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<document id="8B3FE84C0EE7B50FB12D38350571F78D" ID-DOI="10.11646/zootaxa.4809.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3934048" ID-ZooBank="E697983F-7223-4009-91B4-80A401D0AB8A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594126017644" checkinUser="plazi" docAuthor="Johanson, Kjell Arne, Mary, Nathalie Jeanne, Sjöberg, Tin &amp; Malm, Tobias" docDate="2020" docId="03C0C513FFD8FFECD99707B294BC34D1" docLanguage="en" docName="zootaxa.4809.2.1.pdf" docOrigin="Zootaxa 4809 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oecetis variabilis Johanson &amp; Mary &amp; Sjöberg &amp; Malm 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="231" masterDocId="FFF9BD6BFFC4FFF2D900023795403622" masterDocTitle="Eighteen new species of Oecetis McLachlan 1877 (Trichoptera, Leptoceridae) from New Caledonia" masterLastPageNumber="240" masterPageNumber="201" pageNumber="229" updateTime="1698778874990" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="11B1F1A337165EC325636801F2368627">Eighteen new species of Oecetis McLachlan 1877 (Trichoptera, Leptoceridae) from New Caledonia</mods:title>
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<mods:namePart id="47A3F94E2D3059A3ED7194129F4ADC4D">Johanson, Kjell Arne</mods:namePart>
<mods:affiliation id="79CC2E1A14733808316632E2D3969CBC">* kjell. arne. johanson @ nrm. se; https: // orcid. org / 0000 - 0002 - 1893 - 3429 &amp; * kjell. arne. johanson @ nrm. se; https: // orcid. org / 0000 - 0002 - 1893 - 3429</mods:affiliation>
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<mods:namePart id="C9A5D5EEF0D5459CCF67A2003A031457">Mary, Nathalie Jeanne</mods:namePart>
<mods:affiliation id="C29C1A5EFD094EAF17AE94F5E026DE96">ETHYCO, B. P. 13821, 98 803 Nouméa Cedex, Nouvelle-Calédonie. &amp; ethyco 2005 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8250 - 7929</mods:affiliation>
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<mods:namePart id="C1B6C1CBEBF985442FD512DBC00D2750">Sjöberg, Tin</mods:namePart>
<mods:affiliation id="94280CA41B0E6DCA17F38B47F346EFAF">* kjell. arne. johanson @ nrm. se; https: // orcid. org / 0000 - 0002 - 1893 - 3429 &amp; tin. sjoberg @ nrm. se; https: // orcid. org / 0000 - 0002 - 2813 - 9325</mods:affiliation>
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<mods:namePart id="222F1400AF55627EA485FD7C410E822D">Malm, Tobias</mods:namePart>
<mods:affiliation id="DE0563AD57C44105709DAAADA33DA9A6">* kjell. arne. johanson @ nrm. se; https: // orcid. org / 0000 - 0002 - 1893 - 3429 &amp; tobias. malm @ nrm. se; https: // orcid. org / 0000 - 0003 - 4610 - 4086</mods:affiliation>
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<mods:date id="F06AD6E78139EA08067D73A14543860F">2020</mods:date>
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<mods:number id="DCD83E55CED4DC6B72974C8BED1A4FE7">4809</mods:number>
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<paragraph id="8BD67405FFD8FFEED99707B2948C3382" blockId="28.[151,460,1413,1476]" box="[151,460,1413,1440]" pageId="28" pageNumber="229">
<heading id="D09EC369FFD8FFEED99707B2948C3382" bold="true" box="[151,460,1413,1440]" fontSize="11" level="1" pageId="28" pageNumber="229" reason="1">
<emphasis id="B91DA817FFD8FFEED99707B2948C3382" bold="true" box="[151,460,1413,1440]" pageId="28" pageNumber="229">
<taxonomicName id="4C690F86FFD8FFEED99707B294263382" authority="Johanson &amp; Mary &amp; Sjöberg &amp; Malm, 2020" authorityName="Johanson &amp; Mary &amp; Sjöberg &amp; Malm" authorityYear="2020" box="[151,358,1413,1440]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B91DA817FFD8FFEED99707B294263382" bold="true" box="[151,358,1413,1440]" italics="true" pageId="28" pageNumber="229">Oecetis variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E156CFFD8FFEED86E07B1948C3382" box="[366,460,1414,1440]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BD67405FFD8FFEED997079E945533E6" blockId="28.[151,460,1413,1476]" box="[151,277,1449,1476]" pageId="28" pageNumber="229">
<figureCitation id="13526880FFD8FFEED997079E945533E6" box="[151,277,1449,1476]" captionStart="FIGURES 8387" captionStartId="29.[152,270,1980,2004]" captionTargetBox="[191,1392,192,1942]" captionTargetId="figure@29.[184,1404,181,1954]" captionTargetPageId="29" captionText="FIGURES 8387. Genitalia of Oecetis variabilis sp. nov., male holotype. 83, left lateral; 84, dorsal; 85, ventral; 86, phallus, left lateral; 87, phallus, ventral." figureDoi="http://doi.org/10.5281/zenodo.3934088" httpUri="https://zenodo.org/record/3934088/files/figure.png" pageId="28" pageNumber="229">Figs 8387</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373278EFFD8FFEED99707C6919C309D" pageId="28" pageNumber="229" type="diagnosis">
<paragraph id="8BD67405FFD8FFEED99707C6919C309D" blockId="28.[151,1437,1521,2016]" pageId="28" pageNumber="229">
<emphasis id="B91DA817FFD8FFEED99707C694523029" bold="true" box="[151,274,1521,1547]" pageId="28" pageNumber="229">Diagnosis.</emphasis>
This species is distinguished from many of the Australasian
<taxonomicName id="4C690F86FFD8FFEEDABA07C5914E302E" authorityName="McLachlan" authorityYear="1877" box="[954,1038,1522,1548]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA817FFD8FFEEDABA07C5914E302E" box="[954,1038,1522,1548]" italics="true" pageId="28" pageNumber="229">Oecetis</emphasis>
</taxonomicName>
species by the absence of modified tergites on segment VIII and forward. The species is similar to several species distributed in Australasia. The coxopodite of
<taxonomicName id="4C690F86FFD8FFEED9F6040D94C43071" authority="Johanson &amp; Mary &amp; Sjöberg &amp; Malm, 2020" authorityName="Johanson &amp; Mary &amp; Sjöberg &amp; Malm" authorityYear="2020" box="[246,388,1593,1620]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B91DA817FFD8FFEED9F6040D94C43071" box="[246,388,1593,1620]" italics="true" pageId="28" pageNumber="229">O. variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E156CFFD8FFEED88C040D94A43076" box="[396,484,1594,1620]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="4C690F86FFD8FFEEDBC5040D91643076" authority="Tillyard 1925" authorityName="Tillyard" authorityYear="1925" box="[709,1060,1593,1620]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="chathamensis">
<emphasis id="B91DA817FFD8FFEEDBC5040D96C03071" box="[709,896,1593,1620]" italics="true" pageId="28" pageNumber="229">O. chathamensis</emphasis>
<bibRefCitation id="EFF809F4FFD8FFEEDA88040E91643076" author="Tillyard, R. J." box="[904,1060,1593,1620]" pageId="28" pageNumber="229" pagination="277 - 284" refId="ref19908" refString="Tillyard, R. J. (1925) Caddis-flies (order Trichoptera) from the Chatham Islands. Records of the Canterbury Museum, 2, 277 - 284." type="journal article" year="1925">Tillyard 1925</bibRefCitation>
</taxonomicName>
, from the
<collectingCountry id="F37E3495FFD8FFEEDD98040D906F3071" box="[1176,1327,1593,1620]" name="New Zealand" pageId="28" pageNumber="229">New Zealand</collectingCountry>
<collectingRegion id="49ADBAE7FFD8FFEEDC37040E95A83055" country="New Zealand" name="Chatham Islands" pageId="28" pageNumber="229">Chatham islands</collectingRegion>
, but they also appear similar to those of the Australian
<taxonomicName id="4C690F86FFD8FFEEDA5504699125305A" authority="Neboiss 1977" authorityName="Neboiss" authorityYear="1977" box="[853,1125,1629,1656]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="umbra">
<emphasis id="B91DA817FFD8FFEEDA55046996813055" box="[853,961,1629,1656]" italics="true" pageId="28" pageNumber="229">O. umbra</emphasis>
<bibRefCitation id="EFF809F4FFD8FFEEDAC8046A9125305A" author="Neboiss, A." box="[968,1125,1629,1656]" pageId="28" pageNumber="229" pagination="1 - 208" refId="ref19830" refString="Neboiss, A. (1977) A taxonomic and zoogeographic study of Tasmanian caddis flies. Memoires of the National Museum of Victoria, 38, 1 - 208. https: // doi. org / 10.24199 / j. mmv. 1977.38.01" type="journal article" year="1977">Neboiss 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C690F86FFD8FFEEDD71046990D83055" authority="Wells 2006" authorityName="Wells" authorityYear="2006" box="[1137,1432,1629,1656]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="magelensis">
<emphasis id="B91DA817FFD8FFEEDD710469904F3055" box="[1137,1295,1629,1656]" italics="true" pageId="28" pageNumber="229">O. magelensis</emphasis>
<bibRefCitation id="EFF809F4FFD8FFEEDC17046A90D83055" author="Wells, A." box="[1303,1432,1629,1655]" pageId="28" pageNumber="229" pagination="107 - 128" refId="ref20025" refString="Wells, A. (2006) A review of Australian long-horned caddisflies in the Oecetis pechana - Group (Trichoptera: Leptoceridae), with descriptions of thirteen new species. Memoirs of Museum Victoria, 63 (2), 107 - 128. https: // doi. org / 10.24199 / j. mmv. 2006.63.13" type="journal article" year="2006">Wells 2006</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C690F86FFD8FFEED9C504B594A030BE" authority="Wells 2004" authorityName="Wells" authorityYear="2004" box="[197,480,1665,1692]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="crosslandi">
<emphasis id="B91DA817FFD8FFEED9C504B5941A30B9" box="[197,346,1665,1692]" italics="true" pageId="28" pageNumber="229">O. crosslandi</emphasis>
<bibRefCitation id="EFF809F4FFD8FFEED85F04B694A030BE" author="Wells, A." box="[351,480,1665,1692]" pageId="28" pageNumber="229" pagination="85 - 110" refId="ref19966" refString="Wells, A. (2004) The long-horned caddisfly genus Oecetis (Trichoptera: Leptoceridae) in Australia: Two new species groups and 17 new species. Memoires of the Museum of Victoria, 61 (1), 85 - 110. https: // doi. org / 10.24199 / j. mmv. 2004.61.7" type="journal article" year="2004">Wells 2004</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C690F86FFD8FFEED8E904B597EE30B9" authority="Johanson &amp; Mary &amp; Sjöberg &amp; Malm, 2020" authorityName="Johanson &amp; Mary &amp; Sjöberg &amp; Malm" authorityYear="2020" box="[489,686,1665,1692]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B91DA817FFD8FFEED8E904B597EE30B9" box="[489,686,1665,1692]" italics="true" pageId="28" pageNumber="229">Oecetis variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E156CFFD8FFEEDBB404B5964930BE" box="[692,777,1666,1692]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
is separated from these species in its shorter posterior spine of tergum X, the shape of the basoventral process of each coxopodite, and the very broad phallus.
</paragraph>
</subSubSection>
<subSubSection id="C373278EFFD8FFECD9C704FE946D3489" lastPageId="30" lastPageNumber="231" pageId="28" pageNumber="229" type="description">
<paragraph id="8BD67405FFD8FFECD9C704FE9587341D" blockId="28.[151,1437,1521,2016]" lastBlockId="30.[151,1437,153,755]" lastPageId="30" lastPageNumber="231" pageId="28" pageNumber="229">
<emphasis id="B91DA817FFD8FFEED9C704FE94E130C6" bold="true" box="[199,417,1737,1764]" pageId="28" pageNumber="229">Description, male.</emphasis>
Length of each forewing:
<quantity id="4C91D9E0FFD8FFEEDBCE04FD966530C6" box="[718,805,1738,1764]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="28" pageNumber="229" unit="mm" value="7.3">7.3 mm</quantity>
(N = 1). Genitalia (
<figureCitation id="13526880FFD8FFEEDD0504FE91C630C6" box="[1029,1158,1737,1764]" captionStart="FIGURES 8387" captionStartId="29.[152,270,1980,2004]" captionTargetBox="[191,1392,192,1942]" captionTargetId="figure@29.[184,1404,181,1954]" captionTargetPageId="29" captionText="FIGURES 8387. Genitalia of Oecetis variabilis sp. nov., male holotype. 83, left lateral; 84, dorsal; 85, ventral; 86, phallus, left lateral; 87, phallus, ventral." figureDoi="http://doi.org/10.5281/zenodo.3934088" httpUri="https://zenodo.org/record/3934088/files/figure.png" pageId="28" pageNumber="229">Figs 8387</figureCitation>
) slightly longer than tall in lateral view. Segment IX oriented vertically, narrowly drop-shaped in lateral view, ventral part much longer than dorsal part; in lateral view each side with anterior margin weakly convex; ventral half of posterior margin with tall, short, rounded lower posterolateral process of segment IX; upper posterolateral process of segment IX triangular, situated immediately below superior appendages; longitudinal apodeme located evident anteriorly (but not posteriorly) as continuation of lower margin of coxopodite; acrotergite of segment IX forming single uniform cluster of punctures, about as wide as distance between superior appendages in dorsal view; dorsal pair of semimembranous processes of segment IX short, posteriorly oriented lobes situated closely to each other; membranous areas absent. Main branch of tergum X forming pair of semimembranous lobes, about twice as long as superior appendages, oriented posteriorly and curving slightly ventrad near apex, apex with two long setae; in dorsal view well separated from half-length, oriented posteriorly. Posterior spine of tergum X originating basally on main branch, slightly longer than superior appendages, almost straight, oriented posteriorly; in lateral view slender along its length, basally wide in dorsal view before narrowing at mid-point. Superior appendages about half as long as main branch of tergum X, oblong-rectangular in lateral view; oval in dorsal view. Pair of coxopodites each forming tall plate for basal half, sharply narrowing at mid-length into slender process originating from lower edge; ventral process originating basally, immediately posterior of segment IX; asymmetrical with left and right ventral processes of different length. Phallus about as long as coxopodites, almost parallel-sided in lateral view, almost round (onion-shaped) in ventral view; long sclerotized spine originating from right-dorsal part of anterior end, curving ventrad and to left before sharply angling to the right before posterior end; short second sclerotized process located dorsally near posterior end.
</paragraph>
<caption id="DF16248DFFD9FFEFD998058B949531DA" ID-DOI="http://doi.org/10.5281/zenodo.3934088" ID-Zenodo-Dep="3934088" httpUri="https://zenodo.org/record/3934088/files/figure.png" pageId="29" pageNumber="230" startId="29.[152,270,1980,2004]" targetBox="[191,1392,192,1942]" targetPageId="29">
<paragraph id="8BD67405FFD9FFEFD998058B949531DA" blockId="29.[151,1436,1980,2041]" pageId="29" pageNumber="230">
<emphasis id="B91DA817FFD9FFEFD998058B941B31F7" bold="true" box="[152,347,1980,2005]" pageId="29" pageNumber="230">FIGURES 8387.</emphasis>
Genitalia of
<taxonomicName id="4C690F86FFD9FFEFD8E3058A97D731F6" authority="Johanson &amp; Mary &amp; Sjöberg &amp; Malm, 2020" authorityName="Johanson &amp; Mary &amp; Sjöberg &amp; Malm" authorityYear="2020" box="[483,663,1980,2004]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B91DA817FFD9FFEFD8E3058A97D731F6" box="[483,663,1980,2004]" italics="true" pageId="29" pageNumber="230">Oecetis variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E156CFFD9FFEFDB9E058A97B031F6" box="[670,752,1981,2004]" pageId="29" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype.
<emphasis id="B91DA817FFD9FFEFDA99058A96F231F7" bold="true" box="[921,946,1981,2005]" pageId="29" pageNumber="230">83</emphasis>
, left lateral;
<emphasis id="B91DA817FFD9FFEFDD37058A911031F7" bold="true" box="[1079,1104,1981,2005]" pageId="29" pageNumber="230">84</emphasis>
, dorsal;
<emphasis id="B91DA817FFD9FFEFDDAA058A918331F7" bold="true" box="[1194,1219,1981,2005]" pageId="29" pageNumber="230">85</emphasis>
, ventral;
<emphasis id="B91DA817FFD9FFEFDC26058A907F31F7" bold="true" box="[1318,1343,1981,2005]" pageId="29" pageNumber="230">86</emphasis>
, phallus, left lateral;
<emphasis id="B91DA817FFD9FFEFD80C05D6946531DB" bold="true" box="[268,293,2017,2041]" pageId="29" pageNumber="230">87</emphasis>
, phallus, ventral.
</paragraph>
</caption>
<paragraph id="8BD67405FFDAFFECD9C7007E946D3489" blockId="30.[151,1437,153,755]" pageId="30" pageNumber="231">
<emphasis id="B91DA817FFDAFFECD9C7007E94443441" bold="true" box="[199,260,585,611]" pageId="30" pageNumber="231">Male</emphasis>
<materialsCitation id="3B017E58FFDAFFECD80C007E94693489" collectingDate="2003-12-18" collectorName="K. A. Johanson" country="New Caledonia" latitude="-20.530333" location="Mont Panie" longLatPrecision="1" longitude="164.7836" municipality="Wan Pwe On Stream" pageId="30" pageNumber="231" specimenCount="1" stateProvince="Province Nord" typeStatus="holotype">
<emphasis id="B91DA817FFDAFFECD80C007E943C3441" bold="true" box="[268,380,585,611]" pageId="30" pageNumber="231">
<typeStatus id="54D2CAA7FFDAFFECD80C007E94373441" box="[268,375,585,611]" pageId="30" pageNumber="231" type="holotype">holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F37E3495FFDAFFECD883007D97743441" box="[387,564,585,612]" name="New Caledonia" pageId="30" pageNumber="231">New Caledonia</collectingCountry>
,
<collectingRegion id="49ADBAE7FFDAFFECDB40007E97A73441" box="[576,743,585,611]" country="New Caledonia" name="Province Nord" pageId="30" pageNumber="231">Province Nord</collectingRegion>
,
<collectingMunicipality id="6BB2EE7FFFDAFFECDBF3007D96A23441" box="[755,994,585,612]" pageId="30" pageNumber="231">Wan Pwé On Stream</collectingMunicipality>
, draining NNE side of
<location id="8EB622DEFFDAFFECDDE8007D902D3441" LSID="urn:lsid:plazi:treatment:03C0C513FFD8FFECD99707B294BC34D1:8EB622DEFFDAFFECDDE8007D902D3441" box="[1256,1389,585,612]" country="New Caledonia" latitude="-20.530333" longLatPrecision="1" longitude="164.7836" municipality="Wan Pwe On Stream" name="Mont Panie" pageId="30" pageNumber="231" stateProvince="Province Nord">Mont Panié</location>
,
<locationDeviation id="9A646D27FFDAFFECDC79007E94F034AA" location="Cascade de Tao" pageId="30" pageNumber="231">
<quantity id="4C91D9E0FFDAFFECDC79007E95FB34A5" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="30" pageNumber="231" unit="km" value="3.9">3.9 km</quantity>
NW Cascade de Tao
</locationDeviation>
,
<geoCoordinate id="EE5D12C2FFDAFFECD8BD005A970C34A5" box="[445,588,621,647]" degrees="20" direction="south" minutes="31.820" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="-20.530333">20°31.820S</geoCoordinate>
,
<geoCoordinate id="EE5D12C2FFDAFFECDB5A005A97B934A5" box="[602,761,621,647]" degrees="164" direction="east" minutes="47.016" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="164.7836">164°47.016E</geoCoordinate>
,
<date id="FFD752C5FFDAFFECDA07005A96C834A5" box="[775,904,621,647]" pageId="30" pageNumber="231" value="2003-12-18">
<collectingDate id="EF93AB2DFFDAFFECDA07005A96C834A5" box="[775,904,621,647]" pageId="30" pageNumber="231" value="2003-12-18">18.xii.2003</collectingDate>
</date>
,
<collectingMethod id="52280C12FFDAFFECDA96005A914034AA" box="[918,1024,621,648]" pageId="30" pageNumber="231">light trap</collectingMethod>
, loc#085, leg.
<collectorName id="269C11D3FFDAFFECDDAA005A901434A5" box="[1194,1364,621,647]" pageId="30" pageNumber="231">K.A. Johanson</collectorName>
, DNA voucher JK1
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373278EFFDAFFECD9C7008294BC34D1" pageId="30" pageNumber="231" type="etymology">
<paragraph id="8BD67405FFDAFFECD9C7008294BC34D1" blockId="30.[151,1437,153,755]" pageId="30" pageNumber="231">
<emphasis id="B91DA817FFDAFFECD9C70082940E34ED" bold="true" box="[199,334,693,719]" pageId="30" pageNumber="231">Etymology.</emphasis>
The Latin adjective
<taxonomicName id="4C690F86FFDAFFECDB36008297E134ED" authorityName="Johanson &amp; Mary &amp; Sjöberg &amp; Malm" authorityYear="2020" box="[566,673,693,719]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B91DA817FFDAFFECDB36008297E134ED" box="[566,673,693,719]" italics="true" pageId="30" pageNumber="231">variabilis</emphasis>
</taxonomicName>
, means “variable,” referring to the shape of the ventral processes of the coxopodites in ventral view.
</paragraph>
</subSubSection>
</treatment>
</document>