treatments-xml/data/03/C9/88/03C9885D8F21FFD7FF25F9964A1B25E9.xml

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<mods:title id="C66A7EB5B656546ED6FA7A885D99BA96">Taxonomic revision of Drymoluber Amaral, 1930 (Serpentes: Colubridae)</mods:title>
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<mods:namePart id="9723F01C375C8792A8FE7513ACDDB578">Costa, Henrique Caldeira</mods:namePart>
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<treatment id="03C9885D8F21FFD7FF25F9964A1B25E9" ID-DOI="http://doi.org/10.5281/zenodo.6154465" ID-GBIF-Taxon="119580330" ID-Zenodo-Dep="6154465" LSID="urn:lsid:plazi:treatment:03C9885D8F21FFD7FF25F9964A1B25E9" httpUri="http://treatment.plazi.org/id/03C9885D8F21FFD7FF25F9964A1B25E9" lastPageId="32" lastPageNumber="381" pageId="25" pageNumber="374">
<subSubSection id="C37A6AC08F21FFEEFF25F99648BB20AE" pageId="25" pageNumber="374" type="nomenclature">
<paragraph id="8BDF394B8F21FFEEFF25F9964B0F200F" blockId="25.[151,567,1556,1582]" box="[151,567,1556,1582]" pageId="25" pageNumber="374">
<heading id="D0978E278F21FFEEFF25F9964B0F200F" bold="true" box="[151,567,1556,1582]" fontSize="11" level="1" pageId="25" pageNumber="374" reason="1">
<taxonomicName id="4C6042C88F21FFEEFF25F9964B0F200F" authority="Gomes, 1918" authorityName="Gomes" authorityYear="1918" box="[151,567,1556,1582]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F21FFEEFF25F9964B0F200F" bold="true" box="[151,567,1556,1582]" pageId="25" pageNumber="374">
<emphasis id="B914E5598F21FFEEFF25F9964845200F" bold="true" box="[151,381,1556,1582]" italics="true" pageId="25" pageNumber="374">Drymoluber brazili</emphasis>
(Gomes, 1918)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDF394B8F21FFEEFF25F9D848BB20AE" blockId="25.[151,1436,1626,1802]" pageId="25" pageNumber="374">
<taxonomicName id="4C6042C88F21FFEEFF25F9D848D82051" authority="Gomes, 1918" authorityName="Gomes" authorityYear="1918" box="[151,480,1626,1648]" class="Reptilia" family="Colubridae" genus="Drymobius" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F21FFEEFF25F9D848772051" box="[151,335,1626,1648]" italics="true" pageId="25" pageNumber="374">Drymobius brazili</emphasis>
Gomes, 1918
</taxonomicName>
. Memórias do Instituto Butantan, 1, p. 81. 
<typeStatus id="54DB87E98F21FFEEFC22F9D84ACA2051" box="[912,1010,1626,1648]" pageId="25" pageNumber="374" type="holotype">Holotype</typeStatus>
: IBSP 696 (probably destroyed during the fire of 
<date id="FFDE1F8B8F21FFEEFEB9F9F8484F20AE" box="[267,375,1657,1679]" pageId="25" pageNumber="374" value="2010-05">May 2010</date>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A6AC08F21FFEEFF25F91A4BC621FE" pageId="25" pageNumber="374" type="description">
<paragraph id="8BDF394B8F21FFEEFF25F91A4BFA20ED" blockId="25.[151,1436,1626,1802]" pageId="25" pageNumber="374">
<taxonomicName id="4C6042C88F21FFEEFF25F91A4B2C208F" authority="Amaral, 1923" authorityName="Amaral" authorityYear="1923" box="[151,532,1688,1710]" class="Reptilia" family="Colubridae" genus="Drymobius" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B914E5598F21FFEEFF25F91A484C208F" box="[151,372,1688,1710]" italics="true" pageId="25" pageNumber="374">Drymobius rubriceps</emphasis>
Amaral, 1923
</taxonomicName>
. Proceedings of the New 
<collectingCountry id="F37779DB8F21FFEEFC83F91A4ABD208F" box="[817,901,1688,1710]" name="United Kingdom" pageId="25" pageNumber="374">England</collectingCountry>
Zoological Club, 8, p. 85. 
<typeStatus id="54DB87E98F21FFEEFB05F91A4C21208F" box="[1207,1305,1688,1710]" pageId="25" pageNumber="374" type="holotype">Holotype</typeStatus>
: IBSP 1844 (probably destroyed during the fire of 
<date id="FFDE1F8B8F21FFEEFDF9F9354B8E20ED" box="[587,694,1718,1740]" pageId="25" pageNumber="374" value="2010-05">May 2010</date>
).
</paragraph>
<paragraph id="8BDF394B8F21FFEEFF25F9574D2320CA" blockId="25.[151,1436,1626,1802]" box="[151,1051,1749,1771]" pageId="25" pageNumber="374">
<taxonomicName id="4C6042C88F21FFEEFF25F957484820CA" box="[151,368,1749,1771]" class="Reptilia" family="Colubridae" genus="Drymobius" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="boddaerti">
<emphasis id="B914E5598F21FFEEFF25F957484820CA" box="[151,368,1749,1771]" italics="true" pageId="25" pageNumber="374">Drymobius boddaerti</emphasis>
</taxonomicName>
(partim.)—Amaral 1929. Memórias do Instituto Butantan, 4, p. 11.
</paragraph>
<paragraph id="8BDF394B8F21FFEEFF25F9764C10212B" blockId="25.[151,1436,1626,1802]" box="[151,1320,1780,1802]" pageId="25" pageNumber="374">
<emphasis id="B914E5598F21FFEEFF25F9764849212B" box="[151,369,1780,1802]" italics="true" pageId="25" pageNumber="374">
<taxonomicName id="4C6042C88F21FFEEFF25F976482B212B" box="[151,275,1780,1802]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="genus">Drymoluber</taxonomicName>
brazili—
</emphasis>
Stuart 1932. Occasional Papers of the Museum of Zoology, University of Michigan, 236, p. 4.
</paragraph>
<paragraph id="8BDF394B8F21FFEEFF25F8B44BC621FE" blockId="25.[151,1436,1846,2015]" pageId="25" pageNumber="374">
The 
<typeStatus id="54DB87E98F21FFEEFF79F8B5483E216E" box="[203,262,1847,1871]" pageId="25" pageNumber="374">types</typeStatus>
of 
<taxonomicName id="4C6042C88F21FFEEFE9CF8B44B30216E" box="[302,520,1846,1871]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F21FFEEFE9CF8B44B30216E" box="[302,520,1846,1871]" italics="true" pageId="25" pageNumber="374">Drymoluber brazili</emphasis>
</taxonomicName>
and its junior synonym, 
<taxonomicName id="4C6042C88F21FFEEFC97F8B44D2D216E" box="[805,1045,1846,1871]" class="Reptilia" family="Colubridae" genus="Drymobius" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B914E5598F21FFEEFC97F8B44D2D216E" box="[805,1045,1846,1871]" italics="true" pageId="25" pageNumber="374">Drymobius rubriceps</emphasis>
</taxonomicName>
, were personally examined by the senior author in 2009. On 
<date id="FFDE1F8B8F21FFEEFE74F8D94B052152" box="[454,573,1883,1907]" pageId="25" pageNumber="374" value="2010-05">May 2010</date>
, the collections of 
<emphasis id="B914E5598F21FFEEFCA5F8DE4ADB2152" box="[791,995,1884,1907]" italics="true" pageId="25" pageNumber="374">Instituto Butantan</emphasis>
in São Paulo, 
<collectingCountry id="F37779DB8F21FFEEFB3CF8D84DED2152" box="[1166,1237,1882,1907]" name="Brazil" pageId="25" pageNumber="374">Brazil</collectingCountry>
, where those and most known specimens of 
<taxonomicName id="4C6042C88F21FFEEFE0DF8FD4B1221B6" box="[447,554,1918,1943]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F21FFEEFE0DF8FD4B1221B6" box="[447,554,1918,1943]" italics="true" pageId="25" pageNumber="374">D. brazili</emphasis>
</taxonomicName>
were maintained were consumed in a fire (Warrell 
<emphasis id="B914E5598F21FFEEFBD6F8FD4DA421B6" box="[1124,1180,1918,1943]" italics="true" pageId="25" pageNumber="374">et al.</emphasis>
2010; Franco 2012). It is estimated that 80% of the snake collection was destroyed, but there is still no information if the 
<typeStatus id="54DB87E98F21FFEEFA8CF8214C41219A" box="[1342,1401,1955,1979]" pageId="25" pageNumber="374">types</typeStatus>
of 
<taxonomicName id="4C6042C88F21FFEEFF25F844484821FE" box="[151,368,1990,2015]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="374" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F21FFEEFF25F844484821FE" box="[151,368,1990,2015]" italics="true" pageId="25" pageNumber="374">Drymoluber brazili</emphasis>
</taxonomicName>
were lost (F. Franco, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C37A6AC08F22FFEDFF75FF1548612441" pageId="26" pageNumber="375" type="materials_examined">
<paragraph id="8BDF394B8F22FFEDFF75FF154DC526D9" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
<emphasis id="B914E5598F22FFEDFF75FF1548052691" bold="true" box="[199,317,151,176]" pageId="26" pageNumber="375">
<typeStatus id="54DB87E98F22FFEDFF75FF1548002691" box="[199,312,151,176]" pageId="26" pageNumber="375" type="holotype">Holotype</typeStatus>
:
</emphasis>
<emphasis id="B914E5598F22FFEDFEF5FF1B4B2D2691" box="[327,533,153,176]" italics="true" pageId="26" pageNumber="375">Instituto Butantan</emphasis>
, São Paulo, IBSP 696, adult male, SVL 
<quantity id="4C9894AE8F22FFEDFC47FF1A4D5D268E" box="[1013,1125,152,176]" metricMagnitude="0" metricUnit="m" metricValue="1.09" pageId="26" pageNumber="375" unit="mm" value="1090.0">1090 mm</quantity>
, TL 
<quantity id="4C9894AE8F22FFEDFB12FF1A4C3A268E" box="[1184,1282,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="4.7299999999999995" pageId="26" pageNumber="375" unit="mm" value="473.0">473 mm</quantity>
, collected on 
<date id="FFDE1F8B8F22FFEDFF25FF3E486826F5" box="[151,336,188,213]" pageId="26" pageNumber="375" value="1914-09">
September 
<quantity id="4C9894AE8F22FFEDFEAAFF3E486826F5" box="[280,336,188,212]" metricMagnitude="1" metricUnit="m" metricValue="4.86156" pageId="26" pageNumber="375" unit="in" value="1914.0">1914</quantity>
</date>
in the 
<emphasis id="B914E5598F22FFEDFE14FF3F4AAA26F4" box="[422,914,188,213]" italics="true" pageId="26" pageNumber="375">Estação Ferroviária de Engenheiro Lisboa</emphasis>
(currently inactive), near the municipality of Uberaba (-19.80, -47.60), Minas Gerais, 
<collectingCountry id="F37779DB8F22FFEDFDD2FF5D4B9026D9" box="[608,680,223,248]" name="Brazil" pageId="26" pageNumber="375">Brazil</collectingCountry>
. Specimen personally examined by the senior author.
</paragraph>
<paragraph id="8BDF394B8F22FFEDFF75FE8648612441" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
<emphasis id="B914E5598F22FFEDFF75FE864870273C" bold="true" box="[199,328,260,285]" pageId="26" pageNumber="375">
<typeStatus id="54DB87E98F22FFEDFF75FE86487B273C" box="[199,323,260,285]" pageId="26" pageNumber="375" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<emphasis id="B914E5598F22FFEDFEE1FE874B1A273D" box="[339,546,261,284]" italics="true" pageId="26" pageNumber="375">Instituto Butantan</emphasis>
, São Paulo, IBSP 383, adult male, SVL 
<quantity id="4C9894AE8F22FFEDFBB4FE864D50273D" box="[1030,1128,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="8.629999999999999" pageId="26" pageNumber="375" unit="mm" value="863.0">863 mm</quantity>
, TL 
<quantity id="4C9894AE8F22FFEDFB16FE864C3F273D" box="[1188,1287,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="3.94" pageId="26" pageNumber="375" unit="mm" value="394.0">394 mm</quantity>
, collected in 
<date id="FFDE1F8B8F22FFEDFF25FEA548062761" box="[151,318,295,320]" pageId="26" pageNumber="375" value="1913-02">February 1913</date>
, in the 
<emphasis id="B914E5598F22FFEDFE22FEAB4A182761" box="[400,800,295,320]" italics="true" pageId="26" pageNumber="375">Estação Ferroviária Santa Eudóxia</emphasis>
(currently inactive), municipality of São Carlos (-21.77, -47.78), São Paulo, 
<collectingCountry id="F37779DB8F22FFEDFEC5FECE48872744" box="[375,447,332,357]" name="Brazil" pageId="26" pageNumber="375">Brazil</collectingCountry>
; IBSP 573, adult female, SVL 
<quantity id="4C9894AE8F22FFEDFCACFECE4A452745" box="[798,893,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="8.54" pageId="26" pageNumber="375" unit="mm" value="854.0">854 mm</quantity>
, TL 
<quantity id="4C9894AE8F22FFEDFC01FECE4D2B2745" box="[947,1043,332,356]" metricMagnitude="-1" metricUnit="m" metricValue="4.22" pageId="26" pageNumber="375" unit="mm" value="422.0">422 mm</quantity>
, collected in 
<date id="FFDE1F8B8F22FFEDFB1BFECE4C682745" box="[1193,1360,332,357]" pageId="26" pageNumber="375" value="1914-02">February 1914</date>
, in the 
<emphasis id="B914E5598F22FFEDFF25FEF34B7527A9" box="[151,589,367,392]" italics="true" pageId="26" pageNumber="375">Estação Ferroviária de Sampaio Vidal</emphasis>
(currently inactive), municipality of Ribeirão Bonito (-22.07, -48.18), São Paulo, 
<collectingCountry id="F37779DB8F22FFEDFF57FE164815278C" box="[229,301,404,429]" name="Brazil" pageId="26" pageNumber="375">Brazil</collectingCountry>
; IBSP 574, adult male, SVL 
<quantity id="4C9894AE8F22FFEDFDC5FE164BEE278D" box="[631,726,404,429]" metricMagnitude="-1" metricUnit="m" metricValue="8.62" pageId="26" pageNumber="375" unit="mm" value="862.0">862 mm</quantity>
, TL 268+N mm (broken tail), without locality data; IBSP 741, adult male, SVL 
<quantity id="4C9894AE8F22FFEDFEEBFE3A48FE27EE" box="[345,454,440,464]" metricMagnitude="0" metricUnit="m" metricValue="1.1199999999999999" pageId="26" pageNumber="375" unit="mm" value="1120.0">1120 mm</quantity>
, TL 162+N mm (broken tail), collected in 
<date id="FFDE1F8B8F22FFEDFC1EFE354D5A27F1" box="[940,1122,439,464]" pageId="26" pageNumber="375" value="1914-12">December 1914</date>
, in the 
<emphasis id="B914E5598F22FFEDFB04FE3B49F427D5" italics="true" pageId="26" pageNumber="375">
Estação Ferroviária 
<taxonomicName id="4C6042C88F22FFEDFF25FE5F49F427D5" box="[151,204,477,500]" class="Insecta" family="Pompilidae" genus="Java" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="375" phylum="Arthropoda" rank="genus">Java</taxonomicName>
</emphasis>
(currently inactive), municipality of Boa Esperança do Sul (-21.99, -48.39), São Paulo, 
<collectingCountry id="F37779DB8F22FFEDFB71FE5E4C3327D4" box="[1219,1291,476,501]" name="Brazil" pageId="26" pageNumber="375">Brazil</collectingCountry>
; IBSP 1286, adult female, SVL 
<quantity id="4C9894AE8F22FFEDFEDEFD8248F32436" box="[364,459,512,536]" metricMagnitude="-1" metricUnit="m" metricValue="8.98" pageId="26" pageNumber="375" unit="mm" value="898.0">898 mm</quantity>
, TL 
<quantity id="4C9894AE8F22FFEDFDB2FD824B672436" box="[512,607,512,536]" metricMagnitude="-1" metricUnit="m" metricValue="4.02" pageId="26" pageNumber="375" unit="mm" value="402.0">402 mm</quantity>
, collected in 
<date id="FFDE1F8B8F22FFEDFD41FD824A502439" box="[755,872,512,536]" pageId="26" pageNumber="375" value="1917-05">May 1917</date>
, in the 
<emphasis id="B914E5598F22FFEDFC0BFD834C1E2439" box="[953,1318,511,536]" italics="true" pageId="26" pageNumber="375">Estação Ferroviária Pedregulho</emphasis>
(currently inactive), municipality of Pedregulho (-20.26, -47.48), São Paulo, 
<collectingCountry id="F37779DB8F22FFEDFC3CFDA64AEE241C" box="[910,982,548,573]" name="Brazil" pageId="26" pageNumber="375">Brazil</collectingCountry>
. All specimens personally examined by the senior author.
</paragraph>
</subSubSection>
<subSubSection id="C37A6AC08F22FFEDFF75FDEE48C32279" pageId="26" pageNumber="375" type="description">
<paragraph id="8BDF394B8F22FFEDFF75FDEE4D182585" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
<emphasis id="B914E5598F22FFEDFF75FDEE48CA24A4" bold="true" box="[199,498,620,645]" pageId="26" pageNumber="375">
About the 
<typeStatus id="54DB87E98F22FFEDFEE3FDEE48BD24A4" box="[337,389,620,645]" pageId="26" pageNumber="375">type</typeStatus>
locality:
</emphasis>
In the 20th century the 
<emphasis id="B914E5598F22FFEDFC93FDEF4ACB24A5" box="[801,1011,621,644]" italics="true" pageId="26" pageNumber="375">Instituto Butantan</emphasis>
often received snakes from several Brazilian localities, sent by collectors to the institute via the railroads that crossed some regions of the country at that time. Thus, the accuracy of localities of specimens from railroads (like the 
<typeStatus id="54DB87E98F22FFEDFB91FD374D6B24EC" box="[1059,1107,693,717]" pageId="26" pageNumber="375">type</typeStatus>
series of 
<taxonomicName id="4C6042C88F22FFEDFB76FD374C0924EC" box="[1220,1329,692,717]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFB76FD374C0924EC" box="[1220,1329,692,717]" italics="true" pageId="26" pageNumber="375">D. brazili</emphasis>
</taxonomicName>
) must be treated with caution (Pereira 
<emphasis id="B914E5598F22FFEDFE57FD5B4B2724D1" box="[485,543,727,752]" italics="true" pageId="26" pageNumber="375">et al.</emphasis>
2007). In the specific case of 
<taxonomicName id="4C6042C88F22FFEDFC33FD5B4AD624D1" box="[897,1006,727,752]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFC33FD5B4AD624D1" box="[897,1006,727,752]" italics="true" pageId="26" pageNumber="375">D. brazili</emphasis>
</taxonomicName>
, the 
<emphasis id="B914E5598F22FFEDFB98FD5B49DC2534" italics="true" pageId="26" pageNumber="375">Estação Ferroviária Engenheiro Lisboa</emphasis>
(
<typeStatus id="54DB87E98F22FFEDFF40FD7E481C2535" box="[242,292,764,788]" pageId="26" pageNumber="375">type</typeStatus>
locality), was located at km 555 of the 
<emphasis id="B914E5598F22FFEDFD6EFD7E4AB62534" box="[732,910,764,789]" italics="true" pageId="26" pageNumber="375">Tronco-Catalão</emphasis>
railroad, which, at the time of the collection of the 
<typeStatus id="54DB87E98F22FFEDFF72FC9D481A2519" box="[192,290,799,824]" pageId="26" pageNumber="375" type="holotype">holotype</typeStatus>
(
<date id="FFDE1F8B8F22FFEDFE82FC9D48D22519" box="[304,490,799,824]" pageId="26" pageNumber="375" value="1914-09">September 1914</date>
) went from Campinas, São Paulo (-22.09, -47.05) to Ipameri, Goiás (-17.72, -48.15) (Giesbrecht 2009; Cavalcanti 2010). Although it is hard to identify the exact locality where the 
<typeStatus id="54DB87E98F22FFEDFB41FCC64C6D257C" box="[1267,1365,836,861]" pageId="26" pageNumber="375" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C6042C88F22FFEDFA33FCC749D825A1" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFA33FCC749D825A1" italics="true" pageId="26" pageNumber="375">D. brazili</emphasis>
</taxonomicName>
was collected, we believe it really was in the proximities of the 
<emphasis id="B914E5598F22FFEDFC70FCE54D4C25A1" box="[962,1140,871,896]" italics="true" pageId="26" pageNumber="375">Tronco-Catalão</emphasis>
railroad, since the road is located in areas with other confirmed records of the species (see map in 
<figureCitation id="135B25CE8F22FFEDFC72FC0E4D2A2585" box="[960,1042,908,933]" captionStart="FIGURE 12" captionStartId="14.[151,250,1764,1786]" captionTargetBox="[152,1433,625,1738]" captionTargetId="figure@14.[151,1436,621,1742]" captionTargetPageId="14" captionText="FIGURE 12. Geographical distribution of Drymoluber Amaral, 1930. Black symbols represent localities with specimens examined, and white symbols represent literature records. Circles = Drymoluber dichrous (Peters, 1863); Triangles = Drymoluber brazili (Gomes, 1918); Lozenges = Drymoluber apurimacensis Lehr, Carrillo &amp; Hocking, 2004. Tropical and subtropical moist forests; Tropical and subtropical savannas; Deserts and xeric formations; Mangroves; Floodplains;" httpUri="https://zenodo.org/record/249307/files/figure.png" pageId="26" pageNumber="375">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BDF394B8F22FFEDFF75FC2D48C32279" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
Similar locality problems exist with the 
<typeStatus id="54DB87E98F22FFEDFD14FC324A2925E9" box="[678,785,944,968]" pageId="26" pageNumber="375" type="paratype">paratypes</typeStatus>
that purportedly also are from rail stations. The 
<emphasis id="B914E5598F22FFEDFAEEFC3249CC25CC" italics="true" pageId="26" pageNumber="375">Santa Eudóxia</emphasis>
, 
<emphasis id="B914E5598F22FFEDFEB4FC57489625CC" box="[262,430,980,1005]" italics="true" pageId="26" pageNumber="375">Sampaio Vidal</emphasis>
and 
<taxonomicName id="4C6042C88F22FFEDFE5EFC574B1925CD" box="[492,545,981,1004]" class="Insecta" family="Pompilidae" genus="Java" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B914E5598F22FFEDFE5EFC574B1925CD" box="[492,545,981,1004]" italics="true" pageId="26" pageNumber="375">Java</emphasis>
</taxonomicName>
stations, were part of a wide railroad network belonging to the 
<emphasis id="B914E5598F22FFEDFAABFC5648CA2231" italics="true" pageId="26" pageNumber="375">Companhia Paulista de Estradas de Ferro</emphasis>
that crossed the state of São Paulo from the southeast to the north and northwest (Giesbrecht 2009; Cavalcanti 2010). On the other hand, the 
<emphasis id="B914E5598F22FFEDFC83FB9F4DA52214" box="[817,1181,1052,1077]" italics="true" pageId="26" pageNumber="375">Estação Ferroviária Pedregulho</emphasis>
was located at km 455 on the 
<emphasis id="B914E5598F22FFEDFF51FBBD48AD2279" box="[227,405,1087,1112]" italics="true" pageId="26" pageNumber="375">Tronco-Catalão</emphasis>
railroad.
</paragraph>
</subSubSection>
<subSubSection id="C37A6AC08F22FFECFF75FBE64A492584" lastPageId="27" lastPageNumber="376" pageId="26" pageNumber="375" type="diagnosis">
<paragraph id="8BDF394B8F22FFEDFF75FBE64C6E22E4" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
<emphasis id="B914E5598F22FFEDFF75FBE6487B225C" bold="true" box="[199,323,1124,1149]" pageId="26" pageNumber="375">Diagnosis:</emphasis>
<taxonomicName id="4C6042C88F22FFEDFEF9FBE64B1D225C" box="[331,549,1124,1149]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFEF9FBE64B1D225C" box="[331,549,1124,1149]" italics="true" pageId="26" pageNumber="375">Drymoluber brazili</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C6042C88F22FFEDFCEAFBE74D26225D" box="[856,1054,1125,1148]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFCEAFBE74D26225D" box="[856,1054,1125,1148]" italics="true" pageId="26" pageNumber="375">D. apurimacensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6042C88F22FFEDFBE4FBE74DE2225C" box="[1110,1242,1124,1149]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F22FFEDFBE4FBE74DE2225C" box="[1110,1242,1124,1149]" italics="true" pageId="26" pageNumber="375">D. dichrous</emphasis>
</taxonomicName>
by the following combination of characters: 
<emphasis id="B914E5598F22FFEDFE78FB0548D82281" bold="true" box="[458,480,1159,1184]" pageId="26" pageNumber="375">a)</emphasis>
17-17-15 dorsal scale rows with two apical pits; 
<emphasis id="B914E5598F22FFEDFBBFFB054D1E2281" bold="true" box="[1037,1062,1159,1184]" pageId="26" pageNumber="375">b)</emphasis>
182–200 ventrals in males, 
<quantity id="4C9894AE8F22FFEDFAD6FB0549E622E4" metricMagnitude="0" metricUnit="m" metricValue="4.9149" metricValueMax="5.1308" metricValueMin="4.699" pageId="26" pageNumber="375" unit="in" value="193.5" valueMax="202.0" valueMin="185.0">185– 202 in</quantity>
females; 
<emphasis id="B914E5598F22FFEDFEF8FB2E486722E4" bold="true" box="[330,351,1196,1221]" pageId="26" pageNumber="375">c)</emphasis>
109–127 subcaudals in males, 
<quantity id="4C9894AE8F22FFEDFD0CFB2E4A0522E4" box="[702,829,1196,1221]" metricMagnitude="0" metricUnit="m" metricValue="2.9844999999999997" metricValueMax="3.2004" metricValueMin="2.7686" pageId="26" pageNumber="375" unit="in" value="117.5" valueMax="126.0" valueMin="109.0">109–126 in</quantity>
females; 
<emphasis id="B914E5598F22FFEDFC18FB2E4AFA22E4" bold="true" box="[938,962,1196,1221]" pageId="26" pageNumber="375">d)</emphasis>
19–25 maxillary teeth. See 
<tableCitation id="C6E20CF08F22FFEDFB4FFB2E4C6A22E4" box="[1277,1362,1196,1221]" captionStart="TABLE 5" captionStartId="21.[125,215,498,521]" captionText="TABLE 5. Summary of geographic distribution and morphological variation of the three species of Drymoluber Amaral, 1930. Most common (over 50 % of frequency) character states are represented in bold." pageId="26" pageNumber="375">Table 5</tableCitation>
.
</paragraph>
<paragraph id="8BDF394B8F22FFEDFF75FB4D48522359" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
<emphasis id="B914E5598F22FFEDFF75FB4D485722C9" bold="true" box="[199,367,1231,1256]" pageId="26" pageNumber="375">Comparisons:</emphasis>
<taxonomicName id="4C6042C88F22FFEDFECBFB4D4B9722C9" box="[377,687,1231,1256]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFECBFB4D4B9722C9" box="[377,687,1231,1256]" italics="true" pageId="26" pageNumber="375">Drymoluber apurimacensis</emphasis>
</taxonomicName>
has 13-13-13 dorsal scales rows, and 
<taxonomicName id="4C6042C88F22FFEDFBDCFB534DCD22C9" box="[1134,1269,1231,1256]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F22FFEDFBDCFB534DCD22C9" box="[1134,1269,1231,1256]" italics="true" pageId="26" pageNumber="375">D. dichrous</emphasis>
</taxonomicName>
has 15-15-15. Apical pits are absent in 
<taxonomicName id="4C6042C88F22FFEDFE1BFB774B56232D" box="[425,622,1269,1292]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFE1BFB774B56232D" box="[425,622,1269,1292]" italics="true" pageId="26" pageNumber="375">D. apurimacensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C6042C88F22FFEDFDC9FB764A95232D" box="[635,941,1268,1293]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFDC9FB764A95232D" box="[635,941,1268,1293]" italics="true" pageId="26" pageNumber="375">Drymoluber apurimacensis</emphasis>
</taxonomicName>
has 158–164 ventrals in males and 
<quantity id="4C9894AE8F22FFEDFA88FB7649952311" metricMagnitude="0" metricUnit="m" metricValue="4.4196" metricValueMax="4.6228" metricValueMin="4.2164" pageId="26" pageNumber="375" unit="in" value="174.0" valueMax="182.0" valueMin="166.0">166–182 in</quantity>
females, 84–93 subcaudals in males and 
<quantity id="4C9894AE8F22FFEDFDCDFA9A4BDA2311" box="[639,738,1303,1328]" metricMagnitude="0" metricUnit="m" metricValue="2.2606" metricValueMax="2.3114" metricValueMin="2.2098" pageId="26" pageNumber="375" unit="in" value="89.0" valueMax="91.0" valueMin="87.0">87–91 in</quantity>
females. 
<taxonomicName id="4C6042C88F22FFEDFCFDFA954D782311" box="[847,1088,1303,1328]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F22FFEDFCFDFA954D782311" box="[847,1088,1303,1328]" italics="true" pageId="26" pageNumber="375">Drymoluber dichrous</emphasis>
</taxonomicName>
has 157–173 ventrals in males and 
<quantity id="4C9894AE8F22FFEDFF75FABE487F2374" box="[199,327,1340,1365]" metricMagnitude="0" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="26" pageNumber="375" unit="in" value="170.0" valueMax="180.0" valueMin="160.0">160–180 in</quantity>
females, 87–110 subcaudals in males and 
<quantity id="4C9894AE8F22FFEDFC9BFABE4AA32374" box="[809,923,1340,1365]" metricMagnitude="0" metricUnit="m" metricValue="2.4765" metricValueMax="2.7686" metricValueMin="2.1844" pageId="26" pageNumber="375" unit="in" value="97.5" valueMax="109.0" valueMin="86.0">86–109 in</quantity>
females. 
<taxonomicName id="4C6042C88F22FFEDFBBBFABE4C052375" box="[1033,1341,1340,1365]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFBBBFABE4C052375" box="[1033,1341,1340,1365]" italics="true" pageId="26" pageNumber="375">Drymoluber apurimacensis</emphasis>
</taxonomicName>
has 14– 16 maxillary teeth.
</paragraph>
<paragraph id="8BDF394B8F22FFEDFF75FA064867200D" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
Small specimens of 
<taxonomicName id="4C6042C88F22FFEDFE1BFA064BB923BC" box="[425,641,1412,1437]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFE1BFA064BB923BC" box="[425,641,1412,1437]" italics="true" pageId="26" pageNumber="375">Drymoluber brazili</emphasis>
</taxonomicName>
have dark crossbands 2–6 scales wide (mean 3.6) and light interspaces 0.5–5 scales wide (mean 1.6), while in a single specimen of 
<taxonomicName id="4C6042C88F22FFEDFCF4FA2B4D3523E1" box="[838,1037,1449,1472]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFCF4FA2B4D3523E1" box="[838,1037,1449,1472]" italics="true" pageId="26" pageNumber="375">D. apurimacensis</emphasis>
</taxonomicName>
the dark crossbands are 1–2 scales wide and the light interspaces 2–3 scales wide. Young specimens of 
<taxonomicName id="4C6042C88F22FFEDFC26FA4F4D2023C4" box="[916,1048,1484,1509]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F22FFEDFC26FA4F4D2023C4" box="[916,1048,1484,1509]" italics="true" pageId="26" pageNumber="375">D. dichrous</emphasis>
</taxonomicName>
have dark crossbands with similar width to those of 
<taxonomicName id="4C6042C88F22FFEDFED7FA7348EB2029" box="[357,467,1519,1544]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFED7FA7348EB2029" box="[357,467,1519,1544]" italics="true" pageId="26" pageNumber="375">D. brazili</emphasis>
</taxonomicName>
(1.5–7 scales, mean 3.6), but the pale interspaces are on average narrower (0.5–2.5 scales, mean 0.8).
</paragraph>
<paragraph id="8BDF394B8F22FFEDFF75F9B54B93209C" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="375">
The hemipenes of 
<taxonomicName id="4C6042C88F22FFEDFE01F9B54BAE2071" box="[435,662,1591,1616]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F22FFEDFE01F9B54BAE2071" box="[435,662,1591,1616]" italics="true" pageId="26" pageNumber="375">Drymoluber brazili</emphasis>
</taxonomicName>
tend to have fewer calyces than that of 
<taxonomicName id="4C6042C88F22FFEDFB18F9BB4C0F2071" box="[1194,1335,1591,1616]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F22FFEDFB18F9BB4C0F2071" box="[1194,1335,1591,1616]" italics="true" pageId="26" pageNumber="375">D. dichrous</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6042C88F22FFEDFA33F9BB48022055" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFA33F9BB48022055" italics="true" pageId="26" pageNumber="375">D. apurimacensis</emphasis>
</taxonomicName>
, larger spinulate flounces, and spines in the lobular region. The walls of the sulcus spermaticus are less ornamented. The spines of the asulcate face in general are smaller than those of 
<taxonomicName id="4C6042C88F22FFEDFB00F9034C0420B9" box="[1202,1340,1663,1688]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F22FFEDFB00F9034C0420B9" box="[1202,1340,1663,1688]" italics="true" pageId="26" pageNumber="375">D. dichrous</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6042C88F22FFEDFA33F9034802209D" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="375" phylum="Chordata" rank="species" species="apurimacensis">
<emphasis id="B914E5598F22FFEDFA33F9034802209D" italics="true" pageId="26" pageNumber="375">D. apurimacensis</emphasis>
</taxonomicName>
, especially those most proximal.
</paragraph>
<paragraph id="8BDF394B8F22FFECFF75F9454B1C278C" blockId="26.[151,1437,151,2013]" lastBlockId="27.[151,1436,151,2006]" lastPageId="27" lastPageNumber="376" pageId="26" pageNumber="375">
<emphasis id="B914E5598F22FFEDFF75F9454B9A20C1" bold="true" box="[199,674,1735,1760]" pageId="26" pageNumber="375">
Redescription of the 
<typeStatus id="54DB87E98F22FFEDFE76F9454B1320C1" box="[452,555,1735,1760]" pageId="26" pageNumber="375" type="holotype">holotype</typeStatus>
(
<figureCitation id="135B25CE8F22FFEDFD89F9454BAB20C1" box="[571,659,1735,1760]" captionStart="FIGURE 18" captionStartId="28.[151,250,1951,1973]" captionTargetBox="[153,1430,369,1926]" captionTargetId="figure@28.[150,1435,365,1929]" captionTargetPageId="28" captionText="FIGURE 18. Holotype of Drymoluber brazili (Gomes, 1918) (IBSP 696). A) dorsal view of body; B) ventral view of body; C) right side of the head; D) left side of the head; E) dorsum of head; F) gular region. Photos: Henrique C. Costa (A – D, F) and Ana Bárbara Barros (E)." httpUri="https://zenodo.org/record/249312/files/figure.png" pageId="26" pageNumber="375">Fig. 18</figureCitation>
):
</emphasis>
Snout-vent length 
<quantity id="4C9894AE8F22FFEDFCCFF94A4AD120FE" box="[893,1001,1736,1760]" metricMagnitude="0" metricUnit="m" metricValue="1.09" pageId="26" pageNumber="375" unit="mm" value="1090.0">1090 mm</quantity>
and tail length 
<quantity id="4C9894AE8F22FFEDFB2CF94A4DC720FE" box="[1182,1279,1736,1760]" metricMagnitude="-1" metricUnit="m" metricValue="4.7299999999999995" pageId="26" pageNumber="375" unit="mm" value="473.0">473 mm</quantity>
; head distinct from the body, 
<quantity id="4C9894AE8F22FFEDFEF1F96E489F2125" box="[323,423,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="26" pageNumber="375" unit="mm" value="35.1">35.1 mm</quantity>
long (3.2% of the SVL); greatest width of head 
<quantity id="4C9894AE8F22FFEDFC75F96E4D012125" box="[967,1081,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="26" pageNumber="375" unit="mm" value="15.2">15.20 mm</quantity>
(43% of head length); width of head at the supraoculars 
<quantity id="4C9894AE8F22FFEDFE1CF8924B2D2106" box="[430,533,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="26" pageNumber="375" unit="mm" value="11.3">11.3 mm</quantity>
; internasal distance 
<quantity id="4C9894AE8F22FFEDFD4BF88D4A6A2106" box="[761,850,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="26" pageNumber="375" unit="mm" value="6.1">6.1 mm</quantity>
; eye diameter 6.0 mm; eye-nostril distance 
<quantity id="4C9894AE8F22FFEDFA8DF88D4CA02106" box="[1343,1432,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="26" pageNumber="375" unit="mm" value="5.9">5.9 mm</quantity>
. Smooth dorsal scales in 17-17-15 rows with two apical pits; 190 ventrals and 1 preventral (
<emphasis id="B914E5598F22FFEDFB0EF8B74DC2216D" box="[1212,1274,1845,1868]" italics="true" pageId="26" pageNumber="375">sensu</emphasis>
Peters 1964); cloacal shield entire; tail intact with 117 divided subcaudals and one terminal spine; rostral wider than high, visible from above; internasals and prefrontals slightly wider than long; each prefrontal contacting the frontal, supraocular, internasals, posterior nasal, preocular and loreal; frontal about 1.5 times longer than wide; supraoculars longer than wide; parietals about 1.5 times longer than wide; nasal divided above and below the naris, mainly in contact with the first supralabial, but also with the second; loreal slightly longer than high, (vaguely resembling a parallelogram), contacting the second and third supralabials; one preocular; two postoculars, the upper one larger than the lower; two anterior temporals (one upper and one lower) and two posterior temporals (one upper and one lower) (1/1+1/1) on both sides of the head; eight supralabials, the fourth and the fifth contacting the eye; mental triangular, wider than long; nine infralabials, the first pair in contact behind the mental; first to fifth infralabials in contact with the first pair of chinshields; fifth infralabial in contact with the second pair of chinshields; fifth to ninth infralabials contacting the gulars; first pair of chinshields about the half the length of the second; 23 maxillary teeth, increasing in size posteriorly.
</paragraph>
<paragraph id="8BDF394B8F23FFECFF75FE354DF924ED" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">
There are some small differences between the redescription of the 
<typeStatus id="54DB87E98F23FFECFC74FE354D1027F1" box="[966,1064,439,464]" pageId="27" pageNumber="376" type="holotype">holotype</typeStatus>
presented here (inside brackets) and the original description. Gomes (1918) decribed the 
<typeStatus id="54DB87E98F23FFECFCA0FE5E4A4C27D4" box="[786,884,476,501]" pageId="27" pageNumber="376" type="holotype">holotype</typeStatus>
as having 22 maxillary teeth, internasals as wide as long, 10 infralabials, 191 ventrals, SVL 
<quantity id="4C9894AE8F23FFECFDC7FD824BE52436" box="[629,733,512,536]" metricMagnitude="0" metricUnit="m" metricValue="1.1099999999999999" pageId="27" pageNumber="376" unit="mm" value="1110.0">1110 mm</quantity>
and TL = 
<quantity id="4C9894AE8F23FFECFCF8FD824A932436" box="[842,939,512,536]" metricMagnitude="-1" metricUnit="m" metricValue="4.8" pageId="27" pageNumber="376" unit="mm" value="480.0">480 mm</quantity>
. The difference in the number of infralabials may be explained on the basis of the method used in both studies. In the present work, the posterior boundary of the infralabial row is defined as the last scale completely covered by the last supralabial (Peters 1964) (
<figureCitation id="135B25CE8F23FFECFA97FDC54C422441" box="[1317,1402,583,608]" captionStart="FIGURE 18" captionStartId="28.[151,250,1951,1973]" captionTargetBox="[153,1430,369,1926]" captionTargetId="figure@28.[150,1435,365,1929]" captionTargetPageId="28" captionText="FIGURE 18. Holotype of Drymoluber brazili (Gomes, 1918) (IBSP 696). A) dorsal view of body; B) ventral view of body; C) right side of the head; D) left side of the head; E) dorsum of head; F) gular region. Photos: Henrique C. Costa (A – D, F) and Ana Bárbara Barros (E)." httpUri="https://zenodo.org/record/249312/files/figure.png" pageId="27" pageNumber="376">Fig. 18</figureCitation>
C), which is not the definition seemingly used by Gomes (1918). The differences in SVL and TL may be due to a small error in the original measuring, or a reduction in size caused by fixation and preservation over time time (Vervust 
<emphasis id="B914E5598F23FFECFA3AFD13498C24EC" italics="true" pageId="27" pageNumber="376">et al.</emphasis>
2009; Guimarães 
<emphasis id="B914E5598F23FFECFE30FD37488224EC" box="[386,442,692,717]" italics="true" pageId="27" pageNumber="376">et al.</emphasis>
2010). This situation is also applicable to the 
<typeStatus id="54DB87E98F23FFECFC0CFD374D1124EC" box="[958,1065,693,717]" pageId="27" pageNumber="376" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C6042C88F23FFECFBFCFD374D8124EC" box="[1102,1209,692,717]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="376" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F23FFECFBFCFD374D8124EC" box="[1102,1209,692,717]" italics="true" pageId="27" pageNumber="376">D. brazili</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF394B8F23FFECFF75FD554A492584" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">
<emphasis id="B914E5598F23FFECFF75FD554B3724D1" bold="true" box="[199,527,727,752]" pageId="27" pageNumber="376">
Coloration of the 
<typeStatus id="54DB87E98F23FFECFE2DFD554B3224D1" box="[415,522,727,752]" pageId="27" pageNumber="376" type="holotype">holotype</typeStatus>
:
</emphasis>
Gomes (1918) described the 
<typeStatus id="54DB87E98F23FFECFCD1FD554AFD24D1" box="[867,965,727,752]" pageId="27" pageNumber="376" type="holotype">holotype</typeStatus>
in life as having olive-green color in the head and on the anterior part of the body, becoming reddish-brown in the posterior part of body and on the tail. The venter was yellowish-white, while the lateral edges of the ventral plates were the same color as the dorsum. When the senior author examined the 
<typeStatus id="54DB87E98F23FFECFE44FCC64B63257C" box="[502,603,836,861]" pageId="27" pageNumber="376" type="holotype">holotype</typeStatus>
, it was grayish-brown on the dorsum of head and anterior half of the body, becoming yellowish-brown on the posterior half of the body and tail. Supralabials, gular region and venter were yellowish-brown and similar to or slightly paler than the dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C37A6AC08F23FFD7FF75FC2D4D642489" lastPageId="32" lastPageNumber="381" pageId="27" pageNumber="376" type="description">
<paragraph id="8BDF394B8F23FFECFF75FC2D4A642215" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">
<emphasis id="B914E5598F23FFECFF75FC2D4B7225E9" bold="true" box="[199,586,943,968]" pageId="27" pageNumber="376">Coloration of preserved adults:</emphasis>
The dorsal coloration of most of the examined adults (n=49; 77.8%) is uniform bluish-gray, bluish-brown or yellowish-brown throughout. A few specimens (n=13; 20.6%) are grayishbrown or bluish-brown on the anterior half of the body and yellowish-brown on the posterior half. One specimen has faint crossbands as are present in the juvenile color pattern.
</paragraph>
<paragraph id="8BDF394B8F23FFECFF75FBBD4A3422E4" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">The venter of most specimens (n=49; 77.8%) is immaculate, yellowish-cream, and the lateral edges of ventrals have the same color as the dorsal scales. In one specimen (1.6%) the dorsal coloration extends into the ventral region; in other specimens (n=13; 20.6%) the venter is yellowish-colored with some dark spots, and the lateral edges of ventrals are the same color as the dorsal scales.</paragraph>
<paragraph id="8BDF394B8F23FFECFF75FB4D4A1C23BD" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">
Dorsally, the tail is the same color as the body, but shows some variations in the subcaudal coloration. In 
<specimenCount id="9D66F2C28F23FFECFA32FB524834232C" pageId="27" pageNumber="376" type="generic">24 specimens</specimenCount>
(38.1%) the subcaudals are yellowish-cream, immaculate, and with their lateral edges the same color as the dorsum. In some snakes, the lateral edges are inconspicuous and the same color as subcaudals (n=5; 7.9%). In a few, the subcaudals have more than half their area dark like the dorsals (n=1; 1.6%), or have black spots throughout their area (n=1; 1.6%). In others, black spots are numerous only in the posterior region of the tail (n=24; 38.1%), or the posterior part of tail is completely black (n=8; 12.7%).
</paragraph>
<paragraph id="8BDF394B8F23FFECFF75FA2B4D672029" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">In all specimens the head is uniform dorsally and the same color as the dorsum of body. The gular region is immaculate, yellowish-cream. The supralabials of most adult specimens have dark lateral edges (mainly in the last scales (n=59; 93.6%), but in some specimens visible markings are absent (n=4; 6.4%).</paragraph>
<paragraph id="8BDF394B8F23FFECFF75F996482B20C1" blockId="27.[151,1436,151,2006]" pageId="27" pageNumber="376">
<emphasis id="B914E5598F23FFECFF75F9964B31200C" bold="true" box="[199,521,1556,1581]" pageId="27" pageNumber="376">Coloration of adults in life:</emphasis>
Unlike 
<taxonomicName id="4C6042C88F23FFECFDD0F9964A6B200C" box="[610,851,1556,1581]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="376" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F23FFECFDD0F9964A6B200C" box="[610,851,1556,1581]" italics="true" pageId="27" pageNumber="376">Drymoluber dichrous</emphasis>
</taxonomicName>
, there is little information about the coloration of 
<taxonomicName id="4C6042C88F23FFECFA33F99749D82071" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="376" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F23FFECFA33F99749D82071" italics="true" pageId="27" pageNumber="376">D. brazili</emphasis>
</taxonomicName>
in life. Gomes (1918) described the the 
<typeStatus id="54DB87E98F23FFECFD77F9BB4BCD2070" box="[709,757,1593,1617]" pageId="27" pageNumber="376">type</typeStatus>
series in life as follows: two specimens have the dorsum completely olive-green, while in the other three, and the 
<typeStatus id="54DB87E98F23FFECFC80F9DE4AAF2054" box="[818,919,1628,1653]" pageId="27" pageNumber="376" type="holotype">holotype</typeStatus>
, the posterior part of the dorsum is reddishbrown. This same pattern is seen in the illustration (
<figureCitation id="135B25CE8F23FFECFD5CF9FD4A7820B9" box="[750,832,1663,1688]" captionStart="FIGURE 19" captionStartId="29.[151,250,780,802]" captionTargetBox="[156,1430,198,746]" captionTargetId="figure@29.[151,1436,175,759]" captionTargetPageId="29" captionText="FIGURE 19. Color in life of some adult specimens of Drymoluber brazili (Gomes, 1918). A) Visconde de Parnaíba, São Paulo, Brazil (Illustration by R. K. Leyer presented in Amaral [1978]); B) UHE Peixe-Angical, Tocantins, Brazil. Photo: Pedro Henrique Bernardo." httpUri="https://zenodo.org/record/249313/files/figure.png" pageId="27" pageNumber="376">Fig. 19</figureCitation>
A) of one specimen from Visconde de Parnaíba, São Paulo (IBSP 4812) published by Amaral (1978). In some specimens, the dorsal color may consist of grayish tones (
<figureCitation id="135B25CE8F23FFECFF2DF94549CA20C1" box="[159,242,1735,1760]" captionStart="FIGURE 19" captionStartId="29.[151,250,780,802]" captionTargetBox="[156,1430,198,746]" captionTargetId="figure@29.[151,1436,175,759]" captionTargetPageId="29" captionText="FIGURE 19. Color in life of some adult specimens of Drymoluber brazili (Gomes, 1918). A) Visconde de Parnaíba, São Paulo, Brazil (Illustration by R. K. Leyer presented in Amaral [1978]); B) UHE Peixe-Angical, Tocantins, Brazil. Photo: Pedro Henrique Bernardo." httpUri="https://zenodo.org/record/249313/files/figure.png" pageId="27" pageNumber="376">Fig. 19</figureCitation>
B).
</paragraph>
<paragraph id="8BDF394B8F23FFEBFF75F96E4B0926D7" blockId="27.[151,1436,151,2006]" lastBlockId="28.[151,1436,151,316]" lastPageId="28" lastPageNumber="377" pageId="27" pageNumber="376">
<emphasis id="B914E5598F23FFECFF75F96E4B582124" bold="true" box="[199,608,1772,1797]" pageId="27" pageNumber="376">Coloration of preserved juveniles:</emphasis>
The color pattern of preserved juveniles of 
<taxonomicName id="4C6042C88F23FFECFBE0F96E4C142124" box="[1106,1324,1772,1797]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="376" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F23FFECFBE0F96E4C142124" box="[1106,1324,1772,1797]" italics="true" pageId="27" pageNumber="376">Drymoluber brazili</emphasis>
</taxonomicName>
is similar to that of the other species of the genus, consisting of dark crossbands separated by pale interspaces. Only two specimens (10%) have crossbands (47 and 54) visible throughout the body (MZUSP 9596 and IBSP 29221). Crossbands on the tail were not visible in any specimen. The dark crossbands varied in width between 2–6 vertebral/paravertebral scales (mean 3.6; SD=0.89; n=62 crossbands). The last crossband anterior to the cloaca is the narrowest and two scales wide (n=2; 10%), while the first crossband on the body is the widest, 2–6 scales wide (mean 4; SD=0.95; n=20; 100%). The widths of the pale interspaces vary between 0.5–5 scales (mean 1.6; SD=0.97; n=62 interespaces). The interspace anterior to the fifth crossband anterior to the cloaca is the narrowest, with 0.5 or 1.5 scales (n=2; 10%). The interspace posterior to the first crossband is the widest, with 1–5 scales (mean 2.32; SD=0.90; n=20; 100%).
</paragraph>
<paragraph id="8BDF394B8F24FFEBFF75FE824B69271D" blockId="28.[151,1436,151,316]" pageId="28" pageNumber="377">The venter of young specimens is always yellowish-cream and immaculate, and the lateral edges of ventral plates are the same color as the dorsals.</paragraph>
<caption id="DF1F69C38F24FFEBFF25F81D486621D3" httpUri="https://zenodo.org/record/249312/files/figure.png" pageId="28" pageNumber="377" targetBox="[153,1430,369,1926]" targetPageId="28">
<paragraph id="8BDF394B8F24FFEBFF25F81D486621D3" blockId="28.[151,1436,1951,2034]" pageId="28" pageNumber="377">
<emphasis id="B914E5598F24FFEBFF25F81D48272194" bold="true" box="[151,287,1951,1973]" pageId="28" pageNumber="377">FIGURE 18.</emphasis>
Holotype of 
<taxonomicName id="4C6042C88F24FFEBFE17F81D4A3F2194" authority="Gomes, 1918" authorityName="Gomes" authorityYear="1918" box="[421,775,1951,1973]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="377" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F24FFEBFE17F81D4B512194" box="[421,617,1951,1973]" italics="true" pageId="28" pageNumber="377">Drymoluber brazili</emphasis>
(Gomes, 1918)
</taxonomicName>
(IBSP 696). A) dorsal view of body; B) ventral view of body; C) right side of the head; D) left side of the head; E) dorsum of head; F) gular region. Photos: Henrique C. Costa (A–D, F) and Ana Bárbara Barros (E).
</paragraph>
</caption>
<caption id="DF1F69C38F25FFEAFF25FC8E48592541" httpUri="https://zenodo.org/record/249313/files/figure.png" pageId="29" pageNumber="378" targetBox="[156,1430,198,746]" targetPageId="29">
<paragraph id="8BDF394B8F25FFEAFF25FC8E48592541" blockId="29.[151,1436,780,864]" pageId="29" pageNumber="378">
<emphasis id="B914E5598F25FFEAFF25FC8E48272503" bold="true" box="[151,287,780,802]" pageId="29" pageNumber="378">FIGURE 19.</emphasis>
Color in life of some adult specimens of 
<taxonomicName id="4C6042C88F25FFEAFD0FFC8E4D192503" authority="Gomes, 1918" authorityName="Gomes" authorityYear="1918" box="[701,1057,780,802]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="378" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F25FFEAFD0FFC8E4AB82503" box="[701,896,780,802]" italics="true" pageId="29" pageNumber="378">Drymoluber brazili</emphasis>
(Gomes, 1918)
</taxonomicName>
. A) Visconde de Parnaíba, São Paulo, Brazil (Illustration by R. K. Leyer presented in Amaral [1978]); B) 
<emphasis id="B914E5598F25FFEAFCC1FCAE4D792560" box="[883,1089,811,833]" italics="true" pageId="29" pageNumber="378">UHE Peixe-Angical</emphasis>
, Tocantins, Brazil. Photo: Pedro Henrique Bernardo.
</paragraph>
</caption>
<caption id="DF1F69C38F25FFEAFF25FA0A4D96239C" httpUri="https://zenodo.org/record/249314/files/figure.png" pageId="29" pageNumber="378" targetBox="[160,1430,919,1389]" targetPageId="29">
<paragraph id="8BDF394B8F25FFEAFF25FA0A4D96239C" blockId="29.[151,1436,1416,1469]" pageId="29" pageNumber="378">
<emphasis id="B914E5598F25FFEAFF25FA0A481823BF" bold="true" box="[151,288,1416,1438]" pageId="29" pageNumber="378">FIGURE 20.</emphasis>
Color in life of juvenile specimens of 
<taxonomicName id="4C6042C88F25FFEAFD15FA0A4D3623BF" authority="Gomes, 1918" authorityName="Gomes" authorityYear="1918" box="[679,1038,1416,1438]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="378" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F25FFEAFD15FA0A4A1B23BF" box="[679,803,1416,1438]" italics="true" pageId="29" pageNumber="378">Drymoluber</emphasis>
<emphasis id="B914E5598F25FFEAFC98FA0A4A5423BF" box="[810,876,1416,1438]" italics="true" pageId="29" pageNumber="378">brazili</emphasis>
(Gomes, 1918)
</taxonomicName>
. A) 
<emphasis id="B914E5598F25FFEAFB8AFA0B4C7523BF" box="[1080,1357,1416,1438]" italics="true" pageId="29" pageNumber="378">Parque Nacional das Emas</emphasis>
, Goiás, Brazil (IBSP 62682); B) Unknown locality data. Photos: Paula H. Valdujo (A) and Eduardo Santos (B).
</paragraph>
</caption>
<paragraph id="8BDF394B8F25FFEAFF75FA6B4BE02007" blockId="29.[151,1436,1513,2006]" pageId="29" pageNumber="378">The subcaudals are cream colored, with the lateral edges darkened (n=9; 45%), cream (n=6; 30%), or cream with only the posterior edges darkened (n=5; 25%).</paragraph>
<paragraph id="8BDF394B8F25FFEAFF75F9B34D7D2097" blockId="29.[151,1436,1513,2006]" pageId="29" pageNumber="378">
The head has light internasals, light pre-frontals with darkened posterior edges, and a dark frontal and supraoculars with light anterior edges. A light stripe in the parietal region can be present and immaculate (n=9; 45%) or maculate (n=4; 20%). In the remaining specimens (n=7; 35%) the parietals are dark (
<figureCitation id="135B25CE8F25FFEAFB18F9FB4DC420B3" box="[1194,1276,1657,1682]" captionStart="FIGURE 11" captionStartId="13.[151,250,1403,1425]" captionTargetBox="[245,1340,695,1383]" captionTargetId="figure@13.[243,1344,694,1385]" captionTargetPageId="13" captionText="FIGURE 11. Variation in color of the dorsum of the head in young specimens of Drymoluber. A) Presence of a light parietal stripe (MZUSP 8494, D. dichrous, Alto Paraíso, Rondônia, Brazil); B) Light stripe present, although marked with two large spots (IBSP 16499, D. brazili, Turiba do Sul, São Paulo, Brazil); C) Light stripe absent (MZUSP 14298, D. brazili, UHE Luís Eduardo Magalhães, Tocantins, Brazil). Photos: Henrique C. Costa." httpUri="https://zenodo.org/record/249306/files/figure.png" pageId="29" pageNumber="378">Fig. 11</figureCitation>
). 
<collectingCountry id="F37779DB8F25FFEAFAA3F9F84C0820B3" box="[1297,1328,1658,1682]" name="American Samoa" pageId="29" pageNumber="378">As</collectingCountry>
occurs in other species of the genus, lighter shades on the head darken as the specimens grow.
</paragraph>
<paragraph id="8BDF394B8F25FFEAFF75F94048C020DF" blockId="29.[151,1436,1513,2006]" pageId="29" pageNumber="378">
<collectingCountry id="F37779DB8F25FFEAFF75F94049DE20FB" box="[199,230,1730,1754]" name="American Samoa" pageId="29" pageNumber="378">As</collectingCountry>
in adults, the gular region of the juveniles is pale and immaculate. The supralabials have faint dark markings on their lateral edges.
</paragraph>
<paragraph id="8BDF394B8F25FFEAFF75F88B4DD4214B" blockId="29.[151,1436,1513,2006]" pageId="29" pageNumber="378">
<emphasis id="B914E5598F25FFEAFF75F88B4B0A2103" bold="true" box="[199,562,1801,1826]" pageId="29" pageNumber="378">Coloration of juveniles in life:</emphasis>
In life, the light colored regions in preserved juveniles are shades of white, cream and light-brown, except in the head and neck, where an orange-red color prevails. The dark colored regions (including crossbands) vary in shades of brown, olive-green and dark-gray (
<figureCitation id="135B25CE8F25FFEAFC5BF8D34D04214B" box="[1001,1084,1873,1898]" captionStart="FIGURE 20" captionStartId="29.[151,250,1416,1438]" captionTargetBox="[160,1430,919,1389]" captionTargetId="figure@29.[151,1436,914,1395]" captionTargetPageId="29" captionText="FIGURE 20. Color in life of juvenile specimens of Drymoluber brazili (Gomes, 1918). A) Parque Nacional das Emas, Goiás, Brazil (IBSP 62682); B) Unknown locality data. Photos: Paula H. Valdujo (A) and Eduardo Santos (B)." httpUri="https://zenodo.org/record/249314/files/figure.png" pageId="29" pageNumber="378">Fig. 20</figureCitation>
; Amaral 1923).
</paragraph>
<paragraph id="8BDF394B8F25FFEAFF75F8F74CBD21F4" blockId="29.[151,1436,1513,2006]" pageId="29" pageNumber="378">
Apparently, only 
<taxonomicName id="4C6042C88F25FFEAFE3AF8F548CB21AF" box="[392,499,1909,1934]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="378" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F25FFEAFE3AF8F548CB21AF" box="[392,499,1909,1934]" italics="true" pageId="29" pageNumber="378">D. brazili</emphasis>
</taxonomicName>
juveniles have this reddish coloration on the head. However, since those colors tend to become white in preserved specimens, and because photographs and information about the coloration of live juveniles of 
<taxonomicName id="4C6042C88F25FFEAFE93F83F489221F7" box="[289,426,1981,2006]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="378" phylum="Chordata" rank="genus">
<emphasis id="B914E5598F25FFEAFE93F83F489221F7" box="[289,426,1981,2006]" italics="true" pageId="29" pageNumber="378">Drymoluber</emphasis>
</taxonomicName>
are uncommon, it is difficult to know whether this character is diagnostic for 
<taxonomicName id="4C6042C88F25FFEAFAA1F83D4C4521F7" box="[1299,1405,1981,2006]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="378" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F25FFEAFAA1F83D4C4521F7" box="[1299,1405,1981,2006]" italics="true" pageId="29" pageNumber="378">D. brazili</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75FF154A79241C" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">
<emphasis id="B914E5598F26FFE9FF75FF154B902691" bold="true" box="[199,680,151,176]" pageId="30" pageNumber="379">
Hemipenial morphology (n=6) (
<figureCitation id="135B25CE8F26FFE9FDF0FF154BA22691" box="[578,666,151,176]" captionStart="FIGURE 21" captionStartId="31.[151,250,1955,1977]" captionTargetBox="[353,1243,176,1942]" captionTargetId="figure@31.[353,1244,165,1943]" captionTargetPageId="31" captionText="FIGURE 21. Hemipenis of Drymoluber brazili. IBSP 33668, Goiânia, Goiás, Brazil, A) Sulcate face; B) Asulcate face. MCNR 1736, Usina Hidrelétrica de Irapé, Minas Gerais, Brazil, C) Sulcate face; D) Asulcate face. Scale bar = 1 cm. Photos: Henrique C. Costa." httpUri="https://zenodo.org/record/249315/files/figure.png" pageId="30" pageNumber="379">Fig. 21</figureCitation>
):
</emphasis>
Hemipenis single, subcylindrical, not capitate. Sulcus spermaticus single and centrolinear. Lobe about half of the hemipenis length, with papillate calyces (5–10 papillae per calyx, most calyces deep) and spines bordering the sulcus spermaticus. The calyces are gradually replaced by spinulate flounces and spines proximally. The hemipenial body is covered by spines arranged in more or less transverse rows (about 60 or a few more spines in total). Walls of the sulcus spermaticus weakly ornamented, usually with few papillae and spinules. These walls are also bordered on both sides by a longitudinal row of 8–13 spines (mean 8; SD=1.97). The spines from the right side tend to increase in size toward the proximal region, while the spines from the left side increase in size up to the middle of the row, then decrease in size toward the proximal region (n=4). In the rest of the examined specimens (n=2), spines from both sides increase toward the proximal region of the hemipenis. A basal hook is present at the end of each row of spines bordering the sulcus, with the left hook more proximally than the right hook. None (n=2), or three to four small spines (n=2) or spinules (n=2) may be present between the left hook and the wall of the sulcus spermaticus.
</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75FDCA48782534" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">One (n=2) or two (n=2) lateral spines, larger than the hooks, may be present to the left of the sulcus spermaticus, or two large lateral spines may be present on both sides of the sulcus (n=1). The asulcate face is formed by spines arranged in 6–7 more or less transverse rows (counted from proximal to distal region), with largest spines in the median rows, and proximal spines smaller than distal spines. In two specimens (MCNR 1736 and UFMT 6970), those spines are very small. The base of the hemipenis is smooth, with some grooves and several sparse spinules</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75FC9D4CBF25E9" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">
<emphasis id="B914E5598F26FFE9FF75FC9D48792519" bold="true" box="[199,321,799,824]" pageId="30" pageNumber="379">Variation:</emphasis>
Largest male with SVL 
<quantity id="4C9894AE8F26FFE9FDD2FCA24BF62516" box="[608,718,800,824]" metricMagnitude="0" metricUnit="m" metricValue="1.178" pageId="30" pageNumber="379" unit="mm" value="1178.0">1178 mm</quantity>
, TL 410+N mm (IBSP 34369); largest female with SVL 
<quantity id="4C9894AE8F26FFE9FAD6FC9D49FD257D" metricMagnitude="0" metricUnit="m" metricValue="1.035" pageId="30" pageNumber="379" unit="mm" value="1035.0">1035 mm</quantity>
, TL 400+N mm (IBSP 17019). Twenty eight (28) specimens examined (33.7%) had broken tail. The tail of 24 of those specimens (28.9% of the total examined) was healed, suggesting the presence of pseudoautotomy in this species. Tail length of specimens with complete tail is 38.1–62.7% of the SVL in males (mean 44.2%; SD=4.23; n=29) and 41.11–56% (mean 45.7%; SD=3.5; n=26) in females. For variation in meristic characters, see 
<tableCitation id="C6E20CF08F26FFE9FA9EFC2D4CBA25E9" box="[1324,1410,943,968]" captionStart="TABLE 5" captionStartId="21.[125,215,498,521]" captionText="TABLE 5. Summary of geographic distribution and morphological variation of the three species of Drymoluber Amaral, 1930. Most common (over 50 % of frequency) character states are represented in bold." pageId="30" pageNumber="379">Table 5</tableCitation>
.
</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75FC564C0722E4" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">
<emphasis id="B914E5598F26FFE9FF75FC5648CF25CC" bold="true" box="[199,503,980,1005]" pageId="30" pageNumber="379">Geographic distribution:</emphasis>
<taxonomicName id="4C6042C88F26FFE9FDB6FC564BDA25CC" box="[516,738,980,1005]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FDB6FC564BDA25CC" box="[516,738,980,1005]" italics="true" pageId="30" pageNumber="379">Drymoluber brazili</emphasis>
</taxonomicName>
is distributed mainly in 
<collectingCountry id="F37779DB8F26FFE9FBA3FC564D6025CC" box="[1041,1112,980,1005]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
, with a single record from 
<collectingCountry id="F37779DB8F26FFE9FF25FC7A49C72231" box="[151,255,1016,1040]" name="Paraguay" pageId="30" pageNumber="379">Paraguay</collectingCountry>
(Canindeyú province). The species inhabits the Cerrado domain, with additional records from the Caatinga, Atlantic Forest and its transitional areas with the Cerrado (
<figureCitation id="135B25CE8F26FFE9FC10FB9E4ACE2215" box="[930,1014,1052,1077]" captionStart="FIGURE 12" captionStartId="14.[151,250,1764,1786]" captionTargetBox="[152,1433,625,1738]" captionTargetId="figure@14.[151,1436,621,1742]" captionTargetPageId="14" captionText="FIGURE 12. Geographical distribution of Drymoluber Amaral, 1930. Black symbols represent localities with specimens examined, and white symbols represent literature records. Circles = Drymoluber dichrous (Peters, 1863); Triangles = Drymoluber brazili (Gomes, 1918); Lozenges = Drymoluber apurimacensis Lehr, Carrillo &amp; Hocking, 2004. Tropical and subtropical moist forests; Tropical and subtropical savannas; Deserts and xeric formations; Mangroves; Floodplains;" httpUri="https://zenodo.org/record/249307/files/figure.png" pageId="30" pageNumber="379">Fig. 12</figureCitation>
; Appendices II and III). 
<collectingCountry id="F37779DB8F26FFE9FABDFB9F4C632215" box="[1295,1371,1053,1076]" name="France" pageId="30" pageNumber="379">França</collectingCountry>
<emphasis id="B914E5598F26FFE9FAD1FB9F4CA42214" box="[1379,1436,1052,1077]" italics="true" pageId="30" pageNumber="379">et al.</emphasis>
(2006) reported the species in Vilhena, Rondônia, 
<collectingCountry id="F37779DB8F26FFE9FD61FBBD4A2F2279" box="[723,791,1087,1112]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
(-12.74, -60.15), in Amazonian savannas. However, that specimen (CHUNB 12791) is actually a 
<taxonomicName id="4C6042C88F26FFE9FDC3FBE74BCF225C" box="[625,759,1124,1149]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F26FFE9FDC3FBE74BCF225C" box="[625,759,1124,1149]" italics="true" pageId="30" pageNumber="379">D. dichrous</emphasis>
</taxonomicName>
. There is a record from Aripuanã, Mato Grosso, 
<collectingCountry id="F37779DB8F26FFE9FA88FBE64C46225C" box="[1338,1406,1124,1149]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
(- 10.15, -59.45) in the MPEG collection, but the specimen (MPEG 10419) could not be found, and we do not consider this record further. Therefore, the presence of 
<taxonomicName id="4C6042C88F26FFE9FCB2FB2F4A5322E4" box="[768,875,1196,1221]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FCB2FB2F4A5322E4" box="[768,875,1196,1221]" italics="true" pageId="30" pageNumber="379">D. brazili</emphasis>
</taxonomicName>
in the Amazonian region is unconfirmed.
</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75FB524A9A2375" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">
In their material examined section, Lehr 
<emphasis id="B914E5598F26FFE9FD3EFB534BFC22C9" box="[652,708,1231,1256]" italics="true" pageId="30" pageNumber="379">et al.</emphasis>
(2004) listed three specimens of 
<taxonomicName id="4C6042C88F26FFE9FB87FB4D4C3622C9" box="[1077,1294,1231,1256]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FB87FB4D4C3622C9" box="[1077,1294,1231,1256]" italics="true" pageId="30" pageNumber="379">Drymoluber brazili</emphasis>
</taxonomicName>
deposited in MZUSP that were misidentified: MZUSP 5329 (from Morro Branco, Ceará, 
<collectingCountry id="F37779DB8F26FFE9FBFDFB764DAD232C" box="[1103,1173,1268,1293]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
) actually is a young 
<taxonomicName id="4C6042C88F26FFE9FF25FA95484B2311" box="[151,371,1303,1328]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="corais">
<emphasis id="B914E5598F26FFE9FF25FA95484B2311" box="[151,371,1303,1328]" italics="true" pageId="30" pageNumber="379">Drymarchon corais</emphasis>
</taxonomicName>
; MZUSP 11263 (from Gaúcha do Norte, Mato Grosso, 
<collectingCountry id="F37779DB8F26FFE9FC5FFA954D0B2311" box="[1005,1075,1303,1328]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
) and MZUSP 11442 (from Vila Rica, Mato Grosso, 
<collectingCountry id="F37779DB8F26FFE9FEC5FABE48852374" box="[375,445,1340,1365]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
) are specimens of the genus 
<taxonomicName id="4C6042C88F26FFE9FCB3FABE4AA32374" box="[769,923,1340,1365]" class="Reptilia" family="Colubridae" genus="Mastigodryas" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="genus">
<emphasis id="B914E5598F26FFE9FCB3FABE4AA32374" box="[769,923,1340,1365]" italics="true" pageId="30" pageNumber="379">Mastigodryas</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75FADD485D20B8" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">
The records of 
<taxonomicName id="4C6042C88F26FFE9FEC6FADD4B752359" box="[372,589,1375,1400]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FEC6FADD4B752359" box="[372,589,1375,1400]" italics="true" pageId="30" pageNumber="379">Drymoluber brazili</emphasis>
</taxonomicName>
from the Atlantic Forest, in the municipalities of Colatina (-19.54, -40.64) and Baixo Guandú (-19.52, -41.02) state of Espírito Santo, and in Aimorés (-19.50, -41.06), state of Minas Gerais, suggest relictual populations. The lower Doce River region is relatively arid with notable occurrences of rocky outcrops that resemble Caatinga areas, different from the Atlantic Forest of the nearest regions (Jackson 1978; Ribon 1995). Modelings of the Atlantic Forest range under past climatic scenarios of 6000 and 21,000 years ago suggest that much of the area south of the Doce River was not predicted to retain a large, stable forest refuge, leading to an eastward expansion of the Cerrado (Carnaval &amp; Moritz 2008). Thus, it is possible that in one of these times and benefited by the increasing of non-forested areas in southeastern 
<collectingCountry id="F37779DB8F26FFE9FB9DF9DE4D4E2054" box="[1071,1142,1628,1653]" name="Brazil" pageId="30" pageNumber="379">Brazil</collectingCountry>
, 
<taxonomicName id="4C6042C88F26FFE9FB3AF9DF4DC22054" box="[1160,1274,1628,1653]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FB3AF9DF4DC22054" box="[1160,1274,1628,1653]" italics="true" pageId="30" pageNumber="379">D. brazili</emphasis>
</taxonomicName>
expanded its distribution range.
</paragraph>
<paragraph id="8BDF394B8F26FFE9FF75F9264BD821B4" blockId="30.[151,1436,151,1941]" pageId="30" pageNumber="379">
<emphasis id="B914E5598F26FFE9FF75F92648B2209C" bold="true" box="[199,394,1700,1725]" pageId="30" pageNumber="379">Natural history:</emphasis>
<taxonomicName id="4C6042C88F26FFE9FE20F9264B54209C" box="[402,620,1700,1725]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FE20F9264B54209C" box="[402,620,1700,1725]" italics="true" pageId="30" pageNumber="379">Drymoluber brazili</emphasis>
</taxonomicName>
inhabits open areas (Rodrigues 1996; 
<collectingCountry id="F37779DB8F26FFE9FB91F9274D57209D" box="[1059,1135,1701,1724]" name="France" pageId="30" pageNumber="379">França</collectingCountry>
&amp; Araújo 2006; 
<collectingCountry id="F37779DB8F26FFE9FA81F9274C47209D" box="[1331,1407,1701,1724]" name="France" pageId="30" pageNumber="379">França</collectingCountry>
<emphasis id="B914E5598F26FFE9FA3AF927498C20C1" italics="true" pageId="30" pageNumber="379">et al.</emphasis>
2008), and seems to be absent from altered habitats (
<collectingCountry id="F37779DB8F26FFE9FCBAF94A4A6E20C1" box="[776,854,1736,1760]" name="France" pageId="30" pageNumber="379">França</collectingCountry>
&amp; Araújo 2006). Moreira 
<emphasis id="B914E5598F26FFE9FB32F94B4D8020C1" box="[1152,1208,1735,1760]" italics="true" pageId="30" pageNumber="379">et al.</emphasis>
(2009) recorded the species inhabiting termite mounds, a common behavior in Cerrado snakes. 
<taxonomicName id="4C6042C88F26FFE9FC52F96E4D812124" box="[992,1209,1772,1797]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FC52F96E4D812124" box="[992,1209,1772,1797]" italics="true" pageId="30" pageNumber="379">Drymoluber brazili</emphasis>
</taxonomicName>
has diurnal activity, and is probably terrestrial (
<collectingCountry id="F37779DB8F26FFE9FE78F8924B202109" box="[458,536,1808,1832]" name="France" pageId="30" pageNumber="379">França</collectingCountry>
&amp; Araújo 2006; 
<collectingCountry id="F37779DB8F26FFE9FD6CF8934A122109" box="[734,810,1809,1832]" name="France" pageId="30" pageNumber="379">França</collectingCountry>
<emphasis id="B914E5598F26FFE9FC80F8934A542109" box="[818,876,1807,1832]" italics="true" pageId="30" pageNumber="379">et al.</emphasis>
2008). Gomes (1918), however, observed one of the specimens from the 
<typeStatus id="54DB87E98F26FFE9FE1AF8B748E0216C" box="[424,472,1845,1869]" pageId="30" pageNumber="379">type</typeStatus>
series climbing in arboreal structures in captivity. Amaral (1978) and Pavan &amp; Dixo (2004) cited 
<taxonomicName id="4C6042C88F26FFE9FE9CF8DB48A52151" box="[302,413,1879,1904]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="379" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F26FFE9FE9CF8DB48A52151" box="[302,413,1879,1904]" italics="true" pageId="30" pageNumber="379">D. brazili</emphasis>
</taxonomicName>
as arboriculous, but they did not state whether their data were obtained from previous published references or represent field observations.
</paragraph>
<caption id="DF1F69C38F27FFE8FF25F82149CD21D7" httpUri="https://zenodo.org/record/249315/files/figure.png" pageId="31" pageNumber="380" targetBox="[353,1243,176,1942]" targetPageId="31">
<paragraph id="8BDF394B8F27FFE8FF25F82149CD21D7" blockId="31.[151,1436,1955,2038]" pageId="31" pageNumber="380">
<emphasis id="B914E5598F27FFE8FF25F82148272198" bold="true" box="[151,287,1955,1977]" pageId="31" pageNumber="380">FIGURE 21.</emphasis>
Hemipenis of 
<taxonomicName id="4C6042C88F27FFE8FE01F8214B4E2198" box="[435,630,1955,1977]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="380" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F27FFE8FE01F8214B4E2198" box="[435,630,1955,1977]" italics="true" pageId="31" pageNumber="380">Drymoluber brazili</emphasis>
</taxonomicName>
. IBSP 33668, Goiânia, Goiás, Brazil, A) Sulcate face; B) Asulcate face. MCNR 1736, 
<emphasis id="B914E5598F27FFE8FF67F84048D621F6" box="[213,494,1985,2007]" italics="true" pageId="31" pageNumber="380">Usina Hidrelétrica de Irapé</emphasis>
, Minas Gerais, Brazil, C) Sulcate face; D) Asulcate face. Scale bar = 1 cm. Photos: Henrique C. Costa.
</paragraph>
</caption>
<paragraph id="8BDF394B8F18FFD7FF75FF154AE926D6" blockId="32.[151,1436,151,968]" pageId="32" pageNumber="381">
There is a report of an unidentified 
<taxonomicName id="4C6042C88F18FFD7FDD4FF154A7C2691" box="[614,836,151,176]" class="Reptilia" family="Gymnophthalmidae" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="family">Gymnophthalmidae</taxonomicName>
as prey of 
<taxonomicName id="4C6042C88F18FFD7FC79FF154D9F2691" box="[971,1191,151,176]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FC79FF154D9F2691" box="[971,1191,151,176]" italics="true" pageId="32" pageNumber="381">Drymoluber brazili</emphasis>
</taxonomicName>
(
<collectingCountry id="F37779DB8F18FFD7FB0AFF1A4C3E2691" box="[1208,1286,152,176]" name="France" pageId="32" pageNumber="381">França</collectingCountry>
<emphasis id="B914E5598F18FFD7FABDFF1B4C722691" box="[1295,1354,151,176]" italics="true" pageId="32" pageNumber="381">et al.</emphasis>
2008). Additionally, Pavan &amp; Dixo (2004) cited anurans as the main prey of this snake, probably based on uncited literature references, since they collected only one specimen of 
<taxonomicName id="4C6042C88F18FFD7FCEDFF634AF226D9" box="[863,970,223,248]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FCEDFF634AF226D9" box="[863,970,223,248]" italics="true" pageId="32" pageNumber="381">D. brazili</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF394B8F18FFD7FF75FE874A0D2761" blockId="32.[151,1436,151,968]" pageId="32" pageNumber="381">
<collectingCountry id="F37779DB8F18FFD7FF75FE87482B273D" box="[199,275,261,284]" name="France" pageId="32" pageNumber="381">França</collectingCountry>
&amp; Araújo (2006) recorded less than five eggs per clutch of 
<taxonomicName id="4C6042C88F18FFD7FBB3FE864DD8273C" box="[1025,1248,260,285]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FBB3FE864DD8273C" box="[1025,1248,260,285]" italics="true" pageId="32" pageNumber="381">Drymoluber brazili</emphasis>
</taxonomicName>
. One specimen examined in the present study (MZUFV 780) has four eggs.
</paragraph>
<paragraph id="8BDF394B8F18FFD7FF75FECE4AF027D4" blockId="32.[151,1436,151,968]" pageId="32" pageNumber="381">
Gomes (1918) recorded some observations of the defensive behavior of 
<taxonomicName id="4C6042C88F18FFD7FC49FECE4DED2744" box="[1019,1237,332,357]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FC49FECE4DED2744" box="[1019,1237,332,357]" italics="true" pageId="32" pageNumber="381">Drymoluber brazili</emphasis>
</taxonomicName>
: “When handled, the specimens I had examined did not try to bite; however, when bothered by repeated light hits on the dorsum, they assumed a strike posture similar to 
<taxonomicName id="4C6042C88F18FFD7FDECFE174BC9278C" box="[606,753,404,429]" class="Reptilia" family="Colubridae" genus="Drymobius" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="bifossatus">
<emphasis id="B914E5598F18FFD7FDECFE174BC9278C" box="[606,753,404,429]" italics="true" pageId="32" pageNumber="381">D. bifossatus</emphasis>
</taxonomicName>
and other species of close related genera (
<taxonomicName id="4C6042C88F18FFD7FB61FE164C16278C" box="[1235,1326,404,429]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="genus">
<emphasis id="B914E5598F18FFD7FB61FE164C16278C" box="[1235,1326,404,429]" italics="true" pageId="32" pageNumber="381">Coluber</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6042C88F18FFD7FA8EFE164CAD278C" box="[1340,1429,404,429]" class="Reptilia" family="Colubridae" genus="Spilotes" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="genus">
<emphasis id="B914E5598F18FFD7FA8EFE164CAD278C" box="[1340,1429,404,429]" italics="true" pageId="32" pageNumber="381">Spilotes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6042C88F18FFD7FF25FE35480927F1" box="[151,305,439,464]" class="Squamata" family="Colubridae" genus="Herpetodryas" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="381" phylum="Chordata" rank="genus">
<emphasis id="B914E5598F18FFD7FF25FE35480927F1" box="[151,305,439,464]" italics="true" pageId="32" pageNumber="381">Herpetodryas</emphasis>
</taxonomicName>
), also rapidly shaking the tail.” (translation by the authors). Brodie III &amp; Brodie Jr. (2004) suggested that young cross-banded specimens might mimic coralsnakes (
<taxonomicName id="4C6042C88F18FFD7FCE1FE5E4A8427D4" box="[851,956,476,501]" class="Reptilia" family="Elapidae" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="family">Elapidae</taxonomicName>
).
</paragraph>
<paragraph id="8BDF394B8F18FFD7FF75FE7D4DF5247E" blockId="32.[151,1436,151,968]" pageId="32" pageNumber="381">
The long tail of 
<taxonomicName id="4C6042C88F18FFD7FE3BFE7D4B5D2439" box="[393,613,511,536]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FE3BFE7D4B5D2439" box="[393,613,511,536]" italics="true" pageId="32" pageNumber="381">Drymoluber brazili</emphasis>
</taxonomicName>
(mean 44.2% of the SVL in males and 45.7% in females), with a tail beakage ratio of 28.9% (considering only those specimens where the breakage certainly occurred before the collection) is ample evidence of pseudoautotomy (Slowinski &amp; Savage 1995), as in 
<taxonomicName id="4C6042C88F18FFD7FBF1FDCB4DFE2441" box="[1091,1222,583,608]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B914E5598F18FFD7FBF1FDCB4DFE2441" box="[1091,1222,583,608]" italics="true" pageId="32" pageNumber="381">D. dichrous</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF394B8F18FFD7FF75FDEE4D642489" blockId="32.[151,1436,151,968]" pageId="32" pageNumber="381">
Although morphometric data did not show the presence of sexual size dimorphism in 
<taxonomicName id="4C6042C88F18FFD7FB1DFDEF4C2624A4" box="[1199,1310,620,645]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FB1DFDEF4C2624A4" box="[1199,1310,620,645]" italics="true" pageId="32" pageNumber="381">D. brazili</emphasis>
</taxonomicName>
, examined males has on average longer SVL than females (mean 
<quantity id="4C9894AE8F18FFD7FD4BFD124A6E2486" box="[761,854,656,680]" metricMagnitude="-1" metricUnit="m" metricValue="8.7" pageId="32" pageNumber="381" unit="mm" value="870.0">870 mm</quantity>
<emphasis id="B914E5598F18FFD7FCEEFD134A9B2489" box="[860,931,657,680]" italics="true" pageId="32" pageNumber="381">versus</emphasis>
mean 
<quantity id="4C9894AE8F18FFD7FC5DFD0D4D692489" box="[1007,1105,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="8.559999999999999" pageId="32" pageNumber="381" unit="mm" value="856.0">856 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A6AC08F18FFD7FF75FD364A1B25E9" pageId="32" pageNumber="381" type="etymology">
<paragraph id="8BDF394B8F18FFD7FF75FD364A1B25E9" blockId="32.[151,1436,151,968]" pageId="32" pageNumber="381">
<emphasis id="B914E5598F18FFD7FF75FD36486924EC" bold="true" box="[199,337,692,717]" pageId="32" pageNumber="381">Etymology:</emphasis>
The specific name 
<taxonomicName id="4C6042C88F18FFD7FDFEFD364BAD24EC" box="[588,661,692,717]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FDFEFD364BAD24EC" box="[588,661,692,717]" italics="true" pageId="32" pageNumber="381">brazili</emphasis>
</taxonomicName>
honors Vital 
<collectingCountry id="F37779DB8F18FFD7FCF7FD364AB124EC" box="[837,905,692,717]" name="Brazil" pageId="32" pageNumber="381">Brazil</collectingCountry>
Mineiro da Campanha, well known as the discoverer of the species-specificity of antivenoms, founder of the 
<emphasis id="B914E5598F18FFD7FC34FD5B4D6A24D1" box="[902,1106,729,752]" italics="true" pageId="32" pageNumber="381">Instituto Butantan</emphasis>
and its director when Gomes (1918) described the species. Vital 
<collectingCountry id="F37779DB8F18FFD7FDAFFD7E4B592534" box="[541,609,764,789]" name="Brazil" pageId="32" pageNumber="381">Brazil</collectingCountry>
developed a system by which the 
<emphasis id="B914E5598F18FFD7FC6CFD7F4D7C2535" box="[990,1092,765,788]" italics="true" pageId="32" pageNumber="381">Butantan</emphasis>
sent antivenom kits to farmers and ranchers in exchange for live snakes that were freely shipped to the institute by the railroad lines (Adler 2007). This system provided in a few years a snake collection of thousand of specimens to 
<emphasis id="B914E5598F18FFD7FBECFCC74DFC257D" box="[1118,1220,837,860]" italics="true" pageId="32" pageNumber="381">Butantan</emphasis>
including the 
<typeStatus id="54DB87E98F18FFD7FADEFCC74CA4257C" box="[1388,1436,837,861]" pageId="32" pageNumber="381">type</typeStatus>
series of 
<taxonomicName id="4C6042C88F18FFD7FF4EFCE548EE25A1" box="[252,470,871,896]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="381" phylum="Chordata" rank="species" species="brazili">
<emphasis id="B914E5598F18FFD7FF4EFCE548EE25A1" box="[252,470,871,896]" italics="true" pageId="32" pageNumber="381">Drymoluber brazili</emphasis>
</taxonomicName>
. One disadvantage was the fact that the specific locality of collected specimens could not be certified, and they were labeled as being from the railway stations from where they were sent to the institute (see information about the 
<typeStatus id="54DB87E98F18FFD7FE76FC2D4B1E25E9" box="[452,550,943,968]" pageId="32" pageNumber="381" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54DB87E98F18FFD7FDEFFC324BF125E9" box="[605,713,944,968]" pageId="32" pageNumber="381" type="paratype">paratypes</typeStatus>
above).
</paragraph>
</subSubSection>
</treatment>
</document>