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<document id="BAED1237F5F5739621D456ACFAC68A37" ID-DOI="10.37520/aemnp.2020.036" ID-ISSN="1804-6487" ID-Zenodo-Dep="5177635" ID-ZooBank="269CC302-9827-479D-8C09-76757F09AD95" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628618413284" checkinUser="carolina" docAuthor="Yin, Zi-Wei, Hlaváč, Peter &amp; Cuccodoro, Giulio" docDate="2020" docId="03BE384E6D6EFF9EFC00FC775AA69774" docLanguage="en" docName="ActaEntMusNatPrag.60.2.537-544.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 60 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2020.036" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Archiclaviger gaofani Yin &amp; Hlaváč &amp; Cuccodoro 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="543" masterDocId="FF8740366D6CFF98FFC6FF9E5832920F" masterDocTitle="The first record of Archiclaviger in continental Asia, with description of a new species from China (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="544" masterPageNumber="537" pageNumber="539" updateTime="1699047778035" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="52DF50DC16CA484D28A56B459F081D69">The first record of Archiclaviger in continental Asia, with description of a new species from China (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="E1A10F408F30DC11577329F8FD37E3BF">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="366849C460921FB9D776DC74FD6BC115">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China; e-mail: pselaphinae @ gmail. com; https: // orcid. org / 0000 - 0001 - 6659 - 9448</mods:affiliation>
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<mods:namePart id="58A366C293599A8170EC0EA6DECBD289">Hlaváč, Peter</mods:namePart>
<mods:affiliation id="6182E1AE5FA156D411968839D091F557">Department of Entomology, National Museum, Natural History Museum, Cirkusová 1740, CZ- 193 00 Praha</mods:affiliation>
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<mods:namePart id="5ECC253569628A90CA145CE433DE77B0">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="5C232C36E522EEF13AA18C4EAE482147">Muséum dhistoire naturelle, Case postale 6434 CH- 1211 Genève</mods:affiliation>
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<mods:title id="A5EFFDC2DEF74EECB08BBA687BC37CDB">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="B7E3389A04F569DE49AC2385FC9C1BA9">2020</mods:date>
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<mods:number id="84DA1246BA3F3868F8B37A22A0B829B5">2020-10-07</mods:number>
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<subSubSection id="C30DDAD36D6EFF9AFC00FC775D21960E" box="[966,1299,1001,1025]" pageId="2" pageNumber="539" type="nomenclature">
<paragraph id="8BA889586D6EFF9AFC00FC775D21960E" blockId="2.[966,1299,1001,1025]" box="[966,1299,1001,1025]" pageId="2" pageNumber="539">
<heading id="D0E03E346D6EFF9AFC00FC775D21960E" box="[966,1299,1001,1025]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="539" reason="3">
<emphasis id="B963554A6D6EFF9AFC00FC775D21960E" bold="true" box="[966,1299,1001,1025]" pageId="2" pageNumber="539">
<taxonomicName id="4C17F2DB6D6EFF9AFC00FC775C84960E" ID-CoL="8XV2K" authority="Yin &amp; Hlaváč &amp; Cuccodoro, 2020" authorityName="Yin &amp; Hlaváč &amp; Cuccodoro" authorityYear="2020" box="[966,1206,1001,1025]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="539" phylum="Arthropoda" rank="species" species="gaofani" status="sp. nov.">
<emphasis id="B963554A6D6EFF9AFC00FC775C84960E" bold="true" box="[966,1206,1001,1025]" italics="true" pageId="2" pageNumber="539">Archiclaviger gaofani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A250E8316D6EFF9AFB7BFC745D21960E" box="[1213,1299,1002,1025]" pageId="2" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30DDAD36D6EFF9AFBF3FB975D5E96EA" pageId="2" pageNumber="539" type="description">
<paragraph id="8BA889586D6EFF9AFBF3FB975C919613" blockId="2.[828,1437,1033,1253]" box="[1077,1187,1033,1052]" pageId="2" pageNumber="539">
(
<figureCitation id="132C95DD6D6EFF9AFBFDFB975CBA9613" box="[1083,1160,1033,1052]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[151,182,1757,1776]" captionStartId-1="3.[150,181,1747,1766]" captionStartId-2="4.[151,182,1753,1772]" captionTargetBox-0="[176,1406,193,1739]" captionTargetBox-1="[242,1343,189,1735]" captionTargetBox-2="[220,1367,189,1730]" captionTargetId-0="figure-15@1.[166,1419,174,1746]" captionTargetId-1="figure-15@3.[242,1344,189,1735]" captionTargetId-2="figure-13@4.[220,1367,189,1735]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Archiclaviger gaofani sp. nov. AC habitus (A dorsal, male; B lateral, male; C dorsal, female); D frontoventral view of antennal bases, showing antennal fossae and scapes.Abbreviations: a13 = antennomere 13. Scale bars: 0.5 mm in AC; 0.2 mm in D." captionText-1="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." captionText-2="Fig. 3. Morphology of Archiclaviger gaofani sp. nov. A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E." figureDoi-0="http://doi.org/10.5281/zenodo.5177637" figureDoi-1="http://doi.org/10.5281/zenodo.5177639" figureDoi-2="http://doi.org/10.5281/zenodo.5177641" httpUri-0="https://zenodo.org/record/5177637/files/figure.png" httpUri-1="https://zenodo.org/record/5177639/files/figure.png" httpUri-2="https://zenodo.org/record/5177641/files/figure.png" pageId="2" pageNumber="539">Figs 13</figureCitation>
;
<figureCitation id="132C95DD6D6EFF9AFB57FB975CAE9613" box="[1169,1180,1033,1052]" captionStart="Fig" captionStartId="6.[151,182,1017,1036]" captionTargetBox="[151,1436,190,999]" captionTargetId="figure-13@6.[151,1436,190,1001]" captionTargetPageId="6" captionText="Fig. 5. Collection site, distribution and host ant of Archiclaviger. A environment of the collection site of A. gaofani sp. nov.; B the collection site of A. gaofani sp. nov. is located in a highly industrial area of Jintan District, Changzhou; C distribution of Archiclaviger species in Asia; D a living adult of A. gaofani sp. nov. was discovered in a nest of Nylanderia ants after a concrete block had been turned over (photo by F. Gao)." figureDoi="http://doi.org/10.5281/zenodo.5177645" httpUri="https://zenodo.org/record/5177645/files/figure.png" pageId="2" pageNumber="539">5</figureCitation>
)
</paragraph>
<paragraph id="8BA889586D6EFF9AFC71FBBD5D12963B" blockId="2.[828,1437,1033,1253]" box="[951,1312,1055,1078]" pageId="2" pageNumber="539">
Chinese common name:
<emphasis id="B963554A6D6EFF9AFB4DFB815D12963B" bold="true" box="[1163,1312,1055,1076]" pageId="2" pageNumber="539">Ŕ氏Ŝ寡Ṫȃm</emphasis>
</paragraph>
<paragraph id="8BA889586D6EFF9AFCFAFBD35D5E96EA" blockId="2.[828,1437,1033,1253]" pageId="2" pageNumber="539">
<emphasis id="B963554A6D6EFF9AFCFAFBD35B8E966F" bold="true" box="[828,956,1101,1120]" pageId="2" pageNumber="539">Type material.</emphasis>
Hඈඅඈඍඒඉൾ:
<emphasis id="B963554A6D6EFF9AFBE6FBD35C43966F" bold="true" box="[1056,1137,1101,1120]" pageId="2" pageNumber="539">
<collectingCountry id="F300C9C86D6EFF9AFBE6FBD35C5E966F" box="[1056,1132,1101,1120]" name="China" pageId="2" pageNumber="539">CHINA</collectingCountry>
:
</emphasis>
J,
<collectingCountry id="F300C9C86D6EFF9AFB56FBD35CFA966F" box="[1168,1224,1101,1120]" name="China" pageId="2" pageNumber="539">China</collectingCountry>
:
<collectingRegion id="49D347BA6D6EFF9AFB08FBD35D3C966F" box="[1230,1294,1101,1120]" country="China" name="Jiangsu" pageId="2" pageNumber="539">Jiangsu</collectingRegion>
,Changzhou, Jintan Dist., nr. cross of S240/S340 HWY,
<geoCoordinate id="EE23EF9F6D6EFF9AFB4AFBF65D3E9674" box="[1164,1292,1128,1147]" degrees="31" direction="north" minutes="45" orientation="latitude" pageId="2" pageNumber="539" precision="1" seconds="45.64" value="31.762678">31º4545.64″N</geoCoordinate>
,
<geoCoordinate id="EE23EF9F6D6EFF9AFAD2FBF65DAF9674" box="[1300,1437,1128,1147]" degrees="119" direction="north" minutes="36" orientation="latitude" pageId="2" pageNumber="539" precision="1" seconds="38.40" value="119.610664">119º3638.40″N</geoCoordinate>
E,
<quantity id="4CEF24BD6D6EFF9AFC93FB1C5B48969A" box="[853,890,1154,1173]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="539" unit="m" value="2.0">2 m</quantity>
,
<date id="FFA9AF986D6EFF9AFC42FB1C5BD0969A" box="[900,994,1154,1173]" pageId="2" pageNumber="539" value="2020-03-18">18.iii.2020</date>
, F. Gao leg.,
<emphasis id="B963554A6D6EFF9AFB9DFBE05C82969C" bold="true" box="[1115,1200,1150,1171]" pageId="2" pageNumber="539">常州ȑ坛</emphasis>
S240/S340
<emphasis id="B963554A6D6EFF9AFAC8FBE05D51969C" bold="true" box="[1294,1379,1150,1171]" pageId="2" pageNumber="539">交叉ñ口</emphasis>
(
<specimenCount id="9D1142D16D6EFF9AFAB1FB1C5BB396BF" pageId="2" pageNumber="539" type="worker">four workers</specimenCount>
of the host ant are pinned under the specimen) (SNUC). Pൺඋൺ- ඍඒඉൾ:
<emphasis id="B963554A6D6EFF9AFCB6FB265BF096C4" bold="true" box="[880,962,1208,1227]" pageId="2" pageNumber="539">
<collectingCountry id="F300C9C86D6EFF9AFCB6FB265B8C96C4" box="[880,958,1208,1227]" name="China" pageId="2" pageNumber="539">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D1142D16D6EFF9AFC01FB265BD596C5" box="[967,999,1201,1227]" pageId="2" pageNumber="539" type="female">1 ♀</specimenCount>
, same collection data as
<typeStatus id="54AC37FA6D6EFF9AFB79FB265D3896C4" box="[1215,1290,1208,1227]" pageId="2" pageNumber="539" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D1142D16D6EFF9AFAD3FB265D4596C4" box="[1301,1399,1208,1227]" pageId="2" pageNumber="539" type="worker">one worker</specimenCount>
and
<specimenCount id="9D1142D16D6EFF9AFCFAFB4C5BBB96EA" box="[828,905,1234,1253]" pageId="2" pageNumber="539" type="male">one male</specimenCount>
of the host ant are pinned under the specimen) (SNUC).
</paragraph>
</subSubSection>
<subSubSection id="C30DDAD36D6EFF9DFCFAFB60596E95AA" lastPageId="5" lastPageNumber="542" pageId="2" pageNumber="539" type="diagnosis">
<paragraph id="8BA889586D6EFF9DFCFAFB60596E95AA" blockId="2.[828,1437,1278,2069]" lastBlockId="5.[151,760,1838,2085]" lastPageId="5" lastPageNumber="542" pageId="2" pageNumber="539">
<emphasis id="B963554A6D6EFF9AFCFAFB605C7F971A" bold="true" box="[828,1101,1278,1301]" pageId="2" pageNumber="539">Comparative diagnosis.</emphasis>
The female of the new species can be readily separated from
<taxonomicName id="4C17F2DB6D6EFF9AFB51FA805D10973A" authorityName="Heller, Cl. Besuchet" authorityYear="1936" box="[1175,1314,1310,1333]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="539" phylum="Arthropoda" rank="species" species="overbecki">
<emphasis id="B963554A6D6EFF9AFB51FA805C95973A" box="[1175,1191,1310,1333]" italics="true" pageId="2" pageNumber="539">A</emphasis>
.
<emphasis id="B963554A6D6EFF9AFB7EFA805D10973A" box="[1208,1314,1310,1333]" italics="true" pageId="2" pageNumber="539">overbecki</emphasis>
</taxonomicName>
from
<collectingRegion id="49D347BA6D6EFF9AFAAAFA805DAE973A" box="[1388,1436,1310,1333]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="539">Java</collectingRegion>
(known from a single female) by the relatively much shorter antennomeres 45, the presence of a pair of admesal carinae on the gular plate of the head (lacking in
<taxonomicName id="4C17F2DB6D6EFF9AFA43FAE05B9797BA" authorityName="Heller, Cl. Besuchet" authorityYear="1936" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="539" phylum="Arthropoda" rank="species" species="overbecki">
<emphasis id="B963554A6D6EFF9AFA43FAE05DA7979A" box="[1413,1429,1406,1429]" italics="true" pageId="2" pageNumber="539">A</emphasis>
.
<emphasis id="B963554A6D6EFF9AFCFAFA005B9797BA" box="[828,933,1438,1461]" italics="true" pageId="2" pageNumber="539">overbecki</emphasis>
</taxonomicName>
), the posterior margin of the elytra and the base of the abdomen with only few setae (with row of dense, long golden setae in
<taxonomicName id="4C17F2DB6D6EFF9AFBDCFA405C9397FA" authorityName="Heller, Cl. Besuchet" authorityYear="1936" box="[1050,1185,1502,1525]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="539" phylum="Arthropoda" rank="species" species="overbecki">
<emphasis id="B963554A6D6EFF9AFBDCFA405C1897FA" box="[1050,1066,1502,1525]" italics="true" pageId="2" pageNumber="539">A</emphasis>
.
<emphasis id="B963554A6D6EFF9AFBF1FA405C9397FA" box="[1079,1185,1502,1525]" italics="true" pageId="2" pageNumber="539">overbecki</emphasis>
</taxonomicName>
), the more convex disc of the abdomen with fewer long, thick setae, and the relatively longer sternite 2 (IV) in relation to sternite 3 (V).
<emphasis id="B963554A6D6EFF9AFCFAF9A05BFB945A" bold="true" box="[828,969,1598,1621]" pageId="2" pageNumber="539">Description.</emphasis>
[
<figureCitation id="132C95DD6D6EFF9AFC13F9A05C2B945A" box="[981,1049,1598,1621]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="2" pageNumber="539">Figs 2</figureCitation>
and
<figureCitation id="132C95DD6D6EFF9AFB8DF9A05C59945A" box="[1099,1131,1598,1621]" captionStart="Fig" captionStartId="4.[151,182,1753,1772]" captionTargetBox="[220,1367,189,1730]" captionTargetId="figure-13@4.[220,1367,189,1735]" captionTargetPageId="4" captionText="Fig. 3. Morphology of Archiclaviger gaofani sp. nov. A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E." figureDoi="http://doi.org/10.5281/zenodo.5177641" httpUri="https://zenodo.org/record/5177641/files/figure.png" pageId="2" pageNumber="539">3B</figureCitation>
are based on a female specimen, but the morphology is the same in both sexes.]
<emphasis id="B963554A6D6EFF9AFAA2F9C35DAF947A" bold="true" box="[1380,1437,1629,1653]" italics="true" pageId="2" pageNumber="539">Male</emphasis>
(
<figureCitation id="132C95DD6D6EFF9AFC85F9E05B8D949A" box="[835,959,1662,1685]" captionStart="Fig" captionStartId="1.[151,182,1757,1776]" captionTargetBox="[176,1406,193,1739]" captionTargetId="figure-15@1.[166,1419,174,1746]" captionTargetPageId="1" captionText="Fig. 1. Archiclaviger gaofani sp. nov. AC habitus (A dorsal, male; B lateral, male; C dorsal, female); D frontoventral view of antennal bases, showing antennal fossae and scapes.Abbreviations: a13 = antennomere 13. Scale bars: 0.5 mm in AC; 0.2 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5177637" httpUri="https://zenodo.org/record/5177637/files/figure.png" pageId="2" pageNumber="539">Figs 1A, B</figureCitation>
). Body length (from anterior margin of clypeus to apex of abdomen)
<quantity id="4CEF24BD6D6EFF9AFB96F9005C8394BA" box="[1104,1201,1694,1717]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="2" pageNumber="539" unit="mm" value="2.27">2.27 mm</quantity>
; reddish-brown. Head (
<figureCitation id="132C95DD6D6EFF9AFC85F9205BD294DA" box="[835,992,1726,1749]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="2" pageNumber="539">Figs 2BD, C</figureCitation>
) longer than wide, length from anterior margin of clypeus to head base along midline (excluding neck region)
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, width across eyes
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; anteri- or margin of clypeus protuberant and slightly arcuate, gular plate (
<figureCitation id="132C95DD6D6EFF9AFC66F8A05BC0955A" box="[928,1010,1854,1877]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="2" pageNumber="539">Fig. 2C</figureCitation>
) with pair of admesal carinae extending from posterior tentorial pits (gular foveae) to mouthpart; each eye composed of approximately 30 facets. Antenna (
<figureCitation id="132C95DD6D6EFF9AFC82F8005BA595BA" box="[836,919,1950,1973]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="2" pageNumber="539">Fig. 2A</figureCitation>
) clubbed, length of exposed part
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; scape and pedicel short, antennomere 3 elongate, slightly and evenly broadening distally, antennomere 4 shorter and slightly broader than 3, elongate, antennomere 5 shorter and at apex slightly broader than 4, broadening from base toward apex, antennomere 6 much broader than 5, shorter than 4 and 5 combined, with truncate and densely setose apex. Pronotum (
<figureCitation id="132C95DD6D6FFF9BFE8AF83659AF95B0" box="[332,413,1960,1983]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="3" pageNumber="540">Fig. 2B</figureCitation>
) subglobose, slightly transverse, length along midline
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, maximum width
<quantity id="4CEF24BD6D6FFF9BFD55F8565AC195D0" box="[659,755,1992,2015]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="540" unit="mm" value="0.41">0.41 mm</quantity>
, with broad median longitudinal impression. Prosternum (
<figureCitation id="132C95DD6D6FFF9BFF59F79658C79A10" box="[159,245,2056,2079]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="3" pageNumber="540">Fig. 2C</figureCitation>
) with weak median carina, anterior margin of procoxal cavities slightly carinate. Elytra (
<figureCitation id="132C95DD6D6FFF9BFAD1F8D65DA59550" box="[1303,1431,1864,1887]" captionStart="Fig" captionStartId="1.[151,182,1757,1776]" captionTargetBox="[176,1406,193,1739]" captionTargetId="figure-15@1.[166,1419,174,1746]" captionTargetPageId="1" captionText="Fig. 1. Archiclaviger gaofani sp. nov. AC habitus (A dorsal, male; B lateral, male; C dorsal, female); D frontoventral view of antennal bases, showing antennal fossae and scapes.Abbreviations: a13 = antennomere 13. Scale bars: 0.5 mm in AC; 0.2 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5177637" httpUri="https://zenodo.org/record/5177637/files/figure.png" pageId="3" pageNumber="540">Figs 1A, B</figureCitation>
;
<figureCitation id="132C95DD6D6FFF9BFCFAF8F65B6F9570" box="[828,861,1896,1919]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="3" pageNumber="540">2D</figureCitation>
) roundly trapezoidal, much wider than long, length along suture
<quantity id="4CEF24BD6D6FFF9BFC02F8165C169590" box="[964,1060,1928,1951]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="540" unit="mm" value="0.61">0.61 mm</quantity>
, maximum width
<quantity id="4CEF24BD6D6FFF9BFB24F8165D769590" box="[1250,1348,1928,1951]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="3" pageNumber="540" unit="mm" value="0.92">0.92 mm</quantity>
; lacking trichomes at posterior margins. Mesoventrite (
<figureCitation id="132C95DD6D6FFF9BFA86F8365DA795B0" box="[1344,1429,1960,1983]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="3" pageNumber="540">Fig. 2E</figureCitation>
) with golden setae along midline, with short carinae at base; metaventrite (
<figureCitation id="132C95DD6D6FFF9BFBD0F8765C5995F0" box="[1046,1131,2024,2047]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="3" pageNumber="540">Fig. 2E</figureCitation>
) medially strongly convex, with scattered long, golden setae, with microsculpture in area posterior to coxal cavities. Legs short; mesofemur (
<figureCitation id="132C95DD6D68FF9CFF59F8F458C4958E" box="[159,246,1898,1921]" captionStart="Fig" captionStartId="4.[151,182,1753,1772]" captionTargetBox="[220,1367,189,1730]" captionTargetId="figure-13@4.[220,1367,189,1735]" captionTargetPageId="4" captionText="Fig. 3. Morphology of Archiclaviger gaofani sp. nov. A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E." figureDoi="http://doi.org/10.5281/zenodo.5177641" httpUri="https://zenodo.org/record/5177641/files/figure.png" pageId="4" pageNumber="541">Fig. 3A</figureCitation>
) with large, curved ventral projection at base, apex of projection bifid; mesotibia (
<figureCitation id="132C95DD6D68FF9CFDE5F8145A4895AE" box="[547,634,1930,1953]" captionStart="Fig" captionStartId="4.[151,182,1753,1772]" captionTargetBox="[220,1367,189,1730]" captionTargetId="figure-13@4.[220,1367,189,1735]" captionTargetPageId="4" captionText="Fig. 3. Morphology of Archiclaviger gaofani sp. nov. A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E." figureDoi="http://doi.org/10.5281/zenodo.5177641" httpUri="https://zenodo.org/record/5177641/files/figure.png" pageId="4" pageNumber="541">Fig. 3A</figureCitation>
) with large triangular preapical spine in apical fourth.Abdomen (
<figureCitation id="132C95DD6D68FF9CFD0AF834588A95EE" captionStart="Fig" captionStartId="1.[151,182,1757,1776]" captionTargetBox="[176,1406,193,1739]" captionTargetId="figure-15@1.[166,1419,174,1746]" captionTargetPageId="1" captionText="Fig. 1. Archiclaviger gaofani sp. nov. AC habitus (A dorsal, male; B lateral, male; C dorsal, female); D frontoventral view of antennal bases, showing antennal fossae and scapes.Abbreviations: a13 = antennomere 13. Scale bars: 0.5 mm in AC; 0.2 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5177637" httpUri="https://zenodo.org/record/5177637/files/figure.png" pageId="4" pageNumber="541">Fig. 1A</figureCitation>
) rounded, approximately as long as wide, and slightly narrower than elytra, length along midline
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, maximum width
<quantity id="4CEF24BD6D68FF9CFEF5F79459AA9A2E" box="[307,408,2058,2081]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="541" unit="mm" value="0.86">0.86 mm</quantity>
, disc (
<figureCitation id="132C95DD6D68FF9CFE27F7945A079A2E" box="[481,565,2058,2081]" captionStart="Fig" captionStartId="1.[151,182,1757,1776]" captionTargetBox="[176,1406,193,1739]" captionTargetId="figure-15@1.[166,1419,174,1746]" captionTargetPageId="1" captionText="Fig. 1. Archiclaviger gaofani sp. nov. AC habitus (A dorsal, male; B lateral, male; C dorsal, female); D frontoventral view of antennal bases, showing antennal fossae and scapes.Abbreviations: a13 = antennomere 13. Scale bars: 0.5 mm in AC; 0.2 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5177637" httpUri="https://zenodo.org/record/5177637/files/figure.png" pageId="4" pageNumber="541">Fig. 1B</figureCitation>
) strongly convex dorsally; composite tergite (
<figureCitation id="132C95DD6D68FF9CFBADF8D45C8D956E" box="[1131,1215,1866,1889]" captionStart="Fig" captionStartId="3.[150,181,1747,1766]" captionTargetBox="[242,1343,189,1735]" captionTargetId="figure-15@3.[242,1344,189,1735]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Archiclaviger gaofani sp. nov. A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E." figureDoi="http://doi.org/10.5281/zenodo.5177639" httpUri="https://zenodo.org/record/5177639/files/figure.png" pageId="4" pageNumber="541">Fig. 2D</figureCitation>
) broadly and deeply concave at base, lacking basolateral trichomes; both paratergites of first pair bearing group of linear trichome. Sternite 1 (III) and basal margin of sternite 2 (IV) (
<figureCitation id="132C95DD6D68FF9CFAB6F8345B6E95EE" captionStart="Fig" captionStartId="4.[151,182,1753,1772]" captionTargetBox="[220,1367,189,1730]" captionTargetId="figure-13@4.[220,1367,189,1735]" captionTargetPageId="4" captionText="Fig. 3. Morphology of Archiclaviger gaofani sp. nov. A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E." figureDoi="http://doi.org/10.5281/zenodo.5177641" httpUri="https://zenodo.org/record/5177641/files/figure.png" pageId="4" pageNumber="541">Fig. 3B</figureCitation>
) with rhombic microsculpture, sternite 2 (IV) along middle 1.35 times as long as sternite 3 (V), sternites 3 (V) and 4 (VI) along middle each approximately twice as long as their next segment. Aedeagus (
<figureCitation id="132C95DD6D69FF9DFDD5F8B05ABF954A" box="[531,653,1838,1861]" captionStart="Fig" captionStartId="4.[151,182,1753,1772]" captionTargetBox="[220,1367,189,1730]" captionTargetId="figure-13@4.[220,1367,189,1735]" captionTargetPageId="4" captionText="Fig. 3. Morphology of Archiclaviger gaofani sp. nov. A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E." figureDoi="http://doi.org/10.5281/zenodo.5177641" httpUri="https://zenodo.org/record/5177641/files/figure.png" pageId="5" pageNumber="542">Figs 3CE</figureCitation>
) strongly sclerotized, symmetrical, length
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; median lobe undivided, apex pointed; endophallus composed of two elongate sclerites.
</paragraph>
</subSubSection>
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<paragraph id="8BA889586D6FFF9BFF50F94D5CC0950E" blockId="3.[150,1435,1747,1793]" pageId="3" pageNumber="540">
Fig. 2. Morphology of
<taxonomicName id="4C17F2DB6D6FFF9BFE9CF94D5A2794E9" authority="Yin &amp; Hlaváč &amp; Cuccodoro, 2020" authorityName="Yin &amp; Hlaváč &amp; Cuccodoro" authorityYear="2020" box="[346,533,1747,1766]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="gaofani" status="sp. nov.">
<emphasis id="B963554A6D6FFF9BFE9CF94D5A2794E9" box="[346,533,1747,1766]" italics="true" pageId="3" pageNumber="540">Archiclaviger gaofani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A250E8316D6FFF9BFDDDF94D5A6C94E9" box="[539,606,1747,1766]" pageId="3" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
A antenna; B head and pronotum; C head venter and prosternum; D elytral apex and base of abdomen; E meso- and metaventrite. Abbreviations: a26 = antennomere 26. Scale bars: 0.2 mm in A, D; 0.3 mm in B, C, E.
</paragraph>
</caption>
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<paragraph id="8BA889586D68FF9CFF51F9475A199509" blockId="4.[151,1437,1753,1798]" pageId="4" pageNumber="541">
Fig. 3. Morphology of
<taxonomicName id="4C17F2DB6D68FF9CFE9DF9475A2794E3" authority="Yin &amp; Hlaváč &amp; Cuccodoro, 2020" authorityName="Yin &amp; Hlaváč &amp; Cuccodoro" authorityYear="2020" box="[347,533,1753,1772]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="541" phylum="Arthropoda" rank="species" species="gaofani" status="sp. nov.">
<emphasis id="B963554A6D68FF9CFE9DF9475A2794E3" box="[347,533,1753,1772]" italics="true" pageId="4" pageNumber="541">Archiclaviger gaofani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A250E8316D68FF9CFDDDF9475A6C94E3" box="[539,606,1753,1772]" pageId="4" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
A middle leg; B abdominal sternites; CE aedeagus, dorsal (C), lateral (D), and ventral (E). Scale bars: 0.3 mm in A, B; 0.2 mm in C, D, E.
</paragraph>
</caption>
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<paragraph id="8BA889586D69FF9DFF51F9545B5F94F8" blockId="5.[151,1436,1738,1783]" pageId="5" pageNumber="542">
Fig. 4. Lectotype of
<taxonomicName id="4C17F2DB6D69FF9DFE82F9545AB294D2" authority="Heller, 1936" authorityName="Heller, Cl. Besuchet" authorityYear="1936" box="[324,640,1738,1757]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="overbecki">
<emphasis id="B963554A6D69FF9DFE82F9545A2294D2" box="[324,528,1738,1757]" italics="true" pageId="5" pageNumber="542">Archiclaviger overbecki</emphasis>
<bibRefCitation id="EF86F4A96D69FF9DFDD3F9545AB294D2" author="HELLER K. M." box="[533,640,1738,1757]" pageId="5" pageNumber="542" pagination="64 - 76" refId="ref3763" refString="HELLER K. M. 1936: New taxa. In: HELLER K. M. &amp; GUNTHER K.: Ueber einige von H. Overbeck 1933 / 34 auf Java gesammelte Insekten. Tijdschrift voor Entomologie 79: 64 - 76." type="journal article" year="1936">Heller, 1936</bibRefCitation>
</taxonomicName>
.AD habitus, in dorsal (A), ventral (B), lateral (C), and axial (D) view; E head venter and prosternum; F antenna; G type label. Scale bars: 0.5 mm in AD; 0.2 mm in E, F.
</paragraph>
</caption>
<subSubSection id="C30DDAD36D69FF9DFF73F8335D3C956A" pageId="5" pageNumber="542" type="description">
<paragraph id="8BA889586D69FF9DFF73F8335D3C956A" blockId="5.[151,760,1838,2085]" lastBlockId="5.[828,1437,1838,2085]" pageId="5" pageNumber="542">
<emphasis id="B963554A6D69FF9DFF73F833593B95CA" bold="true" box="[181,265,1965,1989]" italics="true" pageId="5" pageNumber="542">Female</emphasis>
(
<figureCitation id="132C95DD6D69FF9DFEDAF830594195CA" box="[284,371,1966,1989]" captionStart="Fig" captionStartId="1.[151,182,1757,1776]" captionTargetBox="[176,1406,193,1739]" captionTargetId="figure-15@1.[166,1419,174,1746]" captionTargetPageId="1" captionText="Fig. 1. Archiclaviger gaofani sp. nov. AC habitus (A dorsal, male; B lateral, male; C dorsal, female); D frontoventral view of antennal bases, showing antennal fossae and scapes.Abbreviations: a13 = antennomere 13. Scale bars: 0.5 mm in AC; 0.2 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5177637" httpUri="https://zenodo.org/record/5177637/files/figure.png" pageId="5" pageNumber="542">Fig. 1C</figureCitation>
). Similar to male in external morphology; body size slightly larger; each eye composed of approximately 30 facets. Measurements (as for male): body length
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, length/width of head 0.41/
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, pronotum 0.38/
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, elytra 0.60/
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, abdomen 0.85/
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, length of antenna
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.
</paragraph>
</subSubSection>
<subSubSection id="C30DDAD36D69FF9DFCFAF8F05BB895EA" pageId="5" pageNumber="542" type="etymology">
<paragraph id="8BA889586D69FF9DFCFAF8F05BB895EA" blockId="5.[828,1437,1838,2085]" pageId="5" pageNumber="542">
<emphasis id="B963554A6D69FF9DFCFAF8F05B8D958A" bold="true" box="[828,959,1902,1925]" pageId="5" pageNumber="542">Etymology.</emphasis>
The new species is named after Fan Gao (Nanjing University,
<collectingRegion id="49D347BA6D69FF9DFBE6F8105C4195AA" box="[1056,1139,1934,1957]" country="China" name="Jiangsu" pageId="5" pageNumber="542">Jiangsu</collectingRegion>
,
<collectingCountry id="F300C9C86D69FF9DFBB9F8105CF195AA" box="[1151,1219,1934,1957]" name="China" pageId="5" pageNumber="542">China</collectingCountry>
), who collected this interesting species, and kindly donated the material to the SNUC.
</paragraph>
</subSubSection>
<subSubSection id="C30DDAD36D69FF9EFCFAF87059099754" lastPageId="6" lastPageNumber="543" pageId="5" pageNumber="542" type="biology_ecology">
<paragraph id="8BA889586D69FF9EFCFAF87059099754" blockId="5.[828,1437,1838,2085]" lastBlockId="6.[151,760,1188,1403]" lastPageId="6" lastPageNumber="543" pageId="5" pageNumber="542">
<emphasis id="B963554A6D69FF9DFCFAF8705C049A0A" bold="true" box="[828,1078,2030,2053]" pageId="5" pageNumber="542">Biology and host ant.</emphasis>
Both individuals were collected in March with a
<taxonomicName id="4C17F2DB6D69FF9DFC3EF7905C429A2A" authorityName="Emery" authorityYear="1906" box="[1016,1136,2062,2085]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B963554A6D69FF9DFC3EF7905C429A2A" box="[1016,1136,2062,2085]" italics="true" pageId="5" pageNumber="542">Nylanderia</emphasis>
</taxonomicName>
ant nesting under concrete blocks in a small green space in a highly industrial area of Jintan City (
<figureCitation id="132C95DD6D6AFF9EFEFEFB5A59E196D4" box="[312,467,1220,1243]" captionStart="Fig" captionStartId="6.[151,182,1017,1036]" captionTargetBox="[151,1436,190,999]" captionTargetId="figure-13@6.[151,1436,190,1001]" captionTargetPageId="6" captionText="Fig. 5. Collection site, distribution and host ant of Archiclaviger. A environment of the collection site of A. gaofani sp. nov.; B the collection site of A. gaofani sp. nov. is located in a highly industrial area of Jintan District, Changzhou; C distribution of Archiclaviger species in Asia; D a living adult of A. gaofani sp. nov. was discovered in a nest of Nylanderia ants after a concrete block had been turned over (photo by F. Gao)." figureDoi="http://doi.org/10.5281/zenodo.5177645" httpUri="https://zenodo.org/record/5177645/files/figure.png" pageId="6" pageNumber="543">Figs 5A, B, D</figureCitation>
). An attempt to find additional specimens in the same locality by the first author on
<date id="FFA9AF986D6AFF9EFF51FA9A597E9714" box="[151,332,1284,1307]" pageId="6" pageNumber="543" value="2020-08-24">August 24, 2020</date>
failed, possibly because it was already too late in season, but the presence of the same ant species was confirmed.
</paragraph>
</subSubSection>
<caption id="DF68D9D06D6AFF9EFF51FC675CD2964D" ID-DOI="http://doi.org/10.5281/zenodo.5177645" ID-Zenodo-Dep="5177645" httpUri="https://zenodo.org/record/5177645/files/figure.png" pageId="6" pageNumber="543" startId="6.[151,182,1017,1036]" targetBox="[151,1436,190,999]" targetPageId="6">
<paragraph id="8BA889586D6AFF9EFF51FC675CD2964D" blockId="6.[151,1437,1017,1090]" pageId="6" pageNumber="543">
Fig. 5. Collection site, distribution and host ant of
<taxonomicName id="4C17F2DB6D6AFF9EFD83FC675A8B9603" authorityName="Heller" authorityYear="1936" box="[581,697,1017,1036]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B963554A6D6AFF9EFD83FC675A8B9603" box="[581,697,1017,1036]" italics="true" pageId="6" pageNumber="543">Archiclaviger</emphasis>
</taxonomicName>
. A environment of the collection site of
<taxonomicName id="4C17F2DB6D6AFF9EFBE7FC675C4B9603" authority="Yin &amp; Hlaváč &amp; Cuccodoro, 2020" authorityName="Yin &amp; Hlaváč &amp; Cuccodoro" authorityYear="2020" box="[1057,1145,1017,1036]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="gaofani" status="sp. nov.">
<emphasis id="B963554A6D6AFF9EFBE7FC675C1C9603" box="[1057,1070,1017,1036]" italics="true" pageId="6" pageNumber="543">A</emphasis>
.
<emphasis id="B963554A6D6AFF9EFBFEFC675C4B9603" box="[1080,1145,1017,1036]" italics="true" pageId="6" pageNumber="543">gaofani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A250E8316D6AFF9EFBB9FC675CF69603" box="[1151,1220,1017,1036]" pageId="6" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
; B the collection site of
<taxonomicName id="4C17F2DB6D6AFF9EFF51FB8A58DD9628" authority="Yin &amp; Hlaváč &amp; Cuccodoro, 2020" authorityName="Yin &amp; Hlaváč &amp; Cuccodoro" authorityYear="2020" box="[151,239,1044,1063]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="gaofani" status="sp. nov.">
<emphasis id="B963554A6D6AFF9EFF51FB8A58969628" box="[151,164,1044,1063]" italics="true" pageId="6" pageNumber="543">A</emphasis>
.
<emphasis id="B963554A6D6AFF9EFF68FB8A58DD9628" box="[174,239,1044,1063]" italics="true" pageId="6" pageNumber="543">gaofani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A250E8316D6AFF9EFF32FB8A59049628" box="[244,310,1044,1063]" pageId="6" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
is located in a highly industrial area of Jintan District, Changzhou; C distribution of
<taxonomicName id="4C17F2DB6D6AFF9EFBCCFB8A5CF39628" box="[1034,1217,1044,1063]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B963554A6D6AFF9EFBCCFB8A5C4D9628" box="[1034,1151,1044,1063]" italics="true" pageId="6" pageNumber="543">Archiclaviger</emphasis>
species
</taxonomicName>
in Asia; D a living adult of
<taxonomicName id="4C17F2DB6D6AFF9EFF68FBB15935964D" authority="Yin &amp; Hlaváč &amp; Cuccodoro, 2020" authorityName="Yin &amp; Hlaváč &amp; Cuccodoro" authorityYear="2020" box="[174,263,1071,1090]" class="Insecta" family="Staphylinidae" genus="Archiclaviger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="gaofani" status="sp. nov.">
<emphasis id="B963554A6D6AFF9EFF68FBB15889964D" box="[174,187,1071,1090]" italics="true" pageId="6" pageNumber="543">A</emphasis>
.
<emphasis id="B963554A6D6AFF9EFF00FBB15935964D" box="[198,263,1071,1090]" italics="true" pageId="6" pageNumber="543">gaofani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A250E8316D6AFF9EFECAFBB1597D964D" box="[268,335,1071,1090]" pageId="6" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
was discovered in a nest of
<taxonomicName id="4C17F2DB6D6AFF9EFDF9FBB15AAD964D" authorityName="Emery" authorityYear="1906" box="[575,671,1071,1090]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B963554A6D6AFF9EFDF9FBB15AAD964D" box="[575,671,1071,1090]" italics="true" pageId="6" pageNumber="543">Nylanderia</emphasis>
</taxonomicName>
ants after a concrete block had been turned over (photo by F. Gao).
</paragraph>
</caption>
<subSubSection id="C30DDAD36D6AFF9EFF51FAFA5AA69774" box="[151,660,1380,1403]" pageId="6" pageNumber="543" type="distribution">
<paragraph id="8BA889586D6AFF9EFF51FAFA5AA69774" blockId="6.[151,760,1188,1403]" box="[151,660,1380,1403]" pageId="6" pageNumber="543">
<emphasis id="B963554A6D6AFF9EFF51FAFA59189774" bold="true" box="[151,298,1380,1403]" pageId="6" pageNumber="543">Distribution.</emphasis>
Eastern
<collectingCountry id="F300C9C86D6AFF9EFE40FAFA59F89774" box="[390,458,1380,1403]" name="China" pageId="6" pageNumber="543">China</collectingCountry>
:
<collectingRegion id="49D347BA6D6AFF9EFE12FAFA5A169774" box="[468,548,1380,1403]" country="China" name="Jiangsu" pageId="6" pageNumber="543">Jiangsu</collectingRegion>
(
<figureCitation id="132C95DD6D6AFF9EFDF5FAFA5AB59774" box="[563,647,1380,1403]" captionStart="Fig" captionStartId="6.[151,182,1017,1036]" captionTargetBox="[151,1436,190,999]" captionTargetId="figure-13@6.[151,1436,190,1001]" captionTargetPageId="6" captionText="Fig. 5. Collection site, distribution and host ant of Archiclaviger. A environment of the collection site of A. gaofani sp. nov.; B the collection site of A. gaofani sp. nov. is located in a highly industrial area of Jintan District, Changzhou; C distribution of Archiclaviger species in Asia; D a living adult of A. gaofani sp. nov. was discovered in a nest of Nylanderia ants after a concrete block had been turned over (photo by F. Gao)." figureDoi="http://doi.org/10.5281/zenodo.5177645" httpUri="https://zenodo.org/record/5177645/files/figure.png" pageId="6" pageNumber="543">Fig. 5C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>