treatments-xml/data/03/C8/7F/03C87F64FFA2FF948ED64C50FB3DA5CA.xml

<document id="3440702057AFF19B4E83760B67EB8B8C" ID-CLB-Dataset="26436" ID-DOI="10.5852/ejt.2019.532" ID-GBIF-Dataset="fbea3203-a10b-4e4f-b5f3-3d1387bc84c3" ID-Zenodo-Dep="3251569" ID-ZooBank="1301752E-3FD5-4F3E-A4F3-6766D18C709A" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561102878928" checkinUser="plazi" docAuthor="Yassin, Amir, Suwalski, Arnaud &amp; Raveloson Ravaomanarivo, Lala H." docDate="2019" docId="03C87F64FFA2FF948ED64C50FB3DA5CA" docLanguage="en" docName="ejt-532_yassin_suwalski_ravaomanariva.pdf" docOrigin="European Journal of Taxonomy 532" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Drosophila (Sophophora) chocolata Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo 2019, sp. nov." docType="treatment" docUuid="F34BF7C9-1143-4093-80E5-47A239D5C86C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="16" masterDocId="FFF1071CFFACFF848F794F3BFFC0A445" masterDocTitle="Resolving the synonymy and polyphyly of the ‘ Drosophila bakoue species complex’ (Diptera: Drosophilidae: ‘ D. montium species group’) with descriptions of two new species from Madagascar" masterLastPageNumber="26" masterPageNumber="1" pageNumber="15" updateTime="1698735486195" updateUser="ExternalLinkService">
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<mods:title id="6264FB2915842CBC119C8BEC5D7BAB64">Resolving the synonymy and polyphyly of the ‘ Drosophila bakoue species complex’ (Diptera: Drosophilidae: ‘ D. montium species group’) with descriptions of two new species from Madagascar</mods:title>
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<mods:namePart id="093622A1E960AE872056EC4604924634">Yassin, Amir</mods:namePart>
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<mods:namePart id="B1A5E6C63865EA71F2AD9E1446129645">Suwalski, Arnaud</mods:namePart>
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<mods:namePart id="33868AD2AC698DE7B21A0EEFC44E8739">Raveloson Ravaomanarivo, Lala H.</mods:namePart>
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<mods:title id="52111C190F0D86D4C54F7F46E7E149B8">European Journal of Taxonomy</mods:title>
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<subSubSection id="C37B9DF9FFA2FF8A8ED64C50FB7EA7EF" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BDECE72FFA2FF8A8ED64C50FB45A7C3" blockId="14.[374,1214,875,974]" box="[431,1157,875,902]" pageId="14" pageNumber="15">
<taxonomicName id="4C61B5F1FFA2FF8A8ED64C50FBE4A7C3" ID-CoL="8BXPF" authority="Yassin &amp; David" authorityName="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo" authorityYear="2019" box="[431,1060,875,902]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chocolata" status="sp. nov." subGenus="Sophophora">
<emphasis id="B9151260FFA2FF8A8ED64C50FCA2A7C3" bold="true" box="[431,866,875,902]" pageId="14" pageNumber="15">
<emphasis id="B9151260FFA2FF8A8ED64C50FDF6A7C3" bold="true" box="[431,566,875,902]" italics="true" pageId="14" pageNumber="15">Drosophila</emphasis>
(
<emphasis id="B9151260FFA2FF8A8D3E4C50FD1BA7C3" bold="true" box="[583,731,875,902]" italics="true" pageId="14" pageNumber="15">Sophophora</emphasis>
) 
<emphasis id="B9151260FFA2FF8A8D944C50FCA2A7C3" bold="true" box="[749,866,875,902]" italics="true" pageId="14" pageNumber="15">chocolata</emphasis>
</emphasis>
Yassin &amp; David
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<taxonomicNameLabel id="A226AF1BFFA2FF8A8B524C57FB45A7C3" box="[1067,1157,876,902]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C37B9DF9FFA2FF8A8D104C88FC0AA788" box="[617,970,947,974]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BDECE72FFA2FF8A8D104C88FC0AA788" blockId="14.[374,1214,875,974]" box="[617,970,947,974]" pageId="14" pageNumber="15">
<figureCitation id="135AD2F7FFA2FF8A8D104C88FD11A78B" box="[617,721,947,974]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1813,1839]" captionStartId-1="5.[189,232,1713,1739]" captionTargetBox-0="[306,1276,844,1746]" captionTargetBox-1="[206,1387,908,1645]" captionTargetId-0="figure@4.[281,1305,819,1760]" captionTargetId-1="figure@5.[189,1400,879,1658]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 1. Bayesian phylogeny of 19 species from the ‘D. montium species group’ Da Lage, 2007 inferred from complete mitochondrial genomes. Nodes supported by 100% posterior probability (pp) are not labeled, those with pp&gt; 90% or 60% are labeled with gray and white circles, respectively. Species belonging to the ‘D. bocqueti species complex’ and the ‘D. bakoue species complex’ are highlighted in orange and green, respectively. ‘K’ and ‘B’ refer to the strains of D. aff tsacasi Bock &amp; Wheeler, 1972 from Kenya and Bioko Island, respectively." captionText-1="Fig. 2. Comparison of Bayesian phylogenies of the ‘D. seguyi species subgroup’ Yassin, 2018 inferred from mitochondrial COI (left) and nuclear Amyrel (right) genes. Nodes supported by&gt;95% posterior probability (pp) are labeled with black circles. An asterisk refers to species sequenced here, whereas two asterisks refer to the sequence of D. vulcana Graber, 1957 of Da Lage et al. (2007), which turned out to belong to D. seguyi Smart, 1945 from Kenya (see the text). Note the discrepancy in position of different geographical strains of D. bakoue Tsacas &amp; Lachaise, 1974 (in red) between the two genes. Species belonging to ‘D. bocqueti species complex’ Tsacas &amp; Lachaise, 1974, ‘D. bakoue species complex’ Rafael, 1984, ‘D. nikananu species complex’ Tsacas &amp; Chassagnard, 1992, and ‘D. megapyga species complex’ Lachaise &amp; Tsacas, 2001 are highlighted in orange, green, yellow and red, respectively." httpUri-0="https://zenodo.org/record/3251571/files/figure.png" httpUri-1="https://zenodo.org/record/3251573/files/figure.png" pageId="14" pageNumber="15">Figs 1–2</figureCitation>
, 
<figureCitation id="135AD2F7FFA2FF8A8DA44C8FFD2CA78B" box="[733,748,948,974]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="14" pageNumber="15">4</figureCitation>
G–H, 5D–F, 6G–H
</paragraph>
</subSubSection>
<subSubSection id="C37B9DF9FFA2FF8A8FC44CCEFCCAA0B6" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BDECE72FFA2FF8A8FC44CCEFEF6A04A" blockId="14.[189,310,1013,1039]" box="[189,310,1013,1039]" pageId="14" pageNumber="15">
<heading id="D096791EFFA2FF8A8FC44CCEFEF6A04A" bold="true" box="[189,310,1013,1039]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9151260FFA2FF8A8FC44CCEFEF6A04A" bold="true" box="[189,310,1013,1039]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8FC44B1EFCCAA0B6" blockId="14.[189,1399,1061,1267]" pageId="14" pageNumber="15">
Male and female body pigmentation brown, halters white, with male femurs darker on all legs (
<figureCitation id="135AD2F7FFA2FF8A8A774B1EFA92A07A" box="[1294,1362,1061,1087]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
G– H); male abdominal tergites T2–T4 entirely dark brown, T5 and T6 entirely black (
<figureCitation id="135AD2F7FFA2FF8A8BFA4B72FB21A026" box="[1155,1249,1097,1123]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="14" pageNumber="15">Fig. 4G</figureCitation>
); dorsalmost surstylus prensiseta on the same axis with remaining prensisetae (
<figureCitation id="135AD2F7FFA2FF8A8CD64B56FC31A0CD" box="[943,1009,1133,1160]" captionStart="Fig" captionStartId="10.[189,232,1918,1944]" captionTargetBox="[213,1356,343,1838]" captionTargetId="figure@10.[189,1400,300,1865]" captionTargetPageId="10" captionText="Fig. 5. Male terminalia of the ‘D. seguyi species complex’. A. D. seguyi Smart, 1945 (MNHN). B. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). D–F. D. chocolata Yassin &amp; David sp. nov. Scale bars: A–D = 50 µm; E–F = 100 µm.." httpUri="https://zenodo.org/record/3251579/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
D–E); hypandrial median process lobate; aedeagus spatulate (
<figureCitation id="135AD2F7FFA2FF8A8D7C4BAAFD46A0EE" box="[517,646,1169,1196]" captionStart="Fig" captionStartId="10.[189,232,1918,1944]" captionTargetBox="[213,1356,343,1838]" captionTargetId="figure@10.[189,1400,300,1865]" captionTargetPageId="10" captionText="Fig. 5. Male terminalia of the ‘D. seguyi species complex’. A. D. seguyi Smart, 1945 (MNHN). B. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). D–F. D. chocolata Yassin &amp; David sp. nov. Scale bars: A–D = 50 µm; E–F = 100 µm.." httpUri="https://zenodo.org/record/3251579/files/figure.png" pageId="14" pageNumber="15">Fig. 5D, F</figureCitation>
); female T2–T7 entirely dark brown (
<figureCitation id="135AD2F7FFA2FF8A8B354BAAFB6DA0EE" box="[1100,1197,1169,1195]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="14" pageNumber="15">Fig. 4H</figureCitation>
); oviscapt fourth posterior peg-like outer ovisensillum not on the same axis with the third and fifth ovisensilla (
<figureCitation id="135AD2F7FFA2FF8A8A734B8EFAA9A095" box="[1290,1385,1205,1232]" captionStart="Fig" captionStartId="11.[189,232,1889,1915]" captionTargetBox="[232,1361,297,1817]" captionTargetId="figure@11.[204,1383,265,1834]" captionTargetPageId="11" captionText="Fig. 6. Female oviscapts of the ‘D. seguyi species complex’. A–B. D. seguyi Smart, 1945 (MNHN). C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). Scale bars: A, C, E, G = 100 µm; B, D, F, H = 50 µm." httpUri="https://zenodo.org/record/3251581/files/figure.png" pageId="14" pageNumber="15">Fig. 6G</figureCitation>
), with anterior ovisensilla short and thick (
<figureCitation id="135AD2F7FFA2FF8A8DE74BE2FD3DA0B6" box="[670,765,1240,1267]" captionStart="Fig" captionStartId="11.[189,232,1889,1915]" captionTargetBox="[232,1361,297,1817]" captionTargetId="figure@11.[204,1383,265,1834]" captionTargetPageId="11" captionText="Fig. 6. Female oviscapts of the ‘D. seguyi species complex’. A–B. D. seguyi Smart, 1945 (MNHN). C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). Scale bars: A, C, E, G = 100 µm; B, D, F, H = 50 µm." httpUri="https://zenodo.org/record/3251581/files/figure.png" pageId="14" pageNumber="15">Fig. 6H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37B9DF9FFA2FF8A8FC44A20FE34A121" box="[189,500,1307,1381]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BDECE72FFA2FF8A8FC44A20FEFEA170" blockId="14.[189,318,1307,1333]" box="[189,318,1307,1333]" pageId="14" pageNumber="15">
<heading id="D096791EFFA2FF8A8FC44A20FEFEA170" bold="true" box="[189,318,1307,1333]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9151260FFA2FF8A8FC44A20FEFEA170" bold="true" box="[189,318,1307,1333]" pageId="14" pageNumber="15">Etimology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8FC44A70FE34A121" blockId="14.[189,500,1354,1381]" box="[189,500,1354,1381]" pageId="14" pageNumber="15">In reference to body color.</paragraph>
</subSubSection>
<subSubSection id="C37B9DF9FFA2FF8A8FC44AB7FB97A2AF" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BDECE72FFA2FF8A8FC44AB7FEACA1E3" blockId="14.[189,364,1420,1446]" box="[189,364,1420,1446]" pageId="14" pageNumber="15">
<heading id="D096791EFFA2FF8A8FC44AB7FEACA1E3" bold="true" box="[189,364,1420,1446]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9151260FFA2FF8A8FC44AB7FEACA1E3" bold="true" box="[189,364,1420,1446]" pageId="14" pageNumber="15">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8F934A87FE9CA193" blockId="14.[189,1399,1468,1566]" box="[234,348,1468,1494]" pageId="14" pageNumber="15">
<typeStatus id="54DA70D0FFA2FF8A8F934A87FE9CA193" box="[234,348,1468,1494]" pageId="14" pageNumber="15" type="holotype">
<emphasis id="B9151260FFA2FF8A8F934A87FE9CA193" bold="true" box="[234,348,1468,1494]" pageId="14" pageNumber="15">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8FC44ADBFDB1A25B" blockId="14.[189,1399,1468,1566]" pageId="14" pageNumber="15">
<materialsCitation id="3B09C42FFFA2FF8A8FC44ADBFDB1A25B" ID-GBIF-Occurrence="2269035521" collectingDate="2008-02-16" collectingDateMax="2014-07" collectingDateMin="2008-02-16" collectionCode="MNHN" collectorName="J. R. David &amp; A. Yassin" country="Madagascar" latitude="-17.333334" location="Andasibe" longLatPrecision="1279" longitude="48.9" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F3768EE2FFA2FF8A8FC44ADBFE4EA1BF" box="[189,398,1504,1530]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
• 
<specimenCount id="9D6705FBFFA2FF8A8EDE4ADBFE00A1BF" box="[423,448,1504,1530]" pageId="14" pageNumber="15">♂</specimenCount>
; 
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<geoCoordinate id="EE55A8B5FFA2FF8A8D304AE5FD70A1BF" box="[585,688,1502,1530]" degrees="17" direction="south" minutes="20" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-17.333334">17°20′ S</geoCoordinate>
, 
<geoCoordinate id="EE55A8B5FFA2FF8A8DC44AE5FCE6A1BF" box="[701,806,1502,1530]" degrees="48" direction="east" minutes="54" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="48.9">48°54′ E</geoCoordinate>
; 
<collectingDate id="EF9B115AFFA2FF8A8C4A4AE4FC3AA1BF" box="[819,1018,1503,1530]" pageId="14" pageNumber="15" value="2008-02-16" valueMax="2008-02-17" valueMin="2008-02-16">
<date id="FFDFE8B2FFA2FF8A8C4A4AE4FC3AA1BF" box="[819,1018,1503,1530]" pageId="14" pageNumber="15" value="2008-02-16" valueMax="2008-02-17" valueMin="2008-02-16">16–17 Feb. 2008</date>
</collectingDate>
(ex-laboratory strain 
<collectingDate id="EF9B115AFFA2FF8A8B804AE4FAAAA1BF" box="[1273,1386,1503,1530]" pageId="14" pageNumber="15" value="2014-07">
<date id="FFDFE8B2FFA2FF8A8B804AE4FAAAA1BF" box="[1273,1386,1503,1530]" pageId="14" pageNumber="15" value="2014-07">Jul. 2014</date>
</collectingDate>
); 
<collectorName id="2694ABA4FFA2FF8A8FC4493FFEFAA258" box="[189,314,1539,1566]" pageId="14" pageNumber="15">J.R. David</collectorName>
&amp; 
<collectorName id="2694ABA4FFA2FF8A8E27493FFE0DA258" box="[350,461,1539,1566]" pageId="14" pageNumber="15">A. Yassin</collectorName>
leg.; 
<collectionCode id="ED7056B7FFA2FF8A8D76493FFDADA25B" box="[527,621,1540,1566]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8F93497DFEA7A225" blockId="14.[189,1135,1606,1668]" box="[234,359,1606,1632]" pageId="14" pageNumber="15">
<typeStatus id="54DA70D0FFA2FF8A8F93497DFEA7A225" box="[234,359,1606,1632]" pageId="14" pageNumber="15" type="paratype">
<emphasis id="B9151260FFA2FF8A8F93497DFEA7A225" bold="true" box="[234,359,1606,1632]" pageId="14" pageNumber="15">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8FC44951FBAFA2C1" blockId="14.[189,1135,1606,1668]" box="[189,1135,1641,1668]" pageId="14" pageNumber="15">
<materialsCitation id="3B09C42FFFA2FF8A8FC44951FBACA2C1" ID-GBIF-Occurrence="2269035523" box="[189,1132,1641,1668]" collectingDate="2008-02-16" collectingDateMax="2014-07" collectingDateMin="2008-02-16" collectionCode="MNHN" collectorName="J. R. David &amp; A. Yassin" country="Madagascar" latitude="-17.333334" location="Andasibe" longLatPrecision="1279" longitude="48.9" pageId="14" pageNumber="15" specimenCount="19" specimenCount-female="10" specimenCount-male="9" typeStatus="paratype">
<collectingCountry id="F3768EE2FFA2FF8A8FC44951FE4EA2C1" box="[189,398,1642,1668]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
• 
<specimenCount id="9D6705FBFFA2FF8A8EDF4951FE2AA2C1" box="[422,490,1642,1668]" pageId="14" pageNumber="15" type="male">9 ♂♂</specimenCount>
, 
<specimenCount id="9D6705FBFFA2FF8A8E8E4951FD89A2C1" box="[503,585,1642,1668]" pageId="14" pageNumber="15" type="female">10 ♀♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED7056B7FFA2FF8A8B774951FBACA2C1" box="[1038,1132,1642,1668]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8F934990FE66A280" blockId="14.[189,1111,1707,1770]" box="[234,422,1707,1733]" pageId="14" pageNumber="15">
<emphasis id="B9151260FFA2FF8A8F934990FE66A280" bold="true" box="[234,422,1707,1733]" pageId="14" pageNumber="15">Other material</emphasis>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8FC449EBFB97A2AF" blockId="14.[189,1111,1707,1770]" box="[189,1111,1743,1770]" pageId="14" pageNumber="15">
<materialsCitation id="3B09C42FFFA2FF8A8FC449EBFB93A2AF" ID-GBIF-Occurrence="2269035524" box="[189,1107,1743,1770]" collectingDate="2008-02-16" collectingDateMax="2014-07" collectingDateMin="2008-02-16" collectionCode="ZUAC" collectorName="J. R. David &amp; A. Yassin" country="Madagascar" latitude="-17.333334" location="Andasibe" longLatPrecision="1279" longitude="48.9" pageId="14" pageNumber="15" specimenCount="7" specimenCount-female="2" specimenCount-male="5">
<collectingCountry id="F3768EE2FFA2FF8A8FC449EBFE4EA2AF" box="[189,398,1744,1770]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
• 
<specimenCount id="9D6705FBFFA2FF8A8EDF49F4FE2AA2AF" box="[422,490,1743,1770]" pageId="14" pageNumber="15" type="male">5 ♂♂</specimenCount>
, 
<specimenCount id="9D6705FBFFA2FF8A8E8E49EBFDFAA2AF" box="[503,570,1744,1770]" pageId="14" pageNumber="15" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED7056B7FFA2FF8A8C8649EBFB93A2AF" box="[1023,1107,1744,1770]" country="Madagascar" httpUri="http://grbio.org/cool/dprk-p82z" name="University of Antananarivo" pageId="14" pageNumber="15">ZUAC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B9DF9FFA2FF8B8FC4482AFAB7A1C1" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="8BDECE72FFA2FF8A8FC4482AFE8EA36E" blockId="14.[189,334,1809,1835]" box="[189,334,1809,1835]" pageId="14" pageNumber="15">
<heading id="D096791EFFA2FF8A8FC4482AFE8EA36E" bold="true" box="[189,334,1809,1835]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9151260FFA2FF8A8FC4482AFE8EA36E" bold="true" box="[189,334,1809,1835]" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDECE72FFA2FF8A8F93487AFEEAA31E" blockId="14.[189,1399,1857,2027]" box="[234,298,1857,1883]" pageId="14" pageNumber="15">
<emphasis id="B9151260FFA2FF8A8F93487AFEEAA31E" bold="true" box="[234,298,1857,1883]" pageId="14" pageNumber="15">Male</emphasis>
</paragraph>
<paragraph id="8BDECE72FFA2FF8B8FC4485EFDF2A528" blockId="14.[189,1399,1857,2027]" lastBlockId="15.[189,1399,267,365]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
HEAD (
<figureCitation id="135AD2F7FFA2FF8A8E75485EFEAAA33A" box="[268,362,1893,1919]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="14" pageNumber="15">Fig. 4G</figureCitation>
). Frons brown, frontal length 
<quantity id="4C996397FFA2FF8A8DBA485EFCEEA33A" box="[707,814,1893,1919]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="15" unit="mm" value="0.38">0.38 mm</quantity>
; frontal index = 1.00, frontal tapering ratio = 1.47. Frontal triangle concolorous; ocellar triangle slightly darker, about 40% of frontal length. Orbital plates shining, apically slightly diverging from eye margin, about 87% of frontal length. Orbital setae black, distance of or3 to or1 = 67% of or3 to vtm, or1/or3 ratio = 1.29, or2/or1 ratio = 0.33, postocellar setae = 73%, ocellar setae = 47%, vibrissal index = 1.00. Face grayish. Carina flat. Cheek index about 12.50. Eye dark red, eye index = 1.39. Antennae dark brown. Arista with five dorsal, two ventral branches, plus terminal fork. Proboscis brown.
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44EADFDD8A6C3" blockId="15.[189,1399,405,647]" pageId="15" pageNumber="16">
THORAX (
<figureCitation id="135AD2F7FFA3FF8B8E544EAEFE51A5EA" box="[301,401,405,431]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="15" pageNumber="16">Fig. 4G</figureCitation>
). Length 
<quantity id="4C996397FFA3FF8B8D704EAEFDABA5F5" box="[521,619,405,432]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="16" unit="mm" value="1.2">1.2 mm</quantity>
. Scutum brown, shining, darker before scutellum and having a darker median stripe on dorsocentral region, six rows of acrostichal setulae. H index = 0.50. Transverse distance of dorsocentral setae 160% of longitudinal distance; dc index = 0.88. Scutellum dark brown; scut index = 0.83. Pleura dark brown, shining. Legs dark brown, sex combs on protarsomeres 1 and 2, with about 21 and 16 peg-like setae, respectively. Wing hyaline, veins reddish, length 
<quantity id="4C996397FFA3FF8B8BCC4D1FFAE1A67A" box="[1205,1313,548,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.02" pageId="15" pageNumber="16" unit="mm" value="2.02">2.02 mm</quantity>
, length to width ratio = 2.18. Indices: C = 2.20, ac = 2.46, hb = 0.52, 4C = 1.88, 
<date id="FFDFE8B2FFA3FF8B8B704D73FBE7A627" box="[1033,1063,584,610]" pageId="15" pageNumber="16">4v</date>
= 3.68, 
<date id="FFDFE8B2FFA3FF8B8BFF4D73FB64A626" box="[1158,1188,584,611]" pageId="15" pageNumber="16">5x</date>
= 0.61, M = 1.38, prox. x = 0.94. Haltere white.
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44D95FB41A68D" blockId="15.[189,1153,686,713]" box="[189,1153,686,713]" pageId="15" pageNumber="16">
ABDOMEN (
<figureCitation id="135AD2F7FFA3FF8B8E424D95FE5AA68D" box="[315,410,686,712]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="15" pageNumber="16">Fig. 4G</figureCitation>
). Entirely brown, shining, tergites T5 and T6 completely black.
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44DCAFDA3A009" blockId="15.[189,1399,752,1100]" pageId="15" pageNumber="16">
TERMINALIA (
<figureCitation id="135AD2F7FFA3FF8B8E2D4DCBFE68A74E" box="[340,424,752,779]" captionStart="Fig" captionStartId="10.[189,232,1918,1944]" captionTargetBox="[213,1356,343,1838]" captionTargetId="figure@10.[189,1400,300,1865]" captionTargetPageId="10" captionText="Fig. 5. Male terminalia of the ‘D. seguyi species complex’. A. D. seguyi Smart, 1945 (MNHN). B. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). D–F. D. chocolata Yassin &amp; David sp. nov. Scale bars: A–D = 50 µm; E–F = 100 µm.." httpUri="https://zenodo.org/record/3251579/files/figure.png" pageId="15" pageNumber="16">Figs. 5</figureCitation>
D–F). Epandrium black, with six setae, the lower most being particularly long; epandrial ventral lobe black with eight bristles. Cercus black; cercal ventral lobe yellow, partially separated from cercus, with a series of three strong, curved spines on the inner margin, and smaller spines along the ventral, outer and dorsal margins, larger dorsally. Surstylus with a regular row of five short, stout peg-like prensisetae, and a ventromedial cluster of prensisetae, the innermost pointing dorsally. Hypandrium black anteriorly, dark posteriorly, as long as broad, with a lobate medial posterior extension bearing two short, divergent thick bristles; posterior margin microtrichose with long fine hairs. Outer paraphyses large, S-curved, transverse, bearing three minute setulae. Inner paraphyses as long as aedeagus, swollen medially, broad and lobate apically. Aedeagus broad, hirsute, subapically narrowed. Aedeagal apodeme black anteriorly.
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8F934B4EFE85A0CA" blockId="15.[189,1009,1141,1203]" box="[234,325,1141,1167]" pageId="15" pageNumber="16">
<emphasis id="B9151260FFA3FF8B8F934B4EFE85A0CA" bold="true" box="[234,325,1141,1167]" pageId="15" pageNumber="16">Female</emphasis>
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44BA2FC31A0F7" blockId="15.[189,1009,1141,1203]" box="[189,1009,1176,1203]" pageId="15" pageNumber="16">
HABITUS (
<figureCitation id="135AD2F7FFA3FF8B8E554BA3FE4BA0F6" box="[300,395,1176,1203]" captionStart="Fig" captionStartId="9.[189,232,1889,1915]" captionTargetBox="[253,1334,292,1806]" captionTargetId="figure@9.[228,1359,267,1833]" captionTargetPageId="9" captionText="Fig. 4. Adult habitus. A–B. D. seguyi Smart, 1945 (MNHN). A. ♂. B. ♀. C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). C. ♂. D. ♀. E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). E. ♂. F. ♀. G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). G. ♂. H. ♀. Only dark morphs of females of species with female-limited color dimorphism are shown. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251577/files/figure.png" pageId="15" pageNumber="16">Fig. 4H</figureCitation>
). Similar to male but with no sex combs on protarsi.
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44BE0FAB7A1C1" blockId="15.[189,1399,1242,1412]" pageId="15" pageNumber="16">
TERMINALIA (
<figureCitation id="135AD2F7FFA3FF8B8E2D4BE1FE5CA0B0" box="[340,412,1242,1269]" captionStart="Fig" captionStartId="11.[189,232,1889,1915]" captionTargetBox="[232,1361,297,1817]" captionTargetId="figure@11.[204,1383,265,1834]" captionTargetPageId="11" captionText="Fig. 6. Female oviscapts of the ‘D. seguyi species complex’. A–B. D. seguyi Smart, 1945 (MNHN). C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). Scale bars: A, C, E, G = 100 µm; B, D, F, H = 50 µm." httpUri="https://zenodo.org/record/3251581/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
G–H). Valve of oviscapt mediodorsally mostly membranous, posteriorly rounded, ventrally slightly concave, with no discal and twelve marginal, peg-like, pointed-tipped, short and thick ovisensilla on the outer surface and one long, straight, subterminal and three tiny (microscopic) trichoid-like ovisensilla on the inner surface. The fourth peg-like ovisensillum characteristically dorsally positioned in respect to the main axis of ovisensilla insertion on the ventral margin of the oviscapt valve.
</paragraph>
</subSubSection>
<subSubSection id="C37B9DF9FFA3FF8B8FC44A97FE11A1B3" box="[189,465,1452,1526]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BDECE72FFA3FF8B8FC44A97FE96A183" blockId="15.[189,342,1452,1478]" box="[189,342,1452,1478]" pageId="15" pageNumber="16">
<heading id="D096791EFFA3FF8B8FC44A97FE96A183" bold="true" box="[189,342,1452,1478]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B9151260FFA3FF8B8FC44A97FE96A183" bold="true" box="[189,342,1452,1478]" pageId="15" pageNumber="16">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44AE0FE11A1B3" blockId="15.[189,465,1499,1526]" box="[189,465,1499,1526]" pageId="15" pageNumber="16">
<collectingCountry id="F3768EE2FFA3FF8B8FC44AE0FE8BA1B0" box="[189,331,1499,1525]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
(endemic).
</paragraph>
</subSubSection>
<subSubSection id="C37B9DF9FFA3FF948FC44926FB3DA5CA" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BDECE72FFA3FF8B8FC44926FEEFA272" blockId="15.[189,303,1565,1591]" box="[189,303,1565,1591]" pageId="15" pageNumber="16">
<heading id="D096791EFFA3FF8B8FC44926FEEFA272" bold="true" box="[189,303,1565,1591]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B9151260FFA3FF8B8FC44926FEEFA272" bold="true" box="[189,303,1565,1591]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDECE72FFA3FF8B8FC44977FC10A378" blockId="15.[189,1399,1612,1854]" pageId="15" pageNumber="16">
Specimens of this species were collected from different localities in 
<collectingCountry id="F3768EE2FFA3FF8B8CB44977FB9FA223" box="[973,1119,1612,1638]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
:Andasibe (
<quantity id="4C996397FFA3FF8B8B9A4977FAB7A222" box="[1251,1399,1612,1639]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="15" pageNumber="16" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">800–1200 m</quantity>
a.s.l.), 
<collectingRegion id="49A50090FFA3FF8B8E75494BFE6CA2CF" box="[268,428,1648,1674]" country="Madagascar" name="Antananarivo" pageId="15" pageNumber="16">Antananarivo</collectingRegion>
(
<quantity id="4C996397FFA3FF8B8EC4494BFDDAA2CE" box="[445,538,1648,1675]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="16" unit="m" value="1300.0">1300 m</quantity>
a.s.l.) and Mandraka (
<quantity id="4C996397FFA3FF8B8C51494BFC44A2CE" box="[808,900,1648,1675]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="16" unit="m" value="1400.0">1400 m</quantity>
a.s.l.) during the 2008 expedition by J.R. David and A. Yassin. It was also collected from Ranomafana (
<quantity id="4C996397FFA3FF8B8CDC49AFFC33A2EB" box="[933,1011,1684,1710]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="m" value="600.0">600 m</quantity>
a.s.l.) in 2009 by J.R. David, V. Debat and A. Yassin. This indicates that 
<taxonomicName id="4C61B5F1FFA3FF8B8DE44982FCF6A297" authority="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo, 2019" authorityName="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo" authorityYear="2019" box="[669,822,1720,1746]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chocolata" status="sp. nov.">
<emphasis id="B9151260FFA3FF8B8DE44982FCF6A297" box="[669,822,1720,1746]" italics="true" pageId="15" pageNumber="16">D. chocolata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A226AF1BFFA3FF8B8C444983FC59A297" box="[829,921,1720,1746]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
is widespread and that, unlike 
<taxonomicName id="4C61B5F1FFA3FF8B8A734982FE96A2B3" authority="Tsacas, 1984" authorityName="Tsacas" authorityYear="1984" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ifestia">
<emphasis id="B9151260FFA3FF8B8A734982FAB7A297" box="[1290,1399,1720,1746]" italics="true" pageId="15" pageNumber="16">D. ifestia</emphasis>
<bibRefCitation id="EFF0B383FFA3FF8B8FC449E7FE96A2B3" author="Tsacas L." box="[189,342,1756,1782]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="15" pageNumber="16" pagination="419 - 438" part="20" refId="ref13148" refString="Tsacas L. 1984. Nouvelles donnees sur la biogeographie et l'evolution du groupe Drosophila melanogaster en Afrique. Description de six nouvelles especes (Diptera, Drosophilidae). Annales de la Societe entomologique de France 20: 419 - 438." title="Nouvelles donnees sur la biogeographie et l'evolution du groupe Drosophila melanogaster en Afrique. Description de six nouvelles especes (Diptera, Drosophilidae)" type="journal article" year="1984">Tsacas, 1984</bibRefCitation>
</taxonomicName>
, is not mountainous. The species can be maintained on a ‘standard 
<taxonomicName id="4C61B5F1FFA3FF8B8B1F49E0FB2BA2B0" authorityName="Fallen" authorityYear="1823" box="[1126,1259,1755,1781]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B9151260FFA3FF8B8B1F49E0FB2BA2B0" box="[1126,1259,1755,1781]" italics="true" pageId="15" pageNumber="16">Drosophila</emphasis>
</taxonomicName>
medium’ in the laboratory.A mutant strain from 
<collectingRegion id="49A50090FFA3FF8B8D1949C4FCC0A35C" box="[608,768,1791,1817]" country="Madagascar" name="Antananarivo" pageId="15" pageNumber="16">Antananarivo</collectingRegion>
was established having a light abdomen in both sexes, but both the mutant and the original strains were subsequently lost.
</paragraph>
<paragraph id="8BDECE72FFA3FF948FC4485EFB3DA5CA" blockId="15.[189,1399,1893,2028]" lastBlockId="16.[189,1399,267,399]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
The species resembles 
<taxonomicName id="4C61B5F1FFA3FF8B8EB7485DFDFBA33A" authorityName="Tsacas" authorityYear="1984" box="[462,571,1893,1919]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ifestia">
<emphasis id="B9151260FFA3FF8B8EB7485DFDFBA33A" box="[462,571,1893,1919]" italics="true" pageId="15" pageNumber="16">D. ifestia</emphasis>
</taxonomicName>
, a species endemic to high mountains in East Africa, in males having entirely dark abdomen. However, 
<taxonomicName id="4C61B5F1FFA3FF8B8D3448B1FD78A3E6" authorityName="Tsacas" authorityYear="1984" box="[589,696,1929,1955]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ifestia">
<emphasis id="B9151260FFA3FF8B8D3448B1FD78A3E6" box="[589,696,1929,1955]" italics="true" pageId="15" pageNumber="16">D. ifestia</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C61B5F1FFA3FF8B8C2A48B1FC2BA3E6" authority="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo, 2019" authorityName="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo" authorityYear="2019" box="[851,1003,1929,1955]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chocolata" status="sp. nov.">
<emphasis id="B9151260FFA3FF8B8C2A48B1FC2BA3E6" box="[851,1003,1929,1955]" italics="true" pageId="15" pageNumber="16">D. chocolata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A226AF1BFFA3FF8B8C8B48B1FB8DA3E1" box="[1010,1101,1930,1956]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
in the color of the thorax being lighter with the halters reddish brown (
<figureCitation id="135AD2F7FFA3FF8B8D9E4896FCF0A382" box="[743,816,1965,1991]" captionStart="Fig" captionStartId="16.[189,232,1958,1984]" captionTargetBox="[198,1390,1336,1891]" captionTargetId="figure@16.[189,1398,1322,1902]" captionTargetPageId="16" captionText="Fig. 7. Photomicrographs of D. ifestia Tsacas, 1984. A. Holotype (P. Vanschuytbroeck and J. Kelenbosch leg.; MNHN). B. Paratype (P. Vanschuytbroeck and J. Kelenbosch leg.; MNHN). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3251583/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
A–B), which are white in 
<taxonomicName id="4C61B5F1FFA3FF8B8B0A4895FACEA382" authority="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo, 2019" authorityName="Yassin &amp; Suwalski &amp; Raveloson Ravaomanarivo" authorityYear="2019" box="[1139,1294,1965,1991]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chocolata" status="sp. nov.">
<emphasis id="B9151260FFA3FF8B8B0A4895FACEA382" box="[1139,1294,1965,1991]" italics="true" pageId="15" pageNumber="16">D. chocolata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A226AF1BFFA3FF8B8A604896FAB7A382" box="[1305,1399,1965,1991]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="135AD2F7FFA3FF8B8FBF48EAFECEA3A9" box="[198,270,2001,2028]" captionStart="Fig" captionStartId="11.[189,232,1889,1915]" captionTargetBox="[232,1361,297,1817]" captionTargetId="figure@11.[204,1383,265,1834]" captionTargetPageId="11" captionText="Fig. 6. Female oviscapts of the ‘D. seguyi species complex’. A–B. D. seguyi Smart, 1945 (MNHN). C–D. D. malagassya Tsacas &amp; Rafael, 1982 (MNHN). E–F. D. curta Chassagnard &amp; Tsacas, 1997 (MNHN). G–H. D. chocolata Yassin &amp; David sp. nov. (MNHN). Scale bars: A, C, E, G = 100 µm; B, D, F, H = 50 µm." httpUri="https://zenodo.org/record/3251581/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
A–B), in the lack of long innermost prensiseta on the surstylus (present in all species of the ‘ 
<taxonomicName id="4C61B5F1FFBCFF948FBE4E37FEF4A560" box="[199,308,268,293]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="seguyi">
<emphasis id="B9151260FFBCFF948FBE4E37FEF4A560" box="[199,308,268,293]" italics="true" pageId="16" pageNumber="17">D. seguyi</emphasis>
</taxonomicName>
species complex’), and in the shape of the hypandrial median process being broadly truncated in 
<taxonomicName id="4C61B5F1FFBCFF948FA34E14FE84A50C" authorityName="Tsacas" authorityYear="1984" box="[218,324,303,329]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ifestia">
<emphasis id="B9151260FFBCFF948FA34E14FE84A50C" box="[218,324,303,329]" italics="true" pageId="16" pageNumber="17">D. ifestia</emphasis>
</taxonomicName>
with the hypandrial bristles extending in parallel to each other (divergent in all species of the ‘ 
<taxonomicName id="4C61B5F1FFBCFF948FBE4E68FEF6A529" box="[199,310,339,364]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="seguyi">
<emphasis id="B9151260FFBCFF948FBE4E68FEF6A529" box="[199,310,339,364]" italics="true" pageId="16" pageNumber="17">D. seguyi</emphasis>
</taxonomicName>
species complex’). We therefore concur with the conclusion of 
<bibRefCitation id="EFF0B383FFBCFF948B534E69FB0EA529" author="Tsacas L." box="[1066,1230,338,364]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="16" pageNumber="17" pagination="419 - 438" part="20" refId="ref13148" refString="Tsacas L. 1984. Nouvelles donnees sur la biogeographie et l'evolution du groupe Drosophila melanogaster en Afrique. Description de six nouvelles especes (Diptera, Drosophilidae). Annales de la Societe entomologique de France 20: 419 - 438." title="Nouvelles donnees sur la biogeographie et l'evolution du groupe Drosophila melanogaster en Afrique. Description de six nouvelles especes (Diptera, Drosophilidae)" type="journal article" year="1984">Tsacas (1984)</bibRefCitation>
that 
<taxonomicName id="4C61B5F1FFBCFF948A734E68FAB7A529" authorityName="Tsacas" authorityYear="1984" box="[1290,1399,338,364]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ifestia">
<emphasis id="B9151260FFBCFF948A734E68FAB7A529" box="[1290,1399,338,364]" italics="true" pageId="16" pageNumber="17">D. ifestia</emphasis>
</taxonomicName>
does not belong to any of the defined species complexes of the ‘ 
<taxonomicName id="4C61B5F1FFBCFF948CD54E4DFBDBA5CA" box="[940,1051,374,399]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="seguyi">
<emphasis id="B9151260FFBCFF948CD54E4DFBDBA5CA" box="[940,1051,374,399]" italics="true" pageId="16" pageNumber="17">D. seguyi</emphasis>
</taxonomicName>
species subgroup’.
</paragraph>
</subSubSection>
</treatment>
</document>