656 lines
100 KiB
XML
656 lines
100 KiB
XML
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<mods:title id="BAAD198B64B2D37D4AFA8764082A631E">A review of the late Cenozoic genus Bohra (Diprotodontia: Macropodidae) and the evolution of tree-kangaroos</mods:title>
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<mods:namePart id="DC8B1D8586A9BAD6DFDB19C52EFDCCE7">Prideaux, Gavin J.</mods:namePart>
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<mods:namePart id="7A1A2C4DEB8D10AF013FEE00ECB558DC">Warburton, Natalie M.</mods:namePart>
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<mods:nameIdentifier id="A9CE9D8809BBAF1B0B1ED2D0AF0B4DF4" type="email">gavin.prideaux@flinders.edu.au</mods:nameIdentifier>
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<mods:title id="15D574112F8A51382C6B3B20F0A62053">Zootaxa</mods:title>
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<mods:date id="B12C7237D42F0A908891F58D654583F8">2023</mods:date>
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<treatment id="03C24E22F63C5621FF01C053C025F3FA" ID-DOI="http://doi.org/10.5281/zenodo.8017938" ID-GBIF-Taxon="209675095" ID-Zenodo-Dep="8017938" LSID="urn:lsid:plazi:treatment:03C24E22F63C5621FF01C053C025F3FA" httpUri="http://treatment.plazi.org/id/03C24E22F63C5621FF01C053C025F3FA" lastPageId="41" lastPageNumber="42" pageId="37" pageNumber="38">
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<subSubSection id="C371ACBFF63C562DFF01C053C62AF38D" box="[151,631,895,922]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8BD4FF34F63C562DFF01C053C62AF38D" blockId="37.[151,631,895,922]" box="[151,631,895,922]" pageId="37" pageNumber="38">
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<heading id="D09C4858F63C562DFF01C053C62AF38D" bold="true" box="[151,631,895,922]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
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<emphasis id="B91F2326F63C562DFF01C053C62AF38D" bold="true" box="[151,631,895,922]" pageId="37" pageNumber="38">
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<taxonomicName id="4C6B84B7F63C562DFF01C053C5B4F38D" authority="(Dawson 2004 b)" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[151,489,895,922]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bila" status="comb. nov.">
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<emphasis id="B91F2326F63C562DFF01C053C544F38D" bold="true" box="[151,281,895,922]" italics="true" pageId="37" pageNumber="38">Bohra bila</emphasis>
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(
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<bibRefCitation id="EFFA82C5F63C562DFEBEC0ACC5BFF38D" author="Dawson L." box="[296,482,896,922]" pageId="37" pageNumber="38" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson 2004b</bibRefCitation>
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)
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</taxonomicName>
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||
<taxonomicNameLabel id="A22C9E5DF63C562DFE67C0ACC62AF38D" box="[497,631,896,922]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C371ACBFF63C562DFF01C0E8C7F9F42E" pageId="37" pageNumber="38" type="reference_group">
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<paragraph id="8BD4FF34F63C562DFF01C0E8C554F3EC" blockId="37.[151,1436,964,1081]" pageId="37" pageNumber="38">
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<taxonomicName id="4C6B84B7F63C562DFF01C0E8C580F3CB" authority="(De Vis, 1895)" baseAuthorityName="De Vis" baseAuthorityYear="1895" box="[151,477,964,989]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="indra">
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<emphasis id="B91F2326F63C562DFF01C0E8C56EF3CB" box="[151,307,964,988]" italics="true" pageId="37" pageNumber="38">Wallabia indra</emphasis>
|
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(
|
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<bibRefCitation id="EFFA82C5F63C562DFED5C0E9C588F3CB" author="De Vis, C. W." box="[323,469,964,989]" pageId="37" pageNumber="38" pagination="75 - 133" refId="ref50079" refString="De Vis, C. W. (1895) A review of the fossil jaws of the Macropodidae in the Queensland Museum. Proceedings of the Linnean Society of New South Wales, Second Series, 10, 75 - 133." type="journal article" year="1895">De Vis, 1895</bibRefCitation>
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||
)
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</taxonomicName>
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:
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<bibRefCitation id="EFFA82C5F63C562DFE7CC0E8C699F3CB" author="Bartholomai, A." box="[490,708,964,988]" pageId="37" pageNumber="38" pagination="373 - 377" refId="ref48102" refString="Bartholomai, A. (1976 b) The genus Wallabia Trouessart (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 17 (3), 373 - 377." type="journal article" year="1976">Bartholomai (1976b)</bibRefCitation>
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||
, pp. 375, 378 (partim), plate 53: figures 1–3. Not
|
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<taxonomicName id="4C6B84B7F63C562DFB77C0E9C55BF3EC" authority="(De Vis, 1895)" baseAuthorityName="De Vis" baseAuthorityYear="1895" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="indra">
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<emphasis id="B91F2326F63C562DFB77C0E9C166F3CB" box="[1249,1339,964,988]" italics="true" pageId="37" pageNumber="38">W. indra</emphasis>
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(
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<bibRefCitation id="EFFA82C5F63C562DFADAC0E9C4A0F3EC" author="De Vis, C. W." pageId="37" pageNumber="38" pagination="75 - 133" refId="ref50079" refString="De Vis, C. W. (1895) A review of the fossil jaws of the Macropodidae in the Queensland Museum. Proceedings of the Linnean Society of New South Wales, Second Series, 10, 75 - 133." type="journal article" year="1895">De Vis, 1895</bibRefCitation>
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)
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</taxonomicName>
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||
.
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||
</paragraph>
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<paragraph id="8BD4FF34F63C562DFF01C72EC7F9F42E" blockId="37.[151,1436,964,1081]" pageId="37" pageNumber="38">
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<treatmentCitationGroup id="AB7BD81AF63C562DFF01C72EC7F9F42E" pageId="37" pageNumber="38">
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<taxonomicName id="4C6B84B7F63C562DFF01C72EC590F40D" authority="Dawson, 2004 b" authorityName="Dawson" authorityYear="2004" box="[151,461,1026,1050]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bila">
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||
<emphasis id="B91F2326F63C562DFF01C72EC540F40D" box="[151,285,1026,1050]" italics="true" pageId="37" pageNumber="38">Silvaroo bila</emphasis>
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||
<bibRefCitation id="EFFA82C5F63C562DFEB1C72EC590F40D" author="Dawson L." box="[295,461,1026,1050]" pageId="37" pageNumber="38" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson, 2004b</bibRefCitation>
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</taxonomicName>
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||
:
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||
<treatmentCitation id="0ACAD925F63C562DFE4CC72EC692F40D" author="Dawson, L." box="[474,719,1026,1050]" page="114" pageId="37" pageNumber="38" year="2006">
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||
<bibRefCitation id="EFFA82C5F63C562DFE4CC72EC621F40D" author="Dawson, L." box="[474,636,1026,1050]" pageId="37" pageNumber="38" pagination="89 - 114" refId="ref49852" refString="Dawson, L. (2006) An ecophysiological approach to the extinction of large marsupial herbivores in middle and late Pleistocene Australia. Alcheringa, 30 (Supplement 1), 89 - 114. https: // doi. org / 10.1080 / 03115510609506857" type="journal article" year="2006">Dawson (2006)</bibRefCitation>
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, p. 114
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||
</treatmentCitation>
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||
.
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||
<treatmentCitation id="0ACAD925F63C562DFD4AC72EC01FF40D" author="Piper, K. J. & Herrmann, N." box="[732,1090,1026,1050]" page="130" pageId="37" pageNumber="38" year="2006">
|
||
<bibRefCitation id="EFFA82C5F63C562DFD4AC72EC7B3F40D" author="Piper, K. J. & Herrmann, N." box="[732,1006,1026,1050]" pageId="37" pageNumber="38" pagination="125 - 131" refId="ref54593" refString="Piper, K. J. & Herrmann, N. (2006) Additions to knowledge of the early Pleistocene wallaby, Baringa nelsonensis Flannery and Hann 1984 (Marsupialia: Macropodinae). Proceedings of the Linnean Society of New South Wales, 127, 125 - 131." type="journal article" year="2006">Piper & Herrmann (2006)</bibRefCitation>
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, p. 130
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||
</treatmentCitation>
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||
.
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||
<bibRefCitation id="EFFA82C5F63C562DFBD9C72EC165F40D" author="Louys, J. & Price, G. J." box="[1103,1336,1026,1050]" pageId="37" pageNumber="38" pagination="551 - 572" refId="ref52682" refString="Louys, J. & Price, G. J. (2015) The Chinchilla Local Fauna: an exceptionally rich and well-preserved Pliocene vertebrate assemblage from fluviatile deposits of south-eastern Queensland, Australia. Acta Palaeontologica Polonica, 60 (3), 551 - 572. https: // doi. org / 10.4202 / app. 00042.2013" type="journal article" year="2015">Louys & Price (2015)</bibRefCitation>
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||
, pp. 565, 567–568, figure
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||
<specimenCount id="9D6D34BDF63C562DFEF9C70DC5C4F42E" box="[367,409,1057,1081]" pageId="37" pageNumber="38" type="generic">8M.</specimenCount>
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<treatmentCitation id="0ACAD925F63C562DFE09C70CC633F42E" author="Piper, K. J." box="[415,622,1056,1081]" page="238" pageId="37" pageNumber="38" year="2016">
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||
<bibRefCitation id="EFFA82C5F63C562DFE09C70CC67CF42F" author="Piper, K. J." box="[415,545,1056,1080]" pageId="37" pageNumber="38" pagination="233 - 253" refId="ref54541" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper (2016)</bibRefCitation>
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, p. 238
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||
</treatmentCitation>
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||
.
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||
<treatmentCitation id="0ACAD925F63C562DFDE1C70CC7FDF42E" author="Eldridge, M. D. & Beck, R. M. D. & Croft, D. A. & Travouillon, K. J. & Fox, B. J." box="[631,928,1056,1081]" page="822" pageId="37" pageNumber="38" year="2019">
|
||
<bibRefCitation id="EFFA82C5F63C562DFDE1C70CC70EF42E" author="Eldridge, M. D. & Beck, R. M. D. & Croft, D. A. & Travouillon, K. J. & Fox, B. J." box="[631,851,1056,1081]" pageId="37" pageNumber="38" pagination="802 - 837" refId="ref50366" refString="Eldridge, M. D., Beck, R. M. D., Croft, D. A., Travouillon, K. J. & Fox, B. J. (2019) An emerging consensus in the evolution, phylogeny, and systematics of marsupials and their fossil relatives (Metatheria). Journal of Mammalogy, 100 (3), 802 - 837. https: // doi. org / 10.1093 / jmammal / gyz 018" type="journal article" year="2019">
|
||
Eldridge
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||
<emphasis id="B91F2326F63C562DFD43C70DC755F42F" box="[725,776,1056,1080]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
|
||
(2019)
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||
</bibRefCitation>
|
||
, p. 822
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371ACBFF63C562DFF01C770C6B5F679" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8BD4FF34F63C562DFF01C770C7E2F461" blockId="37.[151,1437,1116,2043]" box="[151,959,1116,1142]" pageId="37" pageNumber="38">
|
||
<materialsCitation id="3B03F569F63C562DFF01C770C7E2F461" ID-GBIF-Occurrence="4127780305" box="[151,959,1116,1142]" collectionCode="QM" pageId="37" pageNumber="38" specimenCode="F43280" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B91F2326F63C562DFF01C770C556F461" bold="true" box="[151,267,1116,1142]" pageId="37" pageNumber="38">
|
||
<typeStatus id="54D04196F63C562DFF01C770C55BF461" box="[151,262,1116,1142]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED7A67F1F63C562DFE84C770C562F461" box="[274,319,1116,1142]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBCD574FF63C562DFED0C770C5C2F461" box="[326,415,1116,1142]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">F43280</specimenCode>
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||
, partial left adult dentary (preserving p3, m1–4).
|
||
</materialsCitation>
|
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</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C7ACC54EF4F5" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
<emphasis id="B91F2326F63C562DFF51C7ACC534F48D" bold="true" box="[199,361,1152,1178]" pageId="37" pageNumber="38">Type locality.</emphasis>
|
||
<materialsCitation id="3B03F569F63C562DFEE2C7ACC553F4F5" ID-GBIF-Occurrence="4127780314" location="western Darling Downs" municipality="Condamine River" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Queensland">
|
||
Chinchilla Rifle Range (site not specified), north bank of the
|
||
<collectingMunicipality id="6BB0654EF63C562DFBD0C7ACC14FF48D" box="[1094,1298,1152,1178]" pageId="37" pageNumber="38">Condamine River</collectingMunicipality>
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||
, Chinchilla,
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||
<location id="8EB4A9EFF63C562DFF01C789C5EFF4A8" LSID="urn:lsid:plazi:treatment:03C24E22F63C5621FF01C053C025F3FA:8EB4A9EFF63C562DFF01C789C5EFF4A8" box="[151,434,1188,1215]" municipality="Condamine River" name="western Darling Downs" pageId="37" pageNumber="38" stateProvince="Queensland">western Darling Downs</location>
|
||
, southeastern
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||
<collectingRegion id="49AF31D6F63C562DFDC8C788C6BBF4A9" box="[606,742,1188,1214]" country="Australia" name="Queensland" pageId="37" pageNumber="38">Queensland</collectingRegion>
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||
. Chinchilla Sand. Pliocene. Collected by Doris and Cecil Wilkinson
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</materialsCitation>
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||
.
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||
</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C7C0C662F558" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
<materialsCitation id="3B03F569F63C562DFF51C7C0C607F53D" ID-GBIF-Occurrence="4127780316" collectionCode="QM" collectorName="Cecil Wilkinson. A" pageId="37" pageNumber="38" specimenCode="F43276, F43292" specimenCount="2" typeStatus="paratype">
|
||
<emphasis id="B91F2326F63C562DFF51C7C0C51BF511" bold="true" box="[199,326,1260,1286]" pageId="37" pageNumber="38">
|
||
<typeStatus id="54D04196F63C562DFF51C7C0C51CF511" box="[199,321,1260,1286]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED7A67F1F63C562DFED9C7C0C521F511" box="[335,380,1260,1286]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBCD574FF63C562DFE10C7C0C582F511" box="[390,479,1260,1286]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">F43276</specimenCode>
|
||
, partial left maxilla (preserving M1–4). Chinchilla Rifle Range (site not specified). Collected by Doris and
|
||
<collectorName id="269E9AE2F63C562DFE08C63CC607F53D" box="[414,602,1296,1322]" pageId="37" pageNumber="38">Cecil Wilkinson.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B03F569F63C562DFDC8C63CC66AF559" ID-GBIF-Occurrence="4127780330" collectionCode="QM" collectorName="Cecil Wilkinson. A" pageId="37" pageNumber="38" specimenCode="F43276, F43292" specimenCount="2" typeStatus="paratype">
|
||
A second
|
||
<typeStatus id="54D04196F63C562DFD5DC63DC776F53C" box="[715,811,1297,1323]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7A67F1F63C562DFCAFC63CC735F53D" box="[825,872,1296,1322]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBCD574FF63C562DFCF8C63CC79AF53D" box="[878,967,1296,1322]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">F43292</specimenCode>
|
||
, left i1) from Chinchilla is deemed here to be referable to a species of
|
||
<taxonomicName id="4C6B84B7F63C562DFE5FC618C66AF559" authorityName="McNamara" authorityYear="1994" box="[457,567,1332,1358]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F63C562DFE5FC618C66AF559" box="[457,567,1332,1358]" italics="true" pageId="37" pageNumber="38">Congruus</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C674C015F581" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
<emphasis id="B91F2326F63C562DFF51C674C5E6F565" bold="true" box="[199,443,1368,1394]" pageId="37" pageNumber="38">Referred specimens.</emphasis>
|
||
Lion Hill, Chinchilla Rifle Range,
|
||
<collectingRegion id="49AF31D6F63C562DFCD8C674C78BF565" box="[846,982,1368,1394]" country="Australia" name="Queensland" pageId="37" pageNumber="38">Queensland</collectingRegion>
|
||
.
|
||
<collectionCode id="ED7A67F1F63C562DFC74C674C052F565" box="[994,1039,1368,1394]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
F58668, left m1;
|
||
<collectionCode id="ED7A67F1F63C562DFB4BC674C157F565" box="[1245,1290,1368,1394]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
F58669, left juvenile dentary fragment (preserving m3). Collected by Doris and Cecil Wilkinson.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C68CC54EF5C9" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
Chinchilla Sand, Chinchilla Rifle Range (site not specified).
|
||
<collectionCode id="ED7A67F1F63C562DFCF1C68CC7C9F5AD" box="[871,916,1440,1466]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
F58667, left M2. Collected by Doris and Cecil Wilkinson.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C6C4C5AFF631" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
Chinchilla (presumably Chinchilla Sand).
|
||
<collectionCode id="ED7A67F1F63C562DFD30C6C4C68EF615" box="[678,723,1512,1538]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
F4740, partial left adult maxilla (preserving M2–4). Collected by Kendall Broadbent in 1887.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C51CC6B5F679" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
Probably Darling Downs,
|
||
<collectingRegion id="49AF31D6F63C562DFE79C51CC62AF65D" box="[495,631,1584,1610]" country="Australia" name="Queensland" pageId="37" pageNumber="38">Queensland</collectingRegion>
|
||
.
|
||
<collectionCode id="ED7A67F1F63C562DFD14C51CC6F2F65D" box="[642,687,1584,1610]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="37" pageNumber="38" type="Museum">QM</collectionCode>
|
||
F4749, partial left juvenile dentary (preserving dp3, m1 trigonid). No locality details are associated with this specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371ACBFF63C562DFF51C554C72EF7ED" pageId="37" pageNumber="38" type="etymology">
|
||
<paragraph id="8BD4FF34F63C562DFF51C554C606F6A1" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
<emphasis id="B91F2326F63C562DFF51C554C513F685" bold="true" box="[199,334,1656,1682]" pageId="37" pageNumber="38">Etymology.</emphasis>
|
||
In the Wiradjuri language of central
|
||
<collectingRegion id="49AF31D6F63C562DFC9FC555C781F685" box="[777,988,1656,1683]" country="Australia" name="New South Wales" pageId="37" pageNumber="38">New South Wales</collectingRegion>
|
||
,
|
||
<taxonomicName id="4C6B84B7F63C562DFC7CC554C04BF685" authorityName="Dawson" authorityYear="2004" box="[1002,1046,1656,1682]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F63C562DFC7CC554C04BF685" box="[1002,1046,1656,1682]" italics="true" pageId="37" pageNumber="38">bila</emphasis>
|
||
</taxonomicName>
|
||
means ‘river’, a reference to the surmised riparian habitat of this species.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F63C562DFF51C5ECC72EF7ED" blockId="37.[151,1437,1116,2043]" pageId="37" pageNumber="38">
|
||
<emphasis id="B91F2326F63C562DFF51C5ECC5FFF6CD" bold="true" box="[199,418,1728,1754]" pageId="37" pageNumber="38">Revised diagnosis.</emphasis>
|
||
<taxonomicName id="4C6B84B7F63C562DFE3CC5ECC679F6CD" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[426,548,1728,1754]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F63C562DFE3CC5ECC679F6CD" box="[426,548,1728,1754]" italics="true" pageId="37" pageNumber="38">Bohra bila</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all other species of
|
||
<taxonomicName id="4C6B84B7F63C562DFB96C5ECC01BF6CD" authorityName="Flannery & Szalay" authorityYear="1982" box="[1024,1094,1728,1754]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F63C562DFB96C5ECC01BF6CD" box="[1024,1094,1728,1754]" italics="true" pageId="37" pageNumber="38">Bohra</emphasis>
|
||
</taxonomicName>
|
||
by having the palatine extend anteriorly to a position mesial to the M2 protoloph. The upper molars are similar in size to those of
|
||
<taxonomicName id="4C6B84B7F63C562DFB73C5C9C133F6E9" box="[1253,1390,1764,1790]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F63C562DFB73C5C9C133F6E9" box="[1253,1390,1764,1790]" italics="true" pageId="37" pageNumber="38">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B84B7F63C562DFF01C425C576F735" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[151,299,1800,1826]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F63C562DFF01C425C576F735" box="[151,299,1800,1826]" italics="true" pageId="37" pageNumber="38">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
, but its M4 is slightly smaller. They are most similar in morphology to those of the slightly smaller
|
||
<taxonomicName id="4C6B84B7F63C562DFF01C401C4A0F751" authorityName="Prideaux & Warburton" authorityYear="2023" box="[151,253,1836,1862]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="planei">
|
||
<emphasis id="B91F2326F63C562DFF01C401C4A0F751" box="[151,253,1836,1862]" italics="true" pageId="37" pageNumber="38">B. planei</emphasis>
|
||
</taxonomicName>
|
||
. They differ by having the cusp C region of the stylar crest diverge from the postparacrista closer to the paracone apex, and by lacking any sign of the stylar crest on M3. The ventral margin of the dentary is very smoothly curved posteriorly in lateral profile, lacking the digastric eminence of
|
||
<taxonomicName id="4C6B84B7F63C562DFC0FC459C077F799" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[921,1066,1908,1934]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F63C562DFC0FC459C077F799" box="[921,1066,1908,1934]" italics="true" pageId="37" pageNumber="38">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B84B7F63C562DFBCBC459C0A9F799" authorityName="Prideaux & Warburton" authorityYear="2009" box="[1117,1268,1908,1934]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="nullarbora">
|
||
<emphasis id="B91F2326F63C562DFBCBC459C0A9F799" box="[1117,1268,1908,1934]" italics="true" pageId="37" pageNumber="38">B. nullarbora</emphasis>
|
||
</taxonomicName>
|
||
. It differs from that of
|
||
<taxonomicName id="4C6B84B7F63C562DFF72C4B5C531F7A5" box="[228,364,1944,1970]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F63C562DFF72C4B5C531F7A5" box="[228,364,1944,1970]" italics="true" pageId="37" pageNumber="38">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
by being deeper and having a shorter postalveolar shelf, which bears a small mesial process. The lower molars are similar in size and morphology to those of
|
||
<taxonomicName id="4C6B84B7F63C562DFCA6C491C79CF7C1" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[816,961,1980,2006]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F63C562DFCA6C491C79CF7C1" box="[816,961,1980,2006]" italics="true" pageId="37" pageNumber="38">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
, but they lack a distinct premetacristid and the lophids are thinner in buccal view. The p3 is large and robust.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371ACBFF63F5621FF51C3BAC5B0F1BA" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="8BD4FF34F63F562EFF51C3BAC766F19E" blockId="38.[151,1437,150,609]" pageId="38" pageNumber="39">
|
||
<emphasis id="B91F2326F63F562EFF51C3BAC671F0A7" bold="true" box="[199,556,150,177]" pageId="38" pageNumber="39">Description and comparisons.</emphasis>
|
||
Maxilla and palatine. QM F4740 is a chunk of maxilla that retains M2–4 and the complete masseteric process, and has attached to it a thin strip of palatine (
|
||
<figureCitation id="1350E3B1F63F562EFB9EC397C0F3F0C2" box="[1032,1198,187,213]" captionStart="FIGURE 21" captionStartId="38.[151,250,1901,1925]" captionTargetBox="[166,1420,669,1873]" captionTargetId="figure-277@38.[164,1423,666,1875]" captionTargetPageId="38" captionText="FIGURE 21. Maxillary and upper dental specimens of Bohra bila. QM F43276, partial left adult maxilla (A–C).A: lateral view. B: mesio-occlusal view. C: stereo occlusal view (cast). QM F4740, partial left adult maxilla (D–E). D: lateral view. E: stereo occlusal view. QM F58667, left M2 (F–J). F: stereo occlusal view. G: lingual view. H: buccal view. I: anterior view. J: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012716" httpUri="https://zenodo.org/record/8012716/files/figure.png" pageId="38" pageNumber="39">Figure 21D–E</figureCitation>
|
||
). QM F43276 retains M1–4, but the end of its masseteric process is abraded off. It also has a thin strip of palatine fused to it, which extends anteriorly to a position mesial to the M2 anterior root (
|
||
<figureCitation id="1350E3B1F63F562EFCCAC22FC7BCF10A" box="[860,993,259,285]" captionStart="FIGURE 21" captionStartId="38.[151,250,1901,1925]" captionTargetBox="[166,1420,669,1873]" captionTargetId="figure-277@38.[164,1423,666,1875]" captionTargetPageId="38" captionText="FIGURE 21. Maxillary and upper dental specimens of Bohra bila. QM F43276, partial left adult maxilla (A–C).A: lateral view. B: mesio-occlusal view. C: stereo occlusal view (cast). QM F4740, partial left adult maxilla (D–E). D: lateral view. E: stereo occlusal view. QM F58667, left M2 (F–J). F: stereo occlusal view. G: lingual view. H: buccal view. I: anterior view. J: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012716" httpUri="https://zenodo.org/record/8012716/files/figure.png" pageId="38" pageNumber="39">Figure 21B</figureCitation>
|
||
). The masseteric process of QM F4740 is triangular in lateral view. Its tip aligns with the M3 interloph valley (
|
||
<figureCitation id="1350E3B1F63F562EFC26C20BC069F156" box="[944,1076,295,321]" captionStart="FIGURE 21" captionStartId="38.[151,250,1901,1925]" captionTargetBox="[166,1420,669,1873]" captionTargetId="figure-277@38.[164,1423,666,1875]" captionTargetPageId="38" captionText="FIGURE 21. Maxillary and upper dental specimens of Bohra bila. QM F43276, partial left adult maxilla (A–C).A: lateral view. B: mesio-occlusal view. C: stereo occlusal view (cast). QM F4740, partial left adult maxilla (D–E). D: lateral view. E: stereo occlusal view. QM F58667, left M2 (F–J). F: stereo occlusal view. G: lingual view. H: buccal view. I: anterior view. J: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012716" httpUri="https://zenodo.org/record/8012716/files/figure.png" pageId="38" pageNumber="39">Figure 21D</figureCitation>
|
||
), and its leading edge is slightly twisted anterolaterally. Although the tip of the masseteric process is missing in QM F43276, we can extrapolate that it would have been positioned adjacent to the M3 protoloph.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F63F562EFF51C2BFC0F1F276" blockId="38.[151,1437,150,609]" pageId="38" pageNumber="39">
|
||
In
|
||
<taxonomicName id="4C6B84B7F63F562EFF73C2BFC572F1BA" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[229,303,403,429]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F63F562EFF73C2BFC572F1BA" box="[229,303,403,429]" italics="true" pageId="38" pageNumber="39">B. bila</emphasis>
|
||
</taxonomicName>
|
||
, the maxilla–palatine suture extends anteriorly to a position adjacent to the M2 anterior root, but in
|
||
<emphasis id="B91F2326F63F562EFA12C2BFC642F1C6" italics="true" pageId="38" pageNumber="39">
|
||
<taxonomicName id="4C6B84B7F63F562EFA12C2BFC54EF1C6" authorityName="Prideaux & Warburton" authorityYear="2009" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nullarbora">B. nullarbora</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B84B7F63F562EFE8BC29BC5ECF1C6" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[285,433,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="illuminata">B. illuminata</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B84B7F63F562EFE2DC29BC642F1C6" authorityName="Prideaux & Warburton" authorityYear="2023" box="[443,543,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="planei">B. planei</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C6B84B7F63F562EFDC5C29BC686F1C6" box="[595,731,439,465]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F63F562EFDC5C29BC686F1C6" box="[595,731,439,465]" italics="true" pageId="38" pageNumber="39">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
it stops and inflects lingually at or posterior to a position mesial to the anterior edge of M3. The position of the masseteric process in
|
||
<taxonomicName id="4C6B84B7F63F562EFC05C2F7C783F1E2" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[915,990,475,501]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F63F562EFC05C2F7C783F1E2" box="[915,990,475,501]" italics="true" pageId="38" pageNumber="39">B. bila</emphasis>
|
||
</taxonomicName>
|
||
is consistent with that observed in adult specimens of
|
||
<emphasis id="B91F2326F63F562EFEA7C2D3C666F20E" box="[305,571,511,537]" italics="true" pageId="38" pageNumber="39">
|
||
<taxonomicName id="4C6B84B7F63F562EFEA7C2D3C596F20E" authorityName="Prideaux & Warburton" authorityYear="2009" box="[305,459,511,537]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nullarbora">B. nullarbora</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B84B7F63F562EFE40C2D3C666F20E" authorityName="Prideaux & Warburton" authorityYear="2023" box="[470,571,511,537]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="planei">B. planei</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C6B84B7F63F562EFDE7C2D3C6A7F20E" box="[625,762,511,537]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F63F562EFDE7C2D3C6A7F20E" box="[625,762,511,537]" italics="true" pageId="38" pageNumber="39">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
. The form of the masseteric process is a good match for that of
|
||
<taxonomicName id="4C6B84B7F63F562EFF20C10FC514F22A" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[182,329,547,573]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F63F562EFF20C10FC514F22A" box="[182,329,547,573]" italics="true" pageId="38" pageNumber="39">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
, but in the
|
||
<typeStatus id="54D04196F63F562EFE5CC10FC671F22A" box="[458,556,547,573]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
of that species it is instead positioned adjacent to the M2 metaloph. This may be an artefact of its younger ontogenetic age (M4 is in the process of rotating into occlusion).
|
||
</paragraph>
|
||
<caption id="DF14AFBCF63F562EFF01C441C493F7E5" ID-DOI="http://doi.org/10.5281/zenodo.8012716" ID-Zenodo-Dep="8012716" httpUri="https://zenodo.org/record/8012716/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1901,1925]" targetBox="[166,1420,669,1873]" targetPageId="38" targetType="figure">
|
||
<paragraph id="8BD4FF34F63F562EFF01C441C493F7E5" blockId="38.[151,1437,1901,2034]" pageId="38" pageNumber="39">
|
||
<emphasis id="B91F2326F63F562EFF01C441C542F791" bold="true" box="[151,287,1901,1926]" pageId="38" pageNumber="39">FIGURE 21.</emphasis>
|
||
Maxillary and upper dental specimens of
|
||
<taxonomicName id="4C6B84B7F63F562EFD55C442C773F791" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[707,814,1902,1926]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F63F562EFD55C442C773F791" box="[707,814,1902,1926]" italics="true" pageId="38" pageNumber="39">Bohra bila</emphasis>
|
||
</taxonomicName>
|
||
. QM F43276, partial left adult maxilla (A–C). A: lateral view. B: mesio-occlusal view. C: stereo occlusal view (cast). QM F4740, partial left adult maxilla (D–E). D: lateral view. E: stereo occlusal view. QM F58667, left M2 (F–J). F: stereo occlusal view. G: lingual view. H: buccal view. I: anterior view. J: posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF14AFBCF63E562FFF01C5DEC6BDF78D" ID-DOI="http://doi.org/10.5281/zenodo.8012720" ID-Zenodo-Dep="8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1778,1802]" targetBox="[153,1434,183,1751]" targetPageId="39" targetType="figure">
|
||
<paragraph id="8BD4FF34F63E562FFF01C5DEC6BDF78D" blockId="39.[151,1437,1778,1946]" pageId="39" pageNumber="40">
|
||
<emphasis id="B91F2326F63E562FFF01C5DEC57CF71D" bold="true" box="[151,289,1778,1802]" pageId="39" pageNumber="40">FIGURE 22.</emphasis>
|
||
Dentary and lower dental specimens of
|
||
<taxonomicName id="4C6B84B7F63E562FFD52C5DEC76FF71D" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[708,818,1778,1802]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F63E562FFD52C5DEC76FF71D" box="[708,818,1778,1802]" italics="true" pageId="39" pageNumber="40">Bohra bila</emphasis>
|
||
</taxonomicName>
|
||
. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD4FF34F6315620FF51C3BBC7DCF0EE" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
Upper dentition. Previously described in detail for QM F43276 (
|
||
<bibRefCitation id="EFFA82C5F6315620FC27C3BBC03FF0A6" author="Dawson L." box="[945,1122,151,177]" pageId="40" pageNumber="41" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson 2004b</bibRefCitation>
|
||
). The referred unworn M2, QM F58667 (
|
||
<figureCitation id="1350E3B1F6315620FEA4C397C590F0C2" box="[306,461,187,213]" captionStart="FIGURE 21" captionStartId="38.[151,250,1901,1925]" captionTargetBox="[166,1420,669,1873]" captionTargetId="figure-277@38.[164,1423,666,1875]" captionTargetPageId="38" captionText="FIGURE 21. Maxillary and upper dental specimens of Bohra bila. QM F43276, partial left adult maxilla (A–C).A: lateral view. B: mesio-occlusal view. C: stereo occlusal view (cast). QM F4740, partial left adult maxilla (D–E). D: lateral view. E: stereo occlusal view. QM F58667, left M2 (F–J). F: stereo occlusal view. G: lingual view. H: buccal view. I: anterior view. J: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012716" httpUri="https://zenodo.org/record/8012716/files/figure.png" pageId="40" pageNumber="41">Figure 21F–J</figureCitation>
|
||
), is indistinguishable from the M2 of QM F
|
||
<quantity id="4C9352D1F6315620FC54C397C07AF0C2" box="[962,1063,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.0992104" pageId="40" pageNumber="41" unit="in" value="43276.0">43276 in</quantity>
|
||
size and morphology, except for possessing a very slightly weaker cusp C region of the stylar crest.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C22FC029F1E2" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
<taxonomicName id="4C6B84B7F6315620FF51C22FC51CF10A" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[199,321,259,285]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6315620FF51C22FC51CF10A" box="[199,321,259,285]" italics="true" pageId="40" pageNumber="41">Bohra bila</emphasis>
|
||
</taxonomicName>
|
||
is a close match for
|
||
<taxonomicName id="4C6B84B7F6315620FDBCC22FC6E9F10A" box="[554,692,259,285]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F6315620FDBCC22FC6E9F10A" box="[554,692,259,285]" italics="true" pageId="40" pageNumber="41">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B84B7F6315620FD7DC22FC720F10A" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[747,893,259,285]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F6315620FD7DC22FC720F10A" box="[747,893,259,285]" italics="true" pageId="40" pageNumber="41">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
in upper molar size, but it has a slightly smaller M4 (
|
||
<tableCitation id="C6E9CA8FF6315620FF5DC20BC574F156" box="[203,297,295,321]" captionStart="TABLE 1" captionStartId="28.[151,239,483,509]" captionTargetId="graphics-52@28.[151,2042,547,912]" captionTargetPageId="28" captionText="TABLE 1. Cheek-tooth dimensions (mm) of Bohra bandharr. *Anterior end missing enamel; length estimated by extrapolating from anterior edge of anterior root and contours of adjacent enamel." httpUri="http://table.plazi.org/id/DF14AFBCF6055614FF01C2CFC535F236" pageId="40" pageNumber="41" tableUuid="DF14AFBCF6055614FF01C2CFC535F236">Tables 1</tableCitation>
|
||
,
|
||
<tableCitation id="C6E9CA8FF6315620FEA0C20AC539F156" box="[310,356,294,321]" captionStart-0="TABLE 4" captionStart-1="TABLE 5" captionStartId-0="35.[151,239,1007,1033]" captionStartId-1="42.[151,239,728,754]" captionTargetId-0="graphics-215@35.[151,2040,1035,1535]" captionTargetId-1="graphics-33@42.[151,2037,756,1483]" captionTargetPageId-0="35" captionTargetPageId-1="42" captionText-0="TABLE 4. Cheek-tooth dimensions (mm) of Bohra bila. Data for QM F4740 is from Bartholomai (1976a)." captionText-1="TABLE 5. Cheek-tooth dimensions (mm) of Bohra illuminata." httpUri-0="http://table.plazi.org/id/DF14AFBCF63A562BFF01C0C3C152F41E" httpUri-1="http://table.plazi.org/id/DF14AFBCF6335622FF01C1F4C765F2E5" pageId="40" pageNumber="41" tableUuid-0="DF14AFBCF63A562BFF01C0C3C152F41E" tableUuid-1="DF14AFBCF6335622FF01C1F4C765F2E5">4–5</tableCitation>
|
||
). In morphology, including general shape of the crown outline,
|
||
<taxonomicName id="4C6B84B7F6315620FB89C20BC034F156" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[1055,1129,295,321]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6315620FB89C20BC034F156" box="[1055,1129,295,321]" italics="true" pageId="40" pageNumber="41">B. bila</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C6B84B7F6315620FAA7C20BC1C8F156" authorityName="Prideaux & Warburton" authorityYear="2023" box="[1329,1429,295,321]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="planei">
|
||
<emphasis id="B91F2326F6315620FAA7C20BC1C8F156" box="[1329,1429,295,321]" italics="true" pageId="40" pageNumber="41">B. planei</emphasis>
|
||
</taxonomicName>
|
||
, and to a slightly lesser degree,
|
||
<taxonomicName id="4C6B84B7F6315620FE78C267C6D8F172" authorityName="Prideaux & Warburton" authorityYear="2009" box="[494,645,331,357]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="nullarbora">
|
||
<emphasis id="B91F2326F6315620FE78C267C6D8F172" box="[494,645,331,357]" italics="true" pageId="40" pageNumber="41">B. nullarbora</emphasis>
|
||
</taxonomicName>
|
||
, both of which have smaller molars. In contrast to that of
|
||
<taxonomicName id="4C6B84B7F6315620FA9EC267C130F172" authorityName="Prideaux & Warburton" authorityYear="2023" box="[1288,1389,331,357]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="planei">
|
||
<emphasis id="B91F2326F6315620FA9EC267C130F172" box="[1288,1389,331,357]" italics="true" pageId="40" pageNumber="41">B. planei</emphasis>
|
||
</taxonomicName>
|
||
, the cusp C region of the stylar crest diverge from the postparacrista closer to the paracone apex in
|
||
<taxonomicName id="4C6B84B7F6315620FB59C243C146F19E" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[1231,1307,367,393]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6315620FB59C243C146F19E" box="[1231,1307,367,393]" italics="true" pageId="40" pageNumber="41">B. bila</emphasis>
|
||
</taxonomicName>
|
||
rather than further up the paracone. In addition, there is no manifestation of this stylar crest region on M
|
||
<quantity id="4C9352D1F6315620FB35C2BFC091F1BA" box="[1187,1228,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="40" pageNumber="41" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="4C6B84B7F6315620FB45C2BFC140F1BA" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[1235,1309,403,429]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6315620FB45C2BFC140F1BA" box="[1235,1309,403,429]" italics="true" pageId="40" pageNumber="41">B. bila</emphasis>
|
||
</taxonomicName>
|
||
. The upper molars of
|
||
<taxonomicName id="4C6B84B7F6315620FE9EC29BC50FF1C6" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[264,338,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6315620FE9EC29BC50FF1C6" box="[264,338,439,465]" italics="true" pageId="40" pageNumber="41">B. bila</emphasis>
|
||
</taxonomicName>
|
||
differ from those of
|
||
<taxonomicName id="4C6B84B7F6315620FDAFC29BC68CF1C6" authorityName="Prideaux & Warburton" authorityYear="2009" box="[569,721,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="nullarbora">
|
||
<emphasis id="B91F2326F6315620FDAFC29BC68CF1C6" box="[569,721,439,465]" italics="true" pageId="40" pageNumber="41">B. nullarbora</emphasis>
|
||
</taxonomicName>
|
||
by having a cusp C region of the stylar crest on M1–2, incurved postpara- and premeta-cristae, and a broader, more open posterior metaloph face on M4.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C2D3C671F276" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
<materialsCitation id="3B03F569F6315620FF51C2D3C675F276" ID-GBIF-Occurrence="4127780328" collectionCode="QM" pageId="40" pageNumber="41" specimenCode="F43280, F58669" specimenCount="1" typeStatus="holotype">
|
||
Dentary. Previously described in detail for the
|
||
<typeStatus id="54D04196F6315620FD52C2D3C774F20E" box="[708,809,511,537]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectionCode id="ED7A67F1F6315620FCA4C2D3C702F20E" box="[818,863,511,537]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBCD574FF6315620FCF3C2D3C7E7F20E" box="[869,954,511,537]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">F43280</specimenCode>
|
||
(
|
||
<bibRefCitation id="EFFA82C5F6315620FC51C2D3C029F20E" author="Dawson L." box="[967,1140,511,537]" pageId="40" pageNumber="41" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson 2004b</bibRefCitation>
|
||
). The two referred juvenile dentary fragments are too incomplete to add to the knowledge of dentary morphology, but
|
||
<collectionCode id="ED7A67F1F6315620FBEAC10FC0F4F22A" box="[1148,1193,547,573]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBCD574FF6315620FB39C10EC158F22A" box="[1199,1285,546,573]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">F58669</specimenCode>
|
||
does confirm the absence of a digastric eminence
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C147C15EF32E" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
The
|
||
<typeStatus id="54D04196F6315620FF6CC147C501F292" box="[250,348,619,645]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
dentary of
|
||
<taxonomicName id="4C6B84B7F6315620FE4AC147C67AF292" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[476,551,619,645]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6315620FE4AC147C67AF292" box="[476,551,619,645]" italics="true" pageId="40" pageNumber="41">B. bila</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1350E3B1F6315620FDA0C147C687F292" box="[566,730,619,645]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22A–C</figureCitation>
|
||
) is most similar in overall form to that of
|
||
<taxonomicName id="4C6B84B7F6315620FB26C147C115F292" authorityName="Prideaux & Warburton" authorityYear="2009" box="[1200,1352,619,645]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="nullarbora">
|
||
<emphasis id="B91F2326F6315620FB26C147C115F292" box="[1200,1352,619,645]" italics="true" pageId="40" pageNumber="41">B. nullarbora</emphasis>
|
||
</taxonomicName>
|
||
, but its ventral margin has a much more evenly curved profile due to the barely discernible digastric sulcus and absence of a digastric eminence (
|
||
<figureCitation id="1350E3B1F6315620FE3CC19FC60CF2DA" box="[426,593,691,717]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22A–B</figureCitation>
|
||
). It also has a larger and more rugose anterior insertion area for the internal superficial masseter muscle, and a more distinct buccinator sulcus (
|
||
<figureCitation id="1350E3B1F6315620FCF8C1FBC7ACF2E6" box="[878,1009,727,753]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22A</figureCitation>
|
||
).The latter feature is also characteristic of the smaller
|
||
<taxonomicName id="4C6B84B7F6315620FEA3C1D7C5E0F302" box="[309,445,763,789]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F6315620FEA3C1D7C5E0F302" box="[309,445,763,789]" italics="true" pageId="40" pageNumber="41">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
, which is distinguished by its longer, rounded postalveolar shelf. A postalveolar shelf that bears a mesial process is shared with
|
||
<taxonomicName id="4C6B84B7F6315620FDACC033C68FF32E" authorityName="Prideaux & Warburton" authorityYear="2009" box="[570,722,799,825]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="nullarbora">
|
||
<emphasis id="B91F2326F6315620FDACC033C68FF32E" box="[570,722,799,825]" italics="true" pageId="40" pageNumber="41">B. nullarbora</emphasis>
|
||
</taxonomicName>
|
||
, but the process on the latter is more pronounced.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C06FC713F3B2" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
<materialsCitation id="3B03F569F6315620FF51C06FC717F3B2" ID-GBIF-Occurrence="4127780317" collectionCode="QM" pageId="40" pageNumber="41" specimenCode="F4749" specimenCount="1" typeStatus="holotype">
|
||
Lower dentition. The lower cheek teeth of the
|
||
<typeStatus id="54D04196F6315620FD41C06FC764F34A" box="[727,825,835,861]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
have been described (
|
||
<bibRefCitation id="EFFA82C5F6315620FBA0C06FC0B8F34A" author="Dawson L." box="[1078,1253,835,861]" pageId="40" pageNumber="41" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson 2004b</bibRefCitation>
|
||
), but the molars are heavily worn. Here we describe the teeth of three referred specimens, which have sustained little to no wear, one of which (
|
||
<collectionCode id="ED7A67F1F6315620FE9FC0A7C565F3B2" box="[265,312,907,933]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBCD574FF6315620FEA8C0A7C5D5F3B2" box="[318,392,907,933]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">F4749</specimenCode>
|
||
) preserves the previously unknown dp3
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C083C77AF542" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
The dp3 is narrow, especially anteriorly due to the compressed form of the trigonid (
|
||
<figureCitation id="1350E3B1F6315620FB5AC083C105F3DE" box="[1228,1368,943,969]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22D</figureCitation>
|
||
). The protolophid is very narrow and inflects at the protoconid into a paracristid that rises to a point anteriorly that matches the protoconid in height. From there a crest descends lingually to the anterolingual corner of the crown. If this is a continuation of the paracristid, then its terminus is likely the position of the paraconid; however, it is also conceivable that the high point on the paracristid marks the paraconid (
|
||
<figureCitation id="1350E3B1F6315620FC31C713C017F44E" box="[935,1098,1087,1113]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22H, L</figureCitation>
|
||
). The buccal face of the crown is sheer; there is no distinct cingulum, although the crown bulges slightly near to the base. There is no parastylar crest. The metaconid is higher than the protoconid and gives rise to short, fine pre- and post-metacristids (
|
||
<figureCitation id="1350E3B1F6315620FAC7C7ABC491F4D2" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22D</figureCitation>
|
||
). Although broader than protolophid, the hypolophid is quite narrow relative to the m1 protolophid, and sits substantially lower than it in buccal view. Like the protolophid, the hypolophid slopes slightly buccally, resulting in the hypoconid being noticeably lower than the metaconid. The cristid obliqua descends into the interlophid valley and smoothly ascends to the protoconid apex, such that there is no distinct postprotocristid (
|
||
<figureCitation id="1350E3B1F6315620FB46C63BC106F526" box="[1232,1371,1303,1329]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22D</figureCitation>
|
||
). The posterior face of the hypolophid is smooth and featureless.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C673C637F6F6" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
QM F58668 is a complete, isolated, unworn m1 crown (
|
||
<figureCitation id="1350E3B1F6315620FCA5C673C06BF56E" box="[819,1078,1375,1401]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22F–G, J, M–N</figureCitation>
|
||
), whereas QM F4749 is missing the m1 talonid. The only slight difference is that the most posterior portion of the paracristid is slightly inflected in QM F58668 and straight in QM F4749. The m1 protolophid is narrower than the hypolophid, and its crest is straight for much of its length, as opposed to posteriorly convex like the hypolophid. The paracristid trends anterolingually from the protoconid apex before turning lingually, forming a distinct anterior shelf with a rounded lingual border (
|
||
<figureCitation id="1350E3B1F6315620FF09C53FC516F63A" box="[159,331,1555,1581]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22F, M</figureCitation>
|
||
). The small precingulid is at a distinctly lower level than the anterior portion of the paracristid (
|
||
<figureCitation id="1350E3B1F6315620FF09C51BC577F646" box="[159,298,1591,1617]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22M</figureCitation>
|
||
). A short, distinct parametacristid is directed anterobuccally from the metaconid apex, but terminates high on the protolophid anterior face. The cristid obliqua terminates low on the protolophid posterior face, buccal to the tooth midline. There is no distinct premetacristid, but the anterior face of the hypolophid is broadly convex on the lingual side. The posterior face of the hypolophid is smooth and featureless, except for a slight bulge low on the crown on the buccal side (
|
||
<figureCitation id="1350E3B1F6315620FE2AC5EBC600F6F6" box="[444,605,1735,1761]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22F, N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315620FF51C5C7C7DCF75A" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
|
||
The m3 is generally similar in form to the much smaller m1, but differs in the following ways: the lophid sides are more convex, the protolophid is not as narrow relative to the hypolophid, the precingulid is broader, and the parametacristid is very low and barely discernible (
|
||
<figureCitation id="1350E3B1F6315620FD42C41FC72BF75A" box="[724,886,1843,1869]" captionStart="FIGURE 22" captionStartId="39.[151,250,1778,1802]" captionTargetBox="[153,1434,183,1751]" captionTargetId="figure-17@39.[151,1436,181,1754]" captionTargetPageId="39" captionText="FIGURE 22. Dentary and lower dental specimens of Bohra bila. QM F43280 (holotype) left adult dentary (A–C). A: lateral view. B: mesial view. C: stereo occlusal view (cast). QM F4749, partial left juvenile dentary (D, H, L). D: stereo occlusal view. H: lateral view. L: mesial view. QM F58669, left juvenile dentary fragment (E, I, K, O–P). E: stereo occlusal view. I: lateral view. K: mesial view. O: anterior view. P: posterior view. QM F58668, left m1 (F–G, J, M–N). F: stereo occlusal view. G: buccal view. J: lingual view. M: anterior view. N: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012720" httpUri="https://zenodo.org/record/8012720/files/figure.png" pageId="40" pageNumber="41">Figure 22E–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6315621FF51C47BC5B0F1BA" blockId="40.[151,1437,151,2013]" lastBlockId="41.[151,1437,151,1005]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
|
||
The dp3 is not known for any other species of
|
||
<taxonomicName id="4C6B84B7F6315620FD49C47BC778F766" authorityName="Flannery & Szalay" authorityYear="1982" box="[735,805,1879,1905]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6315620FD49C47BC778F766" box="[735,805,1879,1905]" italics="true" pageId="40" pageNumber="41">Bohra</emphasis>
|
||
</taxonomicName>
|
||
. It is closest in overall morphology to that of the much smaller
|
||
<taxonomicName id="4C6B84B7F6315620FF78C457C6BFF782" authority="(Quoy & Gaimard, 1830)" baseAuthorityName="Quoy & Gaimard" baseAuthorityYear="1830" box="[238,738,1915,1941]" class="Mammalia" family="Macropodidae" genus="Setonix" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="brachyurus">
|
||
<emphasis id="B91F2326F6315620FF78C457C598F782" box="[238,453,1915,1941]" italics="true" pageId="40" pageNumber="41">Setonix brachyurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFA82C5F6315620FE47C457C687F782" author="Quoy, J. R. C. & Gaimard, P." box="[465,730,1915,1941]" pageId="40" pageNumber="41" refId="ref55669" refString="Quoy, J. R. C. & Gaimard, P. (1830) Voyage de Decouvertes de l'Astrolabe Execute par Ordre du Roi, Pendant les Annees 1826 - 1827 - 1828 - 1829, sous la Commandement de M. J. Dumond d'Urville. Zoologie. Vol. 1. [1 - 4]. J. Tastu, Paris, 268 pp." type="book" year="1830">Quoy & Gaimard, 1830</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, with these species sharing a very similar trigonid topology. The main difference is that
|
||
<taxonomicName id="4C6B84B7F6315620FE01C4B3C66FF7AE" baseAuthorityName="Quoy & Gaimard" baseAuthorityYear="1830" box="[407,562,1951,1977]" class="Mammalia" family="Macropodidae" genus="Setonix" kingdom="Animalia" order="Diprotodontia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="brachyurus">
|
||
<emphasis id="B91F2326F6315620FE01C4B3C66FF7AE" box="[407,562,1951,1977]" italics="true" pageId="40" pageNumber="41">S. brachyurus</emphasis>
|
||
</taxonomicName>
|
||
lacks the crest that descends lingually to the anterolingual corner of the crown from the anterior high point on the paracristid. In addition, the base of the crown is not as narrow anteriorly relative to posteriorly in
|
||
<taxonomicName id="4C6B84B7F6305621FED8C3BBC5B4F0A6" baseAuthorityName="Quoy & Gaimard" baseAuthorityYear="1830" box="[334,489,151,177]" class="Mammalia" family="Macropodidae" genus="Setonix" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyurus">
|
||
<emphasis id="B91F2326F6305621FED8C3BBC5B4F0A6" box="[334,489,151,177]" italics="true" pageId="41" pageNumber="42">S. brachyurus</emphasis>
|
||
</taxonomicName>
|
||
, and the hypolophid curves more smoothly into the cristid obliqua and premetacristid than in
|
||
<taxonomicName id="4C6B84B7F6305621FF65C397C51CF0C2" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[243,321,187,213]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FF65C397C51CF0C2" box="[243,321,187,213]" italics="true" pageId="41" pageNumber="42">B. bila</emphasis>
|
||
</taxonomicName>
|
||
. In species of
|
||
<taxonomicName id="4C6B84B7F6305621FE64C397C6DEF0C2" box="[498,643,187,213]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FE64C397C6DEF0C2" box="[498,643,187,213]" italics="true" pageId="41" pageNumber="42">Dendrolagus</emphasis>
|
||
</taxonomicName>
|
||
and dorcopsins, there is no distinct protolophid and paracristid, but rather a single crescentic crest (paralophid), which is met halfway along its length by the cristid obliqua. Species of
|
||
<taxonomicName id="4C6B84B7F6305621FF01C22FC575F10A" box="[151,296,259,285]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FF01C22FC575F10A" box="[151,296,259,285]" italics="true" pageId="41" pageNumber="42">Dendrolagus</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<taxonomicName id="4C6B84B7F6305621FEF8C22FC642F10A" authorityName="DeVis" authorityYear="1887" box="[366,543,259,285]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bennettianus">
|
||
<emphasis id="B91F2326F6305621FEF8C22FC642F10A" box="[366,543,259,285]" italics="true" pageId="41" pageNumber="42">D. bennettianus</emphasis>
|
||
</taxonomicName>
|
||
) may also have a distinct parastylar crest, but this is absent from
|
||
<taxonomicName id="4C6B84B7F6305621FB61C22FC11CF10A" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[1271,1345,259,285]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FB61C22FC11CF10A" box="[1271,1345,259,285]" italics="true" pageId="41" pageNumber="42">B. bila</emphasis>
|
||
</taxonomicName>
|
||
. The p3 of
|
||
<taxonomicName id="4C6B84B7F6305621FF20C20BC55FF156" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[182,258,295,321]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FF20C20BC55FF156" box="[182,258,295,321]" italics="true" pageId="41" pageNumber="42">B. bila</emphasis>
|
||
</taxonomicName>
|
||
is large and robust relative to the molars compared with its proportions in
|
||
<taxonomicName id="4C6B84B7F6305621FBD8C20BC0BCF156" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[1102,1249,295,321]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F6305621FBD8C20BC0BCF156" box="[1102,1249,295,321]" italics="true" pageId="41" pageNumber="42">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91F2326F6305621FA8FC20BC16CF157" box="[1305,1329,295,320]" italics="true" pageId="41" pageNumber="42">B.</emphasis>
|
||
sp. cf.
|
||
<taxonomicName id="4C6B84B7F6305621FA12C20BC55CF172" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F6305621FA12C20BC55CF172" italics="true" pageId="41" pageNumber="42">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
, which are the only other species of
|
||
<taxonomicName id="4C6B84B7F6305621FD0EC267C683F172" authorityName="Flannery & Szalay" authorityYear="1982" box="[664,734,331,357]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FD0EC267C683F172" box="[664,734,331,357]" italics="true" pageId="41" pageNumber="42">Bohra</emphasis>
|
||
</taxonomicName>
|
||
for which p3 is known. The lower molars of
|
||
<taxonomicName id="4C6B84B7F6305621FB43C267C17DF172" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[1237,1312,331,357]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FB43C267C17DF172" box="[1237,1312,331,357]" italics="true" pageId="41" pageNumber="42">B. bila</emphasis>
|
||
</taxonomicName>
|
||
are similar in size and morphology to those of
|
||
<taxonomicName id="4C6B84B7F6305621FD88C243C6EDF19E" authorityName=", Prideaux & Warburton" authorityYear="2008" box="[542,688,367,393]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="illuminata">
|
||
<emphasis id="B91F2326F6305621FD88C243C6EDF19E" box="[542,688,367,393]" italics="true" pageId="41" pageNumber="42">B. illuminata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6E9CA8FF6305621FD28C243C766F19E" box="[702,827,366,393]" captionStart-0="TABLE 4" captionStart-1="TABLE 5" captionStartId-0="35.[151,239,1007,1033]" captionStartId-1="42.[151,239,728,754]" captionTargetId-0="graphics-215@35.[151,2040,1035,1535]" captionTargetId-1="graphics-33@42.[151,2037,756,1483]" captionTargetPageId-0="35" captionTargetPageId-1="42" captionText-0="TABLE 4. Cheek-tooth dimensions (mm) of Bohra bila. Data for QM F4740 is from Bartholomai (1976a)." captionText-1="TABLE 5. Cheek-tooth dimensions (mm) of Bohra illuminata." httpUri-0="http://table.plazi.org/id/DF14AFBCF63A562BFF01C0C3C152F41E" httpUri-1="http://table.plazi.org/id/DF14AFBCF6335622FF01C1F4C765F2E5" pageId="41" pageNumber="42" tableUuid-0="DF14AFBCF63A562BFF01C0C3C152F41E" tableUuid-1="DF14AFBCF6335622FF01C1F4C765F2E5">Tables 4–5</tableCitation>
|
||
), but the lophids are thinner in buccal view and the m3 lacks a distinct premetacristid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371ACBFF6305621FF51C29BC025F3FA" pageId="41" pageNumber="42" type="discussion">
|
||
<paragraph id="8BD4FF34F6305621FF51C29BC6D0F22A" blockId="41.[151,1437,151,1005]" pageId="41" pageNumber="42">
|
||
<emphasis id="B91F2326F6305621FF51C29BC566F1C6" bold="true" box="[199,315,439,465]" pageId="41" pageNumber="42">Remarks.</emphasis>
|
||
<taxonomicName id="4C6B84B7F6305621FED4C29BC5E1F1C6" authorityName="Prideaux & Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[322,444,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FED4C29BC5E1F1C6" box="[322,444,439,465]" italics="true" pageId="41" pageNumber="42">Bohra bila</emphasis>
|
||
</taxonomicName>
|
||
is known definitively only from the Pliocene Chinchilla LF of southeastern
|
||
<collectingRegion id="49AF31D6F6305621FA8EC29BC1C1F1C6" box="[1304,1436,439,465]" country="Australia" name="Queensland" pageId="41" pageNumber="42">Queensland</collectingRegion>
|
||
(
|
||
<figureCitation id="1350E3B1F6305621FF09C2F7C4A0F1E2" box="[159,253,475,501]" captionStart="FIGURE 1" captionStartId="5.[151,250,1691,1715]" captionTargetBox="[207,1381,795,1665]" captionTargetId="figure-278@5.[207,1381,795,1665]" captionTargetPageId="5" captionText="FIGURE 1. Map of Australia and New Guinea showing localities that have produced fossils attributable, or potentially attributable, to species of Bohra." figureDoi="http://doi.org/10.5281/zenodo.8012668" httpUri="https://zenodo.org/record/8012668/files/figure.png" pageId="41" pageNumber="42">Figure 1</figureCitation>
|
||
). The referred specimen QM F4749 is from the Darling Downs, and is probably from Chinchilla, judging from the preservation of the bone, teeth and adhering sediment. This species is one of three in the Chinchilla LF, which is the most of any known assemblage.
|
||
</paragraph>
|
||
<paragraph id="8BD4FF34F6305621FF51C16BC025F3FA" blockId="41.[151,1437,151,1005]" pageId="41" pageNumber="42">
|
||
By recognising that
|
||
<taxonomicName id="4C6B84B7F6305621FE32C16BC6BAF276" authority="Dawson, 2004 b" authorityName="Dawson" authorityYear="2004" box="[420,743,583,609]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FE32C16BC66EF276" box="[420,563,583,609]" italics="true" pageId="41" pageNumber="42">Silvaroo bila</emphasis>
|
||
<bibRefCitation id="EFFA82C5F6305621FDAEC16BC6BAF276" author="Dawson L." box="[568,743,583,609]" pageId="41" pageNumber="42" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson, 2004b</bibRefCitation>
|
||
</taxonomicName>
|
||
belongs in
|
||
<taxonomicName id="4C6B84B7F6305621FCF5C16BC09FF276" authority="Flannery & Szalay, 1982" authorityName="Flannery & Szalay" authorityYear="1982" box="[867,1218,583,609]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FCF5C16BC7F4F276" box="[867,937,583,609]" italics="true" pageId="41" pageNumber="42">Bohra</emphasis>
|
||
<bibRefCitation id="EFFA82C5F6305621FC38C16BC09FF276" author="Flannery, T. F. & Szalay, F. S." box="[942,1218,583,609]" pageId="41" pageNumber="42" pagination="83 - 94" refId="ref50999" refString="Flannery, T. F. & Szalay, F. S. (1982) Bohra paulae: a new giant fossil tree kangaroo (Marsupialia: Macropodidae) from New South Wales, Australia. Australian Mammalogy, 5 (2), 83 - 94." type="journal article" year="1982">Flannery & Szalay, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, we render
|
||
<taxonomicName id="4C6B84B7F6305621FAA8C16BC501F292" authority="Dawson, 2004 b a" authorityName="Dawson" authorityYear="2004" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FAA8C16BC1C1F276" box="[1342,1436,583,609]" italics="true" pageId="41" pageNumber="42">Silvaroo</emphasis>
|
||
<bibRefCitation id="EFFA82C5F6305621FF01C147C501F292" author="Dawson L." box="[151,348,619,645]" pageId="41" pageNumber="42" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson, 2004b a</bibRefCitation>
|
||
</taxonomicName>
|
||
junior synonym of
|
||
<taxonomicName id="4C6B84B7F6305621FDA9C147C6D8F292" authorityName="Flannery & Szalay" authorityYear="1982" box="[575,645,619,645]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FDA9C147C6D8F292" box="[575,645,619,645]" italics="true" pageId="41" pageNumber="42">Bohra</emphasis>
|
||
</taxonomicName>
|
||
, because
|
||
<taxonomicName id="4C6B84B7F6305621FD63C147C763F292" authorityName="Dawson" authorityYear="2004" box="[757,830,619,645]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bila">
|
||
<emphasis id="B91F2326F6305621FD63C147C763F292" box="[757,830,619,645]" italics="true" pageId="41" pageNumber="42">S. bila</emphasis>
|
||
</taxonomicName>
|
||
is its
|
||
<typeStatus id="54D04196F6305621FC15C147C7EEF292" box="[899,947,619,645]" pageId="41" pageNumber="42">type</typeStatus>
|
||
species (
|
||
<bibRefCitation id="EFFA82C5F6305621FB8AC147C091F292" author="Dawson L." box="[1052,1228,619,645]" pageId="41" pageNumber="42" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson 2004b</bibRefCitation>
|
||
). Furthermore, we recognise that the species originally named
|
||
<taxonomicName id="4C6B84B7F6305621FD06C1A3C173F2BE" authority="Dawson, Muirhead & Wroe, 1999" authorityName="Dawson, Muirhead & Wroe" authorityYear="1999" box="[656,1326,655,681]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F6305621FD06C1A3C7C6F2BE" box="[656,923,655,681]" italics="true" pageId="41" pageNumber="42">Protemnodon bandharr</emphasis>
|
||
<bibRefCitation id="EFFA82C5F6305621FC32C1A3C173F2BE" author="Dawson, L. & Muirhead, J. & Wroe, S." box="[932,1326,655,681]" pageId="41" pageNumber="42" pagination="265 - 290" refId="ref49949" refString="Dawson, L., Muirhead, J. & Wroe, S. (1999) The Big Sink Local Fauna: a lower Pliocene mammalian fauna from the Wellington Caves complex, Wellington, New South Wales. Records of the Western Australian Museum, Supplement No. 57, 265 - 290." type="book chapter" year="1999">Dawson, Muirhead & Wroe, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, before it was shifted to
|
||
<taxonomicName id="4C6B84B7F6305621FED7C19FC5C2F2DA" authorityName="Dawson" authorityYear="2004" box="[321,415,691,717]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FED7C19FC5C2F2DA" box="[321,415,691,717]" italics="true" pageId="41" pageNumber="42">Silvaroo</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFFA82C5F6305621FE58C19FC6D2F2DA" author="Dawson L." box="[462,655,691,717]" pageId="41" pageNumber="42" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson (2004b)</bibRefCitation>
|
||
, also belongs in
|
||
<taxonomicName id="4C6B84B7F6305621FCC5C19FC7C4F2DA" authorityName="Flannery & Szalay" authorityYear="1982" box="[851,921,691,717]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FCC5C19FC7C4F2DA" box="[851,921,691,717]" italics="true" pageId="41" pageNumber="42">Bohra</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C6B84B7F6305621FC3CC19FC06CF2DA" box="[938,1073,691,717]" class="Mammalia" family="Macropodidae" genus="Silvaroo" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B91F2326F6305621FC3CC19FC055F2DA" box="[938,1032,691,717]" italics="true" pageId="41" pageNumber="42">Silvaroo</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
indet. 1 and
|
||
<taxonomicName id="4C6B84B7F6305621FB5AC19FC109F2DA" box="[1228,1364,691,717]" class="Mammalia" family="Macropodidae" genus="Silvaroo" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B91F2326F6305621FB5AC19FC177F2DA" box="[1228,1322,691,717]" italics="true" pageId="41" pageNumber="42">Silvaroo</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
indet. 2 of
|
||
<bibRefCitation id="EFFA82C5F6305621FF58C1FBC5D6F2E6" author="Dawson L." box="[206,395,727,753]" pageId="41" pageNumber="42" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson (2004b)</bibRefCitation>
|
||
include specimens referable to
|
||
<taxonomicName id="4C6B84B7F6305621FD60C1FBC7DCF2E6" authorityName="Dawson, Muirhead & Wroe" authorityYear="1999" box="[758,897,727,753]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="bandharr">
|
||
<emphasis id="B91F2326F6305621FD60C1FBC7DCF2E6" box="[758,897,727,753]" italics="true" pageId="41" pageNumber="42">B. bandharr</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B84B7F6305621FC2DC1FBC07CF2E6" authorityName="Prideaux & Warburton" authorityYear="2023" box="[955,1057,727,753]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="planei">
|
||
<emphasis id="B91F2326F6305621FC2DC1FBC07CF2E6" box="[955,1057,727,753]" italics="true" pageId="41" pageNumber="42">B. planei</emphasis>
|
||
</taxonomicName>
|
||
, respectively. The one remaining species that was allocated to
|
||
<taxonomicName id="4C6B84B7F6305621FE4AC1D7C667F302" authorityName="Dawson" authorityYear="2004" box="[476,570,763,789]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FE4AC1D7C667F302" box="[476,570,763,789]" italics="true" pageId="41" pageNumber="42">Silvaroo</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="4C6B84B7F6305621FDCDC1D7C059F302" authority="Plane, 1967" authorityName="Plane" authorityYear="1967" box="[603,1028,763,789]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="buloloensis">
|
||
<emphasis id="B91F2326F6305621FDCDC1D7C724F302" box="[603,889,763,789]" italics="true" pageId="41" pageNumber="42">Protemnodon buloloensis</emphasis>
|
||
<bibRefCitation id="EFFA82C5F6305621FC16C1D7C059F302" author="Plane, M. D." box="[896,1028,763,789]" pageId="41" pageNumber="42" pagination="1 - 64" refId="ref54644" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin. Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Pliocene Awe LF of
|
||
<collectingCountry id="F37CBFA4F6305621FACEC1D7C57AF32E" name="Papua New Guinea" pageId="41" pageNumber="42">Papua New Guinea</collectingCountry>
|
||
. Because it lacks the generic attributes of
|
||
<taxonomicName id="4C6B84B7F6305621FD61C033C040F32E" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[759,1053,799,825]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FD61C033C7D3F32E" box="[759,910,799,825]" italics="true" pageId="41" pageNumber="42">Protemnodon</emphasis>
|
||
<bibRefCitation id="EFFA82C5F6305621FC03C033C040F32E" author="Owen, R." box="[917,1053,799,825]" pageId="41" pageNumber="42" pagination="245 - 288" refId="ref54412" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFA82C5F6305621FBBDC033C085F32E" author="Dawson L." box="[1067,1240,799,825]" pageId="41" pageNumber="42" pagination="275 - 290" refId="ref49800" refString="Dawson L. (2004 b) A new fossil genus of forest wallaby (Marsupialia, Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea. Alcheringa, 28 (1), 275 - 290. https: // doi. org / 10.1080 / 03115510408619285" type="journal article" year="2004">Dawson 2004b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFA82C5F6305621FB75C033C48EF34A" author="Prideaux, G. J. & Kerr, I. A. R. & van Zoelen, J. D. & Grun, R. & van der Kaars, S. & Oertle, A. & Douka, K. & Grono, E. & Barron, A. & Mountain, M. J. & Westaway, M. C." pageId="41" pageNumber="42" pagination="223 - 248" refId="ref55040" refString="Prideaux, G. J., Kerr, I. A. R., van Zoelen, J. D., Grun, R., van der Kaars, S., Oertle, A., Douka, K., Grono, E., Barron, A., Mountain, M. J. & Westaway, M. C. (2022) Re-evaluating the evidence for late-surviving megafauna at Nombe rockshelter in the New Guinea highlands. Archaeology in Oceania, 57 (3), 223 - 248. https: // doi. org / 10.1002 / arco. 5274" type="journal article" year="2022">Kerr & Prideaux 2022</bibRefCitation>
|
||
), and because
|
||
<taxonomicName id="4C6B84B7F6305621FE14C06FC5BDF34A" authorityName="Dawson" authorityYear="2004" box="[386,480,835,861]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FE14C06FC5BDF34A" box="[386,480,835,861]" italics="true" pageId="41" pageNumber="42">Silvaroo</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="4C6B84B7F6305621FC94C06FC715F34A" authorityName="Flannery & Szalay" authorityYear="1982" box="[770,840,835,861]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91F2326F6305621FC94C06FC715F34A" box="[770,840,835,861]" italics="true" pageId="41" pageNumber="42">Bohra</emphasis>
|
||
</taxonomicName>
|
||
, the generic allocation of this species is uncertain. Although resolving this problem is outside of the scope of this work, it is worth noting that the form of the p3 is dorcopsin-like: it is markedly elongate relative to the molars and broader anteriorly before constricting just forward of the posterior end of the tooth. It is within the size range of
|
||
<taxonomicName id="4C6B84B7F6305621FCADC083C06FF3DF" box="[827,1074,943,968]" class="Mammalia" family="Macropodidae" genus="Watutia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B91F2326F6305621FCADC083C06FF3DF" box="[827,1074,943,968]" italics="true" pageId="41" pageNumber="42">Watutia novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
, which is likewise from the Awe LF (
|
||
<bibRefCitation id="EFFA82C5F6305621FF50C0FFC5F3F3FA" author="Flannery, T. F. & Hoch, E. & Aplin, K. P." box="[198,430,979,1005]" pageId="41" pageNumber="42" pagination="145 - 152" refId="ref50770" refString="Flannery, T. F., Hoch, E. & Aplin, K. P. (1989) Macropodines from the Pliocene Otibanda Formation, Papua New Guinea. Alcheringa, 13 (2), 145 - 152. https: // doi. org / 10.1080 / 03115518908619048" type="journal article" year="1989">
|
||
Flannery
|
||
<emphasis id="B91F2326F6305621FEA5C0FFC536F3FA" box="[307,363,979,1005]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
|
||
1989
|
||
</bibRefCitation>
|
||
), but it is clearly distinct from it, minimally at the species level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |