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<document id="AEAD2912F23A7E247F4E2BF8A1A0FA15" ID-CLB-Dataset="46025" ID-DOI="10.11646/zootaxa.3786.1.2" ID-GBIF-Dataset="ef8a0c66-3103-4a40-8302-0a1a4055811a" ID-ISSN="1175-5326" ID-Zenodo-Dep="231537" ID-ZooBank="09DC7D2C-B15B-4DC8-9D89-FDC0E97D2776" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460407886034" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2014" docId="03C2A127FFA8F3446380FB418312E9BE" docLanguage="en" docName="zt03786p056.pdf" docOrigin="Zootaxa 3786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Choroterpes (Dilatognathus) cataractae Kluge 2012" docType="treatment" docVersion="9" masterDocId="FFFBD95FFFABF3476317FFF2816EEE71" masterDocTitle="Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla" updateTime="1698300041941" updateUser="ExternalLinkService">
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<mods:title id="9619C0817EAD04E6F20E08704931CF66">Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla</mods:title>
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<mods:namePart id="1315C8D4F99DA45B3898A27B89E2625B">Kluge, Nikita J.</mods:namePart>
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<mods:date id="472CDEF74006740377177F6C23C303B0">2014</mods:date>
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<mods:number id="7FF2F2CD6953DF38C4621BA19D56F452">3786</mods:number>
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<mods:number id="C79EF44C6533494A626F2B39222504E2">1</mods:number>
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<mods:identifier id="EA9A6AE49BD4B3B4C492ECBB4BEBF3C1" type="DOI">10.11646/zootaxa.3786.1.2</mods:identifier>
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<treatment id="03C2A127FFA8F3446380FB418312E9BE" ID-DOI="http://doi.org/10.5281/zenodo.6137920" ID-GBIF-Taxon="119422429" ID-Zenodo-Dep="6137920" LSID="urn:lsid:plazi:treatment:03C2A127FFA8F3446380FB418312E9BE" httpUri="http://treatment.plazi.org/id/03C2A127FFA8F3446380FB418312E9BE" pageId="3">
<subSubSection id="C37143BAFFA8F3446380FB4180AEEA9E" pageId="3" type="nomenclature">
<paragraph id="8BD41031FFA8F3446380FB418265EABC" blockId="3.[151,779,1203,1263]" box="[151,779,1203,1229]" pageId="3">
<heading id="D09CA75DFFA8F3446380FB418265EABC" bold="true" box="[151,779,1203,1229]" fontSize="11" level="1" pageId="3" reason="1">
<taxonomicName id="4C6B6BB2FFA8F3446380FB418265EABC" authority="Kluge 2012" authorityName="Kluge" authorityYear="2012" box="[151,779,1203,1229]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA8F3446380FB418265EABC" bold="true" box="[151,779,1203,1229]" pageId="3">
<emphasis id="B91FCC23FFA8F3446380FB418045EABC" bold="true" box="[151,299,1203,1229]" italics="true" pageId="3">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFA8F344622BFB418087EABC" bold="true" box="[316,489,1203,1229]" italics="true" pageId="3">Dilatognathus</emphasis>
)
<emphasis id="B91FCC23FFA8F34462EDFB418319EABC" bold="true" box="[506,631,1203,1229]" italics="true" pageId="3">cataractae</emphasis>
<bibRefCitation id="EFFA6DC0FFA8F344616AFB418265EABC" author="Kluge" box="[637,779,1203,1229]" pageId="3" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD41031FFA8F3446380FB2480AEEA9E" blockId="3.[151,779,1203,1263]" box="[151,448,1238,1263]" pageId="3">(Figs 115, 30, 32, 36, 37)</paragraph>
</subSubSection>
<subSubSection id="C37143BAFFA8F3446380FAEC83E0E912" pageId="3" type="materials_examined">
<paragraph id="8BD41031FFA8F3446380FAEC82BAEB0F" blockId="3.[151,1436,1310,1999]" pageId="3">
<emphasis id="B91FCC23FFA8F3446380FAEC8191EB46" bold="true" box="[151,255,1310,1335]" pageId="3">Material</emphasis>
(
<typeStatus id="54D0AE93FFA8F3446200FAEC8015EB46" box="[279,379,1310,1335]" pageId="3" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D0AE93FFA8F34462D3FAED835AEB46" box="[452,564,1311,1335]" pageId="3" type="paratype">paratypes</typeStatus>
).
<collectingCountry id="F37C50A1FFA8F3446147FAED8385EB46" box="[592,747,1311,1335]" name="Indonesia" pageId="3">INDONESIA</collectingCountry>
, LOMBOK, Senaru, under Sendang Gila waterfall,
<date id="FFD536F1FFA8F3446665FAED8077EB2A" pageId="3" value="2009-09-19" valueMax="2009-09-26" valueMin="2009-09-19">19 26.IX.2009</date>
, coll. N. Kluge &amp; L. Sheyko: 4 L-S-I♂, 10 L-S-I♀, 1 L-S♂, 1 L-S♀,
<specimenCount id="9D6DDBB8FFA8F3446726FAB085F7EB2A" box="[1073,1177,1346,1371]" pageId="3" type="larva">52 larvae</specimenCount>
and larval exuviae: 12 L n♂, 22 L n♀, 1 L n-
<specimenCount id="9D6DDBB8FFA8F3446271FA8780EFEB0F" box="[358,385,1383,1411]" pageId="3" type="generic">1♂</specimenCount>
(at molting to L n), 11 L n-1♀, 1 L n-
<specimenCount id="9D6DDBB8FFA8F34461E2FA87827DEB0F" box="[757,787,1383,1411]" pageId="3" type="generic">2♂</specimenCount>
, 4 L n-2♀, 1 L n-3♀.
</paragraph>
<paragraph id="8BD41031FFA8F34463D0FA7E8067EB9C" blockId="3.[151,1436,1310,1999]" pageId="3">
<emphasis id="B91FCC23FFA8F34463D0FA7E836DEBD4" bold="true" box="[199,515,1420,1445]" pageId="3">Additional characteristics.</emphasis>
Larva, subimago, imago and eggs are described by
<bibRefCitation id="EFFA6DC0FFA8F3446747FA7E8584EBD4" author="Kluge" box="[1104,1258,1420,1445]" pageId="3" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge (2012)</bibRefCitation>
. Apex of penis, hypodermal coloration of abdomen and legs and denticles on posterior margin of larval terga are figured here (Figs 1, 30, 32).
</paragraph>
<paragraph id="8BD41031FFA8F34463D0FA0B8451E94E" blockId="3.[151,1436,1310,1999]" pageId="3">
Structure of maxillary apex in penultimolarva and previous larval instars differs in males and females (Figs 6 13). In pen-pen-penultimolarva (L n-3) ventro-apical flange and dentiseta are present, and tusk is not expressed (Fig. 13); this structure is observed on a single female larva, and probably is equal in both sexes. In male, larva of next two instars (L n-2 and L n-1) gets a short tusk and retains ventro-apical flange and dentiseta (Figs 7, 8) (this was observed on one larva of each instar).When male larva molts from penultimolarva (L n-1) to ultimolarva (L n), its maxillary structure is greatly changed (
<figureCitation id="13500CB4FFA8F344617BF94783A8E8BF" box="[620,710,1717,1742]" captionStart="FIGURES 14 15" captionStartId="3.[151,264,787,810]" captionTargetBox="[151,1434,193,765]" captionTargetId="figure@3.[151,1436,193,766]" captionTargetPageId="3" captionText="FIGURES 14 15. Choroterpes (Dilatognathus) cataractae, maxilla of penultimolarva before molt to ultimolarva (slides in Canadian balsam): 14, male; 15, female. Abbreviations: af, ventro-apical flange; ds, dentiseta." httpUri="https://zenodo.org/record/231538/files/figure.png" pageId="3">Fig. 14</figureCitation>
). In female, such change takes place earlier. In female penpenultimolarva (L n-2) maxilla has a long tusk and diminished ventro-apical flange; dentiseta is present (Fig. 12), but can be vestigial and nearly indistinguishable among setae (Fig. 11). In female penultimolarva (L n-1) maxilla has no ventro-apical flange and no dentiseta (Fig. 10); it is similar to maxilla of ultimolarva, but shorter (
<figureCitation id="13500CB4FFA8F34467C9F8D4845FE94E" box="[1246,1329,1830,1855]" captionStart="FIGURES 14 15" captionStartId="3.[151,264,787,810]" captionTargetBox="[151,1434,193,765]" captionTargetId="figure@3.[151,1436,193,766]" captionTargetPageId="3" captionText="FIGURES 14 15. Choroterpes (Dilatognathus) cataractae, maxilla of penultimolarva before molt to ultimolarva (slides in Canadian balsam): 14, male; 15, female. Abbreviations: af, ventro-apical flange; ds, dentiseta." httpUri="https://zenodo.org/record/231538/files/figure.png" pageId="3">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BD41031FFA8F34463D0F8BE83E0E912" blockId="3.[151,1436,1310,1999]" box="[199,654,1867,1892]" pageId="3">
<emphasis id="B91FCC23FFA8F34463D0F8BE8026E912" box="[199,328,1868,1891]" italics="true" pageId="3">Dimension.</emphasis>
Fore wing length 7.5
<quantity id="4C93BDD4FFA8F344615FF8B983E7E912" box="[584,649,1867,1892]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" unit="mm" value="8.0">8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37143BAFFA8F34463D0F89C8312E9BE" pageId="3" type="distribution">
<paragraph id="8BD41031FFA8F34463D0F89C8377E9F6" blockId="3.[151,1436,1310,1999]" box="[199,537,1902,1927]" pageId="3">
<emphasis id="B91FCC23FFA8F34463D0F89C8037E9F6" bold="true" box="[199,345,1902,1927]" pageId="3">Distribution</emphasis>
. Lombok Island.
</paragraph>
<paragraph id="8BD41031FFA8F34463D0F8618312E9BE" blockId="3.[151,1436,1310,1999]" pageId="3">
<emphasis id="B91FCC23FFA8F34463D0F8618044E9DD" bold="true" box="[199,298,1939,1964]" pageId="3">Habitat.</emphasis>
All larvae were collected in a single place, under Sendang Gila waterfall. We did not find larvae of this species in other parts of the same river.
</paragraph>
</subSubSection>
</treatment>
</document>