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<document id="B785C731B398684D945E28FDEAABDEE7" ID-DOI="10.5281/zenodo.275935" ID-GBIF-Dataset="2eb79791-0803-42aa-aa73-bb32110be829" ID-ISSN="1175-5326" ID-Zenodo-Dep="275935" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460271267434" checkinUser="plazi" docAuthor="Grismer, Lee, Sumontha, Montri, Cota, Michael, Grismer, Jesse L., Wood, Perry L., Pauwels, Olivier S. G. &amp; Kunya, Kirati" docDate="2010" docId="03CB5E24FFD5CB6D1595FAF6FC047FFD" docLanguage="en" docName="zt02576p055.pdf" docOrigin="Zootaxa 2576" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cnemaspis niyomwanae Grismer, Sumontha, Cota, Grismer, Wood, Pauwels &amp; Kunya, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="49" masterDocId="FFF2265CFFF8CB5C1502FFB4FFF97E4A" masterDocTitle="A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species" masterLastPageNumber="55" masterPageNumber="1" pageNumber="46" updateTime="1698246858311" updateUser="plazi">
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<mods:title id="7183C71B619F4EAC5E540366B12C9ECC">A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species</mods:title>
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<mods:namePart id="3B4C47143E846BD27DD7B18A3369FB29">Grismer, Lee</mods:namePart>
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<mods:namePart id="09826F9665B40F09D8651F005EE3E4E1">Sumontha, Montri</mods:namePart>
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<mods:namePart id="4AB299118CA3689AAE8AE39453A6BE48">Cota, Michael</mods:namePart>
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<mods:namePart id="44D2DED148D9A7FD3F12AD9EA7084850">Grismer, Jesse L.</mods:namePart>
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<mods:namePart id="5A55E70E6B94BF3C03CA393B8FFCFA0E">Wood, Perry L.</mods:namePart>
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<mods:namePart id="E44E1C017A777F58D336C87A0055413B">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="2182D5F4C5328DAA674E9C902B132BB2">Kunya, Kirati</mods:namePart>
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<mods:date id="E28279568DCC2B99D953EC04A53E197D">2010</mods:date>
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<treatment id="03CB5E24FFD5CB6D1595FAF6FC047FFD" ID-DOI="http://doi.org/10.5281/zenodo.6205214" ID-GBIF-Taxon="119395214" ID-Zenodo-Dep="6205214" LSID="urn:lsid:plazi:treatment:03CB5E24FFD5CB6D1595FAF6FC047FFD" httpUri="http://treatment.plazi.org/id/03CB5E24FFD5CB6D1595FAF6FC047FFD" lastPageId="49" lastPageNumber="49" pageId="45" pageNumber="46">
<subSubSection id="C378BCB9FFD5CB711595FAF6FEFE7B80" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BDDEF32FFD5CB711595FAF6FDD97B16" blockId="45.[151,566,1346,1482]" box="[151,544,1346,1372]" pageId="45" pageNumber="46">
<heading id="D095585EFFD5CB711595FAF6FDD97B16" bold="true" box="[151,544,1346,1372]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis id="B9163320FFD5CB711595FAF6FDD97B16" bold="true" box="[151,544,1346,1372]" pageId="45" pageNumber="46">
<taxonomicName id="4C6294B1FFD5CB711595FAF6FE427B16" box="[151,443,1346,1372]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFD5CB711595FAF6FE427B16" bold="true" box="[151,443,1346,1372]" italics="true" pageId="45" pageNumber="46">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFD5CB7114C0FAF6FDD97B16" box="[450,544,1346,1372]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDDEF32FFD5CB711595FAD1FEFE7B80" blockId="45.[151,566,1346,1482]" pageId="45" pageNumber="46">
Niyomwans Rock Gecko Djing Djok Niew Yaow Niyomwan
<figureCitation id="1359F3B7FFD5CB711595FA04FEFE7B80" box="[151,263,1456,1482]" captionStart="FIGURE 21" captionStartId="47.[151,257,1947,1971]" captionTargetBox="[228,1361,194,1930]" captionTargetId="figure@47.[228,1361,194,1930]" captionTargetPageId="47" captionText="FIGURE 21. Upper: adult male Cnemaspis niyomwanae sp. nov. (CUMZ R- 2009,6,24 - 10). Middle: adult female C. niyomwanae sp. nov. (holotype THNHM 15910). Lower: ventral view of uncatalogued female C. niyomwanae sp. nov. (photos by M. Sumontha)." httpUri="https://zenodo.org/record/275955/files/figure.png" pageId="45" pageNumber="46">Figure 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C378BCB9FFD5CB711595FA41FEA078A5" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BDDEF32FFD5CB711595FA41FAE0787E" blockId="45.[151,1437,1525,2036]" pageId="45" pageNumber="46">
<emphasis id="B9163320FFD5CB711595FA41FEE97845" bold="true" box="[151,272,1525,1551]" pageId="45" pageNumber="46">
<typeStatus id="54D95190FFD5CB711595FA41FEF27845" box="[151,267,1525,1551]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult female (
<collectionCode id="ED7377F7FFD5CB7114C1FA41FDC87845" box="[451,561,1525,1551]" pageId="45" pageNumber="46">THNHM</collectionCode>
15910) from Thum Khao Ting, Palean District, Trang Province,
<collectingCountry id="F375AFA2FFD5CB711036FA41FA657845" box="[1332,1436,1525,1551]" name="Thailand" pageId="45" pageNumber="46">Thailand</collectingCountry>
(07°
<geoCoordinate id="EE5689F5FFD5CB7115C7F9AEFED6787E" box="[197,303,1562,1588]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="55" value="9.943">09.943N</geoCoordinate>
99°
<geoCoordinate id="EE5689F5FFD5CB71145FF9AEFE3D787E" box="[349,452,1562,1588]" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="55" value="48.142">48.142E</geoCoordinate>
) at
<quantity id="4C9A42D7FFD5CB7114F2F9AEFDD2787E" box="[496,555,1562,1588]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="45" pageNumber="46" unit="m" value="28.0">28 m</quantity>
in elevation. Collected by M. Sumontha on
<date id="FFDCC9F2FFD5CB711130F9AEFAEC787E" box="[1074,1301,1562,1588]" pageId="45" pageNumber="46" value="2009-12-26">26 December 2009</date>
.
</paragraph>
<paragraph id="8BDDEF32FFD5CB7115C4F98BFEA078A5" blockId="45.[151,1437,1525,2036]" pageId="45" pageNumber="46">
<emphasis id="B9163320FFD5CB7115C4F98BFEB27813" bold="true" box="[198,331,1599,1625]" pageId="45" pageNumber="46">
<typeStatus id="54D95190FFD5CB7115C4F98BFEBF7813" box="[198,326,1599,1625]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Adult male (
<collectionCode id="ED7377F7FFD5CB7114FCF9F4FDAF7810" box="[510,598,1600,1626]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="45" pageNumber="46">ZMKU</collectionCode>
Rep-000315), adult female (PSUZC-RT 2010.56), subadult males (
<collectionCode id="ED7377F7FFD5CB71159DF9D1FF0F7835" box="[159,246,1637,1663]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="45" pageNumber="46">CUMZ</collectionCode>
<collectionCode id="ED7377F7FFD5CB7115FCF9D1FEE57835" box="[254,284,1637,1663]" pageId="45" pageNumber="46">R-</collectionCode>
2009,6,24-10, KZM 008) from Baan Nam Pud, La-ngu District, Satun Province (07°
<geoCoordinate id="EE5689F5FFD5CB711002F9D1FA927835" box="[1280,1387,1637,1663]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="55" value="5.688">05.688N</geoCoordinate>
99°
<geoCoordinate id="EE5689F5FFD5CB711595F93EFF0278EE" box="[151,251,1674,1700]" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="55" value="54.732">54.732E</geoCoordinate>
) at
<quantity id="4C9A42D7FFD5CB711424F93EFE7778EE" box="[294,398,1674,1700]" metricMagnitude="1" metricUnit="m" metricValue="4.2" metricValueMax="4.6" metricValueMin="3.8" pageId="45" pageNumber="46" unit="m" value="42.0" valueMax="46.0" valueMin="38.0">3846 m</quantity>
in elevation.
<collectionCode id="ED7377F7FFD5CB71172EF93EFD7B78EE" box="[556,642,1674,1700]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="45" pageNumber="46">ZMKU</collectionCode>
Rep-000315,
<collectionCode id="ED7377F7FFD5CB71162BF93EFC8778EE" box="[809,894,1674,1700]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="45" pageNumber="46">CUMZ</collectionCode>
<collectionCode id="ED7377F7FFD5CB711687F93EFC5B78EE" box="[901,930,1674,1700]" pageId="45" pageNumber="46">R-</collectionCode>
2009,6,24-10 and KZM 0 0 8 were collected by Siriwat Dangsri on
<date id="FFDCC9F2FFD5CB71149DF904FD677880" box="[415,670,1712,1738]" pageId="45" pageNumber="46" value="2009-10-06" valueMax="2009-10-07" valueMin="2009-10-06">6 and 7 October 2009</date>
. PSUZC-RT 2010.56 was collected by Thanin Kaewmanee on
<date id="FFDCC9F2FFD5CB71107DF904FEAD78A5" pageId="45" pageNumber="46" value="2009-12-24">24 December 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C378BCB9FFD5CB7215C7F94EFD147FD8" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BDDEF32FFD5CB7215C7F94EFD147FD8" blockId="45.[151,1437,1525,2036]" lastBlockId="46.[151,1437,152,1970]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="B9163320FFD5CB7115C7F94EFEBD795E" bold="true" box="[197,324,1786,1812]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
Adult males reaching
<quantity id="4C9A42D7FFD5CB71174EF94EFD4C795E" box="[588,693,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="4.59" pageId="45" pageNumber="46" unit="mm" value="45.9">45.9 mm</quantity>
SVL, adult females reaching
<quantity id="4C9A42D7FFD5CB711113F94EFB83795E" box="[1041,1146,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="5.68" pageId="45" pageNumber="46" unit="mm" value="56.8">56.8 mm</quantity>
SVL; 811 supralabials; 68 infralabials; gulars smooth; forearm scales keeled; subtibials, ventrals, subcaudals smooth; dorsal tubercles keeled; 2631 paravertebral tubercles; enlarged, elongate, laterally compressed, white, isolated tubercles on sides of neck, shoulders, and flanks; dorsolateral, lateral, and ventrolateral caudal tubercles usually present only anteriorly; caudal tubercles do not encircle tail; caudal tubercles absent from lateral, caudal furrow; middorsal caudal furrow absent; median row of smooth, enlarged subcaudals; three porebearing, precloacal scales in males; one or two postcloacal tubercles; shield-like subtibials and enlarged, submetatarsals absent; 3134 subdigital lamellae on fourth toe; yellow and orange alternating bands on forearms and forelimbs in subadult and adult males; paired, yellow, rostral stripes in subadult and adult males; paried, cream to white, circular, occipital blotches; large, cream to white, paravertebral butterfly-shaped dorsal blotches; no dark, longitudinal, gular markings or blotches; head not yellow in adult males; no black neck patch enclosing a white to yellow ocellus; no dark shoulder patch enclosing white to yellow ocellus; no prominent, yellow to white, prescapular crescent or transverse bars on flanks. These differences are summarized across all species in
<tableCitation id="C6E0DA89FFD6CB72171EFECCFD677FD8" box="[540,670,376,402]" captionStart="TABLE 1" captionStartId="9.[151,242,151,175]" captionTargetBox="[151,1436,277,1409]" captionTargetPageId="9" captionText="TABLE 1. Diagnostic characters of Southeast Asian species of Cnemaspis. New species in bold type. / = character not evaluated." httpUri="http://table.plazi.org/id/DF1DBFBAFFF1CB551595FF23FEFD7E85" pageId="46" pageNumber="47" tableUuid="DF1DBFBAFFF1CB551595FF23FEFD7E85">TABLES 1</tableCitation>
and
<tableCitation id="C6E0DA89FFD6CB7217DAFECCFD137FD8" box="[728,746,376,402]" captionStart="TABLE 2" captionStartId="15.[151,241,972,996]" captionTargetBox="[151,1396,1071,2035]" captionTargetPageId="15" captionText="TABLE 2. Diagnostic characteristics of color pattern of the Cnemaspis of Thailand. 1 = character state present; 0 = character state absent. / = character not evaluated." httpUri="http://table.plazi.org/id/DF1DBFBAFFF7CB531595FC78FD537A4E" pageId="46" pageNumber="47" tableUuid="DF1DBFBAFFF7CB531595FC78FD537A4E">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C378BCB9FFD6CB6C15C7FE29FE377FD8" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="8BDDEF32FFD6CB7215C7FE29FADC7D18" blockId="46.[151,1437,152,1970]" pageId="46" pageNumber="47">
<emphasis id="B9163320FFD6CB7215C7FE29FE0E7FFD" bold="true" box="[197,503,413,439]" pageId="46" pageNumber="47">
Description of
<typeStatus id="54D95190FFD6CB721480FE29FE0B7FFD" box="[386,498,413,439]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female; SVL
<quantity id="4C9A42D7FFD6CB7217ECFE29FCA47FFD" box="[750,861,413,439]" metricMagnitude="-2" metricUnit="m" metricValue="5.68" pageId="46" pageNumber="47" unit="mm" value="56.8">56.8 mm</quantity>
; head oblong in dorsal profile, moderate in size (HL/SVL 0.25), somewhat narrow (HW/SVL 0.16), flat (HD/HL 0.44), distinct from neck; snout short (ES/ HL 0.51), concave in lateral profile; postnasal region constricted medially; scales of rostrum low, rounded, juxtaposed, smooth, larger than similarly shaped scales on occiput; distinct, supraorbital ridges; shallow frontonasal sulcus; canthus rostralis smoothly rounded; eye large (ED/HL 0.21); extra-brillar fringe scales small in general but larger anteriorly; pupil round; ear opening oval, taller than wide; rostral concave dorsally, dorsal 75% divided by longitudinal groove; rostral bordered posteriorly by two supranasals contacting medially, and nostrils; bordered laterally by first supralabials; 8R,L raised supralabials of similar size, but smallest posteriorly; 7R,L infralabials, decreasing gradually in size posteriorly; nostrils small, oblong, oriented dorsoposteriorly; bordered posteriorly by small, granular, postnasal scales; mental large, triangular, slightly concave, extending to level of second infralabials, bordered posteriorly by three postmentals, lateral postmentals much larger than center postmental; gular and throat scales smooth, raised, juxtaposed.
</paragraph>
<paragraph id="8BDDEF32FFD6CB7215C7FCE9FC3E7828" blockId="46.[151,1437,152,1970]" pageId="46" pageNumber="47">Body slender, elongate (AG/SVL 0.45); small, raised, smooth, dorsal scales generally equal in size throughout body, intermixed with numerous, moderately sized, keeled, randomly arranged tubercles; enlarged, isolated tubercles on sides of neck, shoulders, and flanks; tubercles extend from occiput to base of tail and are smallest anteriorly; 3134 paravertebral tubercles; pectoral and abdominal scales smooth, flat, imbricate; abdominal scales larger than pectoral scales and dorsals; no pore-bearing, precloacal scales; forelimbs moderately long, slender, dorsal scales keeled; ventral scales of forearm smooth, juxtaposed to subimbricate; palmar scales smooth, raised, juxtaposed; digits long with an inflected joint; claws recurved; subdigital lamellae unnotched; subdigital lamellae wide throughout length of digits, bearing a larger scale at digital inflections; interdigital webbing generally absent; fingers increase in length from first to fourth with fifth same length as fourth; hind limbs longer and thicker than forelimbs; dorsal scales of thigh, keeled, raised, juxtaposed; dorsal scales of foreleg rugose, subimbricate; ventral scales of hind limb smooth, imbricate; plantar scales smooth, flat, subimbricate; no enlarged subtibials or submetatarsals; digits elongate with an inflected joint; claws recurved; subdigital lamellae unnotched; lamellae wide throughout length of digits except at base where scales are more granular; enlarged scale at digital inflections; interdigital webbing absent; toes increase in length from first to fourth with fourth and fifth nearly equal in length; 33R,L subdigital lamellae on fourth toe; caudal scales arranged in segmented whorls, low, smooth, juxtaposed anteriorly; middorsal furrow absent; lateral, caudal furrow weak; median row of enlarged, smooth, subcaudal scales; four scales per caudal segment; other subcaudals smooth; paravertebral rows of large, keeled, caudal tubercles extend length of tail, becoming smaller and smooth posteriorly; dorsolateral row of tubercles present only anteriorly, al other rows absent; caudal tubercles do not encircle tail, absent from lateral, caudal furrow; one or two postcloacal tubercles; tail approximately 1.3% times SVL.</paragraph>
<paragraph id="8BDDEF32FFD6CB7215C7F9D9FA7279F8" blockId="46.[151,1437,152,1970]" pageId="46" pageNumber="47">
<emphasis id="B9163320FFD6CB7215C7F9D9FDEB78CD" bold="true" box="[197,530,1645,1671]" pageId="46" pageNumber="47">
Coloration (in life;
<figureCitation id="1359F3B7FFD6CB7214B6F9D9FDF078CD" box="[436,521,1645,1671]" captionStart="FIGURE 21" captionStartId="47.[151,257,1947,1971]" captionTargetBox="[228,1361,194,1930]" captionTargetId="figure@47.[228,1361,194,1930]" captionTargetPageId="47" captionText="FIGURE 21. Upper: adult male Cnemaspis niyomwanae sp. nov. (CUMZ R- 2009,6,24 - 10). Middle: adult female C. niyomwanae sp. nov. (holotype THNHM 15910). Lower: ventral view of uncatalogued female C. niyomwanae sp. nov. (photos by M. Sumontha)." httpUri="https://zenodo.org/record/275955/files/figure.png" pageId="46" pageNumber="47">Fig. 21</figureCitation>
)
</emphasis>
. Dorsal ground color of head, body, and tail faded green; dorsal ground color of limbs faded brown; top of head bearing a pair of yellow, rostral stripes beginning in interorbital region and terminating in postnasal region; diffuse, light occiptal blotches present; postorbital striping absent; faint, light mottling on sides of head; whitish, medial blotch on nape followed posteriorly by five lightly colored, paravertebral, butterfly-shaped markings between forelimb insertions and base of tail; markings continue onto tail to form lightly colored bands; enlarged, white tubercles on sides of neck, shoulders and flanks; other tubercles on body dark or lightly colored; upper regions of limbs bearing diffuse, light mottling; lower limbs bearing diffuse yellowish and brownish band; digits white bearing broad, brown bands; all ventral surfaces except subcaudal region of uniform beige with fine, dark stippling in some scales; subcaudal region grayish.
</paragraph>
<caption id="DF1DBFBAFFD7CB731595F82FFE4979B8" httpUri="https://zenodo.org/record/275955/files/figure.png" pageId="47" pageNumber="48" targetBox="[228,1361,194,1930]" targetPageId="47">
<paragraph id="8BDDEF32FFD7CB731595F82FFE4979B8" blockId="47.[151,1436,1947,2034]" pageId="47" pageNumber="48">
<emphasis id="B9163320FFD7CB731595F82FFED379F9" bold="true" box="[151,298,1947,1971]" pageId="47" pageNumber="48">FIGURE 21.</emphasis>
Upper: adult male
<taxonomicName id="4C6294B1FFD7CB731702F82FFCF379F8" box="[512,778,1947,1970]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFD7CB731702F82FFCF379F8" box="[512,778,1947,1970]" italics="true" pageId="47" pageNumber="48">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFD7CB731613F82FFC9079F9" box="[785,873,1947,1971]" pageId="47" pageNumber="48" rank="species">
<emphasis id="B9163320FFD7CB731613F82FFC9079F9" bold="true" box="[785,873,1947,1971]" pageId="47" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
(CUMZ R-2009,6,24-10). Middle: adult female
<taxonomicName id="4C6294B1FFD7CB731081F82FFEE77998" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFD7CB731081F82FFEE77998" italics="true" pageId="47" pageNumber="48">C. niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFD7CB731427F80FFE827999" box="[293,379,1979,2003]" pageId="47" pageNumber="48" rank="species">
<emphasis id="B9163320FFD7CB731427F80FFE827999" bold="true" box="[293,379,1979,2003]" pageId="47" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype THNHM 15910). Lower: ventral view of uncatalogued female
<taxonomicName id="4C6294B1FFD7CB73119AF80FFAC67998" box="[1176,1343,1979,2002]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFD7CB73119AF80FFAC67998" box="[1176,1343,1979,2002]" italics="true" pageId="47" pageNumber="48">C. niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFD7CB731044F80FFA657999" box="[1350,1436,1979,2003]" pageId="47" pageNumber="48" rank="species">
<emphasis id="B9163320FFD7CB731044F80FFA657999" bold="true" box="[1350,1436,1979,2003]" pageId="47" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
(photos by M. Sumontha).
</paragraph>
</caption>
<paragraph id="8BDDEF32FFC8CB6C15C7FF23FE377FD8" blockId="48.[151,1437,151,402]" pageId="48" pageNumber="49">
<emphasis id="B9163320FFC8CB6C15C7FF23FEB37EFB" bold="true" box="[197,330,151,177]" pageId="48" pageNumber="49">Variation.</emphasis>
Differences in squamation and morphometrics are presented in
<tableCitation id="C6E0DA89FFC8CB6C1184FF2CFAFD7EF8" box="[1158,1284,152,178]" captionStart="TABLE 9" captionStartId="48.[151,240,450,474]" captionTargetBox="[151,1436,533,1569]" captionTargetPageId="48" captionText="TABLE 9. Descriptive measurements and scale counts of the type series of Cnemaspis niyomwanae. See materials and methods for abbreviations. / = character not evaluated." httpUri="http://table.plazi.org/id/DF1DBFBAFFC8CB6C1595FE76FD247FB3" pageId="48" pageNumber="49" tableUuid="DF1DBFBAFFC8CB6C1595FE76FD247FB3">TABLE 9</tableCitation>
.
<taxonomicName id="4C6294B1FFC8CB6C1011FF2CFED27E9C" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFC8CB6C1011FF2CFED27E9C" italics="true" pageId="48" pageNumber="49">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC8CB6C1431FF09FE687E9D" box="[307,401,189,215]" pageId="48" pageNumber="49" rank="species">
<emphasis id="B9163320FFC8CB6C1431FF09FE687E9D" bold="true" box="[307,401,189,215]" pageId="48" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
shows significant sexual dimorphism with males being more colorful than females. MS 399 has well-defined, light colored dorsal markings and occipital blotches as compared to the more diffuse blotches seen in females. Males also have bright yellow, rostral stripes and alternating bright red-orange and yellow bands on the forelimbs. These markings are much more drab in females. This species also shows a marked light color phase at night (
<figureCitation id="1359F3B7FFC8CB6C1737FEE6FD747F26" box="[565,653,338,364]" captionStart="FIGURE 21" captionStartId="47.[151,257,1947,1971]" captionTargetBox="[228,1361,194,1930]" captionTargetId="figure@47.[228,1361,194,1930]" captionTargetPageId="47" captionText="FIGURE 21. Upper: adult male Cnemaspis niyomwanae sp. nov. (CUMZ R- 2009,6,24 - 10). Middle: adult female C. niyomwanae sp. nov. (holotype THNHM 15910). Lower: ventral view of uncatalogued female C. niyomwanae sp. nov. (photos by M. Sumontha)." httpUri="https://zenodo.org/record/275955/files/figure.png" pageId="48" pageNumber="49">Fig. 21</figureCitation>
) where the colors of both males and females become more faded and much less contrasting.
</paragraph>
</subSubSection>
<caption id="DF1DBFBAFFC8CB6C1595FE76FD247FB3" ID-Table-UUID="DF1DBFBAFFC8CB6C1595FE76FD247FB3" httpUri="http://table.plazi.org/id/DF1DBFBAFFC8CB6C1595FE76FD247FB3" pageId="48" pageNumber="49" targetBox="[151,1436,533,1569]" targetIsTable="true" targetPageId="48">
<paragraph id="8BDDEF32FFC8CB6C1595FE76FD247FB3" blockId="48.[151,1436,450,505]" pageId="48" pageNumber="49">
<emphasis id="B9163320FFC8CB6C1595FE76FEF27F90" bold="true" box="[151,267,450,474]" pageId="48" pageNumber="49">TABLE 9.</emphasis>
Descriptive measurements and scale counts of the type series of
<emphasis id="B9163320FFC8CB6C16C9FE76FB2E7F93" box="[971,1239,450,473]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="4C6294B1FFC8CB6C16C9FE76FB2A7F93" box="[971,1235,450,473]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="niyomwanae">Cnemaspis niyomwanae</taxonomicName>
.
</emphasis>
See materials and methods for abbreviations. / = character not evaluated.
</paragraph>
</caption>
<paragraph id="8BDDEF32FFC8CB6C1723FDA1FAFB786B" pageId="48" pageNumber="49">
<table id="F9621D92FFC834A31595FDA1FA65786B" box="[151,1436,533,1569]" gridcols="6" gridrows="16" pageId="48" pageNumber="49">
<tr id="3552ED70FFC834A31595FDA1FA657C06" box="[151,1436,533,588]" gridrow="0" pageId="48" pageNumber="49" rowspan-0="1">
<th id="7683840CFFC834A31723FDA1FD587C06" box="[545,673,533,588]" gridcol="1" gridrow="0" pageId="48" pageNumber="49">ZMKU Rep 0 0 0 315</th>
<th id="7683840CFFC834A317CCFDA1FCB37C06" box="[718,842,533,588]" gridcol="2" gridrow="0" pageId="48" pageNumber="49">PSUZC-RT 2010.56</th>
<th id="7683840CFFC834A3167EFDA1FC277C06" box="[892,990,533,588]" gridcol="3" gridrow="0" pageId="48" pageNumber="49">THNHM 15910</th>
<th id="7683840CFFC834A31128FDA1FB4E7C06" box="[1066,1207,533,588]" gridcol="4" gridrow="0" pageId="48" pageNumber="49">CUMZ R 2009,6,24-10</th>
<th id="7683840CFFC834A311F9FDA1FA657C06" box="[1275,1436,533,588]" gridcol="5" gridrow="0" pageId="48" pageNumber="49">KZM 0 0 8</th>
</tr>
<tr id="3552ED70FFC834A31595FDD4FA657CEB" box="[151,1436,608,673]" gridrow="1" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FDD4FE6D7CEB" box="[151,404,608,673]" gridcol="0" gridrow="1" pageId="48" pageNumber="49">SVL</th>
<td id="7683840CFFC834A31723FDD4FD587CEB" box="[545,673,608,673]" gridcol="1" gridrow="1" pageId="48" pageNumber="49">paratype 45.9</td>
<td id="7683840CFFC834A317CCFDD4FCB37CEB" box="[718,842,608,673]" gridcol="2" gridrow="1" pageId="48" pageNumber="49">paratype 46.6</td>
<td id="7683840CFFC834A3167EFDD4FC277CEB" box="[892,990,608,673]" gridcol="3" gridrow="1" pageId="48" pageNumber="49">holotype 56.8</td>
<td id="7683840CFFC834A31128FDD4FB4E7CEB" box="[1066,1207,608,673]" gridcol="4" gridrow="1" pageId="48" pageNumber="49">paratype 38.4</td>
<td id="7683840CFFC834A311F9FDD4FA657CEB" box="[1275,1436,608,673]" gridcol="5" gridrow="1" pageId="48" pageNumber="49">paratype 41.1</td>
</tr>
<tr id="3552ED70FFC834A31595FD01FA657C86" box="[151,1436,693,716]" gridrow="2" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FD01FE6D7C86" box="[151,404,693,716]" gridcol="0" gridrow="2" pageId="48" pageNumber="49">Sex</th>
<td id="7683840CFFC834A31723FD01FD587C86" box="[545,673,693,716]" gridcol="1" gridrow="2" pageId="48" pageNumber="49">m</td>
<td id="7683840CFFC834A317CCFD01FCB37C86" box="[718,842,693,716]" gridcol="2" gridrow="2" pageId="48" pageNumber="49">f</td>
<td id="7683840CFFC834A3167EFD01FC277C86" box="[892,990,693,716]" gridcol="3" gridrow="2" pageId="48" pageNumber="49">f</td>
<td id="7683840CFFC834A31128FD01FB4E7C86" box="[1066,1207,693,716]" gridcol="4" gridrow="2" pageId="48" pageNumber="49">m</td>
<td id="7683840CFFC834A311F9FD01FA657C86" box="[1275,1436,693,716]" gridcol="5" gridrow="2" pageId="48" pageNumber="49">M</td>
</tr>
<tr id="3552ED70FFC834A31595FD54FA657D6B" box="[151,1436,736,801]" gridrow="3" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FD54FE6D7D6B" box="[151,404,736,801]" gridcol="0" gridrow="3" pageId="48" pageNumber="49">Supralabials Infralabials</th>
<td id="7683840CFFC834A31723FD54FD587D6B" box="[545,673,736,801]" gridcol="1" gridrow="3" pageId="48" pageNumber="49">8 7</td>
<td id="7683840CFFC834A317CCFD54FCB37D6B" box="[718,842,736,801]" gridcol="2" gridrow="3" pageId="48" pageNumber="49">11 8</td>
<td id="7683840CFFC834A3167EFD54FC277D6B" box="[892,990,736,801]" gridcol="3" gridrow="3" pageId="48" pageNumber="49">8 6</td>
<td id="7683840CFFC834A31128FD54FB4E7D6B" box="[1066,1207,736,801]" gridcol="4" gridrow="3" pageId="48" pageNumber="49">9 7</td>
<td id="7683840CFFC834A311F9FD54FA657D6B" box="[1275,1436,736,801]" gridcol="5" gridrow="3" pageId="48" pageNumber="49">9 7</td>
</tr>
<tr id="3552ED70FFC834A31595FC81FA657D06" box="[151,1436,821,844]" gridrow="4" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FC81FE6D7D06" box="[151,404,821,844]" gridcol="0" gridrow="4" pageId="48" pageNumber="49">Precloacal pores</th>
<td id="7683840CFFC834A31723FC81FD587D06" box="[545,673,821,844]" gridcol="1" gridrow="4" pageId="48" pageNumber="49">3</td>
<td id="7683840CFFC834A317CCFC81FCB37D06" box="[718,842,821,844]" gridcol="2" gridrow="4" pageId="48" pageNumber="49">/</td>
<td id="7683840CFFC834A3167EFC81FC277D06" box="[892,990,821,844]" gridcol="3" gridrow="4" pageId="48" pageNumber="49">/</td>
<td id="7683840CFFC834A31128FC81FB4E7D06" box="[1066,1207,821,844]" gridcol="4" gridrow="4" pageId="48" pageNumber="49">3</td>
<td id="7683840CFFC834A311F9FC81FA657D06" box="[1275,1436,821,844]" gridcol="5" gridrow="4" pageId="48" pageNumber="49">3</td>
</tr>
<tr id="3552ED70FFC834A31595FCD4FA657DEB" box="[151,1436,864,929]" gridrow="5" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FCD4FE6D7DEB" box="[151,404,864,929]" gridcol="0" gridrow="5" pageId="48" pageNumber="49">Paravertebral tubercles 4th toe lamellae</th>
<td id="7683840CFFC834A31723FCD4FD587DEB" box="[545,673,864,929]" gridcol="1" gridrow="5" pageId="48" pageNumber="49">26 34</td>
<td id="7683840CFFC834A317CCFCD4FCB37DEB" box="[718,842,864,929]" gridcol="2" gridrow="5" pageId="48" pageNumber="49">28 31</td>
<td id="7683840CFFC834A3167EFCD4FC277DEB" box="[892,990,864,929]" gridcol="3" gridrow="5" pageId="48" pageNumber="49">31 33</td>
<td id="7683840CFFC834A31128FCD4FB4E7DEB" box="[1066,1207,864,929]" gridcol="4" gridrow="5" pageId="48" pageNumber="49">27 31</td>
<td id="7683840CFFC834A311F9FCD4FA657DEB" box="[1275,1436,864,929]" gridcol="5" gridrow="5" pageId="48" pageNumber="49">28 33</td>
</tr>
<tr id="3552ED70FFC834A31595FC01FA657D86" box="[151,1436,949,972]" gridrow="6" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FC01FE6D7D86" box="[151,404,949,972]" gridcol="0" gridrow="6" pageId="48" pageNumber="49">TL</th>
<td id="7683840CFFC834A31723FC01FD587D86" box="[545,673,949,972]" gridcol="1" gridrow="6" pageId="48" pageNumber="49">70.0</td>
<td id="7683840CFFC834A317CCFC01FCB37D86" box="[718,842,949,972]" gridcol="2" gridrow="6" pageId="48" pageNumber="49">55.7</td>
<td id="7683840CFFC834A3167EFC01FC277D86" box="[892,990,949,972]" gridcol="3" gridrow="6" pageId="48" pageNumber="49">73.6</td>
<td id="7683840CFFC834A31128FC01FB4E7D86" box="[1066,1207,949,972]" gridcol="4" gridrow="6" pageId="48" pageNumber="49">/</td>
<td id="7683840CFFC834A311F9FC01FA657D86" box="[1275,1436,949,972]" gridcol="5" gridrow="6" pageId="48" pageNumber="49">62.0</td>
</tr>
<tr id="3552ED70FFC834A31595FC54FA657A6B" box="[151,1436,992,1057]" gridrow="7" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FC54FE6D7A6B" box="[151,404,992,1057]" gridcol="0" gridrow="7" pageId="48" pageNumber="49">TW FL</th>
<td id="7683840CFFC834A31723FC54FD587A6B" box="[545,673,992,1057]" gridcol="1" gridrow="7" pageId="48" pageNumber="49">4.6 8.6</td>
<td id="7683840CFFC834A317CCFC54FCB37A6B" box="[718,842,992,1057]" gridcol="2" gridrow="7" pageId="48" pageNumber="49">4.0 7.0</td>
<td id="7683840CFFC834A3167EFC54FC277A6B" box="[892,990,992,1057]" gridcol="3" gridrow="7" pageId="48" pageNumber="49">5.8 11.1</td>
<td id="7683840CFFC834A31128FC54FB4E7A6B" box="[1066,1207,992,1057]" gridcol="4" gridrow="7" pageId="48" pageNumber="49">3.9 7.6</td>
<td id="7683840CFFC834A311F9FC54FA657A6B" box="[1275,1436,992,1057]" gridcol="5" gridrow="7" pageId="48" pageNumber="49">4.3 7.9</td>
</tr>
<tr id="3552ED70FFC834A31595FB81FA657A06" box="[151,1436,1077,1100]" gridrow="8" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FB81FE6D7A06" box="[151,404,1077,1100]" gridcol="0" gridrow="8" pageId="48" pageNumber="49">TBL</th>
<td id="7683840CFFC834A31723FB81FD587A06" box="[545,673,1077,1100]" gridcol="1" gridrow="8" pageId="48" pageNumber="49">9.8</td>
<td id="7683840CFFC834A317CCFB81FCB37A06" box="[718,842,1077,1100]" gridcol="2" gridrow="8" pageId="48" pageNumber="49">8.8</td>
<td id="7683840CFFC834A3167EFB81FC277A06" box="[892,990,1077,1100]" gridcol="3" gridrow="8" pageId="48" pageNumber="49">12.5</td>
<td id="7683840CFFC834A31128FB81FB4E7A06" box="[1066,1207,1077,1100]" gridcol="4" gridrow="8" pageId="48" pageNumber="49">9.3</td>
<td id="7683840CFFC834A311F9FB81FA657A06" box="[1275,1436,1077,1100]" gridcol="5" gridrow="8" pageId="48" pageNumber="49">9.3</td>
</tr>
<tr id="3552ED70FFC834A31595FBD4FA657AEB" box="[151,1436,1120,1185]" gridrow="9" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FBD4FE6D7AEB" box="[151,404,1120,1185]" gridcol="0" gridrow="9" pageId="48" pageNumber="49">AG HL</th>
<td id="7683840CFFC834A31723FBD4FD587AEB" box="[545,673,1120,1185]" gridcol="1" gridrow="9" pageId="48" pageNumber="49">21.1 12.4</td>
<td id="7683840CFFC834A317CCFBD4FCB37AEB" box="[718,842,1120,1185]" gridcol="2" gridrow="9" pageId="48" pageNumber="49">18.8 10.4</td>
<td id="7683840CFFC834A3167EFBD4FC277AEB" box="[892,990,1120,1185]" gridcol="3" gridrow="9" pageId="48" pageNumber="49">25.7 14.4</td>
<td id="7683840CFFC834A31128FBD4FB4E7AEB" box="[1066,1207,1120,1185]" gridcol="4" gridrow="9" pageId="48" pageNumber="49">17.5 10.7</td>
<td id="7683840CFFC834A311F9FBD4FA657AEB" box="[1275,1436,1120,1185]" gridcol="5" gridrow="9" pageId="48" pageNumber="49">18.8 10.9</td>
</tr>
<tr id="3552ED70FFC834A31595FB01FA657A86" box="[151,1436,1205,1228]" gridrow="10" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FB01FE6D7A86" box="[151,404,1205,1228]" gridcol="0" gridrow="10" pageId="48" pageNumber="49">HW</th>
<td id="7683840CFFC834A31723FB01FD587A86" box="[545,673,1205,1228]" gridcol="1" gridrow="10" pageId="48" pageNumber="49">7.0</td>
<td id="7683840CFFC834A317CCFB01FCB37A86" box="[718,842,1205,1228]" gridcol="2" gridrow="10" pageId="48" pageNumber="49">6.1</td>
<td id="7683840CFFC834A3167EFB01FC277A86" box="[892,990,1205,1228]" gridcol="3" gridrow="10" pageId="48" pageNumber="49">8.9</td>
<td id="7683840CFFC834A31128FB01FB4E7A86" box="[1066,1207,1205,1228]" gridcol="4" gridrow="10" pageId="48" pageNumber="49">6.4</td>
<td id="7683840CFFC834A311F9FB01FA657A86" box="[1275,1436,1205,1228]" gridcol="5" gridrow="10" pageId="48" pageNumber="49">6.3</td>
</tr>
<tr id="3552ED70FFC834A31595FB54FA657B6B" box="[151,1436,1248,1313]" gridrow="11" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FB54FE6D7B6B" box="[151,404,1248,1313]" gridcol="0" gridrow="11" pageId="48" pageNumber="49">HD ED</th>
<td id="7683840CFFC834A31723FB54FD587B6B" box="[545,673,1248,1313]" gridcol="1" gridrow="11" pageId="48" pageNumber="49">5.0 2.6</td>
<td id="7683840CFFC834A317CCFB54FCB37B6B" box="[718,842,1248,1313]" gridcol="2" gridrow="11" pageId="48" pageNumber="49">4.0 2.5</td>
<td id="7683840CFFC834A3167EFB54FC277B6B" box="[892,990,1248,1313]" gridcol="3" gridrow="11" pageId="48" pageNumber="49">6.4 3.1</td>
<td id="7683840CFFC834A31128FB54FB4E7B6B" box="[1066,1207,1248,1313]" gridcol="4" gridrow="11" pageId="48" pageNumber="49">4.4 2.5</td>
<td id="7683840CFFC834A311F9FB54FA657B6B" box="[1275,1436,1248,1313]" gridcol="5" gridrow="11" pageId="48" pageNumber="49">4.0 2.3</td>
</tr>
<tr id="3552ED70FFC834A31595FA81FA657B06" box="[151,1436,1333,1356]" gridrow="12" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FA81FE6D7B06" box="[151,404,1333,1356]" gridcol="0" gridrow="12" pageId="48" pageNumber="49">EE</th>
<td id="7683840CFFC834A31723FA81FD587B06" box="[545,673,1333,1356]" gridcol="1" gridrow="12" pageId="48" pageNumber="49">3.3</td>
<td id="7683840CFFC834A317CCFA81FCB37B06" box="[718,842,1333,1356]" gridcol="2" gridrow="12" pageId="48" pageNumber="49">2.5</td>
<td id="7683840CFFC834A3167EFA81FC277B06" box="[892,990,1333,1356]" gridcol="3" gridrow="12" pageId="48" pageNumber="49">4.0</td>
<td id="7683840CFFC834A31128FA81FB4E7B06" box="[1066,1207,1333,1356]" gridcol="4" gridrow="12" pageId="48" pageNumber="49">2.8</td>
<td id="7683840CFFC834A311F9FA81FA657B06" box="[1275,1436,1333,1356]" gridcol="5" gridrow="12" pageId="48" pageNumber="49">2.6</td>
</tr>
<tr id="3552ED70FFC834A31595FAD4FA657BEB" box="[151,1436,1376,1441]" gridrow="13" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FAD4FE6D7BEB" box="[151,404,1376,1441]" gridcol="0" gridrow="13" pageId="48" pageNumber="49">ES EN</th>
<td id="7683840CFFC834A31723FAD4FD587BEB" box="[545,673,1376,1441]" gridcol="1" gridrow="13" pageId="48" pageNumber="49">5.9 4.5</td>
<td id="7683840CFFC834A317CCFAD4FCB37BEB" box="[718,842,1376,1441]" gridcol="2" gridrow="13" pageId="48" pageNumber="49">5.1 4.1</td>
<td id="7683840CFFC834A3167EFAD4FC277BEB" box="[892,990,1376,1441]" gridcol="3" gridrow="13" pageId="48" pageNumber="49">7.4 5.7</td>
<td id="7683840CFFC834A31128FAD4FB4E7BEB" box="[1066,1207,1376,1441]" gridcol="4" gridrow="13" pageId="48" pageNumber="49">5.2 3.8</td>
<td id="7683840CFFC834A311F9FAD4FA657BEB" box="[1275,1436,1376,1441]" gridcol="5" gridrow="13" pageId="48" pageNumber="49">5.4 4.9</td>
</tr>
<tr id="3552ED70FFC834A31595FA01FA657B86" box="[151,1436,1461,1484]" gridrow="14" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FA01FE6D7B86" box="[151,404,1461,1484]" gridcol="0" gridrow="14" pageId="48" pageNumber="49">IO</th>
<td id="7683840CFFC834A31723FA01FD587B86" box="[545,673,1461,1484]" gridcol="1" gridrow="14" pageId="48" pageNumber="49">2.5</td>
<td id="7683840CFFC834A317CCFA01FCB37B86" box="[718,842,1461,1484]" gridcol="2" gridrow="14" pageId="48" pageNumber="49">1.9</td>
<td id="7683840CFFC834A3167EFA01FC277B86" box="[892,990,1461,1484]" gridcol="3" gridrow="14" pageId="48" pageNumber="49">3.2</td>
<td id="7683840CFFC834A31128FA01FB4E7B86" box="[1066,1207,1461,1484]" gridcol="4" gridrow="14" pageId="48" pageNumber="49">2.4</td>
<td id="7683840CFFC834A311F9FA01FA657B86" box="[1275,1436,1461,1484]" gridcol="5" gridrow="14" pageId="48" pageNumber="49">1.8</td>
</tr>
<tr id="3552ED70FFC834A31595FA54FA65786B" box="[151,1436,1504,1569]" gridrow="15" pageId="48" pageNumber="49">
<th id="7683840CFFC834A31595FA54FE6D786B" box="[151,404,1504,1569]" gridcol="0" gridrow="15" pageId="48" pageNumber="49">EL IN</th>
<td id="7683840CFFC834A31723FA54FD58786B" box="[545,673,1504,1569]" gridcol="1" gridrow="15" pageId="48" pageNumber="49">2.2 1.3</td>
<td id="7683840CFFC834A317CCFA54FCB3786B" box="[718,842,1504,1569]" gridcol="2" gridrow="15" pageId="48" pageNumber="49">0.8 1.0</td>
<td id="7683840CFFC834A3167EFA54FC27786B" box="[892,990,1504,1569]" gridcol="3" gridrow="15" pageId="48" pageNumber="49">1.5 1.7</td>
<td id="7683840CFFC834A31128FA54FB4E786B" box="[1066,1207,1504,1569]" gridcol="4" gridrow="15" pageId="48" pageNumber="49">0.8 1.1</td>
<td id="7683840CFFC834A311F9FA54FA65786B" box="[1275,1436,1504,1569]" gridcol="5" gridrow="15" pageId="48" pageNumber="49">0.8 /</td>
</tr>
</table>
</paragraph>
<subSubSection id="C378BCB9FFC8CB6C15C4F9D3FE46795D" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BDDEF32FFC8CB6C15C4F9D3FE1E78ED" blockId="48.[151,1437,1639,2002]" pageId="48" pageNumber="49">
<emphasis id="B9163320FFC8CB6C15C4F9D3FE9E78CB" bold="true" box="[198,359,1639,1665]" pageId="48" pageNumber="49">Distribution.</emphasis>
<taxonomicName id="4C6294B1FFC8CB6C146FF9DCFD7078CB" box="[365,649,1640,1665]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFC8CB6C146FF9DCFD7078CB" box="[365,649,1640,1665]" italics="true" pageId="48" pageNumber="49">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC8CB6C1792F9D3FD1478CB" box="[656,749,1639,1665]" pageId="48" pageNumber="49" rank="species">
<emphasis id="B9163320FFC8CB6C1792F9D3FD1478CB" bold="true" box="[656,749,1639,1665]" pageId="48" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the border regions of Trang and Satun Provinces,
<collectingCountry id="F375AFA2FFC8CB6C141BF939FE7878ED" box="[281,385,1677,1703]" name="Thailand" pageId="48" pageNumber="49">Thailand</collectingCountry>
(
<figureCitation id="1359F3B7FFC8CB6C1493F939FE2F78ED" box="[401,470,1677,1703]" captionStart="FIGURE 1" captionStartId="4.[151,255,2000,2024]" captionTargetBox="[175,1419,194,1978]" captionTargetId="figure@4.[175,1419,194,1978]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Cnemaspis in Thailand. Squares represent type localities." httpUri="https://zenodo.org/record/275936/files/figure.png" pageId="48" pageNumber="49">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BDDEF32FFC8CB6C15C4F906FE46795D" blockId="48.[151,1437,1639,2002]" pageId="48" pageNumber="49">
<emphasis id="B9163320FFC8CB6C15C4F906FE687886" bold="true" box="[198,401,1714,1740]" pageId="48" pageNumber="49">Natural history.</emphasis>
<taxonomicName id="4C6294B1FFC8CB6C149FF907FD447886" box="[413,701,1715,1740]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFC8CB6C149FF907FD447886" box="[413,701,1715,1740]" italics="true" pageId="48" pageNumber="49">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC8CB6C17CBF906FCD27886" box="[713,811,1714,1740]" pageId="48" pageNumber="49" rank="species">
<emphasis id="B9163320FFC8CB6C17CBF906FCD27886" bold="true" box="[713,811,1714,1740]" pageId="48" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
is a karst, microhabitat specialist that is abroad at night. During the day, lizards retreat into crevices and caves. All slizards were taken from karst regions in the vicinity of small streams.
</paragraph>
</subSubSection>
<subSubSection id="C378BCB9FFC8CB6D15C4F896FC047FFD" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8BDDEF32FFC8CB6C15C4F896FDBD79CD" blockId="48.[151,1437,1639,2002]" pageId="48" pageNumber="49">
<emphasis id="B9163320FFC8CB6C15C4F896FEAA7976" bold="true" box="[198,339,1826,1852]" pageId="48" pageNumber="49">Etymology.</emphasis>
This species is named in honor of Ms. Piyawan Niyomwan,
<collectingCountry id="F375AFA2FFC8CB6C1152F896FB7F7976" box="[1104,1158,1826,1852]" name="Thailand" pageId="48" pageNumber="49">Thai</collectingCountry>
herpetologist who has worked for many years making significant contributions to our knowledge of the distribution of the amphibians and reptiles of
<collectingCountry id="F375AFA2FFC8CB6C14D6F8D9FDB979CD" box="[468,576,1901,1927]" name="Thailand" pageId="48" pageNumber="49">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8BDDEF32FFC8CB6D15C4F826FC047FFD" blockId="48.[151,1437,1639,2002]" lastBlockId="49.[151,1437,152,439]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="B9163320FFC8CB6C15C4F826FE8A79E6" bold="true" box="[198,371,1938,1964]" pageId="48" pageNumber="49">Comparisons.</emphasis>
<taxonomicName id="4C6294B1FFC8CB6C1481F827FD5E79E6" box="[387,679,1939,1964]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="niyomwanae" status="sp. nov.">
<emphasis id="B9163320FFC8CB6C1481F827FD5E79E6" box="[387,679,1939,1964]" italics="true" pageId="48" pageNumber="49">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC8CB6C17B5F826FCE579E6" box="[695,796,1938,1964]" pageId="48" pageNumber="49" rank="species">
<emphasis id="B9163320FFC8CB6C17B5F826FCE579E6" bold="true" box="[695,796,1938,1964]" pageId="48" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
can be diagnosed from all other Southeast Asian
<taxonomicName id="4C6294B1FFC8CB6C1595F80CFEE0799B" box="[151,281,1976,2001]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B9163320FFC8CB6C1595F80CFEE0799B" box="[151,281,1976,2001]" italics="true" pageId="48" pageNumber="49">Cnemaspis</emphasis>
</taxonomicName>
in having males bearing alternating red-orange and yellow bands on the forelimbs and enlarged, white, isolated tubercles on the sides of the neck, shoulder and flanks. Its lack of white ocelli in a black shoulder or neck patch separates it from
<emphasis id="B9163320FFC9CB6D1770FF09FE9E7EB1" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C6294B1FFC9CB6D1770FF09FD187E9C" box="[626,737,189,214]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="affinis">C. affinis</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D17EEFF09FC7F7E9C" box="[748,902,189,214]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="biocellata">C. biocellata</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1690FF09FBEC7E9C" box="[914,1045,189,214]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="kumpoli">C. kumpoli</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1123FF09FB547E9C" box="[1057,1197,189,214]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="mcguirei">C. mcguirei</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D11BAFF09FA617E9C" box="[1208,1432,189,214]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="pseudomcguirei">C. pseudomcguirei</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1595FF56FE9E7EB1" box="[151,359,226,251]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="puntatonucalis" status="sp. nov.">C. puntatonucalis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2258E5BFFC9CB6D146DFF56FE347EB6" box="[367,461,226,252]" pageId="49" pageNumber="50" rank="species">
<emphasis id="B9163320FFC9CB6D146DFF56FE347EB6" bold="true" box="[367,461,226,252]" pageId="49" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
Its presence of precloacal pores in males further separates it from all other species except
<emphasis id="B9163320FFC9CB6D15ECFEBCFBC27F21" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C6294B1FFC9CB6D15ECFEBCFE987F6B" box="[238,353,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="affinis">C. affinis</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D146CFEBCFE207F6B" box="[366,473,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="argus">C. argus</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D14E4FEBCFD7D7F6B" box="[486,644,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="bayuensis">C. bayuensis</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1790FEBCFCD67F6B" box="[658,815,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="biocellata">C. biocellata</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D163DFEBCFC087F6B" box="[831,1009,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="caudanivea">C. caudanivea</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D16FCFEBCFB0C7F6B" box="[1022,1269,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="chanthaburiensis">C. chanthaburiensis</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1000FEBCFA8A7F6B" box="[1282,1395,264,289]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="dringi">C. dringi</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1082FEBCFEDB7F0C" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="flavigaster">C. flavigaster</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D142DFE99FE0D7F0C" box="[303,500,301,326]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="flavolineata">C. flavolineata</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1703FE99FD557F0C" box="[513,684,301,326]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="karsticola">C. karsticola</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D17B8FE99FCB27F0C" box="[698,843,301,326]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="kumpoli">C. kumpoli</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D165BFE99FC0A7F0C" box="[857,1011,301,326]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="mcguirei">C. mcguirei</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1103FE99FB2B7F0C" box="[1025,1234,301,326]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="monachorum">C. monachorum</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D11E2FE99FA887F0C" box="[1248,1393,301,326]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="nigridia">C. nigridia</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D1082FE99FED47F21" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="nuicamensis">C. nuicamensis</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D143BFEE6FE3A7F21" box="[313,451,338,363]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="paripari">C. paripari</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D14D2FEE6FD4B7F21" box="[464,690,338,363]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="perhentianensis">C. perhentianensis</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D17BCFEE6FC587F21" box="[702,929,338,363]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="pseudomcguirei">C. pseudomcguirei</taxonomicName>
,
<taxonomicName id="4C6294B1FFC9CB6D16ACFEE6FBC27F21" box="[942,1083,338,363]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="chanardi" status="sp. nov.">C. chanardi</taxonomicName>
</emphasis>
<emphasis id="B9163320FFC9CB6D1146FEE6FB5A7F26" bold="true" box="[1092,1187,338,364]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A2258E5BFFC9CB6D1146FEE6FB5A7F26" box="[1092,1187,338,364]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C6294B1FFC9CB6D11B6FEE6FA897F21" box="[1204,1392,338,363]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="vandeventeri" status="sp. nov.">
<emphasis id="B9163320FFC9CB6D11B6FEE6FA897F21" box="[1204,1392,338,363]" italics="true" pageId="49" pageNumber="50">C. vandeventeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC9CB6D107AFEE6FF327FDB" pageId="49" pageNumber="50" rank="species">
<emphasis id="B9163320FFC9CB6D107AFEE6FF327FDB" bold="true" pageId="49" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C6294B1FFC9CB6D15DEFECCFE347FDB" box="[220,461,376,401]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="kamolnorranathi" status="sp. nov.">
<emphasis id="B9163320FFC9CB6D15DEFECCFE347FDB" box="[220,461,376,401]" italics="true" pageId="49" pageNumber="50">C. kamolnorranathi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC9CB6D14D4FEC3FDC17FDB" box="[470,568,375,401]" pageId="49" pageNumber="50" rank="species">
<emphasis id="B9163320FFC9CB6D14D4FEC3FDC17FDB" bold="true" box="[470,568,375,401]" pageId="49" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C6294B1FFC9CB6D174AFECCFCD67FDB" box="[584,815,376,401]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="narathiwatensis" status="sp. nov.">
<emphasis id="B9163320FFC9CB6D174AFECCFCD67FDB" box="[584,815,376,401]" italics="true" pageId="49" pageNumber="50">C. narathiwatensis</emphasis>
</taxonomicName>
<emphasis id="B9163320FFC9CB6D1635FEC3FC607FDB" bold="true" box="[823,921,375,401]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A2258E5BFFC9CB6D1635FEC3FC607FDB" box="[823,921,375,401]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C6294B1FFC9CB6D16A8FECCFBAA7FDB" box="[938,1107,376,401]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="huaseesom" status="sp. nov.">
<emphasis id="B9163320FFC9CB6D16A8FECCFBAA7FDB" box="[938,1107,376,401]" italics="true" pageId="49" pageNumber="50">C. huaseesom</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2258E5BFFC9CB6D1159FEC3FB447FDB" box="[1115,1213,375,401]" pageId="49" pageNumber="50" rank="species">
<emphasis id="B9163320FFC9CB6D1159FEC3FB447FDB" bold="true" box="[1115,1213,375,401]" pageId="49" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<emphasis id="B9163320FFC9CB6D1000FECCFA647FDB" box="[1282,1437,376,401]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C6294B1FFC9CB6D1000FECCFA617FDB" box="[1282,1432,376,401]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="roticanai">C. roticanai</taxonomicName>
.
</emphasis>
Other differences in squamation are scored across all species in
<tableCitation id="C6E0DA89FFC9CB6D1686FE29FC017FFD" box="[900,1016,413,439]" captionStart="TABLE 1" captionStartId="9.[151,242,151,175]" captionTargetBox="[151,1436,277,1409]" captionTargetPageId="9" captionText="TABLE 1. Diagnostic characters of Southeast Asian species of Cnemaspis. New species in bold type. / = character not evaluated." httpUri="http://table.plazi.org/id/DF1DBFBAFFF1CB551595FF23FEFD7E85" pageId="49" pageNumber="50" tableUuid="DF1DBFBAFFF1CB551595FF23FEFD7E85">TABLE 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>