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<document id="49C50F4388DC9ABA52E1F7C2BE2AFFD8" ID-DOI="10.11646/zootaxa.2472.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="6507307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629679035506" checkinUser="felipe" docAuthor="Andersen, Trond, Saether, Ole A. &amp; Mendes, Humberto F." docDate="2010" docId="03C0582D9903FFA4BB8BFC7F072D6AA0" docLanguage="en" docName="zt02472p077.pdf" docOrigin="Zootaxa 2472 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2472.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Allocladius quadrus Andersen &amp; Saether &amp; Mendes 2010, sp. n." docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="FFF920559913FFB6BB1CFFF7045F683C" masterDocTitle="Neotropical Allocladius Kieffer, 1913 and Pseudosmittia Edwards, 1932 (Diptera: Chironomidae) 2472" masterLastPageNumber="77" masterPageNumber="1" pageNumber="17" updateTime="1699094922473" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="7262BA7461F9CEE6C291B24FA72F23B7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9F142E007F44FCEC25260BD389AFD7A5">Neotropical Allocladius Kieffer, 1913 and Pseudosmittia Edwards, 1932 (Diptera: Chironomidae) 2472</mods:title>
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<mods:name id="AE7F2F0549E593BC4D70B1E7E31A24FC" type="personal">
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<mods:namePart id="86EF21BE965BC6425B350EDACFB3ED23">Andersen, Trond</mods:namePart>
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<mods:namePart id="DF271E8303387902CCD03BF4F5CD1F09">Saether, Ole A.</mods:namePart>
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<mods:role id="1652B3A1ABB6FD7D544A191B3154B88A">
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<mods:namePart id="8F0A9AF102A1CA9CE475F3CAC4ACBA1C">Mendes, Humberto F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="43160D6E358806BA4AFF66008E78DA9B">text</mods:typeOfResource>
<mods:relatedItem id="80DA1F3867EF8D5C737DBEF4E58D94E7" type="host">
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<mods:title id="566921581D5C01CAD8AD7884A59FEADD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0F0A93465FD0701268779CEB11142404">
<mods:date id="DB664F401905DC20106FA547A5A728AE">2010</mods:date>
<mods:detail id="21C61C91A48DF737639DADE1A77F0E25" type="pubDate">
<mods:number id="F09E3701EB4E649DB5B6C7283D51BAB9">2010-05-14</mods:number>
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<mods:number id="BB4A87043525FF21F5E82EE0648FA606">2472</mods:number>
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<mods:number id="84960C696B911793842FC9FEEB828E3F">1</mods:number>
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<mods:url id="E09B0400481300E3E855D956878EF45F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2472.1.1</mods:url>
</mods:location>
<mods:classification id="1E48FA41EE80B162B4A5AE0ACDCFD2B9">journal article</mods:classification>
<mods:identifier id="D6EA9146AC8D730A1D9899657EBC514F" type="DOI">10.11646/zootaxa.2472.1.1</mods:identifier>
<mods:identifier id="0AA5B110A729A963B8BBD4EDF317164B" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="A04EB064977A8D8E57427EE5EA9F736B" type="Zenodo-Dep">6507307</mods:identifier>
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<treatment id="03C0582D9903FFA4BB8BFC7F072D6AA0" ID-DOI="http://doi.org/10.5281/zenodo.6509010" ID-Zenodo-Dep="6509010" LSID="urn:lsid:plazi:treatment:03C0582D9903FFA4BB8BFC7F072D6AA0" httpUri="http://treatment.plazi.org/id/03C0582D9903FFA4BB8BFC7F072D6AA0" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C373BAB09903FFA6BB8BFC7F058B6B9E" box="[151,468,904,930]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BD6E93B9903FFA6BB8BFC7F058B6B9E" blockId="16.[151,468,904,965]" box="[151,468,904,930]" pageId="16" pageNumber="17">
<heading id="D09E5E579903FFA6BB8BFC7F058B6B9E" bold="true" box="[151,468,904,930]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B91D35299903FFA6BB8BFC7F058B6B9E" bold="true" box="[151,468,904,930]" pageId="16" pageNumber="17">
<taxonomicName id="4C6992B89903FFA6BB8BFC7F05D56B9E" ID-CoL="89X4P" authority="Andersen &amp; Saether &amp; Mendes, 2010" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" box="[151,394,904,930]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="quadrus" status="sp. nov.">
<emphasis id="B91D35299903FFA6BB8BFC7F05D56B9E" bold="true" box="[151,394,904,930]" italics="true" pageId="16" pageNumber="17">Allocladius quadrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E88529903FFA6BA8EFC7F058B6B9E" box="[402,468,904,930]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373BAB09903FFA6BB8BFC5C05726BF9" box="[151,301,939,965]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BD6E93B9903FFA6BB8BFC5C05726BF9" blockId="16.[151,468,904,965]" box="[151,301,939,965]" pageId="16" pageNumber="17">
(
<figureCitation id="1352F5BE9903FFA6BB83FC5C057A6BF9" box="[159,293,939,965]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Figs 38–44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373BAB09903FFA6BB8BFC02074E6CF6" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD6E93B9903FFA6BB8BFC02052B6C66" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BB8BFC0205086C33" bold="true" box="[151,343,1013,1039]" pageId="16" pageNumber="17">Type material.</emphasis>
<materialsCitation id="3B01E3669903FFA6BA75FC0200C46C09" collectingDate="1999-03-10" collectionCode="ZMBN" collectorName="Lago El Parrillar &amp; T. Andersen &amp; Type No." country="Chile" county="Lago El Parrillar" elevation="246" latitude="-53.4074" location="Region" longLatPrecision="1" longitude="-71.26372" municipality="Punta Arenas" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D257999903FFA6BA75FC0205896C33" box="[361,470,1013,1039]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
male,
<emphasis id="B91D35299903FFA6B926FC02071C6C33" bold="true" box="[570,835,1013,1039]" pageId="16" pageNumber="17">
<collectingCountry id="F37EA9AB9903FFA6B926FC0206D66C32" box="[570,649,1013,1039]" name="Chile" pageId="16" pageNumber="17">CHILE</collectingCountry>
:
<location id="8EB6BFE09903FFA6B9BAFC0206A16C33" LSID="urn:lsid:plazi:treatment:03C0582D9903FFA4BB8BFC7F072D6AA0:8EB6BFE09903FFA6B9BAFC0206A16C33" box="[678,766,1013,1039]" country="Chile" county="Lago El Parrillar" latitude="-53.4074" longLatPrecision="1" longitude="-71.26372" municipality="Punta Arenas" name="Region" pageId="16" pageNumber="17">Región</location>
XII,
</emphasis>
<collectingMunicipality id="6BB273419903FFA6B849FC02005F6C33" box="[853,1024,1013,1039]" pageId="16" pageNumber="17">Punta Arenas</collectingMunicipality>
,
<collectorName id="269C8CED9903FFA6BF0AFC0200AD6C33" box="[1046,1266,1013,1039]" pageId="16" pageNumber="17">
<collectingCounty id="62B791B79903FFA6BF0AFC0200AD6C33" box="[1046,1266,1013,1039]" pageId="16" pageNumber="17">Lago El Parrillar</collectingCounty>
</collectorName>
,
<geoCoordinate id="EE5D8FFC9903FFA6BE14FC0201C76C33" box="[1288,1432,1013,1039]" degrees="53" direction="south" minutes="24.444" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="-53.4074">53º24.444'S</geoCoordinate>
,
<geoCoordinate id="EE5D8FFC9903FFA6BB8BFBEC05706C09" box="[151,303,1051,1077]" degrees="71" direction="west" minutes="15.823" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="-71.26372">71º15.823'W</geoCoordinate>
,
<elevation id="00440E089903FFA6BA21FBEC05936C09" box="[317,460,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="16" pageNumber="17" unit="m" value="246.0">
<quantity id="4C9144DE9903FFA6BA21FBEC05D56C09" box="[317,394,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="16" pageNumber="17" unit="m" value="246.0">246 m</quantity>
a.s.l.
</elevation>
,
<date id="FFD7CFFB9903FFA6BAC0FBEC06026C09" box="[476,605,1051,1077]" pageId="16" pageNumber="17" value="1999-03-10">
<collectingDate id="EF9336139903FFA6BAC0FBEC06026C09" box="[476,605,1051,1077]" pageId="16" pageNumber="17" value="1999-03-10">10.iii.1999</collectingDate>
</date>
,
<collectingMethod id="5228912C9903FFA6B970FBEC06896C09" box="[620,726,1051,1077]" pageId="16" pageNumber="17">hand net</collectingMethod>
,
<collectorName id="269C8CED9903FFA6B9F8FBEC072A6C09" box="[740,885,1051,1077]" pageId="16" pageNumber="17">T. Andersen</collectorName>
(
<collectionCode id="ED7871FE9903FFA6B89BFBEC07816C09" box="[903,990,1051,1077]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="16" pageNumber="17">ZMBN</collectionCode>
<collectorName id="269C8CED9903FFA6B8F4FBEC00086C09" box="[1000,1111,1051,1077]" pageId="16" pageNumber="17">Type No.</collectorName>
424)
</materialsCitation>
.
<materialsCitation id="3B01E3669903FFA6BFB4FBEC052F6C66" collectionCode="ZMBN" country="Chile" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54D257999903FFA6BFB4FBEC014C6C09" box="[1192,1299,1051,1077]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D6F22B29903FFA6BE3CFBEC012A6C09" box="[1312,1397,1051,1077]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
, as
<typeStatus id="54D257999903FFA6BB8BFBB704A26C66" box="[151,253,1088,1114]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7871FE9903FFA6BA11FBB705376C66" box="[269,360,1088,1114]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="16" pageNumber="17">ZMBN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAFB92074E6CF6" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFB9205826C43" bold="true" box="[198,477,1125,1151]" pageId="16" pageNumber="17">Diagnostic characters.</emphasis>
The presence of a square single pars ventralis much wider than long and without setae together with the tongue-like, strongly projecting inferior volsella and the high antennal ratio will separate the species from other species with pars ventralis.
</paragraph>
</subSubSection>
<subSubSection id="C373BAB09903FFA7BBDAFB22051A697B" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BD6E93B9903FFA6BBDAFB2200F86CCC" blockId="16.[151,1436,1013,2013]" box="[198,1191,1237,1264]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFB22050C6CD3" bold="true" box="[198,339,1237,1263]" pageId="16" pageNumber="17">Etymology.</emphasis>
From Latin,
<emphasis id="B91D35299903FFA6BAF2FB21061D6CD3" box="[494,578,1238,1263]" italics="true" pageId="16" pageNumber="17">quadra</emphasis>
, a square, referring to the rectangular pars ventralis.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAFB0D06BD6D06" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFB0D055A6D28" bold="true" box="[198,261,1274,1300]" pageId="16" pageNumber="17">Male</emphasis>
(n = 2). Total length
<quantity id="4C9144DE9903FFA6B915FB0C06E76D29" box="[521,696,1275,1301]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" metricValueMax="2.67" metricValueMin="2.65" pageId="16" pageNumber="17" unit="mm" value="2.66" valueMax="2.67" valueMin="2.65">2.65–2.67 mm</quantity>
. Wing length
<quantity id="4C9144DE9903FFA6B87DFB0C004F6D29" box="[865,1040,1275,1301]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.73" metricValueMin="1.67" pageId="16" pageNumber="17" unit="mm" value="1.7" valueMax="1.73" valueMin="1.67">1.67–1.73 mm</quantity>
. Total length / wing length 1.53– 1.60. Wing length / length of profemur 3.05–3.16.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAFAB105B26D5C" blockId="16.[151,1436,1013,2013]" box="[198,493,1350,1376]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFAB105116D63" box="[198,334,1350,1375]" italics="true" pageId="16" pageNumber="17">Coloration.</emphasis>
Fully brown.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAFA9C064C6E26" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFA9C05516DB8" box="[198,270,1387,1412]" italics="true" pageId="16" pageNumber="17">Head.</emphasis>
AR 1.40–1.44. Ultimate flagellomere 463–491µm long. Temporal setae 6–7, consisting of 3–4 weak inner verticals, 2 strong outer verticals, and 1 postorbital. Clypeus with 9–12 setae. Tentorium, stipes, and cibarial pump as in
<figureCitation id="1352F5BE9903FFA6BAADFA4106796DEC" box="[433,550,1462,1488]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Figure 38</figureCitation>
. Tentorium 135–143 µm long, 34 µm wide. Stipes 150 µm long, 41–49 µm wide. Palpomere (
<figureCitation id="1352F5BE9903FFA6BA76FA2C05E06DC9" box="[362,447,1499,1525]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Fig. 39</figureCitation>
) lengths (in µm): 26–30, 45–53, 94–98, 143–146. Third palpomere with 2 lanceolate sensilla clavata, 19–21 µm long.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAF9D101376E59" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAF9D105446E03" box="[198,283,1574,1599]" italics="true" pageId="16" pageNumber="17">Thorax</emphasis>
(
<figureCitation id="1352F5BE9903FFA6BA32F9D105D56E7C" box="[302,394,1574,1600]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Fig. 40</figureCitation>
). Median antepronotal lobes comparatively well developed, antepronotum with 1 very weak lateral seta. Dorsocentrals 10–11, acrostichals 6, prealars 3–4, supraalar 1. Scutellum with 10 setae.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAF98706C26EE4" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAF987055D6EB5" box="[198,258,1648,1673]" italics="true" pageId="16" pageNumber="17">Wing</emphasis>
(
<figureCitation id="1352F5BE9903FFA6BA0FF98705346EB6" box="[275,363,1648,1674]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Fig. 41</figureCitation>
). VR 1.31–1.32. Anal lobe distinct, slightly projecting. Costal extension absent, but indication of 190–200 µm long false vein. R ending slightly proximal to apex of M
<subScript id="17EDEB7E9903FFA6BF53F95200386E8F" attach="left" box="[1103,1127,1701,1715]" fontSize="6" pageId="16" pageNumber="17">3+4</subScript>
, Cu
<subScript id="17EDEB7E9903FFA6BF87F95200FC6E8F" attach="left" box="[1179,1187,1701,1715]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
sinuous. Brachiolum with 1 seta, R with 2 setae, other veins bare.
</paragraph>
<paragraph id="8BD6E93B9903FFA6B978F95206226E8F" blockId="16.[151,1436,1013,2013]" box="[612,637,1701,1715]" pageId="16" pageNumber="17">
<subScript id="17EDEB7E9903FFA6B978F95206226E8F" attach="left" box="[612,637,1701,1715]" fontSize="6" pageId="16" pageNumber="17">4+5</subScript>
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAF91401716F74" blockId="16.[151,1436,1013,2013]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAF91404A26EC0" box="[198,253,1763,1788]" italics="true" pageId="16" pageNumber="17">Legs</emphasis>
. Spur of fore tibia 53 µm long, spurs of mid tibia 23–26 µm and 15 µm long, of hind tibia 45–49 µm and 23–26 µm long. Width at apex of fore- and mid tibia 30–36 µm, of hind tibia 45–49 µm. Comb of 14–15 setae, longest 38–45 µm long, shortest 23–30 µm long. Length and proportions of legs as in
<tableCitation id="C6EBDC809903FFA6BFCDF8D901746F74" box="[1233,1323,1838,1864]" captionStart="TABLE 6" captionStartId="18.[151,241,162,186]" captionText="TABLE 6. Length (in µm) and proportions of legs of Allocladius quadrus sp. n., male (n = 1–2)." pageId="16" pageNumber="17">Table 6</tableCitation>
.
</paragraph>
<paragraph id="8BD6E93B9903FFA7BBDAF8A4051A697B" blockId="16.[151,1436,1013,2013]" lastBlockId="17.[151,1436,152,327]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAF8A405036F50" box="[198,348,1875,1900]" italics="true" pageId="16" pageNumber="17">Hypopygium</emphasis>
(
<figureCitation id="1352F5BE9903FFA6BA71F8A405A76F51" box="[365,504,1875,1901]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Figs 42–43</figureCitation>
). Anal point 45–49 µm long, including 34 µm long basal sclerotized part with 10–14 setae and 11–15 µm long bare, hyaline distal part. Tergite IX with 8–9 setae in addition to those on anal point. Laterosternite IX with 4–6 setae. Phallapodeme 71–75 µm long. Transverse sternapodeme 45–60 µm long, with weak oral projections. Virga 8–11 µm long, 11 µm wide, thumbnail-like. Pars ventralis (
<figureCitation id="1352F5BE9903FFA6BE20F83401CB6FE1" box="[1340,1428,1987,2013]" captionStart="FIGURES 38–44" captionStartId="17.[151,269,1949,1973]" captionTargetBox="[230,1344,375,1923]" captionTargetId="figure-124@17.[226,1360,353,1941]" captionTargetPageId="17" captionText="FIGURES 38–44. Allocladius quadrus sp. n., male. 38—tentorium, stipes, and cibarial pump; 39—palp; 40—thorax; 41—wing; 42—hypopygium, dorsal aspect; 43—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 44—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) single, subquadrangular, 71–75 µm long, 101–113 µm wide,
<specimenCount id="9D6F22B29902FFA7B89AFF6F0075688E" box="[902,1066,152,178]" pageId="17" pageNumber="18" type="generic">one specimen</specimenCount>
with slight median concavity. Gonocoxite 184–206 µm long. Superior volsella barely indicated as dorsal swelling of inner margin of gonocoxite, reaching to 0.41–0.45 gonocoxite length. Inferior volsella well developed, reaching to 0.61–0.64 gonocoxite length; no apparent posterior lobe. Gonostylus 79–83 µm long; megaseta 9–11 µm long. HR 2.33–2.50. HV 3.19–3.39.
</paragraph>
</subSubSection>
<caption id="DF16B9B39902FFA7BB8BF86A06886FC9" ID-DOI="http://doi.org/10.5281/zenodo.6507321" ID-Zenodo-Dep="6507321" httpUri="https://zenodo.org/record/6507321/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,269,1949,1973]" targetBox="[230,1344,375,1923]" targetPageId="17">
<paragraph id="8BD6E93B9902FFA7BB8BF86A06886FC9" blockId="17.[151,1436,1949,2037]" pageId="17" pageNumber="18">
<emphasis id="B91D35299902FFA7BB8BF86A053F6F89" bold="true" box="[151,352,1949,1973]" pageId="17" pageNumber="18">FIGURES 38–44.</emphasis>
<taxonomicName id="4C6992B89902FFA7BA74F869061E6F89" authority="Andersen &amp; Saether &amp; Mendes, 2010" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" box="[360,577,1950,1973]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quadrus" status="sp. nov.">
<emphasis id="B91D35299902FFA7BA74F869061E6F89" box="[360,577,1950,1973]" italics="true" pageId="17" pageNumber="18">Allocladius quadrus</emphasis>
</taxonomicName>
<emphasis id="B91D35299902FFA7B956F86A06D06F89" bold="true" box="[586,655,1949,1973]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A22E88529902FFA7B956F86A06D66F89" box="[586,649,1949,1973]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
male.
<emphasis id="B91D35299902FFA7B9C7F86A06A96F89" bold="true" box="[731,758,1949,1973]" pageId="17" pageNumber="18">38</emphasis>
—tentorium, stipes, and cibarial pump;
<emphasis id="B91D35299902FFA7BFB4F86A009C6F89" bold="true" box="[1192,1219,1949,1973]" pageId="17" pageNumber="18">39</emphasis>
—palp;
<emphasis id="B91D35299902FFA7BE07F86A016A6F89" bold="true" box="[1307,1333,1949,1973]" pageId="17" pageNumber="18">40</emphasis>
—thorax;
<emphasis id="B91D35299902FFA7BB8BF84A04ED6FE9" bold="true" box="[151,178,1981,2005]" pageId="17" pageNumber="18">41</emphasis>
—wing;
<emphasis id="B91D35299902FFA7BA0DF84A05736FE9" bold="true" box="[273,300,1981,2005]" pageId="17" pageNumber="18">42</emphasis>
—hypopygium, dorsal aspect;
<emphasis id="B91D35299902FFA7B964F84A06CC6FE9" bold="true" box="[632,659,1981,2005]" pageId="17" pageNumber="18">43</emphasis>
—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right;
<emphasis id="B91D35299902FFA7B910F82A06786FC9" bold="true" box="[524,551,2013,2037]" pageId="17" pageNumber="18">44</emphasis>
—pars ventralis.
</paragraph>
</caption>
<caption id="DF16B9B39901FFA4BB8BFF5500FF6885" box="[151,1184,162,186]" pageId="18" pageNumber="19" startId="18.[151,241,162,186]">
<paragraph id="8BD6E93B9901FFA4BB8BFF5500FF6885" blockId="18.[151,1184,162,186]" box="[151,1184,162,186]" pageId="18" pageNumber="19">
<emphasis id="B91D35299901FFA4BB8BFF5505536886" bold="true" box="[151,268,162,186]" pageId="18" pageNumber="19">TABLE 6.</emphasis>
Length (in µm) and proportions of legs of
<taxonomicName id="4C6992B89901FFA4B9CBFF5507F16885" authority="Andersen &amp; Saether &amp; Mendes, 2010" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" box="[727,942,162,185]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrus" status="sp. nov.">
<emphasis id="B91D35299901FFA4B9CBFF5507F16885" box="[727,942,162,185]" italics="true" pageId="18" pageNumber="19">Allocladius quadrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E88529901FFA4B8A9FF5507B06885" box="[949,1007,162,185]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
, male (n = 1–2).
</paragraph>
</caption>
<paragraph id="8BD6E93B9901FFA4BA2CFF2F01736A0B" pageId="18" pageNumber="19">
<table id="F9691B9B99010049BBB9FF23011F6A04" box="[165,1344,212,568]" gridcols="7" gridrows="8" pageId="18" pageNumber="19">
<tr id="3559EB7999010049BBB9FF23011F68D3" box="[165,1344,212,239]" gridrow="0" pageId="18" pageNumber="19" rowspan-0="1">
<th id="7688820599010049BA2CFF2305D168D3" box="[304,398,212,239]" gridcol="1" gridrow="0" pageId="18" pageNumber="19">fe</th>
<th id="7688820599010049B91CFF2306D468D3" box="[512,651,212,239]" gridcol="2" gridrow="0" pageId="18" pageNumber="19">ti</th>
<th id="7688820599010049B9D2FF23071A68D3" box="[718,837,212,239]" gridcol="3" gridrow="0" pageId="18" pageNumber="19">ta1</th>
<th id="7688820599010049B881FF23079A68D3" box="[925,965,212,239]" gridcol="4" gridrow="0" pageId="18" pageNumber="19">ta2</th>
<th id="7688820599010049BF10FF2300F568D3" box="[1036,1194,212,239]" gridcol="5" gridrow="0" pageId="18" pageNumber="19">ta3</th>
<th id="7688820599010049BFC0FF23011F68D3" box="[1244,1344,212,239]" gridcol="6" gridrow="0" pageId="18" pageNumber="19">ta4</th>
</tr>
<tr id="3559EB7999010049BBB9FEF4011F6922" box="[165,1344,259,286]" gridrow="1" pageId="18" pageNumber="19">
<th id="7688820599010049BBB9FEF404E56922" box="[165,186,259,286]" gridcol="0" gridrow="1" pageId="18" pageNumber="19">p1</th>
<td id="7688820599010049BA2CFEF405D16922" box="[304,398,259,286]" gridcol="1" gridrow="1" pageId="18" pageNumber="19">548</td>
<td id="7688820599010049B91CFEF406D46922" box="[512,651,259,286]" gridcol="2" gridrow="1" pageId="18" pageNumber="19">699–718</td>
<td id="7688820599010049B9D2FEF4071A6922" box="[718,837,259,286]" gridcol="3" gridrow="1" pageId="18" pageNumber="19">298–312</td>
<td id="7688820599010049B881FEF4079A6922" box="[925,965,259,286]" gridcol="4" gridrow="1" pageId="18" pageNumber="19">198</td>
<td id="7688820599010049BF10FEF400F56922" box="[1036,1194,259,286]" gridcol="5" gridrow="1" pageId="18" pageNumber="19">132–137</td>
<td id="7688820599010049BFC0FEF4011F6922" box="[1244,1344,259,286]" gridcol="6" gridrow="1" pageId="18" pageNumber="19">85–90</td>
</tr>
<tr id="3559EB7999010049BBB9FEC6011F6971" box="[165,1344,305,333]" gridrow="2" pageId="18" pageNumber="19">
<th id="7688820599010049BBB9FEC604E56971" box="[165,186,305,333]" gridcol="0" gridrow="2" pageId="18" pageNumber="19">p2</th>
<td id="7688820599010049BA2CFEC605D16971" box="[304,398,305,333]" gridcol="1" gridrow="2" pageId="18" pageNumber="19">586–605</td>
<td id="7688820599010049B91CFEC606D46971" box="[512,651,305,333]" gridcol="2" gridrow="2" pageId="18" pageNumber="19">680–709</td>
<td id="7688820599010049B9D2FEC6071A6971" box="[718,837,305,333]" gridcol="3" gridrow="2" pageId="18" pageNumber="19">293</td>
<td id="7688820599010049B881FEC6079A6971" box="[925,965,305,333]" gridcol="4" gridrow="2" pageId="18" pageNumber="19">161</td>
<td id="7688820599010049BF10FEC600F56971" box="[1036,1194,305,333]" gridcol="5" gridrow="2" pageId="18" pageNumber="19">113–123</td>
<td id="7688820599010049BFC0FEC6011F6971" box="[1244,1344,305,333]" gridcol="6" gridrow="2" pageId="18" pageNumber="19">76–80</td>
</tr>
<tr id="3559EB7999010049BBB9FE97011F6940" box="[165,1344,352,380]" gridrow="3" pageId="18" pageNumber="19">
<th id="7688820599010049BBB9FE9704E56940" box="[165,186,352,380]" gridcol="0" gridrow="3" pageId="18" pageNumber="19">p3</th>
<td id="7688820599010049BA2CFE9705D16940" box="[304,398,352,380]" gridcol="1" gridrow="3" pageId="18" pageNumber="19">624–659</td>
<td id="7688820599010049B91CFE9706D46940" box="[512,651,352,380]" gridcol="2" gridrow="3" pageId="18" pageNumber="19">718–747</td>
<td id="7688820599010049B9D2FE97071A6940" box="[718,837,352,380]" gridcol="3" gridrow="3" pageId="18" pageNumber="19">369–378</td>
<td id="7688820599010049B881FE9700F56940" box="[925,1194,352,380]" colspan="2" colspanRight="1" gridcol="4" gridrow="3" pageId="18" pageNumber="19">189–194 161–170</td>
<td id="7688820599010049BFC0FE97011F6940" box="[1244,1344,352,380]" gridcol="6" gridrow="3" pageId="18" pageNumber="19">76</td>
</tr>
<tr id="3559EB7999010049BBB9FE78011F6997" box="[165,1344,399,427]" gridrow="4" pageId="18" pageNumber="19" rowspan-0="1" rowspan-4="1">
<td id="7688820599010049BA2CFE7805D16997" box="[304,398,399,427]" gridcol="1" gridrow="4" pageId="18" pageNumber="19">ta5</td>
<td id="7688820599010049B91CFE7806D46997" box="[512,651,399,427]" gridcol="2" gridrow="4" pageId="18" pageNumber="19">LR</td>
<td id="7688820599010049B9D2FE78071A6997" box="[718,837,399,427]" gridcol="3" gridrow="4" pageId="18" pageNumber="19">BV</td>
<td id="7688820599010049BF10FE7800F56997" box="[1036,1194,399,427]" gridcol="5" gridrow="4" pageId="18" pageNumber="19">SV</td>
<td id="7688820599010049BFC0FE78011F6997" box="[1244,1344,399,427]" gridcol="6" gridrow="4" pageId="18" pageNumber="19">BR</td>
</tr>
<tr id="3559EB7999010049BBB9FE49011F69E6" box="[165,1344,446,474]" gridrow="5" pageId="18" pageNumber="19">
<th id="7688820599010049BBB9FE4904E569E6" box="[165,186,446,474]" gridcol="0" gridrow="5" pageId="18" pageNumber="19">p1</th>
<td id="7688820599010049BA2CFE4905D169E6" box="[304,398,446,474]" gridcol="1" gridrow="5" pageId="18" pageNumber="19">61–66</td>
<td id="7688820599010049B91CFE4906D469E6" box="[512,651,446,474]" gridcol="2" gridrow="5" pageId="18" pageNumber="19">0.43</td>
<td id="7688820599010049B9D2FE49079A69E6" box="[718,965,446,474]" colspan="2" colspanRight="1" gridcol="3" gridrow="5" pageId="18" pageNumber="19">3.21–3.24</td>
<td id="7688820599010049BF10FE4900F569E6" box="[1036,1194,446,474]" gridcol="5" gridrow="5" pageId="18" pageNumber="19">4.06–4.19</td>
<td id="7688820599010049BFC0FE49011F69E6" box="[1244,1344,446,474]" gridcol="6" gridrow="5" pageId="18" pageNumber="19">3.0–3.3</td>
</tr>
<tr id="3559EB7999010049BBB9FE1A011F6A35" box="[165,1344,493,521]" gridrow="6" pageId="18" pageNumber="19">
<th id="7688820599010049BBB9FE1A04E56A35" box="[165,186,493,521]" gridcol="0" gridrow="6" pageId="18" pageNumber="19">p2</th>
<td id="7688820599010049BA2CFE1A05D16A35" box="[304,398,493,521]" gridcol="1" gridrow="6" pageId="18" pageNumber="19">57–66</td>
<td id="7688820599010049B91CFE1A06D46A35" box="[512,651,493,521]" gridcol="2" gridrow="6" pageId="18" pageNumber="19">0.41–0.43</td>
<td id="7688820599010049B9D2FE1A079A6A35" box="[718,965,493,521]" colspan="2" colspanRight="1" gridcol="3" gridrow="6" pageId="18" pageNumber="19">3.71–3.86</td>
<td id="7688820599010049BF10FE1A00F56A35" box="[1036,1194,493,521]" gridcol="5" gridrow="6" pageId="18" pageNumber="19">4.32–4.48</td>
<td id="7688820599010049BFC0FE1A011F6A35" box="[1244,1344,493,521]" gridcol="6" gridrow="6" pageId="18" pageNumber="19">2.8–3.6</td>
</tr>
<tr id="3559EB7999010049BBB9FDEB011F6A04" box="[165,1344,540,568]" gridrow="7" pageId="18" pageNumber="19">
<th id="7688820599010049BBB9FDEB04E56A04" box="[165,186,540,568]" gridcol="0" gridrow="7" pageId="18" pageNumber="19">p3</th>
<td id="7688820599010049BA2CFDEB05D16A04" box="[304,398,540,568]" gridcol="1" gridrow="7" pageId="18" pageNumber="19">66</td>
<td id="7688820599010049B91CFDEB06D46A04" box="[512,651,540,568]" gridcol="2" gridrow="7" pageId="18" pageNumber="19">0.51</td>
<td id="7688820599010049B9D2FDEB079A6A04" box="[718,965,540,568]" colspan="2" colspanRight="1" gridcol="3" gridrow="7" pageId="18" pageNumber="19">3.45–3.56</td>
<td id="7688820599010049BF10FDEB00F56A04" box="[1036,1194,540,568]" gridcol="5" gridrow="7" pageId="18" pageNumber="19">3.64–3.71</td>
<td id="7688820599010049BFC0FDEB011F6A04" box="[1244,1344,540,568]" gridcol="6" gridrow="7" pageId="18" pageNumber="19">6.0–7.0</td>
</tr>
</table>
</paragraph>
<subSubSection id="C373BAB09901FFA4BBDAFDAA072D6AA0" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BD6E93B9901FFA4BBDAFDAA072D6AA0" blockId="18.[151,1435,605,668]" pageId="18" pageNumber="19">
<emphasis id="B91D35299901FFA4BBDAFDAA05A06A4B" bold="true" box="[198,511,605,631]" pageId="18" pageNumber="19">Distribution and biology.</emphasis>
The species is known from southernmost
<collectingCountry id="F37EA9AB9901FFA4B8E9FDAA00686A4B" box="[1013,1079,605,631]" name="Chile" pageId="18" pageNumber="19">Chile</collectingCountry>
, where it was netted on a lake shore at about
<quantity id="4C9144DE9901FFA4BA5EFD7505D26AA0" box="[322,397,642,668]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="m" value="250.0">250 m</quantity>
altitude in an area with
<emphasis id="B91D35299901FFA4B9B4FD74077F6AA0" box="[680,800,643,668]" italics="true" pageId="18" pageNumber="19">Notofagus</emphasis>
forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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