treatments-xml/data/03/C0/D4/03C0D450D16B953941ADFF011FB4D117.xml

<document id="6E2430C3362B0D02C5D46BC10C7409A9" ID-CLB-Dataset="290199" ID-DOI="10.11646/zootaxa.5418.5.6" ID-GBIF-Dataset="40fd70f2-37f9-42be-8ae8-97be12bf3b58" ID-ISSN="1175-5326" ID-Zenodo-Dep="10780063" ID-ZooBank="8681F446-C4A0-466F-A763-57F426B14523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1709588019354" checkinUser="plazi" docAuthor="Dhiya’Ulhaq, Naufal Urfi, Dupérré, Nadine, Buchori, Damayanti, Scheu, Stefan &amp; Drescher, Jochen" docDate="2024" docId="03C0D450D16B953941ADFF011FB4D117" docLanguage="en" docName="zootaxa.5418.5.6.pdf" docOrigin="Zootaxa 5418 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.5.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna trisula Dhiya’ulhaq &amp; Dupérré 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="573" masterDocId="FFF9AC28D178952F413AFF961C7CD506" masterDocTitle="Four new species of Utivarachna Kishida, 1940 (Araneae: Trachelidae) from Sumatra" masterLastPageNumber="575" masterPageNumber="551" pageNumber="570" updateTime="1713441516709" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="16D659AC3457C8667427C2DDB57CCD33">Four new species of Utivarachna Kishida, 1940 (Araneae: Trachelidae) from Sumatra</mods:title>
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<mods:namePart id="42D417C90F9DB28761A6C544C36CF31F">Dhiya’Ulhaq, Naufal Urfi</mods:namePart>
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<mods:affiliation id="2727C9A64BF3AE548EA1B9EF89072F14">Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany</mods:affiliation>
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<mods:namePart id="FAAB4BA04E1F913433F903AE726E3A74">Dupérré, Nadine</mods:namePart>
<mods:affiliation id="B5CE0C63F453E4C8ABBCCEFFE617C717">Centre for Taxonomy and Morphology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Museum of Nature Hamburg, Hamburg, Germany &amp; American Museum of Natural History, New York, NY, United States of America</mods:affiliation>
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<mods:namePart id="6EC142A8AAFFB489303D4BDB20E9A9B3">Buchori, Damayanti</mods:namePart>
<mods:affiliation id="5573CD8EA9B648747EF68F835F15867B">Department of Plant Protection, Faculty of Agriculture, IPB University, Bogor, Indonesia &amp; Centre for Transdisciplinary and Sustainability Sciences, IPB University, Bogor, Indonesia</mods:affiliation>
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<mods:namePart id="1BC5E75FEA84708EADB0F5CFB530DC54">Scheu, Stefan</mods:namePart>
<mods:affiliation id="21EABCDEA1D5F36C0F1E572CDA81E686">Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany &amp; Centre of Biodiversity and Sustainable Land Use, Göttingen, Germany</mods:affiliation>
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<mods:affiliation id="36463D244916EF17E56D76A9B6E15D35">Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany</mods:affiliation>
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<emphasis id="B91DB954D16B953C41ADFF011DF8D5B4" bold="true" box="[151,388,151,178]" italics="true" pageId="19" pageNumber="570">Utivarachna trisula</emphasis>
Dhiya’ulhaq &amp; Dupérré
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, 
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<figureCitation id="135279C3D16B953C41ADFF2D1D45D5D3" box="[151,313,187,213]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStart-3="FIGURE 17" captionStartId-0="17.[151,250,1933,1957]" captionStartId-1="18.[151,250,1923,1947]" captionStartId-2="19.[151,250,1912,1936]" captionStartId-3="20.[151,250,1595,1619]" captionTargetBox-0="[151,1436,273,1910]" captionTargetBox-1="[164,1423,453,1893]" captionTargetBox-2="[151,1436,381,1888]" captionTargetBox-3="[151,1436,344,1566]" captionTargetId-0="figure-65@17.[151,1436,273,1910]" captionTargetId-1="figure-110@18.[164,1423,453,1900]" captionTargetId-2="figure-54@19.[151,1436,381,1888]" captionTargetId-3="figure-82@20.[151,1436,344,1572]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURE 14. Utivarachna trisula sp. nov., male holotype. A, Habitus, dorsal view; B, Sternum, ventral view; C, Left leg I, prolateral view. Scale bars A = 1 mm; B, C = 0.5 mm." captionText-1="FIGURE 15. Utivarachna trisula sp. nov., male holotype. A–E Left palp: A, prolateral view; B, D, ventral view; C, E retrolateral view. Scale bars 0.2 mm. Abbreviations: ac, apical claw of RTA; dl, dorsal lobe of RTA; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." captionText-2="FIGURE 16. Utivarachna trisula sp. nov., female paratype. A, B Habitus, dorsal view (A), ventral view (B); C, Left leg I, prolateral view. Scale bars A, B = 1 mm; C = 0.5 mm." captionText-3="FIGURE 17. Utivarachna trisula sp. nov., female paratype. A Epigynum, ventral view; B, C Internal genitalia, cleared, dorsal view. Scale bars 0.2 mm.Abbreviations: at, atrium; b, bursa; cnd, connecting duct; co, copulatory opening; cod, copulatory duct; fd, fertilization duct; s, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.10780106" figureDoi-1="http://doi.org/10.5281/zenodo.10780108" figureDoi-2="http://doi.org/10.5281/zenodo.10780112" figureDoi-3="http://doi.org/10.5281/zenodo.10780116" httpUri-0="https://zenodo.org/record/10780106/files/figure.png" httpUri-1="https://zenodo.org/record/10780108/files/figure.png" httpUri-2="https://zenodo.org/record/10780112/files/figure.png" httpUri-3="https://zenodo.org/record/10780116/files/figure.png" pageId="19" pageNumber="570">Figures 14–17</figureCitation>
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<emphasis id="B91DB954D16B953C41ADFE951D33D41B" bold="true" box="[151,335,259,285]" pageId="19" pageNumber="570">Type material.</emphasis>
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:
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, 
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(2013_HJ1.1_ AraTrac003N_001), canopy fogging in jungle rubber plantation, 01⁰55’41.6”S, 103⁰15’34.2”E, altitude 
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, leg. 
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(
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<emphasis id="B91DB954D16B953C41ADF8EE1D5ED296" bold="true" box="[151,290,1912,1936]" pageId="19" pageNumber="570">FIGURE 16.</emphasis>
<taxonomicName id="4C691EC5D16B953C4010F8EE1D88D296" authority="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher, 2024" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[298,500,1912,1936]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="570" phylum="Arthropoda" rank="species" species="trisula" status="sp. nov.">
<emphasis id="B91DB954D16B953C4010F8EE1D88D296" box="[298,500,1912,1936]" italics="true" pageId="19" pageNumber="570">Utivarachna trisula</emphasis>
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, female paratype. 
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Habitus, dorsal view (
<emphasis id="B91DB954D16B953C450AF8EE183ED296" bold="true" box="[1072,1090,1912,1936]" pageId="19" pageNumber="570">A</emphasis>
), ventral view (
<emphasis id="B91DB954D16B953C45D0F8EE1887D296" bold="true" box="[1258,1275,1912,1936]" pageId="19" pageNumber="570">B</emphasis>
); 
<emphasis id="B91DB954D16B953C4429F8EE1959D296" bold="true" box="[1299,1317,1912,1936]" pageId="19" pageNumber="570">C</emphasis>
, Left leg I, prolateral view. Scale bars A, B = 1 mm; C = 0.5 mm.
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<typeStatus id="54D2DBE4D16C953B41FDFF001D3FD5B6" box="[199,323,150,176]" pageId="20" pageNumber="571" type="paratype">Paratypes</typeStatus>
: 
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: 
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:
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(2013_HJ1.1_AraTrac003N_002), with the same data as holotype (
<collectionCode id="ED78FD83D16C953B41A5FF2C1C9DD5D2" box="[159,225,186,212]" collectionName="MZB" pageId="20" pageNumber="571">MZB</collectionCode>
)
</materialsCitation>
; 
<specimenCount id="9D6FAECFD16C953B41C0FF2C1D61D5D3" box="[250,285,186,213]" count="1" pageId="20" pageNumber="571" type="female">1♀</specimenCount>
, 
<taxonomicName id="4C691EC5D16C953B400AFF2D1DC2D5D3" authority="Baru" authorityName="Baru" box="[304,446,187,213]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="dusun">Dusun Baru</taxonomicName>
, Air Hitam, Sarolangun, (2013_BJ4.1_AraTrac003N_001), canopy fogging in jungle rubber plantation, 02 
<geoCoordinate id="EE5D0381D16C953B40BEFF761D82D5FD" box="[388,510,224,251]" degrees="0" direction="south" minutes="00" orientation="latitude" pageId="20" pageNumber="571" precision="1" seconds="56.8" value="-0.015777778">⁰00’56.8”S</geoCoordinate>
, 102 
<geoCoordinate id="EE5D0381D16C953B430CFF761ECFD5FD" box="[566,691,224,251]" degrees="0" direction="east" minutes="45" orientation="longitude" pageId="20" pageNumber="571" precision="1" seconds="12.6" value="0.7535">⁰45’12.6”E</geoCoordinate>
, altitude 
<quantity id="4C91C8A3D16C953B4223FF481F28D5FF" box="[793,852,222,249]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="20" pageNumber="571" unit="m" value="64.0">64 m</quantity>
, 
<date id="FFD74386D16C953B425AFF481F97D5FE" box="[864,1003,222,248]" pageId="20" pageNumber="571" value="2013-07-14">14.VII.2013</date>
, leg. J. Drescher (ZMH-A0023868).
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD16C953B41FDFE941D0AD446" pageId="20" pageNumber="571" type="etymology">
<paragraph id="8BD66546D16C953B41FDFE941D0AD446" blockId="20.[151,1437,150,321]" pageId="20" pageNumber="571">
<emphasis id="B91DB954D16C953B41FDFE941D32D41A" bold="true" box="[199,334,258,284]" pageId="20" pageNumber="571">Etymology.</emphasis>
The specific name is taken from Indonesian 
<taxonomicName id="4C691EC5D16C953B427FFE941FF2D41A" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[837,910,258,284]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="trisula">
<emphasis id="B91DB954D16C953B427FFE941FF2D41A" box="[837,910,258,284]" italics="true" pageId="20" pageNumber="571">trisula</emphasis>
</taxonomicName>
, meaning “trident”, referring to the trifid RTA. Noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF1635CED16C953B41ADF9AD1E77D39D" ID-DOI="http://doi.org/10.5281/zenodo.10780116" ID-Zenodo-Dep="10780116" httpUri="https://zenodo.org/record/10780116/files/figure.png" pageId="20" pageNumber="571" startId="20.[151,250,1595,1619]" targetBox="[151,1436,344,1566]" targetPageId="20" targetType="figure">
<paragraph id="8BD66546D16C953B41ADF9AD1E77D39D" blockId="20.[151,1436,1595,1691]" pageId="20" pageNumber="571">
<emphasis id="B91DB954D16C953B41ADF9AD1D5CD355" bold="true" box="[151,288,1595,1619]" pageId="20" pageNumber="571">FIGURE 17.</emphasis>
<taxonomicName id="4C691EC5D16C953B401CF9AD1D92D355" authority="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher, 2024" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[294,494,1595,1619]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="trisula" status="sp. nov.">
<emphasis id="B91DB954D16C953B401CF9AD1D92D355" box="[294,494,1595,1619]" italics="true" pageId="20" pageNumber="571">Utivarachna trisula</emphasis>
</taxonomicName>
<emphasis id="B91DB954D16C953B40CEF9AA1E39D352" bold="true" box="[500,581,1596,1620]" pageId="20" pageNumber="571">
<taxonomicNameLabel id="A22E042FD16C953B40CEF9AA1E39D352" box="[500,581,1596,1620]" pageId="20" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype. 
<emphasis id="B91DB954D16C953B423BF9AD1F6FD355" bold="true" box="[769,787,1595,1619]" pageId="20" pageNumber="571">A</emphasis>
Epigynum, ventral view; 
<emphasis id="B91DB954D16C953B4520F9AA1857D352" bold="true" box="[1050,1067,1596,1620]" pageId="20" pageNumber="571">B</emphasis>
, 
<emphasis id="B91DB954D16C953B450DF9AD1835D355" bold="true" box="[1079,1097,1595,1619]" pageId="20" pageNumber="571">C</emphasis>
Internal genitalia, cleared, dorsal view. Scale bars 0.2 mm. Abbreviations: at, atrium; b, bursa; cnd, connecting duct; co, copulatory opening; cod, copulatory duct; fd, fertilization duct; s, spermatheca.
</paragraph>
</caption>
<subSubSection id="C37336CDD16C953A41FDF95F196FD41B" lastPageId="21" lastPageNumber="572" pageId="20" pageNumber="571" type="diagnosis">
<paragraph id="8BD66546D16C953A41FDF95F196FD41B" blockId="20.[151,1437,1737,2016]" lastBlockId="21.[151,1437,151,285]" lastPageId="21" lastPageNumber="572" pageId="20" pageNumber="571">
<emphasis id="B91DB954D16C953B41FDF95F1D3ED3E5" bold="true" box="[199,322,1737,1763]" pageId="20" pageNumber="571">Diagnosis.</emphasis>
This species belongs to the 
<taxonomicName id="4C691EC5D16C953B434CF95C1F6CD3E2" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[630,784,1738,1764]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16C953B434CF95C1F6CD3E2" box="[630,784,1738,1764]" italics="true" pageId="20" pageNumber="571">kinabaluensis</emphasis>
</taxonomicName>
-group based on the following characters: carapace wedge-shaped with undulating lateral margins, posterior end of carapace not produced into an elongated stalk, PER strongly recurved and much longer than AER, palpal tegulum not enlarged and bulbous, bursae curved backwards towards spermathecae. Among the 
<taxonomicName id="4C691EC5D16C953B40F8F8A01E20D256" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[450,604,1846,1872]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16C953B40F8F8A01E20D256" box="[450,604,1846,1872]" italics="true" pageId="20" pageNumber="571">kinabaluensis</emphasis>
</taxonomicName>
-group, males of this species are most similar to those of 
<taxonomicName id="4C691EC5D16C953B45DAF8A019E0D256" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[1248,1436,1846,1872]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16C953B45DAF8A019E0D256" box="[1248,1436,1846,1872]" italics="true" pageId="20" pageNumber="571">U. kinabaluensis</emphasis>
</taxonomicName>
by the RTA ending in a wide arch and an apical claw separated by a membranous area, but can be distinguished by the thumb-shaped RTA-claw with a broad tip (
<figureCitation id="135279C3D16C953B43ECF8E81F22D29E" box="[726,862,1917,1944]" captionStart="FIGURE 15" captionStartId="18.[151,250,1923,1947]" captionTargetBox="[164,1423,453,1893]" captionTargetId="figure-110@18.[164,1423,453,1900]" captionTargetPageId="18" captionText="FIGURE 15. Utivarachna trisula sp. nov., male holotype. A–E Left palp: A, prolateral view; B, D, ventral view; C, E retrolateral view. Scale bars 0.2 mm. Abbreviations: ac, apical claw of RTA; dl, dorsal lobe of RTA; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780108" httpUri="https://zenodo.org/record/10780108/files/figure.png" pageId="20" pageNumber="571">Fig. 15B, E</figureCitation>
) versus curved and with a sickle-shaped tip in 
<taxonomicName id="4C691EC5D16C953B44BBF8E81D4DD2BA" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16C953B44BBF8E81D4DD2BA" italics="true" pageId="20" pageNumber="571">U. kinabaluensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF818B7D16C953B407AF8341E1BD2BA" author="Deeleman-Reinhold, C. L." box="[320,615,1954,1980]" pageId="20" pageNumber="571" refId="ref9947" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
: figs 594–595; 
<bibRefCitation id="EFF818B7D16C953B422FF8341FBAD2BA" author="Yamasaki, T. &amp; Hashimoto, Y. &amp; Endo, T. &amp; Hyodo, F. &amp; Itioka, T. &amp; Mohamed, M. &amp; Meleng, P." box="[789,966,1954,1980]" pageId="20" pageNumber="571" pagination="55 - 73" refId="ref11196" refString="Yamasaki, T., Hashimoto, Y., Endo, T., Hyodo, F., Itioka, T., Mohamed, M. &amp; Meleng, P. (2023) Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species. Zootaxa, 5343 (1), 55 - 73. https: // doi. org / 10.11646 / zootaxa. 5343.1.3" type="journal article" year="2023">Yamasaki 2023</bibRefCitation>
: figs 25A, B, D), and the arch of the RTA extending into a thinner, membranous lobe on the dorsal side. Females of this species are most similar to those of 
<taxonomicName id="4C691EC5D16C953A44BBF8501D4DD5B7" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" lastPageId="21" lastPageNumber="572" order="Araneae" pageId="20" pageNumber="571" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16C953A44BBF8501D4DD5B7" italics="true" lastPageId="21" lastPageNumber="572" pageId="20" pageNumber="571">U. kinabaluensis</emphasis>
</taxonomicName>
in having the copulatory ducts produced into a horizontal helical coil, but can be distinguished by the posteriorly wider bursae versus anteriorly wider in 
<taxonomicName id="4C691EC5D16D953A43EAFF2D1FF0D5D3" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[720,908,187,213]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="572" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16D953A43EAFF2D1FF0D5D3" box="[720,908,187,213]" italics="true" pageId="21" pageNumber="572">U. kinabaluensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF818B7D16D953A42A0FF2D18F5D5D3" author="Yamasaki, T. &amp; Hashimoto, Y. &amp; Endo, T. &amp; Hyodo, F. &amp; Itioka, T. &amp; Mohamed, M. &amp; Meleng, P." box="[922,1161,187,213]" pageId="21" pageNumber="572" pagination="55 - 73" refId="ref11196" refString="Yamasaki, T., Hashimoto, Y., Endo, T., Hyodo, F., Itioka, T., Mohamed, M. &amp; Meleng, P. (2023) Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species. Zootaxa, 5343 (1), 55 - 73. https: // doi. org / 10.11646 / zootaxa. 5343.1.3" type="journal article" year="2023">
Yamasaki 
<emphasis id="B91DB954D16D953A452AFF2D183BD5D3" box="[1040,1095,187,213]" italics="true" pageId="21" pageNumber="572">et al.</emphasis>
2023
</bibRefCitation>
: fig. 27C), and the thick, posteriorly broadening connecting ducts ending in not clearly distinguishable spermatheca (
<figureCitation id="135279C3D16D953A458EFF49193FD5FF" box="[1204,1347,223,249]" captionStart="FIGURE 17" captionStartId="20.[151,250,1595,1619]" captionTargetBox="[151,1436,344,1566]" captionTargetId="figure-82@20.[151,1436,344,1572]" captionTargetPageId="20" captionText="FIGURE 17. Utivarachna trisula sp. nov., female paratype. A Epigynum, ventral view; B, C Internal genitalia, cleared, dorsal view. Scale bars 0.2 mm.Abbreviations: at, atrium; b, bursa; cnd, connecting duct; co, copulatory opening; cod, copulatory duct; fd, fertilization duct; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10780116" httpUri="https://zenodo.org/record/10780116/files/figure.png" pageId="21" pageNumber="572">Figs 17B, C</figureCitation>
) versus clearly distinguishable, oval-shaped spermatheca in 
<taxonomicName id="4C691EC5D16D953A43DAFE951FE0D41B" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[736,924,259,285]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="572" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16D953A43DAFE951FE0D41B" box="[736,924,259,285]" italics="true" pageId="21" pageNumber="572">U. kinabaluensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF818B7D16D953A4290FE9518E0D41B" author="Yamasaki, T. &amp; Hashimoto, Y. &amp; Endo, T. &amp; Hyodo, F. &amp; Itioka, T. &amp; Mohamed, M. &amp; Meleng, P." box="[938,1180,259,285]" pageId="21" pageNumber="572" pagination="55 - 73" refId="ref11196" refString="Yamasaki, T., Hashimoto, Y., Endo, T., Hyodo, F., Itioka, T., Mohamed, M. &amp; Meleng, P. (2023) Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species. Zootaxa, 5343 (1), 55 - 73. https: // doi. org / 10.11646 / zootaxa. 5343.1.3" type="journal article" year="2023">
Yamasaki 
<emphasis id="B91DB954D16D953A451BFE951825D41B" box="[1057,1113,259,285]" italics="true" pageId="21" pageNumber="572">et al.</emphasis>
2023
</bibRefCitation>
: fig. 27C).
</paragraph>
</subSubSection>
<caption id="DF1635CED16D953A41A2FB021D18D1D6" ID-DOI="http://doi.org/10.5281/zenodo.10794102" ID-Zenodo-Dep="10794102" httpUri="https://zenodo.org/record/10794102/files/figure.png" pageId="21" pageNumber="572" startId="21.[152,212,1172,1196]" targetBox="[151,1435,309,1148]" targetPageId="21">
<paragraph id="8BD66546D16D953A41A2FB021D18D1D6" blockId="21.[151,1436,1172,1232]" pageId="21" pageNumber="572">
<emphasis id="B91DB954D16D953A41A2FB021C90D1AA" bold="true" box="[152,236,1172,1196]" pageId="21" pageNumber="572">MAP 1.</emphasis>
Distribution map of 
<taxonomicName id="4C691EC5D16D953A40FBFB021EEFD1AA" box="[449,659,1172,1196]" pageId="21" pageNumber="572">
<emphasis id="B91DB954D16D953A40FBFB021E3ED1AA" box="[449,578,1172,1196]" italics="true" pageId="21" pageNumber="572">Utivarachna</emphasis>
species
</taxonomicName>
collected in the framework of the 2013 EFForTS project in 
<collectingRegion id="49ADABA4D16D953A45C2FB0219E5D1AA" box="[1272,1433,1172,1196]" country="Indonesia" name="Jambi" pageId="21" pageNumber="572">Jambi Province</collectingRegion>
, 
<collectingRegion id="49ADABA4D16D953A41ADFB2E1C93D1D6" box="[151,239,1208,1232]" country="Indonesia" name="Sumatera Utara" pageId="21" pageNumber="572">Sumatra</collectingRegion>
, 
<collectingCountry id="F37E25D6D16D953A41C3FB2E1D1CD1D6" box="[249,352,1208,1232]" name="Indonesia" pageId="21" pageNumber="572">Indonesia</collectingCountry>
.
</paragraph>
</caption>
<subSubSection id="C37336CDD16D953941FDFB681FF6D613" lastPageId="22" lastPageNumber="573" pageId="21" pageNumber="572" type="description">
<paragraph id="8BD66546D16D953A41FDFB681CABD312" blockId="21.[151,1437,1278,2025]" pageId="21" pageNumber="572">
<emphasis id="B91DB954D16D953A41FDFB681E57D01E" bold="true" box="[199,555,1278,1304]" pageId="21" pageNumber="572">
Description. Male (
<typeStatus id="54D2DBE4D16D953A4091FB681E62D01E" box="[427,542,1278,1304]" pageId="21" pageNumber="572" type="holotype">Holotype</typeStatus>
).
</emphasis>
Total length 3.41. Carapace length 1.72; width 1.72. Carapace maroon, almost wedge-shaped, with narrowly truncate posterior end, strongly sclerotized, surface granulated, with each granule ending in long white seta, lateral margins undulating, fovea short but distinct; PER longer than AER, both slightly recurved (
<figureCitation id="135279C3D16D953A4036FAFC1D04D082" box="[268,376,1386,1412]" captionStart="FIGURE 14" captionStartId="17.[151,250,1933,1957]" captionTargetBox="[151,1436,273,1910]" captionTargetId="figure-65@17.[151,1436,273,1910]" captionTargetPageId="17" captionText="FIGURE 14. Utivarachna trisula sp. nov., male holotype. A, Habitus, dorsal view; B, Sternum, ventral view; C, Left leg I, prolateral view. Scale bars A = 1 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780106" httpUri="https://zenodo.org/record/10780106/files/figure.png" pageId="21" pageNumber="572">Fig. 14A</figureCitation>
); clypeus short, with slight projection between chelicerae. Eye diameters: AME 0.09; ALE 0.12; PLE 0.11; PME 0.11. Eye interdistances: AME–AME 0.07; AME–ALE 0.08; ALE–ALE 0.38; PME–PME 0.14; PME–PLE 0.16; ALE–PLE 0.11; AME–PME 0.08; PLE–PLE 0.69. MOA: length 0.28; anterior width 0.25; posterior width 0.35. Clypeus height 0.21. Sternum heart-shaped, colored as carapace, surface granulated (
<figureCitation id="135279C3D16D953A4454FA401CB6D312" captionStart="FIGURE 14" captionStartId="17.[151,250,1933,1957]" captionTargetBox="[151,1436,273,1910]" captionTargetId="figure-65@17.[151,1436,273,1910]" captionTargetPageId="17" captionText="FIGURE 14. Utivarachna trisula sp. nov., male holotype. A, Habitus, dorsal view; B, Sternum, ventral view; C, Left leg I, prolateral view. Scale bars A = 1 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780106" httpUri="https://zenodo.org/record/10780106/files/figure.png" pageId="21" pageNumber="572">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="8BD66546D16D953A41FDF9881D68D3EA" blockId="21.[151,1437,1278,2025]" pageId="21" pageNumber="572">
Abdomen length 1.69; width 1.30. Abdomen oval, pale-colored, with two pairs of dark patches on lateral margin, one anteriorly and one posteriorly, as well as darker-shaded cardiac pattern and pair of small dark patches directly anterior to spinnerets; most of dorsal surface of abdomen covered with orange scutum (
<figureCitation id="135279C3D16D953A45EAF9F01945D386" box="[1232,1337,1638,1664]" captionStart="FIGURE 14" captionStartId="17.[151,250,1933,1957]" captionTargetBox="[151,1436,273,1910]" captionTargetId="figure-65@17.[151,1436,273,1910]" captionTargetPageId="17" captionText="FIGURE 14. Utivarachna trisula sp. nov., male holotype. A, Habitus, dorsal view; B, Sternum, ventral view; C, Left leg I, prolateral view. Scale bars A = 1 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780106" httpUri="https://zenodo.org/record/10780106/files/figure.png" pageId="21" pageNumber="572">Fig. 14A</figureCitation>
); ventral side of abdomen sclerotized on area anterior to epigastric furrow, projecting a short distance posterior to it on sides, four longitudinal rows of rather faintly sclerotized dots present between epigastric furrow and spinnerets (
<figureCitation id="135279C3D16D953A419AF9441D7BD3EA" box="[160,263,1746,1772]" captionStart="FIGURE 14" captionStartId="17.[151,250,1933,1957]" captionTargetBox="[151,1436,273,1910]" captionTargetId="figure-65@17.[151,1436,273,1910]" captionTargetPageId="17" captionText="FIGURE 14. Utivarachna trisula sp. nov., male holotype. A, Habitus, dorsal view; B, Sternum, ventral view; C, Left leg I, prolateral view. Scale bars A = 1 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780106" httpUri="https://zenodo.org/record/10780106/files/figure.png" pageId="21" pageNumber="572">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="8BD66546D16D953A41FDF9611CA6D2C2" blockId="21.[151,1437,1278,2025]" pageId="21" pageNumber="572">
Legs yellow, covered with long white setae, especially on ventral surface, striated with black bands distally and proximally on tibiae I–IV, while on femora I, II only with distal bands. Anterior legs stouter and longer than posterior legs, ventrally filled with leg cuspules from tarsus to tibiae (
<figureCitation id="135279C3D16D953A4297F8A81868D25E" box="[941,1044,1854,1880]" captionStart="FIGURE 14" captionStartId="17.[151,250,1933,1957]" captionTargetBox="[151,1436,273,1910]" captionTargetId="figure-65@17.[151,1436,273,1910]" captionTargetPageId="17" captionText="FIGURE 14. Utivarachna trisula sp. nov., male holotype. A, Habitus, dorsal view; B, Sternum, ventral view; C, Left leg I, prolateral view. Scale bars A = 1 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780106" httpUri="https://zenodo.org/record/10780106/files/figure.png" pageId="21" pageNumber="572">Fig. 14C</figureCitation>
), metatarsi III and IV distally with comb-like structure followed by brush of setae. Leg measurements: leg I 4.31 (1.26, 0.45, 1.15, 0.96, 0.49); leg II 3.94 (1.18, 0.35, 1.05, 0.9, 0.46); leg III 2.97 (0.88, 0.28, 0.69, 0.78, 0.34); leg IV 3.67 (1.00, 0.31, 0.86, 1.06, 0.44).
</paragraph>
<paragraph id="8BD66546D16D953941FDF858192ED5FF" blockId="21.[151,1437,1278,2025]" lastBlockId="22.[151,1437,151,1041]" lastPageId="22" lastPageNumber="573" pageId="21" pageNumber="572">
Male palp (
<figureCitation id="135279C3D16D953A407EF8581DEAD2EF" box="[324,406,1998,2025]" captionStart="FIGURE 15" captionStartId="18.[151,250,1923,1947]" captionTargetBox="[164,1423,453,1893]" captionTargetId="figure-110@18.[164,1423,453,1900]" captionTargetPageId="18" captionText="FIGURE 15. Utivarachna trisula sp. nov., male holotype. A–E Left palp: A, prolateral view; B, D, ventral view; C, E retrolateral view. Scale bars 0.2 mm. Abbreviations: ac, apical claw of RTA; dl, dorsal lobe of RTA; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780108" httpUri="https://zenodo.org/record/10780108/files/figure.png" pageId="21" pageNumber="572">Fig. 15</figureCitation>
): Cymbium and bulb oval, sperm duct U-shaped with rather broad turn. Embolus long, coiled horizontally, looping twice, visible dorsally through cymbium. RTA rather large, extending retrolaterally, ending in wide arch and apical claw, lobes of arch thumb-shaped, dorsal lobe continuing into longer, thinner part; apical claw stout, thumb-shaped with wide base, positioned between arch-lobes, separated by narrow membranous area.
</paragraph>
<paragraph id="8BD66546D16E953941FDFE951DA2D71F" blockId="22.[151,1437,151,1041]" pageId="22" pageNumber="573">
<emphasis id="B91DB954D16E953941FDFE951DCCD41A" bold="true" box="[199,432,258,285]" pageId="22" pageNumber="573">
Female (
<typeStatus id="54D2DBE4D16E9539400BFE941DDFD41A" box="[305,419,258,284]" pageId="22" pageNumber="573" type="paratype">Paratype</typeStatus>
).
</emphasis>
Total length 3.88. Carapace length 1.70; width 1.35. Eye diameters: AME 0.10; ALE 0.12; PLE 0.11; PME 0.11. Eye interdistances: AME–AME 0.06; AME–ALE 0.06; ALE–ALE 0.34; PME–PME 0.15; PME–PLE 0.15; ALE–PLE 0.09; AME–PME 0.06; PLE–PLE 0.63. MOA: length 0.27; anterior width 0.26; posterior width 0.34. Clypeus height 0.18. Abdomen length 2.18; width 1.71. General appearance as in male except abdomen not covered in dorsal scutum, with three pairs of dark lateral patches (
<figureCitation id="135279C3D16E95394509FE0518E2D4AB" box="[1075,1182,403,429]" captionStart="FIGURE 16" captionStartId="19.[151,250,1912,1936]" captionTargetBox="[151,1436,381,1888]" captionTargetId="figure-54@19.[151,1436,381,1888]" captionTargetPageId="19" captionText="FIGURE 16. Utivarachna trisula sp. nov., female paratype. A, B Habitus, dorsal view (A), ventral view (B); C, Left leg I, prolateral view. Scale bars A, B = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780112" httpUri="https://zenodo.org/record/10780112/files/figure.png" pageId="22" pageNumber="573">Fig. 16A</figureCitation>
), longitudinal rows of sclerotized dots absent on venter (
<figureCitation id="135279C3D16E95394322FE211E03D4D7" box="[536,639,439,465]" captionStart="FIGURE 16" captionStartId="19.[151,250,1912,1936]" captionTargetBox="[151,1436,381,1888]" captionTargetId="figure-54@19.[151,1436,381,1888]" captionTargetPageId="19" captionText="FIGURE 16. Utivarachna trisula sp. nov., female paratype. A, B Habitus, dorsal view (A), ventral view (B); C, Left leg I, prolateral view. Scale bars A, B = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780112" httpUri="https://zenodo.org/record/10780112/files/figure.png" pageId="22" pageNumber="573">Fig. 16B</figureCitation>
). Leg cuspules absent (
<figureCitation id="135279C3D16E953942B2FE211F93D4D7" box="[904,1007,439,465]" captionStart="FIGURE 16" captionStartId="19.[151,250,1912,1936]" captionTargetBox="[151,1436,381,1888]" captionTargetId="figure-54@19.[151,1436,381,1888]" captionTargetPageId="19" captionText="FIGURE 16. Utivarachna trisula sp. nov., female paratype. A, B Habitus, dorsal view (A), ventral view (B); C, Left leg I, prolateral view. Scale bars A, B = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780112" httpUri="https://zenodo.org/record/10780112/files/figure.png" pageId="22" pageNumber="573">Fig. 16C</figureCitation>
). Leg measurements: leg I 3.98 (1.14, 0.46, 1.04, 0.83, 0.51); leg II (3.90; 1.18, 0.43, 0.99, 0.91); leg III 2.85 (0.79, 0.26, 0.68, 0.76, 0.36); leg IV 3.50 (0.89, 0.26, 0.84, 1.06, 0.45).
</paragraph>
<paragraph id="8BD66546D16E953941FDFDB51FF6D613" blockId="22.[151,1437,151,1041]" pageId="22" pageNumber="573">
Copulatory organs (
<figureCitation id="135279C3D16E95394095FDB51E7BD73B" box="[431,519,547,573]" captionStart="FIGURE 17" captionStartId="20.[151,250,1595,1619]" captionTargetBox="[151,1436,344,1566]" captionTargetId="figure-82@20.[151,1436,344,1572]" captionTargetPageId="20" captionText="FIGURE 17. Utivarachna trisula sp. nov., female paratype. A Epigynum, ventral view; B, C Internal genitalia, cleared, dorsal view. Scale bars 0.2 mm.Abbreviations: at, atrium; b, bursa; cnd, connecting duct; co, copulatory opening; cod, copulatory duct; fd, fertilization duct; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10780116" httpUri="https://zenodo.org/record/10780116/files/figure.png" pageId="22" pageNumber="573">Fig. 17</figureCitation>
): Atrium wider than long. Copulatory openings located on anterior corners of atrium, roughly midway between pedicel and epigastric furrow. Copulatory ducts initially thick, narrowing anteriorly, extending into broad horizontal coil and then into three smaller helical coils, finally turning into straight tube towards middle of coils. Bursae slightly inflated, pear-shaped, anterior part curved. Connecting ducts thick, positioned in between the copulatory openings and bursae, broadening posteriorly, ending into bulbous, not clearly distinguishable spermathecae, together forming an S-shaped structure. Fertilization ducts extending from bulbous posterior end of spermathecae, as long as the latter, slightly curved.
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD16E953941FDFC881F44D63F" box="[199,824,798,825]" pageId="22" pageNumber="573" type="distribution">
<paragraph id="8BD66546D16E953941FDFC881F44D63F" blockId="22.[151,1437,151,1041]" box="[199,824,798,825]" pageId="22" pageNumber="573">
<emphasis id="B91DB954D16E953941FDFC881D1DD63E" bold="true" box="[199,353,798,824]" pageId="22" pageNumber="573">Distribution.</emphasis>
<collectingRegion id="49ADABA4D16E9539405DFC891E6BD63F" box="[359,535,799,825]" country="Indonesia" name="Jambi" pageId="22" pageNumber="573">Jambi Province</collectingRegion>
, 
<collectingRegion id="49ADABA4D16E95394319FC891EFCD63F" box="[547,640,799,825]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="573">Sumatra</collectingRegion>
(
<figureCitation id="135279C3D16E953943B5FC891E98D63F" box="[655,740,799,825]" captionStart="MAP 1" captionStartId="21.[152,212,1172,1196]" captionTargetBox="[151,1435,309,1148]" captionTargetId="figure-78@21.[151,1435,309,1148]" captionText="MAP 1. Distribution map of Utivarachna species collected in the framework of the 2013 EFForTS project in Jambi Province, Sumatra, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.10794102" httpUri="https://zenodo.org/record/10794102/files/figure.png" pageId="22" pageNumber="573">Maps 1</figureCitation>
and 
<figureCitation id="135279C3D16E95394220FC891F57D63F" box="[794,811,799,825]" captionStart="MAP 2" captionStartId="22.[152,211,1994,2018]" captionTargetBox="[151,1436,1065,1971]" captionTargetId="figure-394@22.[151,1436,1065,1971]" captionText="MAP 2. Global distribution map of the Utivarachna species treated in this paper." figureDoi="http://doi.org/10.5281/zenodo.10794104" httpUri="https://zenodo.org/record/10794104/files/figure.png" pageId="22" pageNumber="573">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD16E953941FDFCD51FB4D117" pageId="22" pageNumber="573" type="discussion">
<paragraph id="8BD66546D16E953941FDFCD51FB4D117" blockId="22.[151,1437,151,1041]" pageId="22" pageNumber="573">
<emphasis id="B91DB954D16E953941FDFCD51D47D65B" bold="true" box="[199,315,835,861]" pageId="22" pageNumber="573">Remarks.</emphasis>
All specimens were collected by fogging in jungle rubber plantations, and are considered arboreal. The genitalic similarity of 
<taxonomicName id="4C691EC5D16E953940F6FCF11E45D687" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[460,569,871,897]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="trisula">
<emphasis id="B91DB954D16E953940F6FCF11E45D687" box="[460,569,871,897]" italics="true" pageId="22" pageNumber="573">U. trisula</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C691EC5D16E9539434FFCF11F4FD687" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[629,819,871,897]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16E9539434FFCF11F4FD687" box="[629,819,871,897]" italics="true" pageId="22" pageNumber="573">U. kinabaluensis</emphasis>
</taxonomicName>
to members of the 
<taxonomicName id="4C691EC5D16E95394523FCF118E3D687" authorityName="Kishida" authorityYear="1940" box="[1049,1183,871,897]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B91DB954D16E95394523FCF118E3D687" box="[1049,1183,871,897]" italics="true" pageId="22" pageNumber="573">fukasawana</emphasis>
</taxonomicName>
-group (particularly in regards to the RTA-claw and coiling of the copulatory ducts) might suggests a closer relationship with them than towards other members of the 
<taxonomicName id="4C691EC5D16E953940CBFC391EF7D6CF" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[497,651,943,969]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16E953940CBFC391EF7D6CF" box="[497,651,943,969]" italics="true" pageId="22" pageNumber="573">kinabaluensis</emphasis>
</taxonomicName>
-group. Currently, the 
<taxonomicName id="4C691EC5D16E953942B2FC39185ED6CF" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[904,1058,943,969]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D16E953942B2FC39185ED6CF" box="[904,1058,943,969]" italics="true" pageId="22" pageNumber="573">kinabaluensis</emphasis>
</taxonomicName>
-group is only distinguished from the 
<taxonomicName id="4C691EC5D16E953941F8FC451D34D6EB" authorityName="Kishida" authorityYear="1940" box="[194,328,979,1005]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B91DB954D16E953941F8FC451D34D6EB" box="[194,328,979,1005]" italics="true" pageId="22" pageNumber="573">fukasawana</emphasis>
</taxonomicName>
-group by a single character, which is the absence of an elongation of the carapace posterior end. Thus, a revision of the grouping of 
<taxonomicName id="4C691EC5D16E9539431EFC611F75D117" box="[548,777,1015,1041]" pageId="22" pageNumber="573">
<emphasis id="B91DB954D16E9539431EFC611ECED117" box="[548,690,1015,1041]" italics="true" pageId="22" pageNumber="573">Utivarachna</emphasis>
species
</taxonomicName>
is likely needed.
</paragraph>
</subSubSection>
</treatment>
</document>