5D" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453C54FE5AFD2B" box="[197,505,620,644]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453C54FE5AFD2B" box="[197,505,620,644]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FE7F3C54FDFBFD2B" bold="true" box="[511,600,620,644]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FE7F3C54FDFBFD2B" box="[511,600,620,644]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD6B3C55FCFFFD2B" box="[747,860,620,644]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B911EAA6FFCEFFD8FD6B3C55FCFFFD2B" box="[747,860,620,644]" italics="true" pageId="6" pageNumber="65">P. elegans</emphasis>
|
||
</taxonomicName>
|
||
by having fewer mid-body dorsal scales (19–21 vs. 23–25), 4th supralabials separated from suboculars by one row of scales (two in
|
||
<taxonomicName id="4C654D37FFCEFFD8FBA03CACFB30FD04" box="[1056,1171,659,683]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B911EAA6FFCEFFD8FBA03CACFB30FD04" box="[1056,1171,659,683]" italics="true" pageId="6" pageNumber="65">P. elegans</emphasis>
|
||
</taxonomicName>
|
||
), two loreals (one in
|
||
<taxonomicName id="4C654D37FFCEFFD8FA073CACFF4FFD60" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B911EAA6FFCEFFD8FA073CACFF4FFD60" italics="true" pageId="6" pageNumber="65">P. elegans</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales keeled throughout (outermost row of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FBEC3C80FB43FD60" box="[1132,1248,695,719]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B911EAA6FFCEFFD8FBEC3C80FB43FD60" box="[1132,1248,695,719]" italics="true" pageId="6" pageNumber="65">P. elegans</emphasis>
|
||
</taxonomicName>
|
||
) and a different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453CC7FE8EFC2D" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453CC7FE5BFCB8" box="[197,504,767,791]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453CC7FE5BFCB8" box="[197,504,767,791]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FE7F3CC7FDFBFCB8" bold="true" box="[511,600,767,791]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FE7F3CC7FDFBFCB8" box="[511,600,767,791]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD6A3D38FC27FCB8" box="[746,900,767,791]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="flavoviridis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD6A3D38FC27FCB8" box="[746,900,767,791]" italics="true" pageId="6" pageNumber="65">P. flavoviridis</emphasis>
|
||
</taxonomicName>
|
||
by having fewer mid-body dorsal scales (19–21 vs. 33–40), two loreals (one in
|
||
<taxonomicName id="4C654D37FFCEFFD8FE7C3D1CFD3BFC94" box="[508,664,803,827]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="flavoviridis">
|
||
<emphasis id="B911EAA6FFCEFFD8FE7C3D1CFD3BFC94" box="[508,664,803,827]" italics="true" pageId="6" pageNumber="65">P. flavoviridis</emphasis>
|
||
</taxonomicName>
|
||
), lower number of ventral scales (186–193 vs.
|
||
<quantity id="4C9D9B51FFCEFFD8FB353D1AFA96FC95" box="[1205,1333,802,826]" metricMagnitude="0" metricUnit="m" metricValue="5.8293" metricValueMax="6.0198" metricValueMin="5.6388" pageId="6" pageNumber="65" unit="in" value="229.5" valueMax="237.0" valueMin="222.0">222–237 in</quantity>
|
||
<emphasis id="B911EAA6FFCEFFD8FABD3D1CFF7CFCF0" italics="true" pageId="6" pageNumber="65">P. f l a v o - viridis</emphasis>
|
||
), dorsal scales keeled throughout (outermost row of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FBC93D70FB47FCF0" box="[1097,1252,839,863]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="flavoviridis">
|
||
<emphasis id="B911EAA6FFCEFFD8FBC93D70FB47FCF0" box="[1097,1252,839,863]" italics="true" pageId="6" pageNumber="65">P. flavoviridis</emphasis>
|
||
</taxonomicName>
|
||
) and a different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453DB7FCE4FC40" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453DB7FE5AFC08" box="[197,505,911,935]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453DB7FE5AFC08" box="[197,505,911,935]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FD813DB7FDF8FC08" bold="true" box="[513,603,911,935]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FD813DB7FDF8FC08" box="[513,603,911,935]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the
|
||
<taxonomicName id="4C654D37FFCEFFD8FC993DA8FC2EFC08" box="[793,909,911,935]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="jerdonii">
|
||
<emphasis id="B911EAA6FFCEFFD8FC993DA8FC2EFC08" box="[793,909,911,935]" italics="true" pageId="6" pageNumber="65">P. jerdonii</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<emphasis id="B911EAA6FFCEFFD8FB853DA8FB2CFC08" box="[1029,1167,911,935]" italics="true" pageId="6" pageNumber="65">
|
||
P. j.
|
||
<taxonomicName id="4C654D37FFCEFFD8FBB73DB7FB2CFC08" box="[1079,1167,911,935]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="jerdonii">jerdonii</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C654D37FFCEFFD8FB1D3DA8FA89FC08" box="[1181,1322,911,935]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="subSpecies" species="jerdoni" subSpecies="bourreti">
|
||
<emphasis id="B911EAA6FFCEFFD8FB1D3DA8FA89FC08" box="[1181,1322,911,935]" italics="true" pageId="6" pageNumber="65">P. j. bourreti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C654D37FFCEFFD8FAB83DA8FF5FFC64" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="subSpecies" species="jerdoni" subSpecies="xanthomelas">
|
||
<emphasis id="B911EAA6FFCEFFD8FAB83DA8FF5FFC64" italics="true" pageId="6" pageNumber="65">P. j. xanthomelas</emphasis>
|
||
</taxonomicName>
|
||
) by its smaller size, slender body, dorsal scales keeled throughout (outer one or two rows of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FE8D3DE0FE23FC40" box="[269,384,983,1007]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="jerdonii">
|
||
<emphasis id="B911EAA6FFCEFFD8FE8D3DE0FE23FC40" box="[269,384,983,1007]" italics="true" pageId="6" pageNumber="65">P. jerdonii</emphasis>
|
||
</taxonomicName>
|
||
) and by a clearly different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453DC3FB09FBF5" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453DC3FE5AFBBC" box="[197,505,1019,1043]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453DC3FE5AFBBC" box="[197,505,1019,1043]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FD803DC3FDFAFBBC" bold="true" box="[512,601,1019,1043]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FD803DC3FDFAFBBC" box="[512,601,1019,1043]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD6D3DC4FCD6FBBC" box="[749,885,1019,1043]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="kaulbacki">
|
||
<emphasis id="B911EAA6FFCEFFD8FD6D3DC4FCD6FBBC" box="[749,885,1019,1043]" italics="true" pageId="6" pageNumber="65">P. kaulbacki</emphasis>
|
||
</taxonomicName>
|
||
by its smaller size, fewer mid-body dorsal scales (19–21 versus 25), dorsal scales keeled throughout (outermost row of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FB3B3A18FAE7FB98" box="[1211,1348,1055,1079]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="kaulbacki">
|
||
<emphasis id="B911EAA6FFCEFFD8FB3B3A18FAE7FB98" box="[1211,1348,1055,1079]" italics="true" pageId="6" pageNumber="65">P. kaulbacki</emphasis>
|
||
</taxonomicName>
|
||
), fewer ventral scales (186–193 vs.
|
||
<quantity id="4C9D9B51FFCEFFD8FE4F3A7AFDEDFBF5" box="[463,590,1090,1114]" metricMagnitude="0" metricUnit="m" metricValue="5.2451" metricValueMax="5.3848" metricValueMin="5.1054" pageId="6" pageNumber="65" unit="in" value="206.5" valueMax="212.0" valueMin="201.0">201–212 in</quantity>
|
||
<taxonomicName id="4C654D37FFCEFFD8FDD43A7CFD7EFBF4" box="[596,733,1091,1115]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="kaulbacki">
|
||
<emphasis id="B911EAA6FFCEFFD8FDD43A7CFD7EFBF4" box="[596,733,1091,1115]" italics="true" pageId="6" pageNumber="65">P. kaulbacki</emphasis>
|
||
</taxonomicName>
|
||
), and by a clearly different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453A5FFBA9FAA0" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453A5FFE5BFBD0" box="[197,504,1127,1151]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453A5FFE5BFBD0" box="[197,504,1127,1151]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FE7F3A5FFDFBFBD0" bold="true" box="[511,600,1127,1151]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FE7F3A5FFDFBFBD0" box="[511,600,1127,1151]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD6B3A50FC12FBD0" box="[747,945,1127,1151]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD6B3A50FC12FBD0" box="[747,945,1127,1151]" italics="true" pageId="6" pageNumber="65">P. mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
by its smaller size (largest TL
|
||
<quantity id="4C9D9B51FFCEFFD8FA883A5FFAC6FBD0" box="[1288,1381,1127,1151]" metricMagnitude="-1" metricUnit="m" metricValue="8.05" pageId="6" pageNumber="65" unit="mm" value="805.0">805 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C654D37FFCEFFD8FA073A50FE80FB0C" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FA073A50FE80FB0C" italics="true" pageId="6" pageNumber="65">P. maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FEAA3AB3FE22FB0D" box="[298,385,1163,1186]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs. largest TL
|
||
<quantity id="4C9D9B51FFCEFFD8FDAA3AB2FD36FB0D" box="[554,661,1162,1186]" metricMagnitude="0" metricUnit="m" metricValue="2.375" pageId="6" pageNumber="65" unit="mm" value="2375.0">2375 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C654D37FFCEFFD8FD3A3AB4FC23FB0C" box="[698,896,1163,1187]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD3A3AB4FC23FB0C" box="[698,896,1163,1187]" italics="true" pageId="6" pageNumber="65">P. mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
), internasals separated by scales (touching each other in
|
||
<taxonomicName id="4C654D37FFCEFFD8FF753A97FE03FB68" box="[245,416,1199,1223]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FF753A97FE03FB68" box="[245,416,1199,1223]" italics="true" pageId="6" pageNumber="65">mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
), two loreals (one in
|
||
<taxonomicName id="4C654D37FFCEFFD8FD123A97FC9EFB68" box="[658,829,1199,1223]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD123A97FC9EFB68" box="[658,829,1199,1223]" italics="true" pageId="6" pageNumber="65">mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
), fewer infralabials, (11–12 vs. 13–16), more subcaudals (74–85 vs. 60–67), fewer mid-body dorsal scales (19–21 vs. 25), dorsal scales keeled throughout (outermost row of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FE733AC0FD19FAA0" box="[499,698,1271,1295]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FE733AC0FD19FAA0" box="[499,698,1271,1295]" italics="true" pageId="6" pageNumber="65">P. mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
) and a different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453B23FC94FA30" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453B23FE5BFA9C" box="[197,504,1307,1331]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453B23FE5BFA9C" box="[197,504,1307,1331]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FE7F3B23FDFBFA9C" bold="true" box="[511,600,1307,1331]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FE7F3B23FDFBFA9C" box="[511,600,1307,1331]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD6A3B24FC61FA9C" box="[746,962,1308,1331]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mucrosquamatus">
|
||
<emphasis id="B911EAA6FFCEFFD8FD6A3B24FC61FA9C" box="[746,962,1308,1331]" italics="true" pageId="6" pageNumber="65">P. mucrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
by its smaller size (largest TL
|
||
<quantity id="4C9D9B51FFCEFFD8FA9A3B22FAD5FA9D" box="[1306,1398,1306,1330]" metricMagnitude="-1" metricUnit="m" metricValue="8.05" pageId="6" pageNumber="65" unit="mm" value="805.0">805 mm</quantity>
|
||
vs. largest TL
|
||
<quantity id="4C9D9B51FFCEFFD8FE923B07FEDEFAF8" box="[274,381,1343,1367]" metricMagnitude="0" metricUnit="m" metricValue="1.31" pageId="6" pageNumber="65" unit="mm" value="1310.0">1310 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C654D37FFCEFFD8FE203B78FDD9FAF8" box="[416,634,1344,1367]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mucrosquamatus">
|
||
<emphasis id="B911EAA6FFCEFFD8FE203B78FDD9FAF8" box="[416,634,1344,1367]" italics="true" pageId="6" pageNumber="65">P. mucrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
), fewer mid-body dorsal scales (19–21 vs. 23–27), fewer ventral scales (186–193 vs.
|
||
<quantity id="4C9D9B51FFCEFFD8FEB73B5AFE1AFAD5" box="[311,441,1378,1402]" metricMagnitude="0" metricUnit="m" metricValue="5.4229" metricValueMax="5.9182" metricValueMin="4.9276" pageId="6" pageNumber="65" unit="in" value="213.5" valueMax="233.0" valueMin="194.0">194–233 in</quantity>
|
||
<taxonomicName id="4C654D37FFCEFFD8FE433B5CFD03FAD4" box="[451,672,1380,1403]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mucrosquamatus">
|
||
<emphasis id="B911EAA6FFCEFFD8FE433B5CFD03FAD4" box="[451,672,1380,1403]" italics="true" pageId="6" pageNumber="65">P. mucrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales keeled throughout (outermost row of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FE8D3BB0FE45FA30" box="[269,486,1416,1439]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="mucrosquamatus">
|
||
<emphasis id="B911EAA6FFCEFFD8FE8D3BB0FE45FA30" box="[269,486,1416,1439]" italics="true" pageId="6" pageNumber="65">P. mucrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
) and a different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF453B93FACEF9A5" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF453B93FE5AFA6C" box="[197,505,1451,1475]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF453B93FE5AFA6C" box="[197,505,1451,1475]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FD813B93FDF8FA6C" bold="true" box="[513,603,1451,1475]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FD813B93FDF8FA6C" box="[513,603,1451,1475]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD6F3B94FCDEFA6C" box="[751,893,1451,1475]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tokarensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD6F3B94FCDEFA6C" box="[751,893,1451,1475]" italics="true" pageId="6" pageNumber="65">P. tokarensis</emphasis>
|
||
</taxonomicName>
|
||
by having fewer mid-body dorsal scales (19–21 vs. 31–32), two loreals (one in
|
||
<taxonomicName id="4C654D37FFCEFFD8FE7B3BE8FD29FA48" box="[507,650,1487,1511]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tokarensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FE7B3BE8FD29FA48" box="[507,650,1487,1511]" italics="true" pageId="6" pageNumber="65">P. tokarensis</emphasis>
|
||
</taxonomicName>
|
||
), fewer ventral scales (186–193 vs.
|
||
<quantity id="4C9D9B51FFCEFFD8FBA83BF7FB0BFA48" box="[1064,1192,1487,1511]" metricMagnitude="0" metricUnit="m" metricValue="5.2197" metricValueMax="5.2832" metricValueMin="5.1562" pageId="6" pageNumber="65" unit="in" value="205.5" valueMax="208.0" valueMin="203.0">203–208 in</quantity>
|
||
<taxonomicName id="4C654D37FFCEFFD8FB303BE8FA9CFA48" box="[1200,1343,1487,1511]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tokarensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FB303BE8FA9CFA48" box="[1200,1343,1487,1511]" italics="true" pageId="6" pageNumber="65">P. tokarensis</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales keeled throughout (outermost row of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FC0F3BCCFBBEF9A4" box="[911,1053,1523,1547]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tokarensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FC0F3BCCFBBEF9A4" box="[911,1053,1523,1547]" italics="true" pageId="6" pageNumber="65">P. tokarensis</emphasis>
|
||
</taxonomicName>
|
||
) and a different color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF45382FFF66F935" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF45382FFE58F980" box="[197,507,1559,1583]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF45382FFE58F980" box="[197,507,1559,1583]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FD83382FFDFDF980" box="[515,606,1559,1583]" pageId="6" pageNumber="65" rank="species">
|
||
<emphasis id="B911EAA6FFCEFFD8FD83382FFDF4F980" bold="true" box="[515,599,1559,1583]" pageId="6" pageNumber="65">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C654D37FFCEFFD8FD763820FC6EF980" box="[758,973,1559,1583]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="xiangchengensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD763820FC6EF980" box="[758,973,1559,1583]" italics="true" pageId="6" pageNumber="65">P. xiangchengensis</emphasis>
|
||
</taxonomicName>
|
||
by having fewer mid-body dorsal scales (19–21 vs. 25), more subcaudals (74–85 vs. 50–66), dorsal scales keeled throughout (outer two rows of dorsal scales smooth in
|
||
<taxonomicName id="4C654D37FFCEFFD8FED73858FD8FF9D8" box="[343,556,1631,1655]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="xiangchengensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FED73858FD8FF9D8" box="[343,556,1631,1655]" italics="true" pageId="6" pageNumber="65">P. xiangchengensis</emphasis>
|
||
</taxonomicName>
|
||
), absence of a dark brown blotch under loreal pit (present in
|
||
<taxonomicName id="4C654D37FFCEFFD8FB5B3858FF17F934" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="xiangchengensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FB5B3858FF17F934" italics="true" pageId="6" pageNumber="65">P. xiangchengensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36B4FFCEFFD8FF45389FFB21F8E0" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="65">
|
||
<taxonomicName id="4C654D37FFCEFFD8FF45389FFE5FF910" box="[197,508,1703,1727]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="maolanensis" status="sp. nov.">
|
||
<emphasis id="B911EAA6FFCEFFD8FF45389FFE5FF910" box="[197,508,1703,1727]" italics="true" pageId="6" pageNumber="65">Protobothrops maolanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EAA6FFCEFFD8FD87389FFDC7F910" bold="true" box="[519,612,1703,1727]" pageId="6" pageNumber="65">
|
||
<taxonomicNameLabel id="A22257DDFFCEFFD8FD87389FFDC7F910" box="[519,612,1703,1727]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is superficially most similar in appearance to
|
||
<taxonomicName id="4C654D37FFCEFFD8FB033890FAFFF910" box="[1155,1372,1703,1727]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="trungkhanhensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FB033890FAFFF910" box="[1155,1372,1703,1727]" italics="true" pageId="6" pageNumber="65">P. trungkhanhensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3727624FFCEFFD8FF1738F2FF5AF94D" box="[151,249,1738,1762]" name="Vietnam" pageId="6" pageNumber="65">Vietnam</collectingCountry>
|
||
, from which it differs by having more dorsal scale rows on the anterior part of the body (23–25 vs. 19), outermost row of dorsal scales on body strongly keeled in males and weakly keeled in females (dorsal scales strongly keeled throughout in both male and female
|
||
<taxonomicName id="4C654D37FFCEFFD8FD67392CFC1EF884" box="[743,957,1811,1835]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="trungkhanhensis">
|
||
<emphasis id="B911EAA6FFCEFFD8FD67392CFC1EF884" box="[743,957,1811,1835]" italics="true" pageId="6" pageNumber="65">P. trungkhanhensis</emphasis>
|
||
</taxonomicName>
|
||
), lower ED/HL ratio (0.15–0.18 vs. 0.23), higher HW/HL ratio (0.68–0.70 vs. 0.65–0.66), and fewer body bands (68–72 vs. 76–84).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |