1068 lines
143 KiB
XML
1068 lines
143 KiB
XML
<document id="0F4925E4B0DA939FB125F6FD58CB2DC5" ID-DOI="10.11646/zootaxa.5228.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7532470" ID-ZooBank="F5D13127-0304-46C1-8E67-32B5DE69A569" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673571880956" checkinUser="plazi" docAuthor="Balete, Danilo S., Heaney, Lawrence R., Rickart, Eric A., Quidlat, Roselyn S., Rowsey, Dakota M. & Olson, Link E." docDate="2023" docId="03C66E4E3A229769FF24FF074C004134" docLanguage="en" docName="zootaxa.5228.3.2.pdf" docOrigin="Zootaxa 5228 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Podogymnura intermedia Balete & Heaney & Rickart & Quidlat & Rowsey & Olson 2023, new species" docType="treatment" docUuid="FF6BC75B-C222-4A1E-8C88-97EC73A7D098" docUuidSource="ZooBank" docVersion="5" lastPageNumber="262" masterDocId="FFFF16363A2C977BFFB3FF904847444D" masterDocTitle="A re-assessment of diversity among Philippine gymnures (Mammalia: Erinaceidae Podogymnura), with a new species from eastern Mindanao" masterLastPageNumber="266" masterPageNumber="244" pageNumber="258" updateTime="1699446712460" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="EBA95F925B351FB068CBC979B03E2E04">
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<mods:title id="7C953FD2C9868DA4C47D98A69A8DDB7A">A re-assessment of diversity among Philippine gymnures (Mammalia: Erinaceidae Podogymnura), with a new species from eastern Mindanao</mods:title>
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<mods:name id="B870CFD81636B029582699CFA8AEBFAA" type="personal">
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<mods:roleTerm id="50982AB7DFDFD3E71767C171108A51A4">Author</mods:roleTerm>
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<mods:namePart id="9557E8260A8343203483B19799EEC593">Balete, Danilo S.</mods:namePart>
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<mods:affiliation id="D702AEA7B1C2F3A9139423892876DC3A">Field Museum of Natural History, 1400 S DuSable Lake Shore Drive, Chicago, IL 60605, USA.</mods:affiliation>
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</mods:name>
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<mods:name id="959C4773827BC980E57899EEA8844C81" type="personal">
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<mods:role id="FA1D7E1D7E434E803216E61B646736DE">
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<mods:roleTerm id="1E7057AE7F05F20FF8443DC91C113D7E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9FA3A8634CAA27ACBAB5FBBAF3CCC2B4">Heaney, Lawrence R.</mods:namePart>
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<mods:affiliation id="7490E2187DCFDCED965D2D26B23166EE">Field Museum of Natural History, 1400 S DuSable Lake Shore Drive, Chicago, IL 60605, USA.</mods:affiliation>
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</mods:name>
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<mods:name id="49855ECAACE86AF1E8E9CAD6D9B1FEC5" type="personal">
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<mods:roleTerm id="C4F8132CFC334C5CC59E7F9D9DE9DAF1">Author</mods:roleTerm>
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<mods:namePart id="D8AE03886930FE34DB60213AC7C2B368">Rickart, Eric A.</mods:namePart>
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<mods:name id="F4A17CC33356A8812A50D56CE045DE8C" type="personal">
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<mods:roleTerm id="C6ECC05237CE2A280E9464DA56C177B1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3952258208F82FEDAB27899EE7F9E679">Quidlat, Roselyn S.</mods:namePart>
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<mods:affiliation id="A98638A7E8A77A7180FCC350FFBD084E">Philippine Eagle Foundation, Malagos, Baguio District, Davao City 8000, Philippines. & National Housing Authority, Regional Office XI, Davao City 8000, Philippines.</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="BB527E4B231AF5268ABE53D66E16C0DB">Rowsey, Dakota M.</mods:namePart>
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<mods:affiliation id="BC8AD204ED7D9C4BD0244ECFA56E82B4">Arizona State University School of Life Sciences, PO Box 87108, Tempe, AZ 85287.</mods:affiliation>
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<mods:roleTerm id="57EDF45058A45853886E976CE76A227F">Author</mods:roleTerm>
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<mods:namePart id="5FFF3EEE8D186D426EAD79E0EB051985">Olson, Link E.</mods:namePart>
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<mods:affiliation id="7B27EFD1C93CC9813E32C8B6117543D9">University of Alaska Museum, 1962 Yukon Drive, Fairbanks, AK 99775, USA.</mods:affiliation>
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<mods:typeOfResource id="D402C6DFF00340350157B4ECDC65894A">text</mods:typeOfResource>
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<mods:title id="C62E8F97471F47B29A14CBA27174F000">Zootaxa</mods:title>
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<mods:part id="1EECE14EF8DA1E05B27C15E2F54E27D3">
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<mods:date id="30AF22A445CEF766A0516D6B5F21C170">2023</mods:date>
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<mods:number id="407B02775122FD73FE7F07D75AEADFE0">2023-01-13</mods:number>
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<mods:detail id="C64AA2C0D0182E84C04F13CDC12FFFEB" type="volume">
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<mods:number id="706920A8DB0B7CB3D8378544977FADE4">5228</mods:number>
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<mods:detail id="084AB034549F5B6D45E0ED75E9FE1AFD" type="issue">
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<mods:number id="B1FF7DF33324AFC4E7DC76BBEF5BAFA2">3</mods:number>
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<mods:start id="6C9C8CA8BEBE9C9C0E367C82C534E250">244</mods:start>
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<mods:classification id="D6C04C6C91B113903A7029370D62225C">journal article</mods:classification>
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<mods:identifier id="70451370E2C4DA8DC10C4F241655EA60" type="DOI">10.11646/zootaxa.5228.3.2</mods:identifier>
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<mods:identifier id="7A801C5220F9BED08B772C84C94FB54C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F6188043080A4757CBC1D26B7DEB4C57" type="ZooBank">F5D13127-0304-46C1-8E67-32B5DE69A569</mods:identifier>
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<treatment id="03C66E4E3A229769FF24FF074C004134" ID-DOI="http://doi.org/10.5281/zenodo.7542099" ID-Zenodo-Dep="7542099" LSID="urn:lsid:zoobank.org:act:FF6BC75B-C222-4A1E-8C88-97EC73A7D098" httpUri="http://treatment.plazi.org/id/03C66E4E3A229769FF24FF074C004134" lastPageId="18" lastPageNumber="262" pageId="14" pageNumber="258">
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<subSubSection id="C3758CD33A229775FF24FF074A2444FC" box="[151,611,151,178]" pageId="14" pageNumber="258" type="nomenclature">
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<paragraph id="8BD0DF583A229775FF24FF074A2444FC" blockId="14.[151,611,151,214]" box="[151,611,151,178]" pageId="14" pageNumber="258">
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<heading id="D09868343A229775FF24FF074A2444FC" bold="true" box="[151,611,151,178]" fontSize="11" level="1" pageId="14" pageNumber="258" reason="1">
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<emphasis id="B91B034A3A229775FF24FF074A2444FC" bold="true" box="[151,611,151,178]" pageId="14" pageNumber="258">
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<taxonomicName id="4C6FA4DB3A229775FF24FF07498B44FF" authority="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson, 2023" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[151,460,151,178]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="258" phylum="Chordata" rank="species" species="intermedia" status="sp. nov.">
|
||
<emphasis id="B91B034A3A229775FF24FF07498B44FF" bold="true" box="[151,460,151,178]" italics="true" pageId="14" pageNumber="258">Podogymnura intermedia</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A228BE313A229775FE60FF074A2444FC" box="[467,611,151,177]" pageId="14" pageNumber="258" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3758CD33A229775FF24FF2B4A504498" box="[151,535,187,214]" pageId="14" pageNumber="258" type="description">
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<paragraph id="8BD0DF583A229775FF24FF2B4A504498" blockId="14.[151,611,151,214]" box="[151,535,187,214]" pageId="14" pageNumber="258">
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<figureCitation id="1354C3DD3A229775FF24FF2B48A2449B" box="[151,229,187,214]" captionStart="FIGURE 5" captionStartId="7.[152,255,905,929]" captionTargetBox="[177,1411,181,881]" captionTargetId="figure-25@7.[177,1411,181,881]" captionTargetPageId="7" captionText="FIGURE 5. Photographs of skulls of Podogymnura intermedia (holotype, FMNH 186805, A-D), P. truei (FMNH 56169, E-H), and P. minima (FMNH 146594; I-L). All skulls approximately life size." figureDoi="http://doi.org/10.5281/zenodo.7532488" httpUri="https://zenodo.org/record/7532488/files/figure.png" pageId="14" pageNumber="258">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="1354C3DD3A229775FF42FF2C4946449B" box="[241,257,188,214]" captionStart="FIGURE 7" captionStartId="15.[152,255,1175,1199]" captionTargetBox="[151,1435,381,1152]" captionTargetId="figure-100@15.[151,1435,381,1152]" captionTargetPageId="15" captionText="FIGURE 7. Podogymnura intermedia (FMNH 186805, holotype), adult male. Photographed 28 July 2005 by D. S. Balete." figureDoi="http://doi.org/10.5281/zenodo.7532496" httpUri="https://zenodo.org/record/7532496/files/figure.png" pageId="14" pageNumber="258">7</figureCitation>
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||
,
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||
<figureCitation id="1354C3DD3A229775FEBEFF2B495B4498" box="[269,284,187,213]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="14" pageNumber="258">8</figureCitation>
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||
, and 9;
|
||
<tableCitation id="C6EDEAE33A229775FEC6FF2B49944498" box="[373,467,187,213]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="14" pageNumber="258" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Tables 1</tableCitation>
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and
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||
<tableCitation id="C6EDEAE33A229775FDBAFF2B4A504498" box="[521,535,187,213]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="14" pageNumber="258" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">2</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3758CD33A229775FF24FE934B564030" pageId="14" pageNumber="258" type="materials_examined">
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<paragraph id="8BD0DF583A229775FF24FE934A6C45E0" blockId="14.[151,1437,259,2014]" pageId="14" pageNumber="258">
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<materialsCitation id="3B07D5053A229775FF24FE934A6C45E0" collectingDate="2005-07-28" collectorName="D. S. Balete" country="Philippines" location="National Museum" pageId="14" pageNumber="258" specimenCount="2" stateProvince="Manila" typeStatus="holotype">
|
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<emphasis id="B91B034A3A229775FF24FE93494C4550" bold="true" box="[151,267,259,285]" pageId="14" pageNumber="258">
|
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<typeStatus id="54D461FA3A229775FF24FE9349414550" box="[151,262,259,285]" pageId="14" pageNumber="258" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
FMNH 186805, adult male, collected on
|
||
<date id="FFD1F9983A229775FD57FE934B304553" box="[740,887,259,286]" pageId="14" pageNumber="258" value="2005-07-28">
|
||
<collectingDate id="EF9500703A229775FD57FE934B304553" box="[740,887,259,286]" pageId="14" pageNumber="258" value="2005-07-28">28 July 2005</collectingDate>
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</date>
|
||
(original number 3769 of
|
||
<collectorName id="269ABA8E3A229775FB10FE944D764550" box="[1187,1329,259,286]" pageId="14" pageNumber="258">D. S. Balete</collectorName>
|
||
); initially fixed in formalin, now preserved in 70% ethyl alcohol with the skull removed; skull in good condition. A sample of fresh muscle tissue was removed from the left thigh and preserved in 90% ethanol; otherwise the body is in good condition. The holotype has been cataloged and is currently housed at FMNH but will be transferred to the
|
||
<location id="8EB089833A229775FA8FFEFF48BE45E3" LSID="urn:lsid:plazi:treatment:03C66E4E3A229769FF24FF074C004134:8EB089833A229775FA8FFEFF48BE45E3" country="Philippines" name="National Museum" pageId="14" pageNumber="258" stateProvince="Manila">National Museum</location>
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of the
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<collectingCountry id="F3789FC83A229775FEFBFE03498E45E0" box="[328,457,403,429]" name="Philippines" pageId="14" pageNumber="258">Philippines</collectingCountry>
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,
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<collectingRegion id="49AB11BA3A229775FE67FE034A6045E0" box="[468,551,403,429]" country="Philippines" name="Manila" pageId="14" pageNumber="258">Manila</collectingRegion>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD0DF583A229775FF74FE274C7D45B8" blockId="14.[151,1437,259,2014]" pageId="14" pageNumber="258">
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<emphasis id="B91B034A3A229775FF74FE274933459C" bold="true" box="[199,372,439,465]" pageId="14" pageNumber="258">Type Locality.</emphasis>
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<materialsCitation id="3B07D5053A229775FECDFE274C7245B8" country="Philippines" elevation="950" latitude="6.707278" location="Mati Municipality" longLatPrecision="1" longitude="126.19522" municipality="Mindanao Island" pageId="14" pageNumber="258" specimenCount="1" stateProvince="Davao Oriental">
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<quantity id="4C9772BD3A229775FECDFE274988459C" box="[382,463,439,465]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="258" unit="km" value="3.7">3.7 km</quantity>
|
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S,
|
||
<locationDeviation id="9A62C67A3A229775FE49FE274B0F459C" box="[506,840,439,466]" location="Mt. Hamiguitan" pageId="14" pageNumber="258">
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<quantity id="4C9772BD3A229775FE49FE274A0C459C" box="[506,587,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="258" unit="km" value="1.4">1.4 km</quantity>
|
||
E of Mt. Hamiguitan
|
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</locationDeviation>
|
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peak,
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<location id="8EB089833A229775FC2AFE274C2D459C" LSID="urn:lsid:plazi:treatment:03C66E4E3A229769FF24FF074C004134:8EB089833A229775FC2AFE274C2D459C" box="[921,1130,439,465]" country="Philippines" latitude="6.707278" longLatPrecision="1" longitude="126.19522" municipality="Mindanao Island" name="Mati Municipality" pageId="14" pageNumber="258" stateProvince="Davao Oriental">Mati Municipality</location>
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,
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<collectingRegion id="49AB11BA3A229775FBCBFE284DDF459C" box="[1144,1432,439,466]" country="Philippines" name="Davao Oriental" pageId="14" pageNumber="258">Davao Oriental Province</collectingRegion>
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,
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<collectingMunicipality id="6BB445223A229775FF24FE4B491045B8" box="[151,343,475,501]" pageId="14" pageNumber="258">Mindanao Island</collectingMunicipality>
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,
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<collectingCountry id="F3789FC83A229775FED1FE4B49A445B8" box="[354,483,475,501]" name="Philippines" pageId="14" pageNumber="258">Philippines</collectingCountry>
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,
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<quantity id="4C9772BD3A229775FE5DFE4B4A7245BB" box="[494,565,475,502]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="258" unit="m" value="950.0">
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<elevation id="0042386B3A229775FE5DFE4B4A7245BB" box="[494,565,475,502]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="258" unit="m" value="950.0">950 m</elevation>
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</quantity>
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elev.,
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<geoCoordinate id="EE5BB99F3A229775FDCDFE4B4B5C45BB" box="[638,795,475,502]" degrees="6" direction="north" minutes="42" orientation="latitude" pageId="14" pageNumber="258" precision="1" seconds="26.2" value="6.707278">
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6
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<superScript id="7C1A72103A229775FD3FFE4B4AD645A4" attach="left" box="[652,657,475,489]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
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42’ 26.2” N
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</geoCoordinate>
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,
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<geoCoordinate id="EE5BB99F3A229775FC95FE4B4B9F45BB" box="[806,984,475,502]" degrees="126" direction="east" minutes="11" orientation="longitude" pageId="14" pageNumber="258" precision="1" seconds="42.8" value="126.19522">
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126
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<superScript id="7C1A72103A229775FCE3FE4B4B1245A4" attach="left" box="[848,853,475,489]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
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11’ 42.8” E
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</geoCoordinate>
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(
|
||
<figureCitation id="1354C3DD3A229775FC54FE4B4C6F45B8" box="[999,1064,475,501]" captionStart="FIGURE 1" captionStartId="1.[152,255,1850,1874]" captionTargetBox="[156,1431,563,1819]" captionTargetId="figure-189@1.[151,1436,558,1824]" captionTargetPageId="1" captionText="FIGURE 1. Map of Mindanao and adjacent islands showing some major topographic features; mountains where Podogymnura truei (Mt.Apo), P. minima (Mt. Kitanglad Range), P. aureospinula (Bucas Grande and Dinagat Islands), and P. intermedia n. sp. (Mt. Hamiguitan and Mt. Kampalili) occur, and other important peaks, are shown." figureDoi="http://doi.org/10.5281/zenodo.7532474" httpUri="https://zenodo.org/record/7532474/files/figure.png" pageId="14" pageNumber="258">Fig. 1</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A229775FF74FE6F4A00475B" blockId="14.[151,1437,259,2014]" pageId="14" pageNumber="258">
|
||
<emphasis id="B91B034A3A229775FF74FE6F498E4654" bold="true" box="[199,457,511,537]" pageId="14" pageNumber="258">Specimens examined.</emphasis>
|
||
<materialsCitation id="3B07D5053A229775FE68FD904A04475B" county="San Isidro Municipality" elevation="1128" latitude="6.7175" location="Mt. Hamiguitan" longLatPrecision="15" longitude="126.18386" municipality="Three" pageId="14" pageNumber="258" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<location id="8EB089833A229775FE68FD904AD34654" LSID="urn:lsid:plazi:treatment:03C66E4E3A229769FF24FF074C004134:8EB089833A229775FE68FD904AD34654" box="[475,660,511,538]" county="San Isidro Municipality" latitude="6.7175" longLatPrecision="15" longitude="126.18386" municipality="Three" name="Mt. Hamiguitan" pageId="14" pageNumber="258">Mt. Hamiguitan</location>
|
||
: (N = 4)
|
||
<collectingMunicipality id="6BB445223A229775FCB3FE6F4B064654" box="[768,833,511,537]" pageId="14" pageNumber="258">Three</collectingMunicipality>
|
||
additional specimens from the type locality (FMNH 190152, 190153, 190167) and one (FMNH 190151) from
|
||
<quantity id="4C9772BD3A229775FC99FDB34BC04670" box="[810,903,547,574]" metricMagnitude="4" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="258" unit="km" value="17.5">17.5 km</quantity>
|
||
S,
|
||
<locationDeviation id="9A62C67A3A229775FC1DFDB34C9A4670" box="[942,1245,547,574]" location="Mt. Hamiguitan" pageId="14" pageNumber="258">
|
||
<quantity id="4C9772BD3A229775FC1DFDB34BAF4670" box="[942,1000,547,573]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="258" unit="km" value="4.0">4 km</quantity>
|
||
E of Mt. Hamiguitan
|
||
</locationDeviation>
|
||
peak,
|
||
<collectingCounty id="62B1A7D43A229775FA9AFDB3496E462C" pageId="14" pageNumber="258">San Isidro Municipality</collectingCounty>
|
||
, ca.
|
||
<quantity id="4C9772BD3A229775FEEDFDD749FC462F" box="[350,443,583,610]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="14" pageNumber="258" unit="m" value="1128.0">
|
||
<elevation id="0042386B3A229775FEEDFDD749FC462F" box="[350,443,583,610]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="14" pageNumber="258" unit="m" value="1128.0">1,128 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE5BB99F3A229775FD89FDD74AFE462F" box="[570,697,583,610]" degrees="6" direction="north" minutes="43" orientation="latitude" pageId="14" pageNumber="258" precision="15" seconds="3" value="6.7175">
|
||
6
|
||
<superScript id="7C1A72103A229775FDFEFDD74A154618" attach="left" box="[589,594,583,597]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
43’3” N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5BB99F3A229775FD75FDD74B34462F" box="[710,883,583,610]" degrees="126" direction="east" minutes="11" orientation="longitude" pageId="14" pageNumber="258" precision="1" seconds="1.9" value="126.18386">
|
||
126
|
||
<superScript id="7C1A72103A229775FD46FDD74ABD4618" attach="left" box="[757,762,583,597]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
11’1.9” E
|
||
</geoCoordinate>
|
||
. These include
|
||
<specimenCount id="9D6914D13A229775FB96FDD84CDE462C" box="[1061,1177,583,610]" pageId="14" pageNumber="258" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount id="9D6914D13A229775FB61FDD84D15462C" box="[1234,1362,583,610]" pageId="14" pageNumber="258" type="female">one female</specimenCount>
|
||
, and a specimen of undetermined sex prepared as a skull only (FMNH 190167). The juvenile male (FMNH 190151), adult male (FMNH 190152), and young adult female (FMNH 190153) have had their skulls removed and cleaned. The bodies of the adult male and young-adult female had partially decomposed prior to preservation, but that of the juvenile male is in good condition. The prepared skulls have broken crania, including one with broken coronoid and condyloid processes (FMNH 190152)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A229775FF74FCB04B564030" blockId="14.[151,1437,259,2014]" pageId="14" pageNumber="258">
|
||
<materialsCitation id="3B07D5053A229775FF74FCB04B4A4030" country="Philippines" county="Mindanao Island" elevation="1900" latitude="7.29522" location="Mt. Kampalili" longLatPrecision="1" longitude="126.31602" municipality="Maragusan Municipality" pageId="14" pageNumber="258" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Compostela">
|
||
<location id="8EB089833A229775FF74FCB0492E4774" LSID="urn:lsid:plazi:treatment:03C66E4E3A229769FF24FF074C004134:8EB089833A229775FF74FCB0492E4774" box="[199,361,799,826]" country="Philippines" county="Mindanao Island" latitude="7.29522" longLatPrecision="1" longitude="126.31602" municipality="Maragusan Municipality" name="Mt. Kampalili" pageId="14" pageNumber="258" stateProvince="Compostela">Mt. Kampalili</location>
|
||
: (N = 12)
|
||
<location id="8EB089833A229775FE64FC8F4A5F4774" LSID="urn:lsid:plazi:treatment:03C66E4E3A229769FF24FF074C004134:8EB089833A229775FE64FC8F4A5F4774" box="[471,536,799,825]" country="Philippines" county="Mindanao Island" latitude="7.29522" longLatPrecision="1" longitude="126.31602" municipality="Maragusan Municipality" name="Three" pageId="14" pageNumber="258" stateProvince="Compostela">Three</location>
|
||
males (FMNH 194750, 194751, 194752) from
|
||
<quantity id="4C9772BD3A229775FB9AFC8F4C264774" box="[1065,1121,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="258" unit="km" value="2.0">2 km</quantity>
|
||
S,
|
||
<locationDeviation id="9A62C67A3A229775FB30FC8F4DDB4774" box="[1155,1436,799,826]" location="Mt. Kampalili" pageId="14" pageNumber="258">
|
||
<quantity id="4C9772BD3A229775FB30FC8F4CFC4774" box="[1155,1211,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="258" unit="km" value="2.0">2 km</quantity>
|
||
W of Mt. Kampalili
|
||
</locationDeviation>
|
||
peak,
|
||
<collectingMunicipality id="6BB445223A229775FF68FCD449B64710" box="[219,497,835,862]" pageId="14" pageNumber="258">Maragusan Municipality</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49AB11BA3A229775FE4FFCD34B794710" box="[508,830,835,861]" country="Philippines" name="Compostela" pageId="14" pageNumber="258">Compostela Valley Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62B1A7D43A229775FCF9FCD34C4D4710" box="[842,1034,835,861]" pageId="14" pageNumber="258">Mindanao Island</collectingCounty>
|
||
,
|
||
<collectingCountry id="F3789FC83A229775FBA5FCD34CD04710" box="[1046,1175,835,861]" name="Philippines" pageId="14" pageNumber="258">Philippines</collectingCountry>
|
||
, ca.
|
||
<quantity id="4C9772BD3A229775FB78FCD34D604713" box="[1227,1319,835,862]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="258" unit="m" value="1900.0">
|
||
<elevation id="0042386B3A229775FB78FCD34D604713" box="[1227,1319,835,862]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="258" unit="m" value="1900.0">1,900 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE5BB99F3A229775FF24FCF7497F47CF" box="[151,312,871,898]" degrees="7" direction="north" minutes="17" orientation="latitude" pageId="14" pageNumber="258" precision="1" seconds="11.7" value="7.2865834">7° 17’ 11.7” N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5BB99F3A229775FEF1FCF749BA47CF" box="[322,509,871,898]" degrees="126" direction="east" minutes="15" orientation="longitude" pageId="14" pageNumber="258" precision="1" seconds="30.9" value="126.25858">126° 15’ 30.9” E</geoCoordinate>
|
||
;
|
||
<specimenCount id="9D6914D13A229775FDBBFCF84AC947CC" box="[520,654,871,898]" pageId="14" pageNumber="258" type="female">two females</specimenCount>
|
||
(FMNH 194748, 194749) from
|
||
<quantity id="4C9772BD3A229775FC47FCF74C0847CC" box="[1012,1103,871,898]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="14" pageNumber="258" unit="km" value="2.75">2.75 km</quantity>
|
||
S,
|
||
<locationDeviation id="9A62C67A3A229775FBC2FCF74DDB47CC" box="[1137,1436,871,898]" location="Mt. Kampalili" pageId="14" pageNumber="258">
|
||
<quantity id="4C9772BD3A229775FBC2FCF74CFA47CC" box="[1137,1213,871,898]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="258" unit="km" value="0.5">0.5 km</quantity>
|
||
W of Mt. Kampalili
|
||
</locationDeviation>
|
||
peak, ca.
|
||
<quantity id="4C9772BD3A229775FEB0FC1B492747EB" box="[259,352,907,934]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="258" unit="m" value="1500.0">
|
||
<elevation id="0042386B3A229775FEB0FC1B492747EB" box="[259,352,907,934]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="258" unit="m" value="1500.0">1,500 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE5BB99F3A229775FE6EFC1B4A1B47EB" box="[477,604,907,934]" degrees="6" direction="north" minutes="17" orientation="latitude" pageId="14" pageNumber="258" precision="15" seconds="39" value="6.2941666">
|
||
6
|
||
<superScript id="7C1A72103A229775FE58FC1B49B747D4" attach="none" box="[491,496,907,921]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
17’39” N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5BB99F3A229775FDDAFC1B4B6447EB" box="[617,803,907,934]" degrees="126" direction="east" minutes="15" orientation="longitude" pageId="14" pageNumber="258" precision="1" seconds="38.4" value="126.260666">
|
||
126
|
||
<superScript id="7C1A72103A229775FD24FC1B4ADB47D4" attach="left" box="[663,668,907,921]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
15’38.4” E
|
||
</geoCoordinate>
|
||
;
|
||
<specimenCount id="9D6914D13A229775FC83FC1C4BFE47E8" box="[816,953,907,934]" pageId="14" pageNumber="258" type="female">two females</specimenCount>
|
||
(FMNH 208700, 208701) and
|
||
<specimenCount id="9D6914D13A229775FAA8FC1B4DDB47E8" box="[1307,1436,907,933]" pageId="14" pageNumber="258" type="male">three males</specimenCount>
|
||
(FMNH 208699, 208702, 208705) at
|
||
<quantity id="4C9772BD3A229775FDF2FC3F4AD94784" box="[577,670,943,970]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="14" pageNumber="258" unit="km" value="2.25">2.25 km</quantity>
|
||
S,
|
||
<locationDeviation id="9A62C67A3A229775FD76FC3F4BB34784" box="[709,1012,943,970]" location="Mt. Kampalili" pageId="14" pageNumber="258">
|
||
<quantity id="4C9772BD3A229775FD76FC3F4B534784" box="[709,788,943,970]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="258" unit="km" value="3.5">3.5 km</quantity>
|
||
E of Mt. Kampalili
|
||
</locationDeviation>
|
||
peak,
|
||
<quantity id="4C9772BD3A229775FBF3FC3F4CDA4787" box="[1088,1181,943,970]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="14" pageNumber="258" unit="m" value="1470.0">
|
||
<elevation id="0042386B3A229775FBF3FC3F4CDA4787" box="[1088,1181,943,970]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="14" pageNumber="258" unit="m" value="1470.0">1,470 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE5BB99F3A229775FAA9FC3F4DDF4787" box="[1306,1432,943,970]" degrees="7.29112" direction="north" orientation="latitude" pageId="14" pageNumber="258" precision="1" value="7.29112">
|
||
7.29112
|
||
<superScript id="7C1A72103A229775FAC7FC3F4D3E47F0" attach="left" box="[1396,1401,943,957]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5BB99F3A229775FF24FC43497747A3" box="[151,304,979,1006]" degrees="126.31520" direction="east" orientation="longitude" pageId="14" pageNumber="258" precision="1" value="126.3152">
|
||
126.31520
|
||
<superScript id="7C1A72103A229775FEBDFC43495447AC" attach="left" box="[270,275,979,993]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
E
|
||
</geoCoordinate>
|
||
; and
|
||
<specimenCount id="9D6914D13A229775FEDCFC4449BF47A0" box="[367,504,979,1006]" pageId="14" pageNumber="258" type="female">two females</specimenCount>
|
||
(FMNH 208703, 208704) at
|
||
<quantity id="4C9772BD3A229775FCF9FC434BEF47A0" box="[842,936,979,1006]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="258" unit="km" value="1.75">1.75 km</quantity>
|
||
S,
|
||
<locationDeviation id="9A62C67A3A229775FC63FC434D5747A0" box="[976,1296,979,1006]" location="Mt. Kampalili" pageId="14" pageNumber="258">
|
||
<quantity id="4C9772BD3A229775FC63FC434C6A47A0" box="[976,1069,979,1006]" metricMagnitude="3" metricUnit="m" metricValue="4.25" pageId="14" pageNumber="258" unit="km" value="4.25">4.25 km</quantity>
|
||
E of Mt. Kampalili
|
||
</locationDeviation>
|
||
peak,
|
||
<quantity id="4C9772BD3A229775FAEEFC4348EA405F" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="14" pageNumber="258" unit="m" value="1640.0">
|
||
<elevation id="0042386B3A229775FAEEFC4348EA405F" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="14" pageNumber="258" unit="m" value="1640.0">1,640 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE5BB99F3A229775FE99FC6749EF405F" box="[298,424,1015,1042]" degrees="7.29522" direction="north" orientation="latitude" pageId="14" pageNumber="258" precision="1" value="7.29522">
|
||
7.29522
|
||
<superScript id="7C1A72103A229775FE36FC6749CD4048" attach="left" box="[389,394,1015,1029]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5BB99F3A229775FE07FC674A0B405F" box="[436,588,1015,1042]" degrees="126.31602" direction="east" orientation="longitude" pageId="14" pageNumber="258" precision="1" value="126.31602">
|
||
126.31602
|
||
<superScript id="7C1A72103A229775FD98FC674A774048" attach="left" box="[555,560,1015,1029]" fontSize="6" pageId="14" pageNumber="258">º</superScript>
|
||
E
|
||
</geoCoordinate>
|
||
. Three females (FMNH 194748, 208700, 208703) and
|
||
<specimenCount id="9D6914D13A229775FB09FC674D7D405C" box="[1210,1338,1015,1041]" pageId="14" pageNumber="258" type="male">three males</specimenCount>
|
||
(FMNH 194750, 194752, 208705) have had their skulls removed and cleaned; others preserved intact. Two females (FMNH 194748, 208700) have crushed crania, and
|
||
<specimenCount id="9D6914D13A229775FD37FBD04AAC4014" box="[644,747,1087,1114]" pageId="14" pageNumber="258" type="male">one male</specimenCount>
|
||
(FMNH 194752) has a broken left zygomatic arch; and
|
||
<specimenCount id="9D6914D13A229775FAC7FBD0488A4030" pageId="14" pageNumber="258" type="male">one male</specimenCount>
|
||
(FMNH 208705) has a short crack on the occipital
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3758CD33A229775FF74FB174DC64088" pageId="14" pageNumber="258" type="distribution">
|
||
<paragraph id="8BD0DF583A229775FF74FB174DC64088" blockId="14.[151,1437,259,2014]" pageId="14" pageNumber="258">
|
||
<emphasis id="B91B034A3A229775FF74FB17491E40EC" bold="true" box="[199,345,1159,1185]" pageId="14" pageNumber="258">Distribution</emphasis>
|
||
. Currently known from montane forest over ultramafic soil between ca. 950 and
|
||
<quantity id="4C9772BD3A229775FAA5FB174D3140EF" box="[1302,1398,1159,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="14" pageNumber="258" unit="m" value="1128.0">1,128 m</quantity>
|
||
on Mount Hamiguitan, and in montane and mossy forest from ca.
|
||
<quantity id="4C9772BD3A229775FCEBFB3B4BF3408B" box="[856,948,1195,1222]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="14" pageNumber="258" unit="m" value="1470.0">1,470 m</quantity>
|
||
to
|
||
<quantity id="4C9772BD3A229775FC64FB3B4C74408B" box="[983,1075,1195,1222]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="258" unit="m" value="1900.0">1,900 m</quantity>
|
||
on Mount Kampalili (
|
||
<figureCitation id="1354C3DD3A229775FA9DFB3B4D284088" box="[1326,1391,1195,1221]" captionStart="FIGURE 1" captionStartId="1.[152,255,1850,1874]" captionTargetBox="[156,1431,563,1819]" captionTargetId="figure-189@1.[151,1436,558,1824]" captionTargetPageId="1" captionText="FIGURE 1. Map of Mindanao and adjacent islands showing some major topographic features; mountains where Podogymnura truei (Mt.Apo), P. minima (Mt. Kitanglad Range), P. aureospinula (Bucas Grande and Dinagat Islands), and P. intermedia n. sp. (Mt. Hamiguitan and Mt. Kampalili) occur, and other important peaks, are shown." figureDoi="http://doi.org/10.5281/zenodo.7532474" httpUri="https://zenodo.org/record/7532474/files/figure.png" pageId="14" pageNumber="258">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3758CD33A229775FF74FB5F4A83417F" pageId="14" pageNumber="258" type="etymology">
|
||
<paragraph id="8BD0DF583A229775FF74FB5F4A83417F" blockId="14.[151,1437,259,2014]" pageId="14" pageNumber="258">
|
||
<emphasis id="B91B034A3A229775FF74FB5F490F40A4" bold="true" box="[199,328,1231,1257]" pageId="14" pageNumber="258">Etymology</emphasis>
|
||
. From the combined Latin
|
||
<emphasis id="B91B034A3A229775FD2DFB404A9440A4" box="[670,723,1232,1257]" italics="true" pageId="14" pageNumber="258">inter</emphasis>
|
||
(between) +
|
||
<emphasis id="B91B034A3A229775FCCEFB5F4B8B40A4" box="[893,972,1231,1257]" italics="true" pageId="14" pageNumber="258">medius</emphasis>
|
||
(middle), to highlight its intermediate morphology between the two smaller species of
|
||
<taxonomicName id="4C6FA4DB3A229775FD08FB634B1D4140" authorityName="Mearns" authorityYear="1905" box="[699,858,1267,1293]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="258" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A229775FD08FB634B1D4140" box="[699,858,1267,1293]" italics="true" pageId="14" pageNumber="258">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
and the large
|
||
<taxonomicName id="4C6FA4DB3A229775FC48FB644CED4140" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1019,1194,1267,1293]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="258" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A229775FC48FB644CED4140" box="[1019,1194,1267,1293]" italics="true" pageId="14" pageNumber="258">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
. We propose eastern Mindanao gymnure as its English common name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3758CD33A229774FF74FAAB49DA44B4" lastPageId="15" lastPageNumber="259" pageId="14" pageNumber="258" type="diagnosis">
|
||
<paragraph id="8BD0DF583A229774FF74FAAB49DA44B4" blockId="14.[151,1437,259,2014]" lastBlockId="15.[151,1437,151,357]" lastPageId="15" lastPageNumber="259" pageId="14" pageNumber="258">
|
||
<emphasis id="B91B034A3A229775FF74FAAB497D4118" bold="true" box="[199,314,1339,1365]" pageId="14" pageNumber="258">Diagnosis</emphasis>
|
||
. A member of the genus
|
||
<taxonomicName id="4C6FA4DB3A229775FDDCFAAB4B494118" authorityName="Mearns" authorityYear="1905" box="[623,782,1339,1365]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="258" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A229775FDDCFAAB4B494118" box="[623,782,1339,1365]" italics="true" pageId="14" pageNumber="258">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
as diagnosed above, of intermediate to small size (Mt. Hamiguitan average HB =
|
||
<quantity id="4C9772BD3A229775FE74FACF4A614137" box="[455,550,1375,1402]" metricMagnitude="-1" metricUnit="m" metricValue="1.71" pageId="14" pageNumber="258" unit="mm" value="171.0">171 mm</quantity>
|
||
, weight
|
||
<quantity id="4C9772BD3A229775FD35FACF4AFD4137" box="[646,698,1375,1402]" metricMagnitude="-2" metricUnit="kg" metricValue="9.6" pageId="14" pageNumber="258" unit="g" value="96.0">96 g</quantity>
|
||
; Mt. Kampalili average HB = 145, weight
|
||
<quantity id="4C9772BD3A229775FB2EFAF04C964137" box="[1181,1233,1376,1402]" metricMagnitude="-2" metricUnit="kg" metricValue="7.7" pageId="14" pageNumber="258" unit="g" value="77.0">77 g</quantity>
|
||
;
|
||
<tableCitation id="C6EDEAE33A229775FB6EFACF4D684134" box="[1245,1327,1375,1401]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="14" pageNumber="258" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
), defined by the following combination of characters. Dorsal pelage generally coarse in Hamiguitan, softer in Kamapalili specimens, with long guard hairs of
|
||
<specimenCount id="9D6914D13A229775FD8AFA384AEC418F" box="[569,683,1448,1474]" pageId="14" pageNumber="258" type="generic">two types</specimenCount>
|
||
: scattered long, black hairs, and many slightly shorter guard hairs having conspicuous golden-yellow tips that produce an appearance of shiny, golden streaks (
|
||
<figureCitation id="1354C3DD3A229775FB19FA5B4CA841A8" box="[1194,1263,1483,1509]" captionStart="FIGURE 7" captionStartId="15.[152,255,1175,1199]" captionTargetBox="[151,1435,381,1152]" captionTargetId="figure-100@15.[151,1435,381,1152]" captionTargetPageId="15" captionText="FIGURE 7. Podogymnura intermedia (FMNH 186805, holotype), adult male. Photographed 28 July 2005 by D. S. Balete." figureDoi="http://doi.org/10.5281/zenodo.7532496" httpUri="https://zenodo.org/record/7532496/files/figure.png" pageId="14" pageNumber="258">Fig. 7</figureCitation>
|
||
). Both types of guard hairs become progressively shorter laterally and anteriorly over the head and snout. Underfur soft, gray, and dense, becoming sparser over head but more conspicuous laterally as guard hairs become sparser laterally and absent ventrally, where silver highlights become apparent. Ventral pelage shorter and paler, lacking golden streaks. Ears short (average =
|
||
<quantity id="4C9772BD3A229775FEE1F9CB49E24238" box="[338,421,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="258" unit="mm" value="20.0">20 mm</quantity>
|
||
), pigmented grayish-brown. Relative to congeners, hindfoot longer in Hamiguitan (average HF =
|
||
<quantity id="4C9772BD3A229775FF6FF9EF497742D4" box="[220,304,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="14" pageNumber="258" unit="mm" value="36.0">36 mm</quantity>
|
||
), shorter in Kampalili (
|
||
<quantity id="4C9772BD3A229775FD8AF9EF4ACB42D4" box="[569,652,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="14" pageNumber="258" unit="mm" value="34.0">34 mm</quantity>
|
||
) but short relative to HB (21–23 % of HB;
|
||
<tableCitation id="C6EDEAE33A229775FB32F9EF4C9342D4" box="[1153,1236,1663,1689]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="14" pageNumber="258" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
); plantar surface, including plantar pads, darkly pigmented or with mottled grayish-brown areas. Tail pigmented medium to dark grayish brown dorsally, and variable ventrally, usually dark but occasionally mottled with white. Tail relatively short compared to length of head and body (average TV =
|
||
<quantity id="4C9772BD3A229775FD47F97B4B36434B" box="[756,881,1771,1798]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" metricValueMax="5.4" metricValueMin="5.3" pageId="14" pageNumber="258" unit="mm" value="53.5" valueMax="54.0" valueMin="53.0">53–54 mm</quantity>
|
||
; 32–37 % of HB;
|
||
<tableCitation id="C6EDEAE33A229775FBF2F97B4CD34348" box="[1089,1172,1771,1797]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="14" pageNumber="258" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
). Skull of Hamiguitan specimens (
|
||
<figureCitation id="1354C3DD3A229775FEA9F89F492E4367" box="[282,361,1807,1834]" captionStart="FIGURE 5" captionStartId="7.[152,255,905,929]" captionTargetBox="[177,1411,181,881]" captionTargetId="figure-25@7.[177,1411,181,881]" captionTargetPageId="7" captionText="FIGURE 5. Photographs of skulls of Podogymnura intermedia (holotype, FMNH 186805, A-D), P. truei (FMNH 56169, E-H), and P. minima (FMNH 146594; I-L). All skulls approximately life size." figureDoi="http://doi.org/10.5281/zenodo.7532488" httpUri="https://zenodo.org/record/7532488/files/figure.png" pageId="14" pageNumber="258">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="1354C3DD3A229775FEC7F89F49C54364" box="[372,386,1807,1833]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="14" pageNumber="258">8</figureCitation>
|
||
) large (
|
||
<typeStatus id="54D461FA3A229775FE64F89F4A7C4364" box="[471,571,1807,1833]" pageId="14" pageNumber="258" type="holotype">holotype</typeStatus>
|
||
CIL =
|
||
<quantity id="4C9772BD3A229775FD39F89F4AB74367" box="[650,752,1807,1834]" metricMagnitude="-2" metricUnit="m" metricValue="4.34" pageId="14" pageNumber="258" unit="mm" value="43.4">43.4 mm</quantity>
|
||
, BBC =
|
||
<quantity id="4C9772BD3A229775FCFCF89F4BF14364" box="[847,950,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="14" pageNumber="258" unit="mm" value="16.8">16.8 mm</quantity>
|
||
), that of Kampalili specimens smaller, (CIL =
|
||
<quantity id="4C9772BD3A229775FF1CF8A349514303" box="[175,278,1843,1870]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="14" pageNumber="258" unit="mm" value="40.6">40.6 mm</quantity>
|
||
, BBC =
|
||
<quantity id="4C9772BD3A229775FEC8F8A349A34300" box="[379,484,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="14" pageNumber="258" unit="mm" value="16.1">16.1 mm</quantity>
|
||
). Sagittal and nuchal crests low but readily visible, rostrum long and deep. Incisive foramina relatively long and wide in Hamiguitan specimens, narrower and shorter in Kampalili specimens. Upper toothrow long (average I1– M3 =
|
||
<quantity id="4C9772BD3A229775FD92F8EB4A9B43D8" box="[545,732,1915,1942]" metricMagnitude="-2" metricUnit="m" metricValue="2.165" metricValueMax="2.2199999999999998" metricValueMin="2.11" pageId="14" pageNumber="258" unit="mm" value="21.65" valueMax="22.2" valueMin="21.1">21.1 – 22.2 mm</quantity>
|
||
). Post-palatal region proportionately long in specimens from Mt. Hamiguitan, average in Kampalili, both samples with paired concavities between the bullae, bisected by a low medial ridge. Mandible long and robust in Hamiguitan (average LMI =
|
||
<quantity id="4C9772BD3A229775FC6FF8534C004390" box="[988,1095,1987,2014]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="14" pageNumber="258" unit="mm" value="33.5">33.5 mm</quantity>
|
||
), shorter and more slender in Kampalili (average LMI =
|
||
<quantity id="4C9772BD3A239774FE76FF074A6A44FC" box="[453,557,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="15" pageNumber="259" unit="mm" value="31.6">31.6 mm</quantity>
|
||
). Lingual lobe of P3 present in Hamiguitan specimens, absent in Kampalili. P4 proportionately large and square-shaped. A small, inconspicuous cusp present at base of talonid on first and second lower molars (
|
||
<figureCitation id="1354C3DD3A239774FE8AFF4F49D644B4" box="[313,401,223,249]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="15" pageNumber="259">Fig. 8E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3758CD33A23976AFF74FE924D744364" lastPageId="17" lastPageNumber="261" pageId="15" pageNumber="259" type="description">
|
||
<paragraph id="8BD0DF583A239774FF74FE9249F44528" blockId="15.[151,1437,151,357]" pageId="15" pageNumber="259">
|
||
<emphasis id="B91B034A3A239774FF74FE924A6B4551" bold="true" box="[199,556,258,285]" pageId="15" pageNumber="259">Description and Comparisons</emphasis>
|
||
. Because specimens from Mt. Hamiguitan (the
|
||
<typeStatus id="54D461FA3A239774FBF0FE934C344550" box="[1091,1139,259,285]" pageId="15" pageNumber="259">type</typeStatus>
|
||
locality) differ from those from Mt. Kampalili, we include comparisons of specimens from these two places along with comparison to other species of
|
||
<taxonomicName id="4C6FA4DB3A239774FEBEFEDB49EB4528" authorityName="Mearns" authorityYear="1905" box="[269,428,331,357]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="15" pageNumber="259" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A239774FEBEFEDB49EB4528" box="[269,428,331,357]" italics="true" pageId="15" pageNumber="259">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF108FD03A239774FF2BFB074DC540FD" ID-DOI="http://doi.org/10.5281/zenodo.7532496" ID-Zenodo-Dep="7532496" box="[152,1410,1175,1200]" httpUri="https://zenodo.org/record/7532496/files/figure.png" pageId="15" pageNumber="259" startId="15.[152,255,1175,1199]" targetBox="[151,1435,381,1152]" targetPageId="15">
|
||
<paragraph id="8BD0DF583A239774FF2BFB074DC540FD" blockId="15.[152,1410,1175,1200]" box="[152,1410,1175,1200]" pageId="15" pageNumber="259">
|
||
<emphasis id="B91B034A3A239774FF2BFB07495F40FD" bold="true" box="[152,280,1175,1200]" pageId="15" pageNumber="259">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C6FA4DB3A239774FEACFB084A6440FD" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[287,547,1176,1200]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="15" pageNumber="259" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A239774FEACFB084A6440FD" box="[287,547,1176,1200]" italics="true" pageId="15" pageNumber="259">Podogymnura intermedia</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 186805, holotype), adult male. Photographed 28 July 2005 by D. S. Balete.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF108FD03A239774FF2BF8124DD743AF" ID-DOI="http://doi.org/10.5281/zenodo.7532502" ID-Zenodo-Dep="7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="15" pageNumber="259" startId="15.[152,255,1922,1946]" targetBox="[151,1436,1224,1899]" targetPageId="15">
|
||
<paragraph id="8BD0DF583A239774FF2BF8124DD743AF" blockId="15.[151,1437,1922,2018]" pageId="15" pageNumber="259">
|
||
<emphasis id="B91B034A3A239774FF2BF812495E43D7" bold="true" box="[152,281,1922,1946]" pageId="15" pageNumber="259">FIGURE 8.</emphasis>
|
||
Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of
|
||
<taxonomicName id="4C6FA4DB3A239774FABEF812494243F3" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="15" pageNumber="259" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A239774FABEF812494243F3" italics="true" pageId="15" pageNumber="259">Podogymnura intermedia</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of
|
||
<taxonomicName id="4C6FA4DB3A239774FF24F85A49DC43AF" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[151,411,1994,2018]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="15" pageNumber="259" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A239774FF24F85A49DC43AF" box="[151,411,1994,2018]" italics="true" pageId="15" pageNumber="259">Podogymnura intermedia</emphasis>
|
||
</taxonomicName>
|
||
holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF108FD03A3C976BFF2BFB304BE94091" ID-DOI="http://doi.org/10.5281/zenodo.7532506" ID-Zenodo-Dep="7532506" httpUri="https://zenodo.org/record/7532506/files/figure.png" pageId="16" pageNumber="260" startId="16.[152,255,1184,1208]" targetBox="[159,1428,181,1160]" targetPageId="16">
|
||
<paragraph id="8BD0DF583A3C976BFF2BFB304BE94091" blockId="16.[151,1436,1184,1244]" pageId="16" pageNumber="260">
|
||
<emphasis id="B91B034A3A3C976BFF2BFB30495D40F5" bold="true" box="[152,282,1184,1208]" pageId="16" pageNumber="260">FIGURE 9.</emphasis>
|
||
Maxillary and mandibular dentition of
|
||
<taxonomicName id="4C6FA4DB3A3C976BFD0AFB304BF840F5" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[697,959,1184,1208]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3C976BFD0AFB304BF840F5" box="[697,959,1184,1208]" italics="true" pageId="16" pageNumber="260">Podogymnura intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Mt. Hamiguitan (A, B; FMNH 186805, holotype); and (C, D; FMNH 194751), from Mt. Kampalili. Scale bar = 5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD0DF583A3C976BFF74FA9B4C02430D" blockId="16.[151,1437,1290,2037]" pageId="16" pageNumber="260">
|
||
A medium-sized Philippine gymnure, Hamiguitan specimens larger and more robust than specimens from Mt. Kampalili,
|
||
<taxonomicName id="4C6FA4DB3A3C976BFEA0FABF49C64105" authorityName="Sanborn" authorityYear="1953" box="[275,385,1327,1352]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3C976BFEA0FABF49C64105" box="[275,385,1327,1352]" italics="true" pageId="16" pageNumber="260">P. minima</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6FA4DB3A3C976BFE08FABF4A4D4105" box="[443,522,1327,1352]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3C976BFE08FABF4A4D4105" box="[443,522,1327,1352]" italics="true" pageId="16" pageNumber="260">P. truei</emphasis>
|
||
</taxonomicName>
|
||
, but smaller and less robust than
|
||
<taxonomicName id="4C6FA4DB3A3C976BFCC7FABF4C674105" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[884,1056,1326,1352]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3C976BFCC7FABF4C674105" box="[884,1056,1326,1352]" italics="true" pageId="16" pageNumber="260">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6EDEAE33A3C976BFB9EFABE4CCD4104" box="[1069,1162,1326,1353]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="16" pageNumber="260" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Tables 1</tableCitation>
|
||
and
|
||
<tableCitation id="C6EDEAE33A3C976BFB0EFABF4C8A4104" box="[1213,1229,1327,1353]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="16" pageNumber="260" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">2</tableCitation>
|
||
,
|
||
<figureCitation id="1354C3DD3A3C976BFB64FABE4D634104" box="[1239,1316,1326,1353]" captionStart="FIGURE 4" captionStartId="6.[152,255,696,720]" captionTargetBox="[312,1275,181,672]" captionTargetId="figure-19@6.[151,1436,181,672]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of skull of Podogymnura aureospinula, holotype, DMNH 4386, approximately life size. Adapted from Heaney & Morgan 1982." figureDoi="http://doi.org/10.5281/zenodo.7532486" httpUri="https://zenodo.org/record/7532486/files/figure.png" pageId="16" pageNumber="260">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="1354C3DD3A3C976BFA9DFABE4D794104" box="[1326,1342,1326,1353]" captionStart="FIGURE 5" captionStartId="7.[152,255,905,929]" captionTargetBox="[177,1411,181,881]" captionTargetId="figure-25@7.[177,1411,181,881]" captionTargetPageId="7" captionText="FIGURE 5. Photographs of skulls of Podogymnura intermedia (holotype, FMNH 186805, A-D), P. truei (FMNH 56169, E-H), and P. minima (FMNH 146594; I-L). All skulls approximately life size." figureDoi="http://doi.org/10.5281/zenodo.7532488" httpUri="https://zenodo.org/record/7532488/files/figure.png" pageId="16" pageNumber="260">5</figureCitation>
|
||
). Dorsal pelage of
|
||
<taxonomicName id="4C6FA4DB3A3C976BFEB4FAC349DA4121" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[263,413,1362,1388]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3C976BFEB4FAC349DA4121" box="[263,413,1362,1388]" italics="true" pageId="16" pageNumber="260">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan is distinctly darker on head and rostrum, venter slightly paler brownishgray (
|
||
<figureCitation id="1354C3DD3A3C976BFF6AFAE6495841DC" box="[217,287,1398,1425]" captionStart="FIGURE 4" captionStartId="6.[152,255,696,720]" captionTargetBox="[312,1275,181,672]" captionTargetId="figure-19@6.[151,1436,181,672]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of skull of Podogymnura aureospinula, holotype, DMNH 4386, approximately life size. Adapted from Heaney & Morgan 1982." figureDoi="http://doi.org/10.5281/zenodo.7532486" httpUri="https://zenodo.org/record/7532486/files/figure.png" pageId="16" pageNumber="260">Fig. 4</figureCitation>
|
||
); coarse overfur consists of long, stiff guard hairs, ca.
|
||
<quantity id="4C9772BD3A3C976BFC36FAE74B9341DC" box="[901,980,1399,1425]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="260" unit="mm" value="20.0">20 mm</quantity>
|
||
on mid-dorsum, longer on the rump, ca.
|
||
<quantity id="4C9772BD3A3C976BFF24FA0A48AF41F8" box="[151,232,1434,1461]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="260" unit="mm" value="25.0">25 mm</quantity>
|
||
. The long, coarse, grayish-brown dorsal pelage contains both long black guard hairs and long golden-yellow guard hairs that produce a golden-streaked appearance. Mt. Kampalili specimens have soft, dark brown dorsal pelage with shorter black guard hairs and many short golden-yellow-tipped hairs that produce a golden-speckled appearance. Dorsal pelage of
|
||
<taxonomicName id="4C6FA4DB3A3C976BFE6DF9974ACC426D" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[478,651,1542,1568]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3C976BFE6DF9974ACC426D" box="[478,651,1542,1568]" italics="true" pageId="16" pageNumber="260">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
is uniformly paler and golden-brown overall and overfur is shorter and bristly; venter brownish-gray.
|
||
<taxonomicName id="4C6FA4DB3A3C976BFE58F9BB4A7B4209" box="[491,572,1579,1604]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3C976BFE58F9BB4A7B4209" box="[491,572,1579,1604]" italics="true" pageId="16" pageNumber="260">P. truei</emphasis>
|
||
</taxonomicName>
|
||
has soft dorsal pelage that is dark brown with small, inconspicuous speckles of golden-brown at the tips, without conspicuous long and stiff guard hairs.
|
||
<taxonomicName id="4C6FA4DB3A3C976BFC6EF9DF4C0B4225" authorityName="Sanborn" authorityYear="1953" box="[989,1100,1615,1640]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3C976BFC6EF9DF4C0B4225" box="[989,1100,1615,1640]" italics="true" pageId="16" pageNumber="260">P. minima</emphasis>
|
||
</taxonomicName>
|
||
has short, soft, reddish-brown dorsal pelage with golden-brown speckles at the tips that are more conspicuous than those of
|
||
<taxonomicName id="4C6FA4DB3A3C976BFB19F9E34CBD42C1" box="[1194,1274,1651,1676]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3C976BFB19F9E34CBD42C1" box="[1194,1274,1651,1676]" italics="true" pageId="16" pageNumber="260">P. truei</emphasis>
|
||
</taxonomicName>
|
||
but are darker and smaller than those of
|
||
<taxonomicName id="4C6FA4DB3A3C976BFE76F9074A1A42FD" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[453,605,1686,1712]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3C976BFE76F9074A1A42FD" box="[453,605,1686,1712]" italics="true" pageId="16" pageNumber="260">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Mt. Hamiguitan or specimens from Mt. Kampalili. Females of all species have two pairs of small, inconspicuous mammae, one pair inguinal and one pair axial; these are well hidden by the fur.Adult males have a swollen area from the base of the tail to the area around the penile sheath; the sheath is small, ca.
|
||
<quantity id="4C9772BD3A3C976BFEB4F892491A4350" box="[263,349,1794,1821]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity>
|
||
wide and long; no scrotum is evident. In specimens from Hamiguitan, the swollen area is covered by typical abdominal fur, but in specimens from Kampalili, the swollen area is bare.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3C976AFF74F8DB4B4D45C4" blockId="16.[151,1437,1290,2037]" lastBlockId="17.[151,1437,151,2013]" lastPageId="17" lastPageNumber="261" pageId="16" pageNumber="260">
|
||
Ears of all
|
||
<taxonomicName id="4C6FA4DB3A3C976BFEFEF8DA49AB4329" authorityName="Mearns" authorityYear="1905" box="[333,492,1866,1892]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3C976BFEFEF8DA49AB4329" box="[333,492,1866,1892]" italics="true" pageId="16" pageNumber="260">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
are short relative to body size, and sparsely covered with short, nearly invisible hairs; ears of specimens from Hamiguitan are pigmented dark gray but from Kampalili are paler.
|
||
<taxonomicName id="4C6FA4DB3A3C976BFB5EF8FF4DDB43C5" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1261,1436,1902,1928]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3C976BFB5EF8FF4DDB43C5" box="[1261,1436,1902,1928]" italics="true" pageId="16" pageNumber="260">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
ears are palest of the known species. Ears of
|
||
<taxonomicName id="4C6FA4DB3A3C976BFD10F8034AB143E1" box="[675,758,1939,1964]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3C976BFD10F8034AB143E1" box="[675,758,1939,1964]" italics="true" pageId="16" pageNumber="260">P. truei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6FA4DB3A3C976BFC82F8034BE443E1" authorityName="Sanborn" authorityYear="1953" box="[817,931,1939,1964]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3C976BFC82F8034BE443E1" box="[817,931,1939,1964]" italics="true" pageId="16" pageNumber="260">P. minima</emphasis>
|
||
</taxonomicName>
|
||
average slightly longer and paler than in
|
||
<taxonomicName id="4C6FA4DB3A3C976BFA3BF8034957439D" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3C976BFA3BF8034957439D" italics="true" pageId="16" pageNumber="260">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6EDEAE33A3C976BFE93F8264933439C" box="[288,372,1974,2001]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="16" pageNumber="260" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
). Length of tail relative to head and body (32%) in
|
||
<taxonomicName id="4C6FA4DB3A3C976BFC71F8274C1F439D" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[962,1112,1974,2000]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3C976BFC71F8274C1F439D" box="[962,1112,1974,2000]" italics="true" pageId="16" pageNumber="260">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan is shortest among all
|
||
<taxonomicName id="4C6FA4DB3A3C976BFEA1F84A49F643B9" authorityName="Mearns" authorityYear="1905" box="[274,433,2010,2036]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="260" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3C976BFEA1F84A49F643B9" box="[274,433,2010,2036]" italics="true" pageId="16" pageNumber="260">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6EDEAE33A3C976BFE72F84A4A5043B8" box="[449,535,2010,2037]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="16" pageNumber="260" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
). Skin of tail of Hamiguitan specimens is uniformly dark gray throughout; on Kampalili specimens it is ventrally pale brown or mottled with white. Tail of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFC45FF074CE344FC" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1014,1188,151,177]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFC45FF074CE344FC" box="[1014,1188,151,177]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
is longer but overlaps in relative length (33% of HB). Tail of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFDFAFF2B4ADE4499" box="[585,665,187,212]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFDFAFF2B4ADE4499" box="[585,665,187,212]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
is substantially longer both absolutely and proportionately (
|
||
<tableCitation id="C6EDEAE33A3D976AFA84FF2B4DCC4498" box="[1335,1419,187,213]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="17" pageNumber="261" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
). Although
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEBBFF4F493044B5" authorityName="Sanborn" authorityYear="1953" box="[264,375,223,248]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3D976AFEBBFF4F493044B5" box="[264,375,223,248]" italics="true" pageId="17" pageNumber="261">P. minima</emphasis>
|
||
</taxonomicName>
|
||
is smaller overall, tail is equal in length and proportionately longer. Hind foot of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFAB4FF4F4DDB44B4" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[1287,1436,223,249]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFAB4FF4F4DDB44B4" box="[1287,1436,223,249]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
is average among
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEDBFE934A404550" authorityName="Mearns" authorityYear="1905" box="[360,519,259,285]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3D976AFEDBFE934A404550" box="[360,519,259,285]" italics="true" pageId="17" pageNumber="261">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
excepting the larger
|
||
<emphasis id="B91B034A3A3D976AFD48FE934BF54550" box="[763,946,259,285]" italics="true" pageId="17" pageNumber="261">
|
||
<taxonomicName id="4C6FA4DB3A3D976AFD48FE934BE84550" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[763,943,259,285]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">P. aureospinula</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but proportionately shorter than all except
|
||
<taxonomicName id="4C6FA4DB3A3D976AFF24FEB7490F450C" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[151,328,295,321]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFF24FEB7490F450C" box="[151,328,295,321]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6EDEAE33A3D976AFEEAFEB749F7450C" box="[345,432,295,321]" captionStart="TABLE 1" captionStartId="7.[152,244,1013,1037]" captionTargetPageId="7" captionText="TABLE 1. External measurements (mm), weight (g) and measurement ratios (%) of adult Podogymnura, including new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average, for sample size of 3 or less. Sample size less than n is indicated in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A2B977CFF2BFC654D15401B" pageId="17" pageNumber="261" tableUuid="DF108FD03A2B977CFF2BFC654D15401B">Table 1</tableCitation>
|
||
). Hindfoot skin is pigmented medium gray dorsally and ventrally, including digits and plantar pads, paler and often mottled with white in those from Kampalili. Hind feet of
|
||
<emphasis id="B91B034A3A3D976AFBD3FEDB4D2A4529" box="[1120,1389,331,357]" italics="true" pageId="17" pageNumber="261">
|
||
<taxonomicName id="4C6FA4DB3A3D976AFBD3FEDB4D554528" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1120,1298,331,357]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">P. aureospinula</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6FA4DB3A3D976AFAAFFEDB4D2A4529" box="[1308,1389,331,356]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">P. truei</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFF24FEFF494045C5" authorityName="Sanborn" authorityYear="1953" box="[151,263,367,392]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3D976AFF24FEFF494045C5" box="[151,263,367,392]" italics="true" pageId="17" pageNumber="261">P. minima</emphasis>
|
||
</taxonomicName>
|
||
are unpigmented, both dorsally and ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3D976AFF74FE034A27462C" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="261">
|
||
Skull of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE9DFE03498245E0" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[302,453,403,429]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFE9DFE03498245E0" box="[302,453,403,429]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354C3DD3A3D976AFE6BFE034A5945E0" box="[472,542,403,429]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8</figureCitation>
|
||
,
|
||
<tableCitation id="C6EDEAE33A3D976AFD9DFE034AC245E0" box="[558,645,403,429]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="17" pageNumber="261" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">Table 2</tableCitation>
|
||
) is smaller than the much larger
|
||
<taxonomicName id="4C6FA4DB3A3D976AFBB9FE034CFB45E0" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1034,1212,403,429]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFBB9FE034CFB45E0" box="[1034,1212,403,429]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354C3DD3A3D976AFB7DFE034D5245E0" box="[1230,1301,403,429]" captionStart="FIGURE 4" captionStartId="6.[152,255,696,720]" captionTargetBox="[312,1275,181,672]" captionTargetId="figure-19@6.[151,1436,181,672]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of skull of Podogymnura aureospinula, holotype, DMNH 4386, approximately life size. Adapted from Heaney & Morgan 1982." figureDoi="http://doi.org/10.5281/zenodo.7532486" httpUri="https://zenodo.org/record/7532486/files/figure.png" pageId="17" pageNumber="261">Fig. 4</figureCitation>
|
||
), but larger than its congeners in most respects, with Mt. Hamiguitan specimens larger than those from Kampalili (
|
||
<figureCitation id="1354C3DD3A3D976AFA9DFE274D35459C" box="[1326,1394,438,465]" captionStart="FIGURE 5" captionStartId="7.[152,255,905,929]" captionTargetBox="[177,1411,181,881]" captionTargetId="figure-25@7.[177,1411,181,881]" captionTargetPageId="7" captionText="FIGURE 5. Photographs of skulls of Podogymnura intermedia (holotype, FMNH 186805, A-D), P. truei (FMNH 56169, E-H), and P. minima (FMNH 146594; I-L). All skulls approximately life size." figureDoi="http://doi.org/10.5281/zenodo.7532488" httpUri="https://zenodo.org/record/7532488/files/figure.png" pageId="17" pageNumber="261">Fig. 5</figureCitation>
|
||
). It is similar to
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE95FE4B493045B9" box="[294,375,475,500]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFE95FE4B493045B9" box="[294,375,475,500]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE03FE4B4A6145B9" box="[432,550,475,500]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="mimima">
|
||
<emphasis id="B91B034A3A3D976AFE03FE4B4A6145B9" box="[432,550,475,500]" italics="true" pageId="17" pageNumber="261">P. mimima</emphasis>
|
||
</taxonomicName>
|
||
in its limited development of the prominent nuchal and sagittal crests that are hallmarks of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE9EFE6F499B4654" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[301,476,511,537]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFE9EFE6F499B4654" box="[301,476,511,537]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
. Maximum height of sagittal crest is
|
||
<quantity id="4C9772BD3A3D976AFC36FE6F4B9B4654" box="[901,988,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="261" unit="mm" value="0.8">0.8 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFBB2FE6F4CD04654" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[1025,1175,511,537]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFBB2FE6F4CD04654" box="[1025,1175,511,537]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
. The nuchal crest of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFA3BFE6F49574670" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFA3BFE6F49574670" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
slants slightly posteriad relative to the cranium but does not project beyond occipital margins (
|
||
<figureCitation id="1354C3DD3A3D976AFAFAFDB34DCA4670" box="[1353,1421,547,573]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8</figureCitation>
|
||
), which it does in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEE3FDD749B8462C" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[336,511,583,609]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFEE3FDD749B8462C" box="[336,511,583,609]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354C3DD3A3D976AFDBDFDD74A14462C" box="[526,595,583,609]" captionStart="FIGURE 4" captionStartId="6.[152,255,696,720]" captionTargetBox="[312,1275,181,672]" captionTargetId="figure-19@6.[151,1436,181,672]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of skull of Podogymnura aureospinula, holotype, DMNH 4386, approximately life size. Adapted from Heaney & Morgan 1982." figureDoi="http://doi.org/10.5281/zenodo.7532486" httpUri="https://zenodo.org/record/7532486/files/figure.png" pageId="17" pageNumber="261">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3D976AFF74FDFB4A094030" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="261">
|
||
In lateral view, skull of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE78FDFB4A1846C8" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[459,607,619,645]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFE78FDFB4A1846C8" box="[459,607,619,645]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan (
|
||
<figureCitation id="1354C3DD3A3D976AFC87FDFB4B3246C8" box="[820,885,619,645]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8</figureCitation>
|
||
) cuts a nearly straight slanting profile from top of braincase to tip of narrow and tapered rostrum, whereas in specimens from Kampalili the profile is slightly concave. Frontals of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEABFD2349814680" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[280,454,691,717]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFEABFD2349814680" box="[280,454,691,717]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
are dorsolaterally inflated, making them only about 9% narrower than braincase, and in dorsal view producing a narrow-waisted hour-glass shape in interorbital region (
|
||
<figureCitation id="1354C3DD3A3D976AFBACFD474C2246BC" box="[1055,1125,727,753]" captionStart="FIGURE 4" captionStartId="6.[152,255,696,720]" captionTargetBox="[312,1275,181,672]" captionTargetId="figure-19@6.[151,1436,181,672]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of skull of Podogymnura aureospinula, holotype, DMNH 4386, approximately life size. Adapted from Heaney & Morgan 1982." figureDoi="http://doi.org/10.5281/zenodo.7532486" httpUri="https://zenodo.org/record/7532486/files/figure.png" pageId="17" pageNumber="261">Fig. 4</figureCitation>
|
||
). In lateral view, prominent frontal swellings in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEC5FD6B4A634758" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[374,548,763,789]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFEC5FD6B4A634758" box="[374,548,763,789]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
produce a convex dorsal profile. All three small-bodied
|
||
<taxonomicName id="4C6FA4DB3A3D976AFB28FD6B4D7D4758" authorityName="Mearns" authorityYear="1905" box="[1179,1338,763,789]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3D976AFB28FD6B4D7D4758" box="[1179,1338,763,789]" italics="true" pageId="17" pageNumber="261">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
share an interorbital region in which there is a relatively broadly-waisted hourglass shape, although lacrimal and interorbital breadths of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEADFCD349F44710" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[286,435,835,861]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFEADFCD349F44710" box="[286,435,835,861]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan are greater than in Kampalili and the other two species (
|
||
<tableCitation id="C6EDEAE33A3D976AFAB6FCD34D1A4710" box="[1285,1373,835,861]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="17" pageNumber="261" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">Table 2</tableCitation>
|
||
), and all lack the temporal ridges that in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFD82FCF74AA447CC" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[561,739,871,897]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFD82FCF74AA447CC" box="[561,739,871,897]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
form a low crest converging at the interorbital region. In
|
||
<taxonomicName id="4C6FA4DB3A3D976AFA3BFCF7495747E8" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFA3BFCF7495747E8" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
and the two other small-bodied species, the parietals are dorsolaterally inflated from the anterior edge of interparietals, forming a cranium that tapers anteriad to the frontal region and flattens posteriad to the occipital region (
|
||
<figureCitation id="1354C3DD3A3D976AFF5DFC43497547A0" box="[238,306,979,1005]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8</figureCitation>
|
||
). Incisive foramina cordate, similar to those of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFCF5FC434BF147A1" authorityName="Sanborn" authorityYear="1953" box="[838,950,979,1004]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3D976AFCF5FC434BF147A1" box="[838,950,979,1004]" italics="true" pageId="17" pageNumber="261">P. minima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFC5FFC434C7A47A1" box="[1004,1085,979,1004]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFC5FFC434C7A47A1" box="[1004,1085,979,1004]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
in size, in contrast to the wider and longer foramina in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE10FC674A14405C" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[419,595,1015,1041]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFE10FC674A14405C" box="[419,595,1015,1041]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFD38FC674B66405C" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[651,801,1015,1041]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFD38FC674B66405C" box="[651,801,1015,1041]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
. Palatal length, width of the posterior palatal ridge, and lingual palatal breadth at M
|
||
<superScript id="7C1A72103A3D976AFE67FB8A499B4065" attach="left" box="[468,476,1050,1064]" fontSize="6" pageId="17" pageNumber="261">3</superScript>
|
||
of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFDB6FB8B4ADB4078" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[517,668,1051,1077]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFDB6FB8B4ADB4078" box="[517,668,1051,1077]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
are large (except in comparison to
|
||
<taxonomicName id="4C6FA4DB3A3D976AFB87FB8B4CA24078" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1076,1253,1051,1077]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFB87FB8B4CA24078" box="[1076,1253,1051,1077]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<tableCitation id="C6EDEAE33A3D976AFB46FB8B4D0B4078" box="[1269,1356,1051,1077]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="17" pageNumber="261" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">Table 2</tableCitation>
|
||
). Postpalatal length of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEE3FBAF49A44014" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[336,483,1087,1113]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFEE3FBAF49A44014" box="[336,483,1087,1113]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan is the greatest among the small
|
||
<taxonomicName id="4C6FA4DB3A3D976AFBBAFBAF4CEF4014" authorityName="Mearns" authorityYear="1905" box="[1033,1192,1087,1113]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3D976AFBBAFBAF4CEF4014" box="[1033,1192,1087,1113]" italics="true" pageId="17" pageNumber="261">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
, with specimens from Kampalili averaging shortest (
|
||
<tableCitation id="C6EDEAE33A3D976AFE5AFBF34A064030" box="[489,577,1123,1149]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="17" pageNumber="261" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3D976AFF74FB1748964134" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="261">
|
||
The basicranium of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE15FB174A7D40EC" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[422,570,1159,1185]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFE15FB174A7D40EC" box="[422,570,1159,1185]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
is similar to its congeners, differing mainly in the large size of the basicranial area, auditory bulla, and paraoccipital process (
|
||
<figureCitation id="1354C3DD3A3D976AFD00FB3B4ABE4088" box="[691,761,1195,1221]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8</figureCitation>
|
||
). Aside from overall size, the basicranium of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFAB5FB3B4DDB4088" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[1286,1436,1195,1221]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFAB5FB3B4DDB4088" box="[1286,1436,1195,1221]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
is similar to that of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEC7FB5F4A6440A4" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[372,547,1231,1257]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFEC7FB5F4A6440A4" box="[372,547,1231,1257]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
(and unlike other
|
||
<taxonomicName id="4C6FA4DB3A3D976AFD41FB5F4BAC40A4" box="[754,1003,1231,1257]" pageId="17" pageNumber="261">
|
||
<emphasis id="B91B034A3A3D976AFD41FB5F4BD640A4" box="[754,913,1231,1257]" italics="true" pageId="17" pageNumber="261">Podogymnura</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
) in having paired, shallow concavities traversed medially by a fine, short ridge that runs parallel to the bullae from its base to the tip of the tympanic wing, although in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE92FA874996417C" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[289,465,1303,1329]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFE92FA874996417C" box="[289,465,1303,1329]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
the depressions are more expansive and the mid-ventral ridge longer and larger. The medial ridge is low and poorly defined in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFDC3FAAB4A864119" box="[624,705,1339,1364]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFDC3FAAB4A864119" box="[624,705,1339,1364]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
and nearly absent in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFC02FAAB4C664119" authorityName="Sanborn" authorityYear="1953" box="[945,1057,1339,1364]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3D976AFC02FAAB4C664119" box="[945,1057,1339,1364]" italics="true" pageId="17" pageNumber="261">P. minima</emphasis>
|
||
</taxonomicName>
|
||
, and the concavities are absent in both.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3D976AFF74FA134AD64260" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="261">
|
||
Mandible of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFEEAFA1349A841D0" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[345,495,1411,1437]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFEEAFA1349A841D0" box="[345,495,1411,1437]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354C3DD3A3D976AFE4CFA134A0441D0" box="[511,579,1411,1437]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8</figureCitation>
|
||
,
|
||
<tableCitation id="C6EDEAE33A3D976AFDE3FA134AE241D0" box="[592,677,1411,1437]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="17" pageNumber="261" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">Table 2</tableCitation>
|
||
) relatively long and thick, except in comparison to
|
||
<taxonomicName id="4C6FA4DB3A3D976AFB5EFA134DDB41D0" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[1261,1436,1411,1437]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFB5EFA134DDB41D0" box="[1261,1436,1411,1437]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354C3DD3A3D976AFF2CFA3748A1418C" box="[159,230,1446,1473]" captionStart="FIGURE 5" captionStartId="7.[152,255,905,929]" captionTargetBox="[177,1411,181,881]" captionTargetId="figure-25@7.[177,1411,181,881]" captionTargetPageId="7" captionText="FIGURE 5. Photographs of skulls of Podogymnura intermedia (holotype, FMNH 186805, A-D), P. truei (FMNH 56169, E-H), and P. minima (FMNH 146594; I-L). All skulls approximately life size." figureDoi="http://doi.org/10.5281/zenodo.7532488" httpUri="https://zenodo.org/record/7532488/files/figure.png" pageId="17" pageNumber="261">Fig. 5</figureCitation>
|
||
).
|
||
<taxonomicName id="4C6FA4DB3A3D976AFF4DFA3749E8418C" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[254,431,1447,1473]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFF4DFA3749E8418C" box="[254,431,1447,1473]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
has larger, wider, and longer angular, coronoid, and condyloid processes than all of its congeners. In
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE8FFA5B499241A8" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[316,469,1483,1509]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFE8FFA5B499241A8" box="[316,469,1483,1509]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan, the angular process is narrower but longer, coronoid wider, and condyloid process longer and more robust than in specimens from Kampalili,
|
||
<taxonomicName id="4C6FA4DB3A3D976AFBBCFA7F4C384245" authorityName="Sanborn" authorityYear="1953" box="[1039,1151,1519,1544]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3D976AFBBCFA7F4C384245" box="[1039,1151,1519,1544]" italics="true" pageId="17" pageNumber="261">P. minima</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFB73FA7F4D564245" box="[1216,1297,1519,1544]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFB73FA7F4D564245" box="[1216,1297,1519,1544]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
. Position of mandibular foramina is similar in all species.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3D976AFF74F9A74D744364" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="261">
|
||
Dental features of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE28F9A74A76421C" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[411,561,1591,1617]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFE28F9A74A76421C" box="[411,561,1591,1617]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354C3DD3A3D976AFDF1F9A74AD5421C" box="[578,658,1591,1617]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Figs. 8</figureCitation>
|
||
,
|
||
<figureCitation id="1354C3DD3A3D976AFD12F9A74AB2421C" box="[673,757,1591,1617]" captionStart="FIGURE 9" captionStartId="16.[152,255,1184,1208]" captionTargetBox="[159,1428,181,1160]" captionTargetId="figure-19@16.[159,1428,181,1160]" captionTargetPageId="16" captionText="FIGURE 9. Maxillary and mandibular dentition of Podogymnura intermedia from Mt. Hamiguitan (A, B; FMNH 186805, holotype); and (C, D; FMNH 194751), from Mt. Kampalili. Scale bar = 5mm." figureDoi="http://doi.org/10.5281/zenodo.7532506" httpUri="https://zenodo.org/record/7532506/files/figure.png" pageId="17" pageNumber="261">9A, 9B</figureCitation>
|
||
,
|
||
<tableCitation id="C6EDEAE33A3D976AFCB1F9A74B10421C" box="[770,855,1591,1617]" captionStart="TABLE 2" captionStartId="9.[152,244,151,175]" captionTargetPageId="9" captionText="TABLE 2. Cranial and dental measurements (mm) of adult Podogymnura from Dinagat and Mindanao Islands, including the new species from eastern Mindanao. Measurements of sample size (n) greater than 3 are given as mean ± 1 SD and their ranges; as average for sample sizes less than 3. Sample size less than n shown in parentheses after the range." httpUri="http://table.plazi.org/id/DF108FD03A259772FF2BFF074D1D44B5" pageId="17" pageNumber="261" tableUuid="DF108FD03A259772FF2BFF074D1D44B5">Table 2</tableCitation>
|
||
) are similar to those of congeners in most respects (see
|
||
<taxonomicName id="4C6FA4DB3A3D976AFF78F9CB492D4238" authorityName="Mearns" authorityYear="1905" box="[203,362,1627,1653]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3D976AFF78F9CB492D4238" box="[203,362,1627,1653]" italics="true" pageId="17" pageNumber="261">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
Diagnosis), usually differing only in relative sizes. The presence of a discernible lingual lobe in P
|
||
<quantity id="4C9772BD3A3D976AFF1BF9EF489642D4" box="[168,209,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="17" pageNumber="261" unit="in" value="3.0">3 in</quantity>
|
||
specimens from Hamiguitan is shared with
|
||
<taxonomicName id="4C6FA4DB3A3D976AFD09F9EF4B2F42D4" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[698,872,1663,1689]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFD09F9EF4B2F42D4" box="[698,872,1663,1689]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
; this lobe is absent among all other
|
||
<emphasis id="B91B034A3A3D976AFB44F9EF4DDA42D4" box="[1271,1437,1663,1689]" italics="true" pageId="17" pageNumber="261">
|
||
<taxonomicName id="4C6FA4DB3A3D976AFB44F9EF4DDE42D4" authorityName="Mearns" authorityYear="1905" box="[1271,1433,1663,1689]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="genus">Podogymnura</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
P4 is proportionately larger and more square-shaped in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFCADF9334B9742F0" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[798,976,1699,1725]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFCADF9334B9742F0" box="[798,976,1699,1725]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFBBFF9334CE342F0" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[1036,1188,1699,1725]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFBBFF9334CE342F0" box="[1036,1188,1699,1725]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C6FA4DB3A3D976AFABAF9334D1B42F1" box="[1289,1372,1699,1724]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFABAF9334D1B42F1" box="[1289,1372,1699,1724]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C6FA4DB3A3D976AFA3BF93348AC42AD" authorityName="Sanborn" authorityYear="1953" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3D976AFA3BF93348AC42AD" italics="true" pageId="17" pageNumber="261">P. minima</emphasis>
|
||
</taxonomicName>
|
||
. A small, inconspicuous cusp at base of talonid on m1 and m2 is present in all specimens of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFAB4F9574DDB42AC" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[1287,1436,1735,1761]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFAB4F9574DDB42AC" box="[1287,1436,1735,1761]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Hamiguitan (
|
||
<figureCitation id="1354C3DD3A3D976AFED4F97B49FC4348" box="[359,443,1771,1797]" captionStart="FIGURE 8" captionStartId="15.[152,255,1922,1946]" captionTargetBox="[151,1436,1224,1899]" captionTargetId="figure-123@15.[151,1436,1224,1899]" captionTargetPageId="15" captionText="FIGURE 8. Dorsal (A), ventral (B) and lateral (C) views of the skull and lateral view of the mandible (D) of Podogymnura intermedia (FMNH 186805, holotype); scale bar refers to these images. E, occlusal view of the second mandibular molar of Podogymnura intermedia holotype; arrow points to a small cusp at the base of talonid, between the entoconid and metaconid." figureDoi="http://doi.org/10.5281/zenodo.7532502" httpUri="https://zenodo.org/record/7532502/files/figure.png" pageId="17" pageNumber="261">Fig. 8E</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE4AF97B4B954348" authority="(Heaney & Morgan 1982)" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[505,978,1771,1797]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3D976AFE4AF97B4AE04348" box="[505,679,1771,1797]" italics="true" pageId="17" pageNumber="261">P. aureospinula</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFEA2A93A3D976AFD06F97B4B8D4348" author="Heaney, L. R. & Morgan, G. S." box="[693,970,1771,1797]" pageId="17" pageNumber="261" pagination="13 - 26" refId="ref16439" refString="Heaney, L. R. & Morgan, G. S. (1982) A new species of gynmure, Podogymnura, (Mammalia: Erinaceidae) from Dinagat Island, Philippines. Proceedings of the Biological Society of Washington, 95, 13 - 26." type="journal article" year="1982">Heaney & Morgan 1982</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; a poorly-developed version is present in a small percentage of
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE3EF89F499A4365" box="[397,477,1807,1832]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3D976AFE3EF89F499A4365" box="[397,477,1807,1832]" italics="true" pageId="17" pageNumber="261">P. truei</emphasis>
|
||
</taxonomicName>
|
||
, and is absent in other populations, including
|
||
<taxonomicName id="4C6FA4DB3A3D976AFC51F89F4C314364" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[994,1142,1807,1833]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFC51F89F4C314364" box="[994,1142,1807,1833]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Kampalili.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3758CD33A3D9769FF74F8A34C004134" lastPageId="18" lastPageNumber="262" pageId="17" pageNumber="261" type="biology_ecology">
|
||
<paragraph id="8BD0DF583A3D976AFF74F8A34B4B433C" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="261">
|
||
<emphasis id="B91B034A3A3D976AFF74F8A349614300" bold="true" box="[199,294,1843,1869]" pageId="17" pageNumber="261">Ecology</emphasis>
|
||
.
|
||
<materialsCitation id="3B07D5053A3D976AFE82F8A34B4F433C" county="Mt. Kampalili" location="We" municipality="Mt. Hamiguitan" pageId="17" pageNumber="261" specimenCount="1">
|
||
<location id="8EB089833A3D976AFE82F8A349114300" LSID="urn:lsid:plazi:treatment:03C66E4E3A229769FF24FF074C004134:8EB089833A3D976AFE82F8A349114300" box="[305,342,1843,1869]" county="Mt. Kampalili" municipality="Mt. Hamiguitan" name="We" pageId="17" pageNumber="261">We</location>
|
||
recorded
|
||
<taxonomicName id="4C6FA4DB3A3D976AFE72F8A34A144300" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[449,595,1843,1869]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFE72F8A34A144300" box="[449,595,1843,1869]" italics="true" pageId="17" pageNumber="261">P.intermedia</emphasis>
|
||
</taxonomicName>
|
||
on
|
||
<collectingMunicipality id="6BB445223A3D976AFDCAF8A34B6D4300" box="[633,810,1843,1869]" pageId="17" pageNumber="261">Mt. Hamiguitan</collectingMunicipality>
|
||
and
|
||
<collectingCounty id="62B1A7D43A3D976AFCEFF8A34BBF4300" box="[860,1016,1843,1869]" pageId="17" pageNumber="261">Mt. Kampalili</collectingCounty>
|
||
in different
|
||
<typeStatus id="54D461FA3A3D976AFBCFF8A34CF04300" box="[1148,1207,1843,1869]" pageId="17" pageNumber="261">types</typeStatus>
|
||
of forest formations, and so we present the ecological information separately
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3D9769FF74F8EB4D3845C4" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,150,1401]" lastPageId="18" lastPageNumber="262" pageId="17" pageNumber="261">
|
||
On Mt. Hamiguitan, we captured
|
||
<taxonomicName id="4C6FA4DB3A3D976AFD88F8EB4A8843D8" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[571,719,1915,1941]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="261" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3D976AFD88F8EB4A8843D8" box="[571,719,1915,1941]" italics="true" pageId="17" pageNumber="261">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
in primary montane forest over ultramafic soil at
|
||
<quantity id="4C9772BD3A3D976AFB45F8EA4D7B43D8" box="[1270,1340,1914,1941]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="261" unit="m" value="950.0">950 m</quantity>
|
||
to
|
||
<quantity id="4C9772BD3A3D976AFAEEF8EB48EA43F4" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="17" pageNumber="261" unit="m" value="1128.0">1,128 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFFEA2A93A3D976AFF08F80F49C343F4" author="Balete, D. S. & Quidlat, R. S. & Ibanez, J. C." box="[187,388,1951,1977]" pageId="17" pageNumber="261" pagination="65 - 80" refId="ref15423" refString="Balete, D. S., Quidlat, R. S. & Ibanez, J. C. (2006) The non-volant small mammals of Mt. Hamiguitan, Eastern Mindanao, Philippines. Banwa, 3, 65 - 80." type="journal article" year="2006">
|
||
Balete
|
||
<emphasis id="B91B034A3A3D976AFEB9F80F497D43F4" box="[266,314,1951,1977]" italics="true" pageId="17" pageNumber="261">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
). We did not record it in lowland forest at
|
||
<quantity id="4C9772BD3A3D976AFCE2F80E4BD043F4" box="[849,919,1950,1977]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="17" pageNumber="261" unit="m" value="525.0">525 m</quantity>
|
||
in 924 trap-nights. At
|
||
<quantity id="4C9772BD3A3D976AFB3CF80E4C9043F4" box="[1167,1239,1950,1977]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="261" unit="m" value="950.0">950 m</quantity>
|
||
, three individuals were caught in 228 trap-nights with earthworm bait (1.3% success) compared to one individual caught in 672 trap-nights with roasted coconut coated with peanut butter (0.15%); none were captured in traps set above ground on vines and trees. On Mt. Kampalili, we recorded this species in upper montane and mossy forest at ca.
|
||
<quantity id="4C9772BD3A3E9769FAF3FF2B4DDB4498" box="[1344,1436,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="18" pageNumber="262" unit="m" value="1470.0">1,470 m</quantity>
|
||
–
|
||
<quantity id="4C9772BD3A3E9769FF1DFF4F494B44B4" box="[174,268,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="18" pageNumber="262" unit="m" value="1900.0">1,900 m</quantity>
|
||
elevation; forested habitats below and above these elevations were not surveyed. Seven
|
||
<taxonomicName id="4C6FA4DB3A3E9769FAB6FF4F4DDB44B4" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[1285,1436,223,249]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3E9769FAB6FF4F4DDB44B4" box="[1285,1436,223,249]" italics="true" pageId="18" pageNumber="262">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
recorded during 2010 were captured in 247 trap-nights with live earthworm bait (2.83 % trap success); none were captured in 506 trap-nights using coconut bait. All were captured on the ground, and none in traps set in trees or on vines, and nearly all were captured at night. On the adjacent peak of Mt. Kangayag (ca.
|
||
<quantity id="4C9772BD3A3E9769FB19FEDB4D4F4528" box="[1194,1288,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="18" pageNumber="262" unit="m" value="1630.0">1,630 m</quantity>
|
||
) of the same mountain range, we captured no gymnures in 2,140 trap-nights, of which 250 were baited with live earthworms.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3E9769FF74FE034D344670" blockId="18.[151,1437,150,1401]" pageId="18" pageNumber="262">
|
||
The apparent absence of
|
||
<taxonomicName id="4C6FA4DB3A3E9769FE57FE034A3D45E0" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[484,634,403,429]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3E9769FE57FE034A3D45E0" box="[484,634,403,429]" italics="true" pageId="18" pageNumber="262">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
in the lowlands and restriction to montane and mossy forest is similar to
|
||
<taxonomicName id="4C6FA4DB3A3E9769FF06FE274961459D" authorityName="Sanborn" authorityYear="1953" box="[181,294,439,464]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3E9769FF06FE274961459D" box="[181,294,439,464]" italics="true" pageId="18" pageNumber="262">P. minima</emphasis>
|
||
</taxonomicName>
|
||
, which occurs in secondary and old-growth montane and mossy forest from ca.
|
||
<quantity id="4C9772BD3A3E9769FB08FE274D5E459C" box="[1211,1305,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="262" unit="m" value="1300.0">1,300 m</quantity>
|
||
to
|
||
<quantity id="4C9772BD3A3E9769FA8CFE274DDB459C" box="[1343,1436,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="262" unit="m" value="2800.0">2,800 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFFEA2A93A3E9769FF2CFE4B493C45B8" author="Heaney, L. R. & Balete, D. S. & Dolar, L. & Alcala, A. C. & Dans, A. & Gonzales, P. C. & Ingle, N. R. & Lepiten, M. & Oliver, W. & Ong, P. & Rickart, E. A. & Tabaranza, B. R. Jr. & Utzurrum, R. C. B." box="[159,379,475,501]" pageId="18" pageNumber="262" pagination="1 - 61" refId="ref16635" refString="Heaney, L. R., Balete, D. S., Dolar, L., Alcala, A. C., Dans, A., Gonzales, P. C., Ingle, N. R., Lepiten, M., Oliver, W., Ong, P., Rickart, E. A., Tabaranza, B. R. Jr. & Utzurrum, R. C. B. (1998) A synopsis of the mammalian fauna of the Philippine Islands. Fieldiana: Zoology, New Series, 88, 1 - 61." type="journal article" year="1998">
|
||
Heaney
|
||
<emphasis id="B91B034A3A3E9769FF4DFE4B497645B8" box="[254,305,475,501]" italics="true" pageId="18" pageNumber="262">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
, 2006), and to
|
||
<taxonomicName id="4C6FA4DB3A3E9769FD95FE4B4A3F45B9" box="[550,632,475,500]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="truei">
|
||
<emphasis id="B91B034A3A3E9769FD95FE4B4A3F45B9" box="[550,632,475,500]" italics="true" pageId="18" pageNumber="262">P. truei</emphasis>
|
||
</taxonomicName>
|
||
, which has been recorded from montane and mossy forest at ca.
|
||
<quantity id="4C9772BD3A3E9769FAEEFE4B48EA4654" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="18" pageNumber="262" unit="m" value="1640.0">1,640 m</quantity>
|
||
to at least
|
||
<quantity id="4C9772BD3A3E9769FE99FE6E49C04654" box="[298,391,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="18" pageNumber="262" unit="m" value="2250.0">2,250 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFFEA2A93A3E9769FE24FE6F4A124654" author="Hoogstraal, H." box="[407,597,510,537]" pageId="18" pageNumber="262" pagination="1 - 33" refId="ref16904" refString="Hoogstraal, H. (1951) Philippine zoological expedition, 1946 - 1947. Narrative and itinerary. Fieldiana: Zoology, 33, 1 - 33." type="journal article" year="1951">Hoogstraal 1951</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFEA2A93A3E9769FDD1FE6F4B434654" author="Sanborn, C. C." box="[610,772,510,537]" pageId="18" pageNumber="262" pagination="1 - 158" refId="ref17972" refString="Sanborn, C. C. (1952) Philippine zoological expedition, 1946 - 1947: Mammals. Fieldiana: Zoology, 33, 1 - 158." type="journal article" year="1952">Sanborn 1952</bibRefCitation>
|
||
). In contrast,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FC2CFE6F4C084654" baseAuthorityName="Heaney & Morgan" baseAuthorityYear="1982" box="[927,1103,511,537]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="aureospinula">
|
||
<emphasis id="B91B034A3A3E9769FC2CFE6F4C084654" box="[927,1103,511,537]" italics="true" pageId="18" pageNumber="262">P. aureospinula</emphasis>
|
||
</taxonomicName>
|
||
occurs in secondary and oldgrowth lowland forest at low elevations (
|
||
<bibRefCitation id="EFFEA2A93A3E9769FDD1FDB34B304670" author="Heaney, L. R. & Morgan, G. S." box="[610,887,547,573]" pageId="18" pageNumber="262" pagination="13 - 26" refId="ref16439" refString="Heaney, L. R. & Morgan, G. S. (1982) A new species of gynmure, Podogymnura, (Mammalia: Erinaceidae) from Dinagat Island, Philippines. Proceedings of the Biological Society of Washington, 95, 13 - 26." type="journal article" year="1982">Heaney & Morgan 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFEA2A93A3E9769FC30FDB34CC54670" author="Heaney, L. R. & Rabor, D. S." box="[899,1154,547,573]" pageId="18" pageNumber="262" pagination="1 - 30" refId="ref16391" refString="Heaney, L. R. & Rabor, D. S. (1982) An annotated checklist of the mammals of Dinagat and Siargao islands, Philippines. Occasional Papers of the Museum of Zoology, University of Michigan, 699, 1 - 30." type="journal article" year="1982">Heaney & Rabor 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFEA2A93A3E9769FB3DFDB34D204670" author="Heaney, L. R. & Dolar, M. L. & Balete, D. S. & Esselstyn, J. A. & Rickart, E. A. & Sedlock, J. L." box="[1166,1383,547,573]" pageId="18" pageNumber="262" refId="ref16746" refString="Heaney, L. R., Dolar, M. L., Balete, D. S., Esselstyn, J. A., Rickart, E. A. & Sedlock, J. L. (2010) Synopsis of Philippine Mammals. Field Museum website, http: // www. fieldmuseum. org / philippine _ mammals /" type="url" year="2010">
|
||
Heaney
|
||
<emphasis id="B91B034A3A3E9769FB58FDB34D5B4670" box="[1259,1308,547,573]" italics="true" pageId="18" pageNumber="262">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3E9769FF74FDD74B0347CC" blockId="18.[151,1437,150,1401]" pageId="18" pageNumber="262">
|
||
Stomach contents of two individuals from Mt. Hamiguitan contained mainly chewed remains of arthropod exoskeletons, including coleopterans and their larvae, as well as an operculum and shell fragments of small land snails. A few pieces that appeared to be partially digested bits of small earthworms were present. Stomach contents of
|
||
<specimenCount id="9D6914D13A3E9769FF07FD2349214680" box="[180,358,691,717]" pageId="18" pageNumber="262" type="generic">three specimens</specimenCount>
|
||
from Mt. Kampalili contained chewed remains of arthropod exoskeletons, including coleopterans and centipedes, and two contained pieces of earthworms. The presence of land snails in the diet of this species is the first record of this food item among
|
||
<taxonomicName id="4C6FA4DB3A3E9769FDD8FD6B4B4D4758" authorityName="Mearns" authorityYear="1905" box="[619,778,763,789]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3E9769FDD8FD6B4B4D4758" box="[619,778,763,789]" italics="true" pageId="18" pageNumber="262">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
. These diet and trapping data indicate that, as with other
|
||
<taxonomicName id="4C6FA4DB3A3E9769FF24FC8F49714774" authorityName="Mearns" authorityYear="1905" box="[151,310,799,825]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3E9769FF24FC8F49714774" box="[151,310,799,825]" italics="true" pageId="18" pageNumber="262">Podogymnura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FEF1FC8F49924774" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[322,469,799,825]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3E9769FEF1FC8F49924774" box="[322,469,799,825]" italics="true" pageId="18" pageNumber="262">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
forages on the ground for leaf-litter invertebrates. These feeding habits may differ from those of
|
||
<taxonomicName id="4C6FA4DB3A3E9769FF45FCD349234711" authorityName="Sanborn" authorityYear="1953" box="[246,356,835,860]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="B91B034A3A3E9769FF45FCD349234711" box="[246,356,835,860]" italics="true" pageId="18" pageNumber="262">P. minima</emphasis>
|
||
</taxonomicName>
|
||
, which feeds more commonly on earthworms than arthropods (
|
||
<bibRefCitation id="EFFEA2A93A3E9769FB90FCD34CBE4710" author="Heaney, L. R. & Tabaranza, B. R. Jr. & Rickart, E. A. & Balete, D. S. & Ingle, N. R." box="[1059,1273,835,861]" pageId="18" pageNumber="262" pagination="1 - 63" refId="ref16812" refString="Heaney, L. R., Tabaranza, B. R. Jr., Rickart, E. A., Balete, D. S. & Ingle, N. R. (2006) The mammals of Mt. Kitanglad Nature Park, Mindanao, Philippines. Fieldiana: Zoology (New Series), 112, 1 - 63. https: // doi. org / 10.3158 / 0015 - 0754 (2006) 186 [1: TMOMKN] 2.0. CO; 2" type="journal article" year="2006">
|
||
Heaney
|
||
<emphasis id="B91B034A3A3E9769FB33FCD34CF74710" box="[1152,1200,835,861]" italics="true" pageId="18" pageNumber="262">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
), although this difference may simply reflect differences in prey availability.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3E9769FF74FC1B4C62405C" blockId="18.[151,1437,150,1401]" pageId="18" pageNumber="262">
|
||
An adult male from Mt. Hamiguitan taken in May had testes measuring 11 x
|
||
<quantity id="4C9772BD3A3E9769FBE8FC1A4CE447E8" box="[1115,1187,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="262" unit="mm" value="5.0">5 mm</quantity>
|
||
; an adult female was nulliparous. The presence of a juvenile in May suggests that some breeding occurred earlier in the year. None of the specimens from Mt. Kampalili captured in February and May (
|
||
<specimenCount id="9D6914D13A3E9769FCD5FC434B9347A0" box="[870,980,979,1005]" pageId="18" pageNumber="262" type="adult">five adult</specimenCount>
|
||
males,
|
||
<specimenCount id="9D6914D13A3E9769FB9DFC434CE147A0" box="[1070,1190,979,1005]" pageId="18" pageNumber="262" type="adult">three adult</specimenCount>
|
||
females, three young adult females and one young adult female) showed signs of reproductive activity.
|
||
</paragraph>
|
||
<paragraph id="8BD0DF583A3E9769FF24FB8B4C004134" blockId="18.[151,1437,150,1401]" pageId="18" pageNumber="262">
|
||
Three species of murid rodents (
|
||
<taxonomicName id="4C6FA4DB3A3E9769FDA2FB8B4ABC4078" authorityName="Balete, Heaney, Rickart, Quidlat & Ibanez" authorityYear="2008" box="[529,763,1051,1077]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="hamiguitan">
|
||
<emphasis id="B91B034A3A3E9769FDA2FB8B4ABC4078" box="[529,763,1051,1077]" italics="true" pageId="18" pageNumber="262">Batomys hamiguitan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FCBFFB8B4B9F4078" authorityName="Mearns" authorityYear="1905" box="[780,984,1051,1077]" class="Mammalia" family="Muridae" genus="Bullimus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="bagobus">
|
||
<emphasis id="B91B034A3A3E9769FCBFFB8B4B9F4078" box="[780,984,1051,1077]" italics="true" pageId="18" pageNumber="262">Bullimus bagobus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6FA4DB3A3E9769FBA8FB8B4C874079" box="[1051,1216,1051,1076]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="B91B034A3A3E9769FBA8FB8B4C874079" box="[1051,1216,1051,1076]" italics="true" pageId="18" pageNumber="262">Rattus everetti</emphasis>
|
||
</taxonomicName>
|
||
) were documented as being sympatric with
|
||
<taxonomicName id="4C6FA4DB3A3E9769FE0AFBAF4A164014" authorityName="Balete & Heaney & Rickart & Quidlat & Rowsey & Olson" authorityYear="2023" box="[441,593,1087,1113]" class="Mammalia" family="Erinaceidae" genus="Podogymnura" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="intermedia">
|
||
<emphasis id="B91B034A3A3E9769FE0AFBAF4A164014" box="[441,593,1087,1113]" italics="true" pageId="18" pageNumber="262">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
on Mt. Hamiguitan along with the Mindanao shrew (
|
||
<taxonomicName id="4C6FA4DB3A3E9769FB7EFBAF4DD34014" authorityName="Miller" authorityYear="1910" box="[1229,1428,1087,1113]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="beatus">
|
||
<emphasis id="B91B034A3A3E9769FB7EFBAF4DD34014" box="[1229,1428,1087,1113]" italics="true" pageId="18" pageNumber="262">Crocidura beatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFEA2A93A3E9769FF24FBF349274030" author="Balete, D. S. & Quidlat, R. S. & Ibanez, J. C." box="[151,352,1123,1149]" pageId="18" pageNumber="262" pagination="65 - 80" refId="ref15423" refString="Balete, D. S., Quidlat, R. S. & Ibanez, J. C. (2006) The non-volant small mammals of Mt. Hamiguitan, Eastern Mindanao, Philippines. Banwa, 3, 65 - 80." type="journal article" year="2006">
|
||
Balete
|
||
<emphasis id="B91B034A3A3E9769FF55FBF349504030" box="[230,279,1123,1149]" italics="true" pageId="18" pageNumber="262">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFEA2A93A3E9769FED8FBF349EF4030" author="Balete, D. S. & Heaney, L. R. & Rickart, E. A. & Quidlat, R. S. & Ibanez, J. C." box="[363,424,1123,1149]" pageId="18" pageNumber="262" pagination="411 - 428" refId="ref15468" refString="Balete, D. S., Heaney, L. R., Rickart, E. A., Quidlat, R. S. & Ibanez, J. C. (2008) A new species of Batomys (Muridae: Murinae) from eastern Mindanao Island, Philippines. Proceedings of the Biological Society of Washington, 121, 411 - 428. https: // doi. org / 10.2988 / 07 - 47.1" type="journal article" year="2008">2008</bibRefCitation>
|
||
). In addition to these native species, we recorded the non-native spiny ricefield rat (
|
||
<taxonomicName id="4C6FA4DB3A3E9769FAE6FBF348AA40EC" baseAuthorityName="Peale" baseAuthorityYear="1848" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="exulans">
|
||
<emphasis id="B91B034A3A3E9769FAE6FBF348AA40EC" italics="true" pageId="18" pageNumber="262">Rattus exulans</emphasis>
|
||
</taxonomicName>
|
||
) at two disturbed sites near the
|
||
<typeStatus id="54D461FA3A3E9769FDF9FB174A3D40EC" box="[586,634,1159,1185]" pageId="18" pageNumber="262">type</typeStatus>
|
||
locality: a narrow patch of non-native cogon grass,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FB04FB174DD140EC" box="[1207,1430,1159,1185]" class="Liliopsida" family="Poaceae" genus="Imperata" kingdom="Plantae" order="Poales" pageId="18" pageNumber="262" phylum="Tracheophyta" rank="species" species="cylindrica">
|
||
<emphasis id="B91B034A3A3E9769FB04FB174DD140EC" box="[1207,1430,1159,1185]" italics="true" pageId="18" pageNumber="262">Imperata cylindrica</emphasis>
|
||
</taxonomicName>
|
||
, surrounding the shoreline and dry lakebed of Tinagong Dagat, a seasonal lake; and in disturbed vegetation along a foot-trail, but not in the nearby forest interior. On Mt. Kampalili this species was associated with
|
||
<taxonomicName id="4C6FA4DB3A3E9769FAA0FB5F4DD040A4" box="[1299,1431,1231,1257]" class="Mammalia" family="Muridae" genus="Apomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B91B034A3A3E9769FAA0FB5F4D2B40A4" box="[1299,1388,1231,1257]" italics="true" pageId="18" pageNumber="262">Apomys</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FF24FB6349264140" authorityName="Mearns" authorityYear="1905" box="[151,353,1267,1293]" class="Mammalia" family="Muridae" genus="Bullimus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="bagobus">
|
||
<emphasis id="B91B034A3A3E9769FF24FB6349264140" box="[151,353,1267,1293]" italics="true" pageId="18" pageNumber="262">Bullimus bagobus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FEDCFB634AD54140" authority=", Baletemys" authorityName="Baletemys" box="[367,658,1267,1293]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="B91B034A3A3E9769FEDCFB634A564141" box="[367,529,1267,1292]" italics="true" pageId="18" pageNumber="262">Rattus everetti</emphasis>
|
||
,
|
||
<emphasis id="B91B034A3A3E9769FDACFB634AD54140" box="[543,658,1267,1293]" italics="true" pageId="18" pageNumber="262">Baletemys</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91B034A3A3E9769FD2AFB634B444140" box="[665,771,1267,1293]" italics="true" pageId="18" pageNumber="262">kampalili</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFEA2A93A3E9769FCA1FB634BB64140" author="Rowsey, D. M. & Duya, M. R. M. & Ibanez, J. C. & Jansa, S. A. & Rickart, E. A. & Heaney, L. R." box="[786,1009,1267,1293]" pageId="18" pageNumber="262" refId="ref17774" refString="Rowsey, D. M., Duya, M. R. M., Ibanez, J. C., Jansa, S. A., Rickart, E. A. & Heaney, L. R. (2022) A new genus and species of shrewlike mouse (Rodentia: Muridae) from a new center of endemism in eastern Mindanao, Philippines. Journal of Mammalogy, 103. https: // doi. org / 10.1093 / jmammal / gyac 057." type="book" year="2022">
|
||
Rowsey
|
||
<emphasis id="B91B034A3A3E9769FCC6FB634BE04140" box="[885,935,1267,1293]" italics="true" pageId="18" pageNumber="262">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
), an undescribed species of
|
||
<taxonomicName id="4C6FA4DB3A3E9769FA98FB634DD24141" authorityName="Mearns" authorityYear="1905" box="[1323,1429,1267,1292]" class="Mammalia" family="Muridae" genus="Tarsomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91B034A3A3E9769FA98FB634DD24141" box="[1323,1429,1267,1292]" italics="true" pageId="18" pageNumber="262">Tarsomys</emphasis>
|
||
</taxonomicName>
|
||
, and one squirrel,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FEE6FA874AC8417C" baseAuthorityName="Waterhouse" baseAuthorityYear="1839" box="[341,655,1303,1329]" class="Mammalia" family="Sciuridae" genus="Sundasciurus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="philippinensis">
|
||
<emphasis id="B91B034A3A3E9769FEE6FA874AC8417C" box="[341,655,1303,1329]" italics="true" pageId="18" pageNumber="262">Sundasciurus philippinensis</emphasis>
|
||
</taxonomicName>
|
||
(specimens in FMNH). The commensal house shrew,
|
||
<taxonomicName id="4C6FA4DB3A3E9769FB56FA874DD2417D" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[1253,1429,1303,1329]" class="Mammalia" family="Soricidae" genus="Suncus" kingdom="Animalia" order="Soricomorpha" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="murinus">
|
||
<emphasis id="B91B034A3A3E9769FB56FA874DD2417D" box="[1253,1429,1303,1329]" italics="true" pageId="18" pageNumber="262">Suncus murinus</emphasis>
|
||
</taxonomicName>
|
||
, was also present. All of the native co-occurring species are endemic to the
|
||
<collectingCountry id="F3789FC83A3E9769FC5CFAAB4C374118" box="[1007,1136,1339,1365]" name="Philippines" pageId="18" pageNumber="262">Philippines</collectingCountry>
|
||
, and all but
|
||
<taxonomicName id="4C6FA4DB3A3E9769FB4DFAAB4D354119" box="[1278,1394,1339,1364]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="262" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="B91B034A3A3E9769FB4DFAAB4D354119" box="[1278,1394,1339,1364]" italics="true" pageId="18" pageNumber="262">R. everetti</emphasis>
|
||
</taxonomicName>
|
||
are restricted to the Mindanao Faunal Region (
|
||
<bibRefCitation id="EFFEA2A93A3E9769FDCBFACF4B564134" author="Heaney, L. R." box="[632,785,1375,1401]" pageId="18" pageNumber="262" pagination="127 - 165" refId="ref16332" refString="Heaney, L. R. (1986) Biogeography of mammals in Southeast Asia: estimates of rates of colonization, extinction, and speciation. Biological Journal of the Linnean Society, 28, 127 - 165. https: // doi. org / 10.1111 / j. 1095 - 8312.1986. tb 01752. x" type="journal article" year="1986">Heaney 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFEA2A93A3E9769FCAEFACF4BB44134" author="Heaney, L. R. & Tabaranza, B. R. Jr. & Rickart, E. A. & Balete, D. S. & Ingle, N. R." box="[797,1011,1375,1401]" pageId="18" pageNumber="262" pagination="1 - 63" refId="ref16812" refString="Heaney, L. R., Tabaranza, B. R. Jr., Rickart, E. A., Balete, D. S. & Ingle, N. R. (2006) The mammals of Mt. Kitanglad Nature Park, Mindanao, Philippines. Fieldiana: Zoology (New Series), 112, 1 - 63. https: // doi. org / 10.3158 / 0015 - 0754 (2006) 186 [1: TMOMKN] 2.0. CO; 2" type="journal article" year="2006">
|
||
Heaney
|
||
<emphasis id="B91B034A3A3E9769FCCAFACF4BED4134" box="[889,938,1375,1401]" italics="true" pageId="18" pageNumber="262">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFEA2A93A3E9769FC4DFACF4C7C4134" author="Heaney, L. R. & Dolar, M. L. & Balete, D. S. & Esselstyn, J. A. & Rickart, E. A. & Sedlock, J. L." box="[1022,1083,1375,1401]" pageId="18" pageNumber="262" refId="ref16746" refString="Heaney, L. R., Dolar, M. L., Balete, D. S., Esselstyn, J. A., Rickart, E. A. & Sedlock, J. L. (2010) Synopsis of Philippine Mammals. Field Museum website, http: // www. fieldmuseum. org / philippine _ mammals /" type="url" year="2010">2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |