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<document id="D6643D78183FDE56B3CE4DC13901441C" ID-DOI="10.5281/zenodo.181365" ID-GBIF-Dataset="054c3e51-58a4-477c-8ed3-cf546e522202" ID-ISSN="1175-5326" ID-Zenodo-Dep="181365" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459832588023" checkinUser="plazi" docAuthor="Grebennikov, Vasily V." docDate="2008" docId="03C2B611FFB4A76FFF3DF9A5FD2CF8CD" docLanguage="en" docName="zt01732p053.pdf" docOrigin="Zootaxa 1732" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Rioneta uluguruensis Johnson 1975" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFFBCE69FFB6A76CFFAAFFD8FF97FF9C" masterDocTitle="A featherwing beetle without wings: re-discovery and second species of Rioneta (Coleoptera: Ptiliidae) from the Uluguru Mountains, Tanzania" masterLastPageNumber="53" masterPageNumber="45" pageNumber="47" updateTime="1698223062435" updateUser="plazi">
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<mods:title id="E90AAF8A90CFFCB7EA5F1B55054D8129">A featherwing beetle without wings: re-discovery and second species of Rioneta (Coleoptera: Ptiliidae) from the Uluguru Mountains, Tanzania</mods:title>
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<mods:namePart id="F5D0C240CA743D0B7F15D05C4A029652">Grebennikov, Vasily V.</mods:namePart>
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<mods:date id="6D44C77C3C47330F4FE2604E9FBB461C">2008</mods:date>
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<treatment id="03C2B611FFB4A76FFF3DF9A5FD2CF8CD" ID-DOI="http://doi.org/10.5281/zenodo.6231926" ID-GBIF-Taxon="119358154" ID-Zenodo-Dep="6231926" LSID="urn:lsid:plazi:treatment:03C2B611FFB4A76FFF3DF9A5FD2CF8CD" httpUri="http://treatment.plazi.org/id/03C2B611FFB4A76FFF3DF9A5FD2CF8CD" lastPageId="3" lastPageNumber="47" pageId="2" pageNumber="47">
<subSubSection id="C371548CFFB4A76EFF3DF9A5FECDF926" pageId="2" pageNumber="47" type="nomenclature">
<paragraph id="8BD40707FFB4A76EFF3DF9A5FDC3F90B" blockId="2.[151,596,1661,1722]" box="[151,596,1661,1687]" pageId="2" pageNumber="47">
<heading id="D09CB06BFFB4A76EFF3DF9A5FDC3F90B" bold="true" box="[151,596,1661,1687]" fontSize="11" level="1" pageId="2" pageNumber="47" reason="1">
<taxonomicName id="4C6B7C84FFB4A76EFF3DF9A5FDC3F90B" ID-CoL="4T4L8" authority="Johnson, 1975" authorityName="Johnson" authorityYear="1975" box="[151,596,1661,1687]" class="Insecta" family="Ptiliidae" genus="Rioneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="uluguruensis">
<emphasis id="B91FDB15FFB4A76EFF3DF9A5FDC3F90B" bold="true" box="[151,596,1661,1687]" pageId="2" pageNumber="47">
<emphasis id="B91FDB15FFB4A76EFF3DF9A5FE0BF90B" bold="true" box="[151,412,1661,1687]" italics="true" pageId="2" pageNumber="47">Rioneta uluguruensis</emphasis>
<bibRefCitation id="EFFA7AF6FFB4A76EFE09F9A5FDC3F90B" author="Johnson" box="[419,596,1661,1687]" pageId="2" pageNumber="53" refString="Johnson, C. (1975) Mission entomologique du Musee Royal de l'Afrique Centrale aux Monts Uluguru, Tanzanie (L. Berger, N. Leleup et J. Debecker, V - VIII. 1971). 5. Coleoptera: Ptiliidae. Revue de Zoologie Africaine, 89, 719 - 722." type="journal article" year="1975">Johnson, 1975</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD40707FFB4A76EFF3DF978FECDF926" blockId="2.[151,596,1661,1722]" box="[151,346,1696,1722]" pageId="2" pageNumber="47">
(
<figureCitation id="13501B82FFB4A76EFF35F978FE9EF926" box="[159,265,1696,1722]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1516,1540]" captionTargetBox="[264,1322,183,1504]" captionTargetId="figure@3.[264,1323,183,1506]" captionTargetPageId="3" captionText="FIGURES 1 4. Habitus images of Rioneta beetles. 1 — R. uluguruensis, female, dorsal; 2 — R. uluguruensis, male, ventral; 3 — R. inexpectata sp. n., male, dorsal; 4 — R. inexpectata sp. n., female, ventral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/181366/files/figure.png" pageId="2" pageNumber="47">Figs 1, 2</figureCitation>
,
<figureCitation id="13501B82FFB4A76EFEBFF978FEC6F926" box="[277,337,1696,1722]" captionStart-0="FIGURES 5 15" captionStart-1="FIGURES 16 22" captionStart-2="FIGURES 23 33" captionStart-3="FIGURES 34 36. 34, 35" captionStartId-0="4.[151,269,1484,1508]" captionStartId-1="5.[151,269,1226,1250]" captionStartId-2="6.[151,269,1490,1514]" captionStartId-3="7.[151,269,1086,1110]" captionTargetBox-0="[151,1436,194,1476]" captionTargetBox-1="[151,1436,194,1202]" captionTargetBox-2="[151,1436,194,1466]" captionTargetBox-3="[264,1323,194,1061]" captionTargetId-0="figure@4.[151,1436,194,1478]" captionTargetId-1="figure@5.[151,1436,194,1202]" captionTargetId-2="figure@6.[151,1436,194,1466]" captionTargetId-3="figure@7.[264,1323,194,1062]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 5 15. SEM images of Rioneta uluguruensis. 5 — habitus, ventral; 6 — head, ventral; 7 — left fore-tarsus, ventral; 8 — two apical antennomeres, arrow showing groups of sensillae, ventral; 9 — internal surface of meso- and metathorax with metendosternite, two arrows show internal part of lateral longitudinal keel on each side of metaventrite, dorsal; 10 — spermatheca and sperm pump, ventral; 11 — pretothorax and abdomen, arrow showing lateral longitudinal keel on left metaventrite, left elytron removed, fronto-lateral; 12 — pterothorax and abdomen, lateral, left elytron removed, three arrows showing spiracles I III, respectively; 13 — internal surface of abdominal tergites V X, double-pointed central arrow showing fused tergites VIII and IX, two lateral arrows showing remnants of internal apodemae of tergite IX, dorsal; 14 — abdominal apex, both elytra removed, dorsal, vertical double-pointed arrow showing fused tergites VIII and IX, horizontal arrows showing spiracles VIII IV; 15 — right part of abdomen, right elytron removed, dorsal, oblique arrow showing posterior edge of terige VII without wing cleaning (= toiletry) device, vertical arrows showing spiracles VIII III." captionText-1="FIGURES 16 22. Rioneta uluguruensis, details of morphology. 16 — habitus, ventral, left legs omitted, aedeagus and metendosternite shown; 18 — head, dorsal, antennomeres 2 11 omitted; 18 — left fore-tarsus and tibia, ventral (separately shown enlarged bifurcate spine on apical part of tibia); 19 — right hind leg, ventral (separately shown enlarged tarsus with two visible apical tarsomeres); 20 — mouthparts (maxillae, mentum, and labium), ventral; 21 — right antenna, dorsal; 22 — clypeus, dorsal." captionText-2="FIGURES 23 33. Rioneta uluguruensis, details of morphology. 23 — mesonotum, mesoscutellum, basal part of metascutellum, mesopleuron and basal part of metapleuron, elytra omitted, dorsal; 24 — pterothorax and abdomen, elytra omitted, dorsal; 25 — left metapleuron, lateral; 26 — apex of abdomen showing location of internal glands, dorsal (separately on the left showing right internal gland, enlarged); 27 — fused abdominal tergites VIII IX and partly fused tergite X, dorsal; 28 — abdominal sternites VIII—IX, male, ventral; 29, 30 — spermatheca and sperm pump, partly oblique; 31, 32 — aedeagus with apex directed upwards, dorsal (31) and ventral (32); 33 — abdominal sternite IX, male, ventral." captionText-3="FIGURES 34 36. 34, 35 — Rioneta uluguruensis, spermatheca, transparent light microscopy images, scale 0.05 mm; 36 — map of Eastern Arc Mountains with arrow showing Uluguru Mountains, the type locality and known area of distribution of both Rioneta species." httpUri-0="https://zenodo.org/record/181367/files/figure.png" httpUri-1="https://zenodo.org/record/181368/files/figure.png" httpUri-2="https://zenodo.org/record/181369/files/figure.png" httpUri-3="https://zenodo.org/record/181370/files/figure.png" pageId="2" pageNumber="47">535</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371548CFFB4A76EFF3DF932FB9AF898" box="[151,1037,1770,1796]" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="8BD40707FFB4A76EFF3DF932FB9AF898" blockId="2.[151,1436,1770,2036]" box="[151,1037,1770,1796]" pageId="2" pageNumber="47">
<emphasis id="B91FDB15FFB4A76EFF3DF932FE80F898" bold="true" box="[151,279,1770,1796]" pageId="2" pageNumber="47">Diagnosis.</emphasis>
Differs from
<taxonomicName id="4C6B7C84FFB4A76EFE12F933FDF7F898" box="[440,608,1771,1796]" class="Insecta" family="Ptiliidae" genus="Rioneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="inexpectata">
<emphasis id="B91FDB15FFB4A76EFE12F933FDF7F898" box="[440,608,1771,1796]" italics="true" pageId="2" pageNumber="47">R. inexpectata</emphasis>
</taxonomicName>
by markedly wider body (
<figureCitation id="13501B82FFB4A76EFC3FF932FC68F898" box="[917,1023,1770,1796]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1516,1540]" captionTargetBox="[264,1322,183,1504]" captionTargetId="figure@3.[264,1323,183,1506]" captionTargetPageId="3" captionText="FIGURES 1 4. Habitus images of Rioneta beetles. 1 — R. uluguruensis, female, dorsal; 2 — R. uluguruensis, male, ventral; 3 — R. inexpectata sp. n., male, dorsal; 4 — R. inexpectata sp. n., female, ventral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/181366/files/figure.png" pageId="2" pageNumber="47">Figs 1, 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C371548CFFB4A76EFF6CF8CAFDA1F8E0" pageId="2" pageNumber="47" type="description">
<paragraph id="8BD40707FFB4A76EFF6CF8CAFDA1F8E0" blockId="2.[151,1436,1770,2036]" pageId="2" pageNumber="47">
<emphasis id="B91FDB15FFB4A76EFF6CF8CAFEC8F8B0" bold="true" box="[198,351,1810,1836]" pageId="2" pageNumber="47">Description.</emphasis>
With the characters of the genus. Body length
<quantity id="4C93AAE2FFB4A76EFCD5F8CAFBBEF8B0" box="[895,1065,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.115" metricValueMax="1.13" metricValueMin="1.1" pageId="2" pageNumber="47" unit="mm" value="1.115" valueMax="1.13" valueMin="1.1">1.101.13 mm</quantity>
(mean
<quantity id="4C93AAE2FFB4A76EFBD5F8CAFB7DF8B0" box="[1151,1258,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="2" pageNumber="47" unit="mm" value="1.12">1.12 mm</quantity>
, n=3), pronotal length
<quantity id="4C93AAE2FFB4A76EFF4DF8E2FE06F8C8" box="[231,401,1850,1876]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="2" pageNumber="47" unit="mm" value="0.3" valueMax="0.31" valueMin="0.29">0.290.31 mm</quantity>
(mean
<quantity id="4C93AAE2FFB4A76EFE4DF8E2FDC4F8C8" box="[487,595,1850,1876]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="47" unit="mm" value="0.3">0.30 mm</quantity>
; n=3); pronotal width 0.580.62 (mean 0.60; n=3); ratio pronotal width/ length 1.931.98 (mean 1.96; n=3).
</paragraph>
</subSubSection>
<subSubSection id="C371548CFFB4A76FFF6CF852FD2CF8CD" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47" type="materials_examined">
<paragraph id="8BD40707FFB4A76FFF6CF852FD2CF8CD" blockId="2.[151,1436,1770,2036]" lastBlockId="3.[151,1436,1660,1873]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
<emphasis id="B91FDB15FFB4A76EFF6CF852FE35F838" bold="true" box="[198,418,1930,1956]" pageId="2" pageNumber="47">Material studied.</emphasis>
<typeStatus id="54D0B9A5FFB4A76EFE04F852FD8CF838" box="[430,539,1930,1956]" pageId="2" pageNumber="47" type="holotype">Holotype</typeStatus>
(dry mounted on card, female) in the collection of the Musée Royal de lAfrique Centrale, Tervuren [
<collectionCode id="ED7A9FC2FFB4A76EFE57F86AFDCAF850" box="[509,605,1970,1996]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="47">MRAC</collectionCode>
]: “
<typeStatus id="54D0B9A5FFB4A76EFDD1F86AFD6CF850" box="[635,763,1970,1996]" pageId="2" pageNumber="47" type="holotype">Holotypus</typeStatus>
” (printed on pink label); “
<collectingCountry id="F37C4797FFB4A76EFB90F86AFB3CF850" box="[1082,1195,1970,1996]" name="Tanzania" pageId="2" pageNumber="47">Tanzanie</collectingCountry>
: Mts Uluguru/ Lukwangule Plateau/ alt.
<quantity id="4C93AAE2FFB4A76EFE39F802FDA0F868" box="[403,567,2010,2036]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="2" pageNumber="47" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">24002600 m</quantity>
./ humus,
<date id="FFD521C7FFB4A76EFD03F802FCAAF868" box="[681,829,2010,2036]" pageId="2" pageNumber="47" value="1971-07-02" valueMax="1971-07-22" valueMin="1971-07-02">222/VII/71</date>
” (printed on white label); “Coll. Mus. Tervuren/ Mission Mts Uluguru/ L. Berger, N. Leleup/ J. Debecker V/
<date id="FFD521C7FFB5A76FFC28F9A4FC42F90A" box="[898,981,1660,1686]" bridgedPair="/" pageId="3" pageNumber="48" value="1971-08">Viii/71</date>
” (printed on white label); “
<typeStatus id="54D0B9A5FFB5A76FFA86F9A4FA0BF90A" box="[1324,1436,1660,1686]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
female/
<taxonomicName id="4C6B7C84FFB5A76FFF5DF97AFEC5F920" box="[247,338,1698,1724]" class="Insecta" family="Ptiliidae" genus="Rioneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Rioneta</taxonomicName>
/
<taxonomicName id="4C6B7C84FFB5A76FFEC9F97AFE6EF920" box="[355,505,1698,1724]" class="Insecta" family="Ptiliidae" genus="Rioneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="uluguruensis" status="sp. nov.">uluguruensis</taxonomicName>
<taxonomicNameLabel id="A22C666EFFB5A76FFDA8F97AFDD7F920" box="[514,576,1698,1724]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
/ det. C. Johnson 1974” (written on white label).
<typeStatus id="54D0B9A5FFB5A76FFB27F97AFB64F920" box="[1165,1267,1698,1724]" pageId="3" pageNumber="48" type="paratype">Paratype</typeStatus>
(dry mounted on card, male): similarly labelled. Additional material: 20 dry mounted, 10 mounted on Euparal microscope slides and about 100 stored in alcohol: “
<collectingCountry id="F37C4797FFB5A76FFDD9F935FD73F89B" box="[627,740,1773,1799]" name="Tanzania" pageId="3" pageNumber="48">Tanzania</collectingCountry>
: Uluguru Mts., W slope between Tchenzema vil. and Lukwangule Plateau, H-
<quantity id="4C93AAE2FFB5A76FFE2CF8CAFE73F8B0" box="[390,484,1810,1836]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="48" unit="m" value="2200.0">2200 m</quantity>
.
<date id="FFD521C7FFB5A76FFE5AF8CAFD57F8B0" box="[496,704,1810,1836]" pageId="3" pageNumber="48" value="2002-10-19" valueMax="2002-10-21" valueMin="2002-10-19">Oct. 1921, 2002</date>
, V. Grebennikov leg.” (
<collectionCode id="ED7A9FC2FFB5A76FFC7CF8CAFBA6F8B0" box="[982,1073,1810,1836]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="3" pageNumber="48">MRAC</collectionCode>
, Canadian National Collection of Insects, Arachnids, and Nematodes [
<collectionCode id="ED7A9FC2FFB5A76FFDCBF8EFFD30F8CD" box="[609,679,1847,1873]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="48">CNC</collectionCode>
]).
</paragraph>
</subSubSection>
</treatment>
</document>