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49 KiB
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404 lines
49 KiB
XML
<document id="22297C87F36B88E177BF8B06C5A11EFD" ID-CLB-Dataset="5948" ID-DOI="10.11646/zootaxa.4980.3.1" ID-GBIF-Dataset="f36e1c1f-1bfa-4fbc-b2dd-5edf6c7173af" ID-ISSN="1175-5326" ID-Zenodo-Dep="4896889" ID-ZooBank="F1FCA8AC-A984-4547-8A05-F1993BDAEE7C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622722708824" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2021" docId="03CC879003273905C5BA440F7D3BAF62" docLanguage="en" docName="zootaxa.4980.3.1.pdf" docOrigin="Zootaxa 4980 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Uokeaster ahi Mah 2021" docType="treatment" docVersion="7" lastPageNumber="416" masterDocId="FFF5FFE8032A390AC52D45617F37AA6F" masterDocTitle="The East Pacific / South Pacific Boundary: New taxa and occurrences from Rapa Nui (Easter Island), New Caledonia and adjacent regions" masterLastPageNumber="450" masterPageNumber="401" pageNumber="414" updateTime="1698999201094" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0164CFC015EBD561BD40FF67D2809737">The East Pacific / South Pacific Boundary: New taxa and occurrences from Rapa Nui (Easter Island), New Caledonia and adjacent regions</mods:title>
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<mods:namePart id="E8C1B154429B060A2CB33DDFB8E5A8AE">Mah, Christopher L.</mods:namePart>
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<mods:title id="1D0E8E9A59832B42A398965A03C3CC4C">Zootaxa</mods:title>
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<mods:date id="CD6EE21C9E88A67BAC3BFBB19B1114B8">2021</mods:date>
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<mods:number id="77F5888A0E84DAA009CF56393C2B25C2">2021-06-02</mods:number>
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<mods:number id="9ED7720060A27F8E0F894D19BBBF4A98">4980</mods:number>
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<treatment id="03CC879003273905C5BA440F7D3BAF62" ID-DOI="http://doi.org/10.5281/zenodo.5041175" ID-GBIF-Taxon="183202805" ID-Zenodo-Dep="5041175" LSID="urn:lsid:plazi:treatment:03CC879003273905C5BA440F7D3BAF62" httpUri="http://treatment.plazi.org/id/03CC879003273905C5BA440F7D3BAF62" lastPageId="15" lastPageNumber="416" pageId="13" pageNumber="414">
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<subSubSection id="C37F650D03273907C5BA440F7ED4ABE6" box="[151,483,366,393]" pageId="13" pageNumber="414" type="nomenclature">
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<paragraph id="8BDA368603273907C5BA440F7ED4ABE6" blockId="13.[151,483,366,429]" box="[151,483,366,393]" pageId="13" pageNumber="414">
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<heading id="D09281EA03273907C5BA440F7ED4ABE6" bold="true" box="[151,483,366,393]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
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<emphasis id="B911EA9403273907C5BA440F7ED4ABE6" bold="true" box="[151,483,366,393]" italics="true" pageId="13" pageNumber="414">
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<taxonomicName id="4C654D0503273907C5BA440F7E77ABE6" ID-CoL="8KPZF" authority="Mah, 2021" authorityName="Mah" authorityYear="2021" box="[151,320,366,393]" class="Asteroidea" family="Asterodiscididae" genus="Uokeaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="441" phylum="Echinodermata" rank="species" species="ahi" status="gen. nov.">Uokeaster ahi</taxonomicName>
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n. gen. n. sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F650D03273907C5BA44F27DEDAB80" pageId="13" pageNumber="414" type="description">
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<paragraph id="8BDA368603273907C5BA44F27E1EABC2" blockId="13.[151,483,366,429]" box="[151,297,402,429]" pageId="13" pageNumber="414">
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<figureCitation id="135E2A0303273907C5BA44F27E1EABC2" box="[151,297,402,429]" captionStart="FIGURE 5" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[157,1430,187,1837]" captionTargetId="figure-30@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 5. Uokeaster ahi n. gen. n. sp. Holotype CASIZ 222370 (Rapa Nui): A. In situ observation (not the holotype). Courtesy A. Mecho. B. Abactinal-lateral view. Scale bar equals 1.0 cm. C. Abactinal surface. Scale bar equals 2.0 cm. D. Actinolateral view showing marginal plate series. IM = intermarginal plates. Scale bar equals 0.75 cm. E. Actinal surface. Scale bar equals 2.0 cm. F. Close-up of actinal surface showing furrow spines. Scale bar equals 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.4896903" httpUri="https://zenodo.org/record/4896903/files/figure.png" pageId="13" pageNumber="414">Figure 5A–E</figureCitation>
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</paragraph>
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||
<paragraph id="8BDA368603273907C5BA44B67DEDAB80" blockId="13.[151,730,471,495]" box="[151,730,471,495]" pageId="13" pageNumber="414">
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<bibRefCitation id="EFF44B7703273907C5BA44B67E53AB80" author="Mecho, A. & Easton, E. E. & Sellanes, J. & Gorny, M. & Mah, C." box="[151,356,471,495]" pageId="13" pageNumber="414" pagination="91" refId="ref30588" refString="Mecho, A., Easton, E. E., Sellanes, J., Gorny, M. & Mah, C. (2019) Unexplored diversity of the mesophotic echinoderm fauna of the Easter Island ecoregion. Marine Biology, 166, 91 https: // doi. org / 10.1007 / s 00227 - 019 - 3537 - x" type="journal article" year="2019">
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Mecho
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<emphasis id="B911EA9403273907C5C944B97E27AB80" box="[228,272,471,495]" italics="true" pageId="13" pageNumber="414">et al</emphasis>
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||
. (2019)
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||
</bibRefCitation>
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||
: 91 (as Family
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<taxonomicName id="4C654D0503273907C4D344B67D9FAB80" authorityName="Rowe" authorityYear="1977" box="[510,680,471,495]" class="Asteroidea" family="Asterodiscididae" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="family">Asterodiscididae</taxonomicName>
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sp.).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F650D03273907C5BA477C7C30A810" pageId="13" pageNumber="414" type="etymology">
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<paragraph id="8BDA368603273907C5BA477C7C30A810" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
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<emphasis id="B911EA9403273907C5BA477C7E29A858" bold="true" box="[151,286,541,567]" pageId="13" pageNumber="414">Etymology.</emphasis>
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The genus is named for the Rapa Nui marine deity
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<emphasis id="B911EA9403273907C67E477C7CB9A858" box="[851,910,541,567]" italics="true" pageId="13" pageNumber="414">Uoke</emphasis>
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, who submerged Rapa Nui into the ocean using a large lever, and “
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<emphasis id="B911EA9403273907C44547237E97A834" box="[360,416,578,603]" italics="true" pageId="13" pageNumber="414">aster</emphasis>
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||
” for star. The species epithet
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<emphasis id="B911EA9403273907C7C747207C39A834" box="[746,782,577,603]" italics="true" pageId="13" pageNumber="414">ahi</emphasis>
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is the Rapa Nui word for “fire” which alludes to the bright orange color of this species. Noun is held in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F650D03273907C5EA47E87B22A884" pageId="13" pageNumber="414" type="diagnosis">
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<paragraph id="8BDA368603273907C5EA47E87B22A884" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
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<emphasis id="B911EA9403273907C5EA47E87E75A8CC" bold="true" box="[199,322,649,675]" pageId="13" pageNumber="414">Diagnosis.</emphasis>
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This species is distinguished from other asterodiscidid forms based on the presence of the distinct continuous dermal layer present from the abactinal, marginal and actinal surface, and the marginal plates forming blunt, conical spines with similar
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<typeStatus id="54DE882403273907C73C47B37D7BA883" box="[529,588,722,748]" pageId="13" pageNumber="414">types</typeStatus>
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of spines covering the abactinal surface.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F650D03273907C5EA47947D03AC00" pageId="13" pageNumber="414" type="discussion">
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||
<paragraph id="8BDA368603273907C5EA47947D93A938" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
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||
<emphasis id="B911EA9403273907C5EA47947E78A960" bold="true" box="[199,335,757,783]" pageId="13" pageNumber="414">Comments.</emphasis>
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<taxonomicName id="4C654D0503273907C47847947EE1A960" authorityName="Verrill" authorityYear="1868" box="[341,470,757,783]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C47847947EE1A960" box="[341,470,757,783]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C654D0503273907C72747947D62A960" authorityName="Gray" authorityYear="1840" box="[522,597,757,783]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C72747947D62A960" box="[522,597,757,783]" italics="true" pageId="13" pageNumber="414">Paulia</emphasis>
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</taxonomicName>
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have abactinal surfaces with a close-set, dense, granular covering. and have fewer, larger, spineless marginal plates which show a strongly convex surface.
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<taxonomicName id="4C654D0503273907C12946787BB2A95C" authorityName="Verrill" authorityYear="1868" box="[1028,1157,793,819]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C12946787BB2A95C" box="[1028,1157,793,819]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
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</taxonomicName>
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displays alternating bald marginal plates with large, conical projections.
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</paragraph>
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<paragraph id="8BDA368603273907C5EA46007E6AAEF4" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
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This species shares the enlarged penultimate superomarginal plates and displays morphology intermediate form between the East Pacific
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<taxonomicName id="4C654D0503273907C49746E47D0CA9F0" authorityName="Verrill" authorityYear="1868" box="[442,571,901,927]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C49746E47D0CA9F0" box="[442,571,901,927]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C654D0503273907C75B46E47DF6A9F0" authorityName="Gray" authorityYear="1840" box="[630,705,901,927]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C75B46E47DF6A9F0" box="[630,705,901,927]" italics="true" pageId="13" pageNumber="414">Paulia</emphasis>
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</taxonomicName>
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and the widely occurring Indo-Pacific
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<taxonomicName id="4C654D0503273907C1AB46E47A1FA9F0" authorityName="A. M. Clark" authorityYear="1974" box="[1158,1320,901,927]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C1AB46E47A1FA9F0" box="[1158,1320,901,927]" italics="true" pageId="13" pageNumber="414">Asterodiscides</emphasis>
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</taxonomicName>
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. Morphological assessments by
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<bibRefCitation id="EFF44B7703273907C48946CB7D03A9AC" author="Rowe, F. W. E." box="[420,564,937,964]" pageId="13" pageNumber="414" pagination="187 - 233" refId="ref31297" refString="Rowe, F. W. E. (1977) A new family of Asteroidea (Echinodermata), with the description of five new species and one new subspecies of Asterodiscides. Records of the Australian Museum, 31 (5), 187 - 233. https: // doi. org / 10.3853 / j. 0067 - 1975.31.1977.209" type="journal article" year="1977">Rowe (1977</bibRefCitation>
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,
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<bibRefCitation id="EFF44B7703273907C76F46C87D4CA9AB" author="Rowe, F. W. E." box="[578,635,937,964]" pageId="13" pageNumber="414" pagination="531 - 577" refId="ref31360" refString="Rowe, F. W. E. (1985) Six new species of Asterodiscides A. M. Clark (Echinodermata, Asteroidea), with a discussion of the origin and distribution of the Asterodiscididae and other ' amphi-Pacific' Echinoderms. Bulletin du Museum national d'Histoire naturelle A, 7 (3), 531 - 577." type="journal article" year="1985">1985</bibRefCitation>
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||
) and phylogenetic analysis by
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<bibRefCitation id="EFF44B7703273907C6CE46C87BDAA9AC" author="Blake, D. B. & Portell, R. W." box="[995,1261,937,963]" pageId="13" pageNumber="414" pagination="25 - 42" refId="ref26678" refString="Blake, D. B. & Portell, R. W. (2011) Kionaster petersonae, n. gen. and sp. (Asteroidea), the first fossil occurrence of the Asterodiscididae, from the Miocene of Florida. Swiss Journal of Paleontology, 130, 25 - 42. https: // doi. org / 10.1007 / s 13358 - 010 - 0005 - 0" type="journal article" year="2011">Blake & Portell (2011)</bibRefCitation>
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||
all support the hypothesis that
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<taxonomicName id="4C654D0503273907C46646AC7EFBA988" authorityName="Verrill" authorityYear="1868" box="[331,460,973,999]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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<emphasis id="B911EA9403273907C46646AC7EFBA988" box="[331,460,973,999]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
|
||
</taxonomicName>
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||
,
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<taxonomicName id="4C654D0503273907C4F646AC7D11A988" authorityName="Gray" authorityYear="1840" box="[475,550,973,999]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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||
<emphasis id="B911EA9403273907C4F646AC7D11A988" box="[475,550,973,999]" italics="true" pageId="13" pageNumber="414">Paulia</emphasis>
|
||
</taxonomicName>
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||
and related asterodiscidid genera are more stemward relative to the more derived
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<emphasis id="B911EA9403273907C5F446907ECAAE64" box="[217,509,1009,1035]" italics="true" pageId="13" pageNumber="414">
|
||
<taxonomicName id="4C654D0503273907C5F446907E48AE64" authorityName="A. M. Clark" authorityYear="1974" box="[217,383,1009,1035]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">Asterodiscides</taxonomicName>
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||
.
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<taxonomicName id="4C654D0503273907C4A746907ECAAE64" authority="Mah, 2021" authorityName="Mah" authorityYear="2021" box="[394,509,1009,1035]" class="Asteroidea" family="Asterodiscididae" genus="Uokeaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="441" phylum="Echinodermata" rank="genus" status="gen. nov.">Uokeaster</taxonomicName>
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||
</emphasis>
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has not previously been included in a cladistic analysis but its marginal plate and body morphology are similar to those of
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<taxonomicName id="4C654D0503273907C74341747DD8AE40" authorityName="Verrill" authorityYear="1868" box="[622,751,1045,1071]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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||
<emphasis id="B911EA9403273907C74341747DD8AE40" box="[622,751,1045,1071]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicName id="4C654D0503273907C7D241747C7DAE40" authorityName="Gray" authorityYear="1840" box="[767,842,1045,1071]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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||
<emphasis id="B911EA9403273907C7D241747C7DAE40" box="[767,842,1045,1071]" italics="true" pageId="13" pageNumber="414">Paulia</emphasis>
|
||
</taxonomicName>
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and
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||
<taxonomicName id="4C654D0503273907C6AE41777CC6AE40" authorityName="Blake & Portell" authorityYear="2011" box="[899,1009,1046,1071]" class="Asteroidea" family="Asterodiscididae" genus="Kionaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
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||
<emphasis id="B911EA9403273907C6AE41777CC6AE40" box="[899,1009,1046,1071]" italics="true" pageId="13" pageNumber="414">Kionaster</emphasis>
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||
</taxonomicName>
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which suggest a more plesiomorphic morphology rather than that of
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<taxonomicName id="4C654D0503273907C4DB41587DAFAE3C" authorityName="A. M. Clark" authorityYear="1974" box="[502,664,1081,1107]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C4DB41587DAFAE3C" box="[502,664,1081,1107]" italics="true" pageId="13" pageNumber="414">Asterodiscides</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C654D0503273907C63741587CBAAE3C" authority="Mah, 2021" authorityName="Mah" authorityYear="2021" box="[794,909,1081,1107]" class="Asteroidea" family="Asterodiscididae" genus="Uokeaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="441" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B911EA9403273907C63741587CBAAE3C" box="[794,909,1081,1107]" italics="true" pageId="13" pageNumber="414">Uokeaster</emphasis>
|
||
</taxonomicName>
|
||
does have disproportionately enlarged penultimate superomarginal plates which are present in
|
||
<taxonomicName id="4C654D0503273907C797413C7C6BAE18" authorityName="A. M. Clark" authorityYear="1974" box="[698,860,1117,1143]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C797413C7C6BAE18" box="[698,860,1117,1143]" italics="true" pageId="13" pageNumber="414">Asterodiscides</emphasis>
|
||
</taxonomicName>
|
||
, but not present in either
|
||
<taxonomicName id="4C654D0503273907C156413C7BCBAE18" authorityName="Verrill" authorityYear="1868" box="[1147,1276,1117,1143]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C156413C7BCBAE18" box="[1147,1276,1117,1143]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C654D0503273907C027413C7A62AE18" authorityName="Gray" authorityYear="1840" box="[1290,1365,1117,1143]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C027413C7A62AE18" box="[1290,1365,1117,1143]" italics="true" pageId="13" pageNumber="414">Paulia</emphasis>
|
||
</taxonomicName>
|
||
or the extinct
|
||
<emphasis id="B911EA9403273907C5C741E37E6AAEF4" box="[234,349,1154,1179]" italics="true" pageId="13" pageNumber="414">
|
||
<taxonomicName id="4C654D0503273907C5C741E37E6EAEF4" authorityName="Blake & Portell" authorityYear="2011" box="[234,345,1154,1179]" class="Asteroidea" family="Asterodiscididae" genus="Kionaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">Kionaster</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA368603273907C5EA41C47EBBAE8B" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
|
||
<bibRefCitation id="EFF44B7703273907C5EA41C47E87AED0" author="Mecho, A. & Easton, E. E. & Sellanes, J. & Gorny, M. & Mah, C." box="[199,432,1189,1216]" pageId="13" pageNumber="414" pagination="91" refId="ref30588" refString="Mecho, A., Easton, E. E., Sellanes, J., Gorny, M. & Mah, C. (2019) Unexplored diversity of the mesophotic echinoderm fauna of the Easter Island ecoregion. Marine Biology, 166, 91 https: // doi. org / 10.1007 / s 00227 - 019 - 3537 - x" type="journal article" year="2019">
|
||
Mecho
|
||
<emphasis id="B911EA9403273907C43241C77E63AED0" box="[287,340,1189,1215]" italics="true" pageId="13" pageNumber="414">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
has observed this species
|
||
<emphasis id="B911EA9403273907C7C541C77C07AED0" box="[744,816,1190,1215]" italics="true" pageId="13" pageNumber="414">in situ</emphasis>
|
||
from Pukao and Apolo islands in Rapa Nui between
|
||
<quantity id="4C9D9B6303273907C5BA41A87E21AE8B" box="[151,278,1225,1252]" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="13" pageNumber="414" unit="m" value="170.0" valueMax="180.0" valueMin="160.0">160–180 m</quantity>
|
||
(
|
||
<figureCitation id="135E2A0303273907C40841A87EB7AE8B" box="[293,384,1225,1252]" captionStart="FIGURE 5" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[157,1430,187,1837]" captionTargetId="figure-30@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 5. Uokeaster ahi n. gen. n. sp. Holotype CASIZ 222370 (Rapa Nui): A. In situ observation (not the holotype). Courtesy A. Mecho. B. Abactinal-lateral view. Scale bar equals 1.0 cm. C. Abactinal surface. Scale bar equals 2.0 cm. D. Actinolateral view showing marginal plate series. IM = intermarginal plates. Scale bar equals 0.75 cm. E. Actinal surface. Scale bar equals 2.0 cm. F. Close-up of actinal surface showing furrow spines. Scale bar equals 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.4896903" httpUri="https://zenodo.org/record/4896903/files/figure.png" pageId="13" pageNumber="414">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA368603273907C5EA418C7C74AF1C" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
|
||
<emphasis id="B911EA9403273907C5EA418C7EEFAF68" bold="true" box="[199,472,1261,1287]" pageId="13" pageNumber="414">Biogeographic Trends.</emphasis>
|
||
<bibRefCitation id="EFF44B7703273907C4CF418C7DDAAF68" author="Blake, D. B. & Portell, R. W." box="[482,749,1261,1287]" pageId="13" pageNumber="414" pagination="25 - 42" refId="ref26678" refString="Blake, D. B. & Portell, R. W. (2011) Kionaster petersonae, n. gen. and sp. (Asteroidea), the first fossil occurrence of the Asterodiscididae, from the Miocene of Florida. Swiss Journal of Paleontology, 130, 25 - 42. https: // doi. org / 10.1007 / s 13358 - 010 - 0005 - 0" type="journal article" year="2011">Blake & Portell (2011)</bibRefCitation>
|
||
and especially
|
||
<bibRefCitation id="EFF44B7703273907C689418F7B0DAF67" author="Rowe, F. W. E." box="[932,1082,1261,1288]" pageId="13" pageNumber="414" pagination="531 - 577" refId="ref31360" refString="Rowe, F. W. E. (1985) Six new species of Asterodiscides A. M. Clark (Echinodermata, Asteroidea), with a discussion of the origin and distribution of the Asterodiscididae and other ' amphi-Pacific' Echinoderms. Bulletin du Museum national d'Histoire naturelle A, 7 (3), 531 - 577." type="journal article" year="1985">Rowe (1985)</bibRefCitation>
|
||
touch upon the biogeographic significance of
|
||
<taxonomicName id="4C654D0503273907C46740707EFCAF44" authorityName="Verrill" authorityYear="1868" box="[330,459,1297,1323]" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C46740707EFCAF44" box="[330,459,1297,1323]" italics="true" pageId="13" pageNumber="414">Amphiaster</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D0503273907C72540707D64AF44" authorityName="Gray" authorityYear="1840" box="[520,595,1297,1323]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C72540707D64AF44" box="[520,595,1297,1323]" italics="true" pageId="13" pageNumber="414">Paulia</emphasis>
|
||
</taxonomicName>
|
||
occurring in the East Pacific relative to
|
||
<taxonomicName id="4C654D0503273907C10340707BE7AF44" authorityName="A. M. Clark" authorityYear="1974" box="[1070,1232,1297,1323]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C10340707BE7AF44" box="[1070,1232,1297,1323]" italics="true" pageId="13" pageNumber="414">Asterodiscides</emphasis>
|
||
</taxonomicName>
|
||
occurring widely throughout the Indo-Pacific. Although
|
||
<bibRefCitation id="EFF44B7703273907C76140577C78AF20" author="Lane, D. J. & Rowe, F. W. E." box="[588,847,1333,1360]" pageId="13" pageNumber="414" pagination="419 - 429" refId="ref29111" refString="Lane, D. J. & Rowe, F. W. E. (2009) A new species of Asterodiscides (Echinodermata, Asteroidea, Asterodiscididae) from the tropical southwest Pacific, and the biogeography of the genus revisited. Zoosystema, 31 (3), 419 - 429. https: // doi. org / 10.5252 / z 2009 n 3 a 2" type="journal article" year="2009">Lane and Rowe (2009)</bibRefCitation>
|
||
have produced a cluster analysis for
|
||
<taxonomicName id="4C654D0503273907C1DE40547AA2AF20" authorityName="A. M. Clark" authorityYear="1974" box="[1267,1429,1333,1359]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C1DE40547AA2AF20" box="[1267,1429,1333,1359]" italics="true" pageId="13" pageNumber="414">Asterodiscides</emphasis>
|
||
</taxonomicName>
|
||
, no phylogeographic analysis for species has been performed.
|
||
</paragraph>
|
||
<paragraph id="8BDA368603273907C5EA401C7D03AC00" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">
|
||
Occurrence data of the asterodiscidid taxa suggests interesting trends across the tropical Eastern Pacific.
|
||
<taxonomicName id="4C654D0503273907C067401C7E1BAFD4" authorityName="Verrill" authorityYear="1868" class="Asteroidea" family="Asterodiscididae" genus="Amphiaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="species" species="insignis">
|
||
<emphasis id="B911EA9403273907C067401C7E1BAFD4" italics="true" pageId="13" pageNumber="414">Amphiaster insignis</emphasis>
|
||
</taxonomicName>
|
||
occurs primarily along the coast of
|
||
<collectingCountry id="F372761603273907C79640C07C26AFD4" box="[699,785,1441,1467]" name="Mexico" pageId="13" pageNumber="414">Mexico</collectingCountry>
|
||
with observations south to Malpelo Island, off the coast of Columbia (
|
||
<bibRefCitation id="EFF44B7703273907C43940A47EA9AFB0" author="Maluf, Y." box="[276,414,1477,1503]" pageId="13" pageNumber="414" pagination="1 - 242" refId="ref30361" refString="Maluf, Y. (1988) Composition and distribution of the central eastern Pacific echinoderms. Natural History Museum of Los Angeles County, Technical Reports, 2, 1 - 242." type="journal article" year="1988">Maluf 1988</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C654D0503273907C49F40A47D6DAFB0" authorityName="Gray" authorityYear="1840" box="[434,602,1477,1503]" class="Asteroidea" family="Asterodiscididae" genus="Paulia" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="species" species="horrida">
|
||
<emphasis id="B911EA9403273907C49F40A47D6DAFB0" box="[434,602,1477,1503]" italics="true" pageId="13" pageNumber="414">Paulia horrida</emphasis>
|
||
</taxonomicName>
|
||
has also been recorded from the coast, but also to the Galapagos and Tres Marinas Islands off
|
||
<collectingCountry id="F372761603273907C45B408B7E9AAC6B" box="[374,429,1514,1540]" name="Peru" pageId="13" pageNumber="414">Peru</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C654D0503273907C49A40887DB3AC6C" baseAuthorityName="Ludwig" baseAuthorityYear="1905" box="[439,644,1513,1539]" class="Bivalvia" family="Kelliellidae" genus="Pauliella" kingdom="Animalia" order="Veneroida" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="aenigma">
|
||
<emphasis id="B911EA9403273907C49A40887DB3AC6C" box="[439,644,1513,1539]" italics="true" pageId="13" pageNumber="414">Pauliella aenigma</emphasis>
|
||
</taxonomicName>
|
||
is recorded from Cocos Island and Rocas Alijos and finally,
|
||
<taxonomicName id="4C654D0503273907C00440887AABAC6C" authority="Mah, 2021" authorityName="Mah" authorityYear="2021" box="[1321,1436,1513,1539]" class="Asteroidea" family="Asterodiscididae" genus="Uokeaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B911EA9403273907C00440887AABAC6C" box="[1321,1436,1513,1539]" italics="true" pageId="13" pageNumber="414">Uokeaster</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EA9403273907C5BA436C7FD2AC48" bold="true" box="[151,229,1549,1575]" pageId="13" pageNumber="414">
|
||
<taxonomicNameLabel id="A22257EF03273907C5BA436C7FD2AC48" box="[151,229,1549,1575]" pageId="13" pageNumber="414" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is recorded, for now, only from Rapa Nui, the westernmost of the asterodiscidid genera relative to
|
||
<emphasis id="B911EA9403273907C007436C7FEFAC24" italics="true" pageId="13" pageNumber="414">
|
||
<taxonomicName id="4C654D0503273907C007436C7FE3AC24" authorityName="A. M. Clark" authorityYear="1974" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">Asterodiscides</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Shared characters between
|
||
<taxonomicName id="4C654D0503273907C72143507D48AC24" authority="Mah, 2021" authorityName="Mah" authorityYear="2021" box="[524,639,1585,1611]" class="Asteroidea" family="Asterodiscididae" genus="Uokeaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="441" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B911EA9403273907C72143507D48AC24" box="[524,639,1585,1611]" italics="true" pageId="13" pageNumber="414">Uokeaster</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D0503273907C79943507C61AC24" authorityName="A. M. Clark" authorityYear="1974" box="[692,854,1585,1611]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="414" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9403273907C79943507C61AC24" box="[692,854,1585,1611]" italics="true" pageId="13" pageNumber="414">Asterodiscides</emphasis>
|
||
</taxonomicName>
|
||
and the biogeographic proximity suggest the former could be the sister taxon to the latter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F650D03273905C5EA43187C03AE5A" lastPageId="15" lastPageNumber="416" pageId="13" pageNumber="414" type="description">
|
||
<paragraph id="8BDA368603273907C5EA43187AA5ACFB" blockId="13.[151,1437,541,2008]" box="[199,1426,1657,1684]" pageId="13" pageNumber="414">
|
||
<emphasis id="B911EA9403273907C5EA43187E6EACFC" bold="true" box="[199,345,1657,1683]" pageId="13" pageNumber="414">Description.</emphasis>
|
||
Body stout, stellate (R/r=2.3) in shape, arms triangular, disk arched with curved interradial arcs.
|
||
</paragraph>
|
||
<paragraph id="8BDA368603273907C5EA43FC7BDAADB7" blockId="13.[151,1437,541,2008]" pageId="13" pageNumber="414">Abactinal plates obscured by soft dermal layer displaying a creased texture and perforated by one to ten small papular pores per dermal segment, which are triangular in shape. Primary structures are large, blunt conical spines, with bare, smooth surface. These spines are large, pointed plates rather than articulated structures as seen on other spiniferous goniasterids. Pointed conical plates present on disk and along radial and adradial regions and vary from pointed to semi-oval (egg shaped) in shape. Approximately four to eight large spines from arm tip to primary circlet on disk. Skin around base of spines and on remainder of surface with spine-free regions interradially adjacent to superomarginal contact. Smaller, spines, apparently incipient, present around periphery of other primary, larger spines. Madreporite triangular in shape, flanked by three large primary pointed plates. No pedicellariae. Papulae prominent, translucent emerging from three to five papular pores per papular region, present on disk and arms.</paragraph>
|
||
<caption id="DF1A660E03243904C5BA42287B7CADA3" ID-DOI="http://doi.org/10.5281/zenodo.4896903" ID-Zenodo-Dep="4896903" httpUri="https://zenodo.org/record/4896903/files/figure.png" pageId="14" pageNumber="415" startId="14.[151,250,1865,1889]" targetBox="[157,1430,187,1837]" targetPageId="14">
|
||
<paragraph id="8BDA368603243904C5BA42287B7CADA3" blockId="14.[151,1437,1864,1997]" pageId="14" pageNumber="415">
|
||
<emphasis id="B911EA9403243904C5BA42287E3AAD0E" bold="true" box="[151,269,1865,1889]" pageId="14" pageNumber="415">FIGURE 5</emphasis>
|
||
.
|
||
<emphasis id="B911EA9403243904C43442297D01AD0E" bold="true" box="[281,566,1864,1889]" pageId="14" pageNumber="415">
|
||
<emphasis id="B911EA9403243904C43442297E9DAD0F" bold="true" box="[281,426,1864,1888]" italics="true" pageId="14" pageNumber="415">
|
||
<taxonomicName id="4C654D0503243904C43442297EB4AD0F" authority="Mah, 2021" authorityName="Mah" authorityYear="2021" box="[281,387,1864,1888]" class="Asteroidea" family="Asterodiscididae" genus="Uokeaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="441" phylum="Echinodermata" rank="genus" status="gen. nov.">Uokeaster</taxonomicName>
|
||
ahi
|
||
</emphasis>
|
||
n. gen. n. sp.
|
||
</emphasis>
|
||
Holotype CASIZ 222370 (Rapa Nui):
|
||
<emphasis id="B911EA9403243904C6ED42287CEFAD0E" bold="true" box="[960,984,1865,1889]" pageId="14" pageNumber="415">A.</emphasis>
|
||
<emphasis id="B911EA9403243904C6F3422B7B29AD0E" box="[990,1054,1866,1889]" italics="true" pageId="14" pageNumber="415">In situ</emphasis>
|
||
observation (not the holotype). Courtesy A. Mecho.
|
||
<emphasis id="B911EA9403243904C416420C7E65ADEA" bold="true" box="[315,338,1901,1925]" pageId="14" pageNumber="415">B.</emphasis>
|
||
Abactinal-lateral view. Scale bar equals 1.0 cm.
|
||
<emphasis id="B911EA9403243904C667420C7C54ADEA" bold="true" box="[842,867,1901,1925]" pageId="14" pageNumber="415">C.</emphasis>
|
||
Abactinal surface. Scale bar equals 2.0 cm.
|
||
<emphasis id="B911EA9403243904C002420C7A70ADEA" bold="true" box="[1327,1351,1901,1925]" pageId="14" pageNumber="415">D.</emphasis>
|
||
Actinolateral view showing marginal plate series. IM = intermarginal plates. Scale bar equals 0.75 cm.
|
||
<emphasis id="B911EA9403243904C15B42F07BB9ADC6" bold="true" box="[1142,1166,1937,1961]" pageId="14" pageNumber="415">E.</emphasis>
|
||
Actinal surface. Scale bar equals 2.0 cm.
|
||
<emphasis id="B911EA9403243904C41D42D47E74ADA2" bold="true" box="[304,323,1973,1997]" pageId="14" pageNumber="415">F.</emphasis>
|
||
Close-up of actinal surface showing furrow spines. Scale bar equals 0.5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA368603253905C5EA45F67D10A876" blockId="15.[151,1437,151,1293]" pageId="15" pageNumber="416">Superomarginals 8 to 10, inferomarginals 16 to 20 per interradius (arm-tip to arm-tip) with dermal covering from abactinal surface extending intermarginally to actinal surface. Superomarginal plates pointed, strongly convex but with smooth bare surface, round in cross-section, widely spaced along the lateral surface. Distalmost superomarginals round, strongly convex and enlarged, approximately two to three times the size of adjacent superomarginal plates, and abutted over midline. Larger individuals (R>3.5) with a single spine emerging from central superomarginal surface. Inferomarginals substantially smaller than superomarginals but similar in shape, surfaces bare, smooth, round to quadrate in cross-section. Smaller specimen with short, nipple-like spines present on the tips of primarily interradial inferomarginal spines continuing to arm tips. Inferomarginals become lower and less pointed distally along series adjacent to the arm tip. Both superomarginal and inferomarginals in continuous, mostly uninterrupted series, but one interradius with absent inferomarginal, covered by dermal tissue. Terminal plate, smooth, triangular in shape with pointed tip.</paragraph>
|
||
<paragraph id="8BDA368603253905C5EA47427E2EA956" blockId="15.[151,1437,151,1293]" pageId="15" pageNumber="416">Actinal surface with approximately three complete plate series and one or two incomplete plate series adjacent to inferomarginals. All actinal plate boundaries obscured by continuous dermal layer as present on abactinal and marginal surfaces. Each plate with a large, cylindrical blunt spine sitting on the center of each plate. Proximal actinal plates with one to four, mostly three smaller spines approximately <20% the size of larger, more abundant actinal spines. The larger actinal spines sit in linear series tracking the more fully developed actinal series along the arm until they end adjacent to the enlarged distalmost convex superomarginal plates. A spine is present on nearly every actinal plate extending out to the contact with the inferomarginals, and are widely spaced from one another across the surface.</paragraph>
|
||
<paragraph id="8BDA368603253905C5EA46227E9AA982" blockId="15.[151,1437,151,1293]" pageId="15" pageNumber="416">Furrow spines two, blunt, cylindrical in cross-section, with blunt, round to quadrate tips. Each adambulacral plate with a single, large cylindrical blunt subambulacral spine identical to the others present on the actinal plate surfaces. Subambulacral spines absent on first adambulacral plate. Each oral plate with two furrow spines and a third blunt, spine, triangular in cross-section projecting into mouth. Remainder of oral plate surface bare, smooth, covered by dermal layer.</paragraph>
|
||
<paragraph id="8BDA368603253905C5EA46967D6FAE7E" blockId="15.[151,1437,151,1293]" box="[199,600,1015,1041]" pageId="15" pageNumber="416">Color in life a bright to dull orange.</paragraph>
|
||
<paragraph id="8BDA368603253905C5EA417B7C03AE5A" blockId="15.[151,1437,151,1293]" box="[199,820,1050,1077]" pageId="15" pageNumber="416">
|
||
<emphasis id="B911EA9403253905C5EA417B7E6DAE5B" bold="true" box="[199,346,1050,1076]" pageId="15" pageNumber="416">Occurrence.</emphasis>
|
||
Rapa Nui,
|
||
<quantity id="4C9D9B6303253905C4F4417A7D7CAE5A" box="[473,587,1051,1077]" metricMagnitude="1" metricUnit="m" metricValue="7.15" metricValueMax="11.0" metricValueMin="3.3" pageId="15" pageNumber="416" unit="m" value="71.5" valueMax="110.0" valueMin="33.0">33–110 m</quantity>
|
||
,
|
||
<quantity id="4C9D9B6303253905C77A417A7DE1AE5A" box="[599,726,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="15" pageNumber="416" unit="m" value="170.0" valueMax="180.0" valueMin="160.0">160–180 m</quantity>
|
||
(
|
||
<emphasis id="B911EA9403253905C7CB417A7C1CAE5B" box="[742,811,1051,1076]" italics="true" pageId="15" pageNumber="416">in situ</emphasis>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F650D03253905C5EA415F7D3BAF62" pageId="15" pageNumber="416" type="materials_examined">
|
||
<paragraph id="8BDA368603253905C5EA415F7CD9AE12" blockId="15.[151,1437,151,1293]" pageId="15" pageNumber="416">
|
||
<emphasis id="B911EA9403253905C5EA415F7E86AE37" bold="true" box="[199,433,1086,1112]" pageId="15" pageNumber="416">Material described.</emphasis>
|
||
<materialsCitation id="3B0D3CDB03253905C494415E7CD9AE12" ID-GBIF-Occurrence="3311713386" collectingDate="2017-03-05" collectionCode="R" collectorName="de Pascua & Coll. Bart Shepherd & Tyler Phelps & Luis Rocha" country="Iceland" elevation="33" location="Moto Nui" pageId="15" pageNumber="416" specimenCode="CASIZ 222370" specimenCount="33" typeStatus="holotype">
|
||
<typeStatus id="54DE882403253905C494415E7D11AE36" box="[441,550,1087,1113]" pageId="15" pageNumber="416" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DBC39EFD03253905C71D415E7DD6AE36" box="[560,737,1087,1113]" collectionCode="CASIZ" pageId="15" pageNumber="416">CASIZ 222370</specimenCode>
|
||
.
|
||
<location id="8EBA605D03253905C7C6415E7C6AAE36" LSID="urn:lsid:plazi:treatment:03CC879003273905C5BA440F7D3BAF62:8EBA605D03253905C7C6415E7C6AAE36" box="[747,861,1087,1113]" country="Iceland" name="Moto Nui" pageId="15" pageNumber="416">Moto Nui</location>
|
||
,
|
||
<collectingCountry id="F372761603253905C645415E7CA7AE36" box="[872,912,1087,1113]" name="Iceland" pageId="15" pageNumber="416">Isla</collectingCountry>
|
||
<collectorName id="2690535003253905C6B5415E7B3FAE36" box="[920,1032,1087,1113]" pageId="15" pageNumber="416">de Pascua</collectorName>
|
||
(Rapa Nui),
|
||
<specimenCount id="9D63FD0F03253905C1B1415E7BEAAE36" box="[1180,1245,1087,1113]" pageId="15" pageNumber="416" type="generic">
|
||
<quantity id="4C9D9B6303253905C1B1415E7BEEAE36" box="[1180,1241,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="15" pageNumber="416" unit="m" value="33.0">
|
||
<elevation id="0048D1B503253905C1B1415E7BEEAE36" box="[1180,1241,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="15" pageNumber="416" unit="m" value="33.0">33 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
<collectorName id="2690535003253905C1C9415E7FFBAE12" pageId="15" pageNumber="416">Coll. Bart Shepherd</collectorName>
|
||
,
|
||
<collectorName id="2690535003253905C5FB41027E54AE12" box="[214,355,1123,1149]" pageId="15" pageNumber="416">Tyler Phelps</collectorName>
|
||
and
|
||
<collectorName id="2690535003253905C4B441027D2BAE12" box="[409,540,1123,1149]" pageId="15" pageNumber="416">Luis Rocha</collectorName>
|
||
,
|
||
<date id="FFDB104603253905C70541037DFFAE12" box="[552,712,1122,1149]" pageId="15" pageNumber="416" value="2017-03-05">
|
||
<collectingDate id="EF9FE9AE03253905C70541037DFFAE12" box="[552,712,1122,1149]" pageId="15" pageNumber="416" value="2017-03-05">5 March 2017</collectingDate>
|
||
</date>
|
||
, 1 wet spec.
|
||
<collectionCode id="ED74AE4303253905C67A41037C5CAE12" box="[855,875,1122,1149]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="416">R</collectionCode>
|
||
=3.5, r=1.8.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDA368603253905C5EA41E67D3BAF62" blockId="15.[151,1437,151,1293]" pageId="15" pageNumber="416">
|
||
<typeStatus id="54DE882403253905C5EA41E67E0FAECE" box="[199,312,1159,1185]" pageId="15" pageNumber="416" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<materialsCitation id="3B0D3CDB03253905C46841E67C29AEAA" ID-GBIF-Occurrence="3311713305" collectingDate="2017-03-06" collectionCode="CASIZ" collectorName="de Pascua & Coll. Bart Shepherd & Tyler Phelps & Luis Rocha" country="Iceland" elevation="82" latitude="-27.153242" location="Isla" longLatPrecision="1" longitude="-109.4479" municipality="Rapa Nui" pageId="15" pageNumber="416" specimenCode="CASIZ 222372" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBC39EFD03253905C46841E67EC0AECE" box="[325,503,1159,1185]" collectionCode="CASIZ" pageId="15" pageNumber="416">CASIZ 222372</specimenCode>
|
||
. Moto Nui,
|
||
<collectingCountry id="F372761603253905C7A941E67D9BAECE" box="[644,684,1159,1185]" name="Iceland" pageId="15" pageNumber="416">Isla</collectingCountry>
|
||
<collectorName id="2690535003253905C79841E67C10AECE" box="[693,807,1159,1185]" pageId="15" pageNumber="416">de Pascua</collectorName>
|
||
(Rapa Nui), ~
|
||
<specimenCount id="9D63FD0F03253905C6E341E67B26AECE" box="[974,1041,1159,1185]" pageId="15" pageNumber="416" type="generic">
|
||
<quantity id="4C9D9B6303253905C6E341E67B3AAECE" box="[974,1037,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="15" pageNumber="416" unit="m" value="82.0">
|
||
<elevation id="0048D1B503253905C6E341E67B3AAECE" box="[974,1037,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="15" pageNumber="416" unit="m" value="82.0">82 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
<collectorName id="2690535003253905C13741E67A36AECE" box="[1050,1281,1159,1185]" pageId="15" pageNumber="416">Coll. Bart Shepherd</collectorName>
|
||
,
|
||
<collectorName id="2690535003253905C02341E67AABAECE" box="[1294,1436,1159,1185]" pageId="15" pageNumber="416">Tyler Phelps</collectorName>
|
||
and
|
||
<collectorName id="2690535003253905C5EA41CA7E7DAEAA" box="[199,330,1195,1221]" pageId="15" pageNumber="416">Luis Rocha</collectorName>
|
||
,
|
||
<date id="FFDB104603253905C47B41CA7EC1AEAA" box="[342,502,1195,1221]" pageId="15" pageNumber="416" value="2017-03-06">
|
||
<collectingDate id="EF9FE9AE03253905C47B41CA7EC1AEAA" box="[342,502,1195,1221]" pageId="15" pageNumber="416" value="2017-03-06">6 March 2017</collectingDate>
|
||
</date>
|
||
, 1 wet spec.
|
||
<collectionCode id="ED74AE4303253905C7AA41CA7DACAEAA" box="[647,667,1195,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="416">R</collectionCode>
|
||
=3.8, r=1.6.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B0D3CDB03253905C60841CA7D3FAF62" ID-GBIF-Occurrence="3311713349" collectingDate="2017-03-06" collectionCode="CASIZ" collectorName="de Pascua & Coll. Bart Shepherd & Tyler Phelps & Luis Rocha" country="Iceland" elevation="82" latitude="-27.153242" location="Isla" longLatPrecision="1" longitude="-109.4479" municipality="Rapa Nui" pageId="15" pageNumber="416" specimenCode="CASIZ 222372" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBC39EFD03253905C60841CA7CE2AEAA" box="[805,981,1194,1221]" collectionCode="CASIZ" pageId="15" pageNumber="416">CASIZ 222375</specimenCode>
|
||
.
|
||
<location id="8EBA605D03253905C6F241CA7B51AEAA" LSID="urn:lsid:plazi:treatment:03CC879003273905C5BA440F7D3BAF62:8EBA605D03253905C6F241CA7B51AEAA" box="[991,1126,1195,1221]" country="Iceland" latitude="-27.153242" longLatPrecision="1" longitude="-109.4479" municipality="Rapa Nui" name="Hanga Piko" pageId="15" pageNumber="416">Hanga Piko</location>
|
||
,
|
||
<collectingCountry id="F372761603253905C15C41CA7BAEAEAA" box="[1137,1177,1195,1221]" name="Iceland" pageId="15" pageNumber="416">Isla</collectingCountry>
|
||
<collectorName id="2690535003253905C18C41CA7A26AEAA" box="[1185,1297,1195,1221]" pageId="15" pageNumber="416">de Pascua</collectorName>
|
||
(
|
||
<collectingMunicipality id="6BBEACFC03253905C03241CA7AA7AEA9" box="[1311,1424,1195,1222]" pageId="15" pageNumber="416">Rapa Nui</collectingMunicipality>
|
||
),
|
||
<geoCoordinate id="EE51504103253905C5BA41B17E19AE84" box="[151,302,1232,1259]" degrees="27" direction="south" minutes="9" orientation="latitude" pageId="15" pageNumber="416" precision="1" seconds="11.67" value="-27.153242">27°9′11.67″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE51504103253905C41741B17EC1AE84" box="[314,502,1232,1259]" degrees="109" direction="west" minutes="26" orientation="longitude" pageId="15" pageNumber="416" precision="1" seconds="52.42" value="-109.4479">109°26′52.42″W</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D63FD0F03253905C72E41AE7D66AE86" box="[515,593,1231,1257]" pageId="15" pageNumber="416" type="generic">
|
||
<quantity id="4C9D9B6303253905C72E41AE7D7AAE86" box="[515,589,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="416" unit="m" value="110.0">
|
||
<elevation id="0048D1B503253905C72E41AE7D7AAE86" box="[515,589,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="416" unit="m" value="110.0">110 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
<collectorName id="2690535003253905C77441AE7C09AE86" box="[601,830,1231,1257]" pageId="15" pageNumber="416">Coll. Bart Shepherd</collectorName>
|
||
,
|
||
<collectorName id="2690535003253905C66741AE7CEFAE86" box="[842,984,1231,1257]" pageId="15" pageNumber="416">Tyler Phelps</collectorName>
|
||
and
|
||
<collectorName id="2690535003253905C13C41AE7BA2AE86" box="[1041,1173,1231,1257]" pageId="15" pageNumber="416">Luis Rocha</collectorName>
|
||
,
|
||
<date id="FFDB104603253905C18F41AE7A66AE86" box="[1186,1361,1231,1257]" pageId="15" pageNumber="416" value="2017-03-11">
|
||
<collectingDate id="EF9FE9AE03253905C18F41AE7A66AE86" box="[1186,1361,1231,1257]" pageId="15" pageNumber="416" value="2017-03-11">11 March 2017</collectingDate>
|
||
</date>
|
||
, 2 wet spec.
|
||
<collectionCode id="ED74AE4303253905C5FA41927FDCAF62" box="[215,235,1267,1293]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="416">R</collectionCode>
|
||
=2.3, r=1.0;
|
||
<collectionCode id="ED74AE4303253905C45841927EBEAF62" box="[373,393,1267,1293]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="416">R</collectionCode>
|
||
=1.4, r=0.6
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |