162 lines
21 KiB
XML
162 lines
21 KiB
XML
<document id="5B7359102C257DB2F64F2B89B5783A3E" ID-DOI="10.5281/zenodo.12521148" ID-ISSN="1407-8619" ID-Zenodo-Dep="12521148" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719252154146" checkinUser="felipe" docAuthor="Rukmane, Anita" docDate="2019" docId="03CC878DFFD0DB41DDC751BEFEB0161D" docLanguage="en" docName="BalticJColeopterol.19.2.141-150.pdf" docOrigin="Baltic Journal of Coleopterology 19 (2)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.3:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="3" docTitle="Pachyrhynchus sagittatus Rukmane, 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="143" masterDocId="FFF5FFF5FFD1DB43DD245341FFCD152A" masterDocTitle="Four new species and two subspecies of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae: Pachyrhynchini) from Luzon Island, Philippines" masterLastPageNumber="150" masterPageNumber="141" pageNumber="142" updateTime="1719254198936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="76A60BAF2332CD79A8773402F33C38DE">Four new species and two subspecies of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae: Pachyrhynchini) from Luzon Island, Philippines</mods:title>
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<mods:namePart id="1C337799831DBA117B42F0FCF014C55B">Rukmane, Anita</mods:namePart>
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<mods:title id="0E21D58B361F75F35C619EC55AAEF667">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="7024F1447A524DF8BE01CD147A240A08">
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<mods:date id="E2F6A1BE4808F3F6A0448190C428B403">2019</mods:date>
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<mods:number id="D5DAA46F6FA842AC5D14412905534600">2019-12-31</mods:number>
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<mods:number id="7D1E21A2E5782BBDEE9173E562BE32CC">19</mods:number>
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<mods:identifier id="7072790930EBBB5D256291C733C3D278" type="DOI">10.5281/zenodo.12521148</mods:identifier>
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<mods:identifier id="1EA8EA5EB49838B7A2233E1824B4EF28" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03CC878DFFD0DB41DDC751BEFEB0161D" LSID="urn:lsid:plazi:treatment:03CC878DFFD0DB41DDC751BEFEB0161D" httpUri="http://treatment.plazi.org/id/03CC878DFFD0DB41DDC751BEFEB0161D" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142">
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<subSubSection id="C37F6510FFD0DB42DDC751BEFD9E163D" box="[227,595,767,791]" pageId="1" pageNumber="142" type="nomenclature">
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<paragraph id="8BDA369BFFD0DB42DDC751BEFD9E163D" blockId="1.[227,595,767,823]" box="[227,595,767,791]" pageId="1" pageNumber="142">
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<heading id="D09281F7FFD0DB42DDC751BEFD9E163D" bold="true" box="[227,595,767,791]" fontSize="10" level="2" pageId="1" pageNumber="142" reason="4">
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<emphasis id="B911EA89FFD0DB42DDC751BEFD9E163D" bold="true" box="[227,595,767,791]" pageId="1" pageNumber="142">
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<taxonomicName id="4C654D18FFD0DB42DDC751BEFE36163D" authorityName="Rukmane" authorityYear="2019" box="[227,507,767,791]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="sagittatus" status="sp. nov.">
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<emphasis id="B911EA89FFD0DB42DDC751BEFE36163D" bold="true" box="[227,507,767,791]" italics="true" pageId="1" pageNumber="142">Pachyrhynchus sagittatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257F2FFD0DB42DF2551BEFD9E163D" box="[513,595,767,791]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F6510FFD0DB42DDC7505EFEBC161D" box="[227,369,799,823]" pageId="1" pageNumber="142" type="description">
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<paragraph id="8BDA369BFFD0DB42DDC7505EFEBC161D" blockId="1.[227,595,767,823]" box="[227,369,799,823]" pageId="1" pageNumber="142">
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<figureCitation id="135E2A1EFFD0DB42DDC7505EFEE3161D" box="[227,302,799,823]" captionStart="Fig" captionStartId="4.[303,344,1548,1572]" captionTargetBox="[334,1313,210,1406]" captionTargetId="figure-14@4.[323,1333,209,1459]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus of the following species: A – P. tetramaculatus sp. nov. male; B – P. tetramaculatus sp. nov. female; C – P.sagittatus sp. nov. male; D – P. kirklayroni sp. nov. male; P. congestus ssp. aedamlayroni subsp. nov. male; E - P.congestus ssp. aedamlayroni subsp. nov female" figureDoi="http://doi.org/10.5281/zenodo.12521150" httpUri="https://zenodo.org/record/12521150/files/figure.png" pageId="1" pageNumber="142">Fig. 1C</figureCitation>
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,
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<figureCitation id="135E2A1EFFD0DB42DC1C505EFE88161D" box="[312,325,799,823]" captionStart="Fig" captionStartId="7.[227,267,1283,1307]" captionTargetBox="[239,1246,229,1199]" captionTargetId="figure-4@7.[234,1313,209,1209]" captionTargetPageId="7" captionText="Fig. 4. A, C, H, J, L – aedeagus in lateral view; B, D, I, K, M – aedeagus in ventral view; A, B – P.disargus sp. nov.; C, D – P.congestus ssp. aedamlayroni subsp. nov.; H, I – P.kirklayroni sp. nov.; J, K – P.sagittatus sp. nov.; L, M – P.tetramaculatus sp.nov.; E, F, G – female genitalia of P.phaleratus ssp. dannylayroni subsp. nov.; E – sternite VIII in ventral view; F – ovipositor in dorsal view; G – spermatheca. Scale bar 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.12521156" httpUri="https://zenodo.org/record/12521156/files/figure.png" pageId="1" pageNumber="142">4</figureCitation>
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J-K.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F6510FFD0DB42DDC7501EFE30113D" pageId="1" pageNumber="142" type="materials_examined">
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<paragraph id="8BDA369BFFD0DB42DDC7501EFE30113D" blockId="1.[227,737,863,1047]" pageId="1" pageNumber="142">
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<emphasis id="B911EA89FFD0DB42DDC7501EFE4D165D" bold="true" box="[227,384,863,887]" pageId="1" pageNumber="142">Type material.</emphasis>
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<materialsCitation id="3B0D3CC6FFD0DB42DCA2501EFD11169D" collectingDate="2015-05" country="Philippines" location="Sierra Madre" municipality="Luzon" pageId="1" pageNumber="142" specimenCount="1" stateProvince="Quirino" typeStatus="holotype">
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<typeStatus id="54DE8839FFD0DB42DCA2501EFE2B165D" box="[390,486,863,887]" pageId="1" pageNumber="142" type="holotype">Holotype</typeStatus>
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. Male. ‘’
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<collectingCountry id="F372760BFFD0DB42DF11501EFD18165D" box="[565,725,863,887]" name="Philippines" pageId="1" pageNumber="142">PHILIPPINES</collectingCountry>
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/
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<collectingMunicipality id="6BBEACE1FFD0DB42DDC750C1FEE516BD" box="[227,296,896,919]" pageId="1" pageNumber="142">Luzon</collectingMunicipality>
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,
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<location id="8EBA6040FFD0DB42DC14503EFE7716BD" LSID="urn:lsid:plazi:treatment:03CC878DFFD0DB41DDC751BEFEB0161D:8EBA6040FFD0DB42DC14503EFE7716BD" box="[304,442,895,919]" country="Philippines" municipality="Luzon" name="Sierra Madre" pageId="1" pageNumber="142" stateProvince="Quirino">Sierra Madre</location>
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,
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<collectingRegion id="49A1F879FFD0DB42DCE7503EFDD916BD" box="[451,532,895,919]" country="Philippines" name="Quirino" pageId="1" pageNumber="142">Quirino</collectingRegion>
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/
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<date id="FFDB105BFFD0DB42DF0750C1FDBF16BD" box="[547,626,895,919]" pageId="1" pageNumber="142" value="2015-05">
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<collectingDate id="EF9FE9B3FFD0DB42DF0750C1FDBF16BD" box="[547,626,895,919]" pageId="1" pageNumber="142" value="2015-05">V. 2015</collectingDate>
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</date>
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/ local collector leg.’’ (white rectangular label, printed)
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</materialsCitation>
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; ‘’
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<typeStatus id="54DE8839FFD0DB42DDCE50FEFEB016FD" box="[234,381,959,983]" pageId="1" pageNumber="142" type="holotype">HOLOTYPE</typeStatus>
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/ Male /
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<taxonomicName id="4C654D18FFD0DB42DCF150FEFD2C16FD" authorityName="Rukmane" authorityYear="2019" box="[469,737,959,983]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="sagittatus">
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<emphasis id="B911EA89FFD0DB42DCF150FEFD2C16FD" box="[469,737,959,983]" italics="true" pageId="1" pageNumber="142">Pachyrhynchus sagittatus</emphasis>
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</taxonomicName>
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/ Rukmane, 2019 / det. Rukmane A. 2019’’ (red rectangular label, printed).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F6510FFD0DB42DDC7577EFD00117D" box="[227,717,1087,1111]" pageId="1" pageNumber="142" type="distribution">
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<paragraph id="8BDA369BFFD0DB42DDC7577EFD00117D" blockId="1.[227,717,1087,1111]" box="[227,717,1087,1111]" pageId="1" pageNumber="142">
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<emphasis id="B911EA89FFD0DB42DDC7577EFEB8117D" bold="true" box="[227,373,1087,1111]" pageId="1" pageNumber="142">Distribution:</emphasis>
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Luzon Island,
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<collectingRegion id="49A1F879FFD0DB42DF34577EFD04117D" box="[528,713,1087,1111]" country="Philippines" name="Quirino" pageId="1" pageNumber="142">Quirino Province</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F6510FFD0DB41DDC7573EFE7617BD" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142" type="description">
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<paragraph id="8BDA369BFFD0DB42DDC7573EFD14119D" blockId="1.[227,738,1151,1335]" pageId="1" pageNumber="142">
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<emphasis id="B911EA89FFD0DB42DDC7573EFEA811BD" bold="true" box="[227,357,1151,1175]" pageId="1" pageNumber="142">Description.</emphasis>
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Dimensions (
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<typeStatus id="54DE8839FFD0DB42DCC9573EFD8411BD" box="[493,585,1151,1175]" pageId="1" pageNumber="142" type="holotype">holotype</typeStatus>
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): LB: 12.1; LE: 7.7; WE: 5.5; LP: 4.1; WP: 4.0; LR: 1.9; WR: 1.8.
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</paragraph>
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<paragraph id="8BDA369BFFD0DB42DDC75781FD93101D" blockId="1.[227,738,1151,1335]" pageId="1" pageNumber="142">
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Integument coperry red, body strongly shiny, underside with weaker luster; body with markings of pale green round to recumbent scales. Dorsal habitus as shown in
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<figureCitation id="135E2A1EFFD0DB42DF2C565EFD97101D" box="[520,602,1311,1335]" captionStart="Fig" captionStartId="4.[303,344,1548,1572]" captionTargetBox="[334,1313,210,1406]" captionTargetId="figure-14@4.[323,1333,209,1459]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus of the following species: A – P. tetramaculatus sp. nov. male; B – P. tetramaculatus sp. nov. female; C – P.sagittatus sp. nov. male; D – P. kirklayroni sp. nov. male; P. congestus ssp. aedamlayroni subsp. nov. male; E - P.congestus ssp. aedamlayroni subsp. nov female" figureDoi="http://doi.org/10.5281/zenodo.12521150" httpUri="https://zenodo.org/record/12521150/files/figure.png" pageId="1" pageNumber="142">Fig. 1C</figureCitation>
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.
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</paragraph>
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<paragraph id="8BDA369BFFD0DB42DDC75621FB4C149D" blockId="1.[227,737,1375,1751]" lastBlockId="1.[774,1285,191,439]" pageId="1" pageNumber="142">Rostrum longer than wide (LR/WR 1.06), pubescent, finely punctured, bulging on apical part, with peak slightly before middle, deep triangular impression on basal half, longitudinal groove from midline to base of forehead, shallow transverse grow at base of forehead; forehead with lanceolate patch of scales medially, strongly wrinkled, nearly two times as wide as eye; lateral parts of rostrum covered with round to oval scales and hair-like scales, most intense along genae; few white hairs before antennal scrobes and long light hairs after antennal scrobes, with longer hairs near apex; antennomers evenly covered with long light hairs, scape covered with pubescence on basal part and long light hairs on apical part; pedicel 1.5 times as long as first antennomer, lon ger th an wide, an ten n omer s II-V subsphereical, sub equal in length. Head glabrous, finely punctured; eyes small, slightly prominent from outline of the head.</paragraph>
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<paragraph id="8BDA369BFFD0DB42DE22529EFB7E161D" blockId="1.[774,1285,479,823]" pageId="1" pageNumber="142">Prothorax subspherical, nearly same length as width (LP/WP 1.03), widest just in the middle; weakly punctured; dorsal contour slightly incurved at apical 1/3, gradually increased to middle, rounded, gradually decreased to subbasal part, straightened to base; with the following markings of pale green scales: 1) arrow-shaped patch medially on disc from subbasal part to slightly after midline; 2) two small spots on disc medially, each redirected laterally; 3) subovate shape patch on each latero-ventral part.</paragraph>
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<paragraph id="8BDA369BFFD0DB42DE225021FBF116DD" blockId="1.[774,1285,863,1015]" pageId="1" pageNumber="142">Legs stout; coxa with pale green roundish scales and hair-like scales; femur with short light hairs on basal part, pubescent in all length and with round pale green scales on apical part; tarsus with long, rare, golden setae.</paragraph>
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<paragraph id="8BDA369BFFD0DB42DE22575EFC86133D" blockId="1.[774,1285,1055,1559]" pageId="1" pageNumber="142">Elytra sub ovate (LE/WE 1.4), intervals slightly pronounced, nearly smooth; each elytron with the following markings: 1) elongated patch at subbasal part along interval III and one along lateral margin; 2) transverse medial stripe from one lateral margin to other, stipe interrupted at interval V and VII forming small round spot; 3) longitudinal line from apical ½ to apex along interval III; 4) longitudinal line along lateral margin from apical 1/3 to basal 1/2; widest just in the middle; on dorsal dimension narrow at the base, gradually increased to middle, widest just in the middle, rounded and gradually decreased to apex up to apical ½, narrowed more strongly in direction to apex; apex sharpened, with weak pubescence.</paragraph>
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<paragraph id="8BDA369BFFD0DB42DE22557EFBA413BD" blockId="1.[774,1286,1599,1687]" pageId="1" pageNumber="142">
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Elytra wider than prothorax (WE/WP 1.38), nearly twice as long as prothorax (LE/LP 1.88). Aedeagus as shown in
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<figureCitation id="135E2A1EFFD0DB42DEDF553EFBF613BD" box="[1019,1083,1663,1687]" captionStart="Fig" captionStartId="7.[227,267,1283,1307]" captionTargetBox="[239,1246,229,1199]" captionTargetId="figure-4@7.[234,1313,209,1209]" captionTargetPageId="7" captionText="Fig. 4. A, C, H, J, L – aedeagus in lateral view; B, D, I, K, M – aedeagus in ventral view; A, B – P.disargus sp. nov.; C, D – P.congestus ssp. aedamlayroni subsp. nov.; H, I – P.kirklayroni sp. nov.; J, K – P.sagittatus sp. nov.; L, M – P.tetramaculatus sp.nov.; E, F, G – female genitalia of P.phaleratus ssp. dannylayroni subsp. nov.; E – sternite VIII in ventral view; F – ovipositor in dorsal view; G – spermatheca. Scale bar 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.12521156" httpUri="https://zenodo.org/record/12521156/files/figure.png" pageId="1" pageNumber="142">Fig. 4</figureCitation>
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J-K.
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</paragraph>
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<paragraph id="8BDA369BFFD0DB42DE2255FEFC1C13FD" blockId="1.[774,977,1727,1751]" box="[774,977,1727,1751]" pageId="1" pageNumber="142">
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<emphasis id="B911EA89FFD0DB42DE2255FEFC9213FD" bold="true" box="[774,863,1727,1751]" pageId="1" pageNumber="142">Female.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="8BDA369BFFD3DB41DC0B53FEFE7617BD" blockId="2.[303,814,191,663]" pageId="2" pageNumber="143">
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<emphasis id="B911EA89FFD3DB41DC0B53FEFDED15FD" bold="true" box="[303,544,191,215]" pageId="2" pageNumber="143">Differential analyses.</emphasis>
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This new species according to its body characteristics and scaly markings on the body is unique enough to be different from any other known species of the genus
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<taxonomicName id="4C654D18FFD3DB41DC0B527EFE1A147D" authorityName="Germar" authorityYear="1824" box="[303,471,319,343]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis id="B911EA89FFD3DB41DC0B527EFE1A147D" box="[303,471,319,343]" italics="true" pageId="2" pageNumber="143">Pachyrhynchus</emphasis>
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||
</taxonomicName>
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that comes from the Luzon Island. Nerveless,
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<taxonomicName id="4C654D18FFD3DB41DCC05221FDA5145D" authorityName="Rukmane" authorityYear="2019" box="[484,616,352,375]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="sagittatus" status="sp. nov.">
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<emphasis id="B911EA89FFD3DB41DCC05221FDA5145D" box="[484,616,352,375]" italics="true" pageId="2" pageNumber="143">P. sagittatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257F2FFD3DB41DF4B5221FD0F145D" box="[623,706,352,375]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
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is similar to newly described
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<taxonomicName id="4C654D18FFD3DB41DF2252C1FD5614BD" authorityName="Rukmane" authorityYear="2019" box="[518,667,383,407]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="kirklayroni" status="sp. nov.">
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<emphasis id="B911EA89FFD3DB41DF2252C1FD5614BD" box="[518,667,383,407]" italics="true" pageId="2" pageNumber="143">P. kirklayroni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257F2FFD3DB41DF8052C1FD3014BD" box="[676,765,384,407]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
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, but differs by the following characters: 1) Prothorax of
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<taxonomicName id="4C654D18FFD3DB41DC695281FE2F14FD" authorityName="Rukmane" authorityYear="2019" box="[333,482,447,471]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="kirklayroni" status="sp. nov.">
|
||
<emphasis id="B911EA89FFD3DB41DC695281FE2F14FD" box="[333,482,447,471]" italics="true" pageId="2" pageNumber="143">P. kirklayroni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F2FFD3DB41DCCE5281FD8D14FD" box="[490,576,448,471]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
|
||
wider, more strongly rounded, dorsal contour at apical ½ without depression; 2) rostrum of
|
||
<taxonomicName id="4C654D18FFD3DB41DF1C5141FD00173D" authorityName="Rukmane" authorityYear="2019" box="[568,717,511,535]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="kirklayroni" status="sp. nov.">
|
||
<emphasis id="B911EA89FFD3DB41DF1C5141FD00173D" box="[568,717,511,535]" italics="true" pageId="2" pageNumber="143">P. kirklayroni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F2FFD3DB41DFF35141FCE0173D" box="[727,813,512,535]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
|
||
wider, with weaker depression on basal part; 3) elytra of
|
||
<taxonomicName id="4C654D18FFD3DB41DCBF5101FDE4177D" authorityName="Rukmane" authorityYear="2019" box="[411,553,576,599]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="sagittatus" status="sp. nov.">
|
||
<emphasis id="B911EA89FFD3DB41DCBF5101FDE4177D" box="[411,553,576,599]" italics="true" pageId="2" pageNumber="143">P. sagittatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F2FFD3DB41DF105101FD5D177D" box="[564,656,576,599]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shorter, with unique scaly markings; 4) male aedeagus of different shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6510FFD3DB41DC0B51FEFEB0161D" pageId="2" pageNumber="143" type="etymology">
|
||
<paragraph id="8BDA369BFFD3DB41DC0B51FEFEB0161D" blockId="2.[303,813,703,823]" pageId="2" pageNumber="143">
|
||
<emphasis id="B911EA89FFD3DB41DC0B51FEFE6317FD" bold="true" box="[303,430,703,727]" pageId="2" pageNumber="143">Etymology.</emphasis>
|
||
This species was named based on its typical marking on the pronotum – arrow shape spot dorsally on disc. Latin word of the arrow is sagitta.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |