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<document id="8BCBB30D244A8DB82C5B71D6C4D51961" ID-DOI="10.37520/aemnp.2021.009" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037330" ID-ZooBank="27EA8466-6B29-40F4-8236-DD49061C523B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624895377019" checkinUser="carolina" docAuthor="Liu, Tian-tian, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2021" docId="03CB023BAC00FF8F4C89F9791502F8D8" docLanguage="en" docName="ActaEntMusNatPra.61.1.163-201.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.009" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Bryothinusa yoshigoui Liu &amp; Ono &amp; Maruyama 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="176" masterDocId="FFF27A43AC0AFF824F48FFFB175FFFCD" masterDocTitle="Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="201" masterPageNumber="163" pageNumber="173" updateTime="1698977316832" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="CF6B02EF846656CBF1E2B2423A8A8E47">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="4A624118AE8D3F91433452C7B47A71BC">Liu, Tian-tian</mods:namePart>
<mods:affiliation id="C14C1DEF6F6E72C43B3FB142D5A2D196">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395, Japan &amp; The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:namePart id="9C37C272B1536FA9ECD7290D744507F8">Ono, Hiroki</mods:namePart>
<mods:affiliation id="1F7D0D5FA99F7979635593BBD4B51ABF">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan</mods:affiliation>
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<mods:namePart id="25610043E59FA7F786CB80E69E27D19B">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="16D325313DF0935AB4CA2A877C9F8577">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<heading id="D0950441AC00FF884C89F9791248F957" box="[961,1303,1666,1690]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="173" reason="3">
<emphasis id="B9166F3FAC00FF884C89F9791248F957" bold="true" box="[961,1303,1666,1690]" pageId="10" pageNumber="173">
<taxonomicName id="4C62C8AEAC00FF884C89F97913E4F957" ID-CoL="8XW4R" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[961,1211,1666,1690]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="173" phylum="Arthropoda" rank="species" species="yoshigoui" status="sp. nov.">
<emphasis id="B9166F3FAC00FF884C89F97913E4F957" bold="true" box="[961,1211,1666,1690]" italics="true" pageId="10" pageNumber="173">Bryothinusa yoshigoui</emphasis>
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<paragraph id="8BDDB32DAC00FF884B5FF959139EF978" blockId="10.[828,1437,1698,1865]" box="[1047,1217,1698,1717]" pageId="10" pageNumber="173">
(
<figureCitation id="1359AFA8AC00FF884B55F959133DF978" box="[1053,1122,1698,1717]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="10" pageNumber="173">Figs 1E</figureCitation>
; 
<figureCitation id="1359AFA8AC00FF884B23F95913D9F978" box="[1131,1158,1698,1717]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="10" pageNumber="173">2E</figureCitation>
; 
<figureCitation id="1359AFA8AC00FF884BC6F95913C4F978" box="[1166,1179,1698,1717]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="10" pageNumber="173">7</figureCitation>
; 
<figureCitation id="1359AFA8AC00FF884BECF95913E5F978" box="[1188,1210,1698,1717]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn &amp; Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="10" pageNumber="173">22</figureCitation>
)
</paragraph>
<paragraph id="8BDDB32DAC00FF884CD3F9471495F884" blockId="10.[828,1437,1698,1865]" pageId="10" pageNumber="173">
(Japanese name: Hiroshima-nagisa-hanekakushi) 
<emphasis id="B9166F3FAC00FF884C74F91D149EF934" bold="true" box="[828,961,1766,1785]" pageId="10" pageNumber="173">Type material.</emphasis>
Hඈඅඈඍඒඉൾ: J, 
<emphasis id="B9166F3FAC00FF884B02F91D13AFF934" bold="true" box="[1098,1264,1766,1785]" pageId="10" pageNumber="173">
<collectingCountry id="F375F3BDAC00FF884B02F91D13CDF934" box="[1098,1170,1766,1785]" name="Japan" pageId="10" pageNumber="173">JAPAN</collectingCountry>
: HඈඇඌΗǕ:
</emphasis>
Mitsuguchi-kaigan, Yasuura-chô, Kure, Hiroshima-ken, 
<date id="FFDC95EDAC00FF884B3EF8FA1386F8D9" box="[1142,1241,1793,1812]" pageId="10" pageNumber="173" value="2019-07-28">28.vii.2019</date>
, H. Yoshigou (KUM). Pൺ*©ൺඍඒඉൾඌ: 
<emphasis id="B9166F3FAC00FF884CD7F8E014ABF8E3" bold="true" box="[927,1012,1819,1838]" pageId="10" pageNumber="173">HඈඇඌΗǕ:</emphasis>
6 JJ 
<specimenCount id="9D6478A4AC00FF884B62F8E01316F8E3" box="[1066,1097,1813,1838]" pageId="10" pageNumber="173" type="female">1 ♀</specimenCount>
, 2 unsexed spec., same data as 
<typeStatus id="54D90D8FAC00FF884A1AF8E012C2F8E3" box="[1362,1437,1819,1838]" pageId="10" pageNumber="173" type="holotype">holotype</typeStatus>
(KUM, NMPC).
</paragraph>
<paragraph id="8BDDB32DAC00FF884C74F89913E9F7D4" blockId="10.[828,1437,1890,2073]" pageId="10" pageNumber="173">
<emphasis id="B9166F3FAC00FF884C74F8991495F8B4" bold="true" box="[828,970,1890,1913]" pageId="10" pageNumber="173">Description.</emphasis>
Body (
<figureCitation id="1359AFA8AC00FF884B69F8991329F8B4" box="[1057,1142,1890,1913]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="10" pageNumber="173">Fig. 1E</figureCitation>
) small about 
<quantity id="4C9A1EC8AC00FF884A5EF89912C4F8B4" box="[1302,1435,1890,1913]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="10" pageNumber="173" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.3–2.4 mm</quantity>
long, covered with minute setae; background colour deep reddish brown, head, pronotum and elytra reddish brown; posteriorly abdominal tergite VI infuscate; antenna and legs similarly reddish-brown coloured, lightly paler than body colour and paler toward apex.
</paragraph>
<caption id="DF1DE3A5AC01FF894FDFF82D16F3F7D3" ID-DOI="http://doi.org/10.5281/zenodo.5037344" ID-Zenodo-Dep="5037344" httpUri="https://zenodo.org/record/5037344/files/figure.png" pageId="11" pageNumber="174" startId="11.[151,182,2006,2025]" targetBox="[153,1406,200,1947]" targetPageId="11">
<paragraph id="8BDDB32DAC01FF894FDFF82D16F3F7D3" blockId="11.[151,1438,2005,2078]" pageId="11" pageNumber="174">
Fig. 6. 
<taxonomicName id="4C62C8AEAC01FF894F9DF82E1570F824" authority="Ahn &amp; Jeon, 2004" authorityName="Ahn &amp; Jeon" authorityYear="2004" box="[213,559,2005,2025]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="174" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="B9166F3FAC01FF894F9DF82E16D7F825" box="[213,392,2005,2024]" italics="true" pageId="11" pageNumber="174">Bryothinusa koreana</emphasis>
<bibRefCitation id="EFF3CEDCAC01FF894EC7F82D1570F824" author="AHN K. J. &amp; JEON M. J." box="[399,559,2006,2025]" pageId="11" pageNumber="174" pagination="29 - 35" refId="ref25135" refString="AHN K. J. &amp; JEON M. J. 2004: The Korean species of the genus Bryothinusa Casey with descriptions of two new species (Coleoptera: Staphylinidae: Aleocharinae). Coleopterists Bulletin 58: 29 - 35." type="journal article" year="2004">Ahn &amp; Jeon, 2004</bibRefCitation>
</taxonomicName>
. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020).
</paragraph>
</caption>
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<paragraph id="8BDDB32DAC06FF8E4FDFF8101267F7D5" blockId="12.[151,1437,2026,2072]" pageId="12" pageNumber="175">
Fig. 7. 
<taxonomicName id="4C62C8AEAC06FF8E4F9BF81116CCF830" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[211,403,2026,2045]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="175" phylum="Arthropoda" rank="species" species="yoshigoui" status="sp. nov.">
<emphasis id="B9166F3FAC06FF8E4F9BF81116CCF830" box="[211,403,2026,2045]" italics="true" pageId="12" pageNumber="175">Bryothinusa yoshigoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC06FF8E4ED0F8101685F833" box="[408,474,2027,2046]" pageId="12" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BDDB32DAC07FF8F4FFDFF451680FE78" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
Head oval, wider than long, longer than pronotum and elytra; eyes (
<figureCitation id="1359AFA8AC07FF8F4E69FF25162EFF38" box="[289,369,222,245]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="13" pageNumber="176">Fig. 2E</figureCitation>
) small, reniform, shorter than 1/2 of head length, with 12–14 facets; antennal length a little shorter than distance from anterior margin of head to posterior margin of elytra; antennal segments all longer than wide; segment I as long as II, 1.2 times longer than III, and shorter than XI; posterior margin of antennal segment III narrower than anterior margin.
</paragraph>
<paragraph id="8BDDB32DAC07FF8F4FFDFE4515A6FCD8" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
Mouthparts (
<figureCitation id="1359AFA8AC07FF8F4E73FE451624FE18" box="[315,379,446,469]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Figs 7</figureCitation>
A-E) with distribution of labral setae as in 
<figureCitation id="1359AFA8AC07FF8F4F9EFE251617FE38" box="[214,328,478,501]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Figure 7A</figureCitation>
; mandibles (
<figureCitation id="1359AFA8AC07FF8F4E9EFE251575FE38" box="[470,554,478,501]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7B</figureCitation>
) elongate triangle, asymmetrical, right with median tooth; median tooth longer than 1/4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (
<figureCitation id="1359AFA8AC07FF8F4D21FDC515E4FD98" box="[617,699,574,597]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7C</figureCitation>
) with second spines very small and not obvious, at tip with two spines not closely interdigitating; 4 setae on inner margin of lacinia; labial palpus (
<figureCitation id="1359AFA8AC07FF8F4ED6FD6516AFFD78" box="[414,496,670,693]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7D</figureCitation>
) with most setae located basally; mentum (
<figureCitation id="1359AFA8AC07FF8F4E05FD4516C6FD18" box="[333,409,702,725]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7E</figureCitation>
) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “v seta” of mentum near the level of anterior margin.
</paragraph>
<paragraph id="8BDDB32DAC07FF8F4FFDFCE5165AFC98" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">Pronotum almost as long as elytra, slightly narrower than head.</paragraph>
<paragraph id="8BDDB32DAC07FF8F4FFDFCA516ADFC58" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">Elytra widened apically, wider than long, almost as wide as pronotum; hind wings absent.</paragraph>
<paragraph id="8BDDB32DAC07FF8F4FFDFC6515A7FBD8" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
Male abdominal sternite VIII (
<figureCitation id="1359AFA8AC07FF8F4D49FC65150EFC78" box="[513,593,926,949]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7F</figureCitation>
) with posterior margin slightly serrate. Median lobe of aedeagus (
<figureCitation id="1359AFA8AC07FF8F4D80FC451790FC38" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Figs 7H, I</figureCitation>
) with some granules on apical lobe, and not clearly visible in dorsal view; paramere elongate, as in 
<figureCitation id="1359AFA8AC07FF8F4DC6FC0515ABFBD8" box="[654,756,1022,1045]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Figure 7J</figureCitation>
.
</paragraph>
<paragraph id="8BDDB32DAC07FF8F4FFDFBE516D4FB58" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
Female abdominal sternite VIII (
<figureCitation id="1359AFA8AC07FF8F4D41FBE51505FBF8" box="[521,602,1054,1077]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7G</figureCitation>
) with posterior margin slightly serrate but smoother than in male. Spermatheca (
<figureCitation id="1359AFA8AC07FF8F4E48FBA51609FBB8" box="[256,342,1118,1141]" captionStart="Fig" captionStartId="12.[151,182,2027,2046]" captionTargetBox="[152,1420,189,1977]" captionTargetId="figure-73@12.[152,1436,189,1977]" captionTargetPageId="12" captionText="Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037346" httpUri="https://zenodo.org/record/5037346/files/figure.png" pageId="13" pageNumber="176">Fig. 7K</figureCitation>
) small, simplified in shape, about half times coiled near base.
</paragraph>
<paragraph id="8BDDB32DAC07FF8F4FDFFB65159FFAD8" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
<emphasis id="B9166F3FAC07FF8F4FDFFB65161AFB78" bold="true" box="[151,325,1182,1205]" pageId="13" pageNumber="176">Measurements.</emphasis>
BL 2.30–2.40, FBL 1.00–1.20, HL 0.38– 0.45, HW 0.43–0.48, PL 0.35–0.38, PW 0.40–0.43, EL 0.35–0.38, EW 0.41–0.43, HTL 0.34–0.39, AL 0.88–0.97, AS-I 0.09–0.10, AS-II 0.09–0.10, AS-III 0.07–0.08.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC07FF8F4FDFFAE516AAF9D8" pageId="13" pageNumber="176" type="diagnosis">
<paragraph id="8BDDB32DAC07FF8F4FDFFAE516AAF9D8" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
<emphasis id="B9166F3FAC07FF8F4FDFFAE516D1FAF8" bold="true" box="[151,398,1310,1333]" pageId="13" pageNumber="176">Differential diagnosis.</emphasis>
In general appearance, the species is closely similar to 
<taxonomicName id="4C62C8AEAC07FF8F4E2EFAC51691FA98" authorityName="" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[358,462,1342,1365]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="tsutsuii">
<emphasis id="B9166F3FAC07FF8F4E2EFAC51691FA98" box="[358,462,1342,1365]" italics="true" pageId="13" pageNumber="176">B. tsutsuii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C62C8AEAC07FF8F4E91FAC51510FA98" authorityName="Sawada" authorityYear="1971" box="[473,591,1341,1365]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="algarum">
<emphasis id="B9166F3FAC07FF8F4E91FAC51510FA98" box="[473,591,1341,1365]" italics="true" pageId="13" pageNumber="176">B. algarum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C62C8AEAC07FF8F4D13FAC51593FA98" authorityName="Ahn &amp; Jeon" authorityYear="2004" box="[603,716,1341,1365]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="B9166F3FAC07FF8F4D13FAC51593FA98" box="[603,716,1341,1365]" italics="true" pageId="13" pageNumber="176">B. koreana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C62C8AEAC07FF8F4FDFFAA5164BFAB8" authorityName="Liu, Ono &amp; Maruyama" authorityYear="2020" box="[151,276,1374,1397]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B9166F3FAC07FF8F4FDFFAA5164BFAB8" box="[151,276,1374,1397]" italics="true" pageId="13" pageNumber="176">B. japonica</emphasis>
</taxonomicName>
, in but easily distinguished from 
<taxonomicName id="4C62C8AEAC07FF8F4DC2FAA515A7FAB8" authorityName="" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[650,760,1374,1397]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="tsutsuii">
<emphasis id="B9166F3FAC07FF8F4DC2FAA515A7FAB8" box="[650,760,1374,1397]" italics="true" pageId="13" pageNumber="176">B. tsutsuii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C62C8AEAC07FF8F4F8BFA851666FA58" authorityName="Sawada" authorityYear="1971" box="[195,313,1405,1429]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="algarum">
<emphasis id="B9166F3FAC07FF8F4F8BFA851666FA58" box="[195,313,1405,1429]" italics="true" pageId="13" pageNumber="176">B. algarum</emphasis>
</taxonomicName>
by the spermathecal shape and the median lobe of the aedeagus with granule-like projections, from 
<taxonomicName id="4C62C8AEAC07FF8F4DA9FA6517B3FA18" authorityName="Ahn &amp; Jeon" authorityYear="2004" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="B9166F3FAC07FF8F4DA9FA6517B3FA18" italics="true" pageId="13" pageNumber="176">B. koreana</emphasis>
</taxonomicName>
at the number of facets and the darker body colour, and from 
<taxonomicName id="4C62C8AEAC07FF8F4E4CFA2516DEFA38" authorityName="Liu, Ono &amp; Maruyama" authorityYear="2020" box="[260,385,1502,1525]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B9166F3FAC07FF8F4E4CFA2516DEFA38" box="[260,385,1502,1525]" italics="true" pageId="13" pageNumber="176">B. japonica</emphasis>
</taxonomicName>
by the apical part of spermatheca without granule-like projections.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC07FF8F4FDFF9E5153AF9F8" box="[151,613,1566,1589]" pageId="13" pageNumber="176" type="distribution">
<paragraph id="8BDDB32DAC07FF8F4FDFF9E5153AF9F8" blockId="13.[151,761,190,1813]" box="[151,613,1566,1589]" pageId="13" pageNumber="176">
<emphasis id="B9166F3FAC07FF8F4FDFF9E51675F9F8" bold="true" box="[151,298,1566,1589]" pageId="13" pageNumber="176">Distribution.</emphasis>
<collectingCountry id="F375F3BDAC07FF8F4E78F9E51632F9F8" box="[304,365,1566,1589]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
(
<emphasis id="B9166F3FAC07FF8F4E34F9E51681F9F8" bold="true" box="[380,478,1566,1589]" pageId="13" pageNumber="176">Honshû:</emphasis>
<collectingRegion id="49A67DCFAC07FF8F4EACF9E51505F9F8" box="[484,602,1566,1589]" country="Japan" name="Hiroshima" pageId="13" pageNumber="176">Hiroshima</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC07FF8F4FDFF9C5164FF958" pageId="13" pageNumber="176" type="etymology">
<paragraph id="8BDDB32DAC07FF8F4FDFF9C5164FF958" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
<emphasis id="B9166F3FAC07FF8F4FDFF9C51647F998" bold="true" box="[151,280,1598,1621]" pageId="13" pageNumber="176">Etymology.</emphasis>
This new species is named after Mr. Hidenori Yoshigou, a famous ichthyologist and the collector of the 
<typeStatus id="54D90D8FAC07FF8F4FDFF985179AF958" box="[151,197,1662,1685]" pageId="13" pageNumber="176">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC07FF8F4FDFF9651502F8D8" pageId="13" pageNumber="176" type="discussion">
<paragraph id="8BDDB32DAC07FF8F4FDFF9651502F8D8" blockId="13.[151,761,190,1813]" pageId="13" pageNumber="176">
<emphasis id="B9166F3FAC07FF8F4FDFF9651659F978" bold="true" box="[151,262,1694,1717]" pageId="13" pageNumber="176">Remarks.</emphasis>
The specimens were collected on the coastline of Setonaikai (inland sea between Honshû and Shikoku), 
<collectingCountry id="F375F3BDAC07FF8F4FDFF9251788F938" box="[151,215,1758,1781]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
. A distribution map of examined specimens of this species in 
<collectingCountry id="F375F3BDAC07FF8F4E4EF905161CF8D8" box="[262,323,1790,1813]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
is illustrated in 
<figureCitation id="1359AFA8AC07FF8F4EA7F9051506F8D8" box="[495,601,1790,1813]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn &amp; Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="13" pageNumber="176">Figure 22</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>